taxonID	type	description	language	source
A14594A8F7BE55589E4AFAC941127A67.taxon	diagnosis	Diagnosis. Quediini (as recently redefined by Brunke et al. (2021)) can be distinguished from other Staphylininae based on the following combination of characters: disc of head and pronotum with microsculpture, at least on lateral part of either head or pronotum; head with frontoclypeal punctures, and with posterior frontal and basal macropunctures (e. g., Fig. 3 C) that are distinguishable from ground punctation by their larger diameter and longer, thicker setae; pronotum shield-shaped, slightly elongate to strongly transverse; profemora without apical row of lateroventral spines; protibiae without distinct subapical notch; all pretarsi with pair of empodial setae; all abdominal segments with only anterior transverse line (no traces of posterior transverse line), this line not encompassing spiracles (e. g., Fig. 3 E, F).	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
E35D008D421E5A10861EFB0A1EF660D5.taxon	description	Figs 2 A, B, 3 C, E, 4 A, B, 6 A, B, E, F, H, I, 9 A, 11 A (map)	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
E35D008D421E5A10861EFB0A1EF660D5.taxon	diagnosis	Diagnosis. Iratiquedius amabilis may be recognized within the genus by a combination of the evenly punctate elytra, the lack of golden setae or impressions at the base of the abdominal tergites and the distinctly impressed micropunctures on the anterior angles of the pronotum. The species most closely resembles I. mutator and can be distinguished from it by the distinct micropunctures of the anterior angles of the pronotum, the shorter apex of the median lobe in lateral view in males or broader, more densely setose apex of female tergite X.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
E35D008D421E5A10861EFB0A1EF660D5.taxon	distribution	Distribution. United States: CA. This species is known only from two rather close localities in the Sierra Nevada of California.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
CE4046D5DAF15D83AC30F5043F1172D7.taxon	description	Figs 2 A-F, 3 A, C-F, 4 A-D, 6 A-P, 7 A-N, 9 A-E	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
CE4046D5DAF15D83AC30F5043F1172D7.taxon	etymology	Etymology. The generic name is a combination of the Latin adjective ' iratus' and Quedius. It refers to the characteristic shape of the eyes, which are strongly convergent anteriad and create a comical, angry appearance.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
CE4046D5DAF15D83AC30F5043F1172D7.taxon	diagnosis	Diagnosis. Within Quediini, Iratiquedius can be distinguished from all other genera of the tribe by the distinctive eyes, which occupy nearly the entire lateral head margin, and are so convergent anteriad that their inner margin forms an obtuse angle with the suprantennal ridge (Fig. 3 A). The global diagnosis is the same as the regional Nearctic diagnosis.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
CE4046D5DAF15D83AC30F5043F1172D7.taxon	description	Description. Medium to small rove beetles, with variable coloration (Fig. 2 A-F). With the character states of Quediini (see Brunke et al. (2021)) and the following: antennomere 3 longer than 2, without dense setation; outer antennomeres (8 - 10) about as long as wide or shorter; antennae inserted close to inner margin of eye, separated by about the width of the antennal sclerite or less; head with eyes large, strongly convex, bulging from and nearly occupying entire lateral head outline, convergent anteriad and with inner margin forming obtuse angle with suprantennal ridge (Fig. 3 A); with basal puncture doubled (at least one side), interocular punctures present in some individuals of some species (I. amabilis, I. mutator) or absent, paraocular punctures absent, genal puncture absent (Fig. 3 C, D); frons not well-developed anterolaterad of antennal insertions; labrum notched medially, creating two lobes; apical maxillary and labial palpi fusiform and glabrous; infraorbital ridge complete to mandibles; gular sutures converging towards neck and narrowly spaced posteriad; mandibles with dorsal lateral groove absent or rudimentary, right mandible with single proximal tooth, tooth simple (Fig. 3 D) or bifid (Fig. 3 C); pronotum transverse to elongate, non-explanate, with three punctures in dorsal row, sublateral row at most reaching large lateral puncture, not extended posteriad; with only single large lateral puncture (e. g., Fig. 4 A-D); hypomeron strongly inflexed, not visible in lateral view; basisternum with pair of macrosetae (reduced in I. seriatus and I. prostans) and well-developed longitudinal carina; scutellum impunctate; elytron with subbasal ridge complete and forming scutellar collar, disc without microsculpture between punctures; row of humeral spines present and well-developed; elytral punctation evenly distributed or in serial rows (I. seriatus); foretibia with lateral spines (reduced in I. seriatus, absent in I. prostans) and apical spurs; metatarsomeres with disc setose; metatibia with at least three spines on outer face; abdominal tergite I with prototergal glands developed as moderately deep impressions, outer margin with row of setae; abdominal tergites not deeply impressed at base but some species with paired median impressions causing a ' pinched' appearance; abdominal sternite III with basal transverse line forming obtuse angle at middle, not produced posteriad; aedeagus with well developed paramere bearing peg setae; at least some species with discrete, paired internal sac sclerites that may be homologous with the ventral paired sclerites described by Brunke et al. (2016) (e. g., I. seriatus, I. uncifer (Fig. 7 H-K )).	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
CE4046D5DAF15D83AC30F5043F1172D7.taxon	distribution	Distribution. Iratiquedius is endemic to western North America.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
79373CA5B5195C8FA0F3D1F0BF6F8C21.taxon	description	Figs 2 C, D, 4 C, D, 6 C, D, G, J, K, 9 B, 11 A (map)	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
79373CA5B5195C8FA0F3D1F0BF6F8C21.taxon	diagnosis	Diagnosis. Iratiquedius mutator may be recognized within the genus by a combination of the evenly punctate elytra, the lack of golden setae or impressions at the base of the abdominal tergites and the indistinct micropunctures on the anterior angles of the pronotum. The species most closely resembles I. amabilis and can be distinguished from it by the indistinct micropunctures of the anterior angles of the pronotum, the longer apex of the median lobe in lateral view in males or sharp, pointed apex of female tergite X.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
79373CA5B5195C8FA0F3D1F0BF6F8C21.taxon	distribution	Distribution. United States: CA. This species is known only from two localities: one in the mountains of the northern Coast Range and one in the Central Valley region.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
5D0E804CB51750B4BEACACDFA7EAD003.taxon	description	Figs 2 E, 3 F, 6 L-P, 9 C, 11 B (map)	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
5D0E804CB51750B4BEACACDFA7EAD003.taxon	diagnosis	Diagnosis. Iratiquedius prostans can be distinguished by a combination of elytra with even punctation, not arranged in rows, and pale pubescence at the bases of the abdominal tergites and sternites.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
5D0E804CB51750B4BEACACDFA7EAD003.taxon	distribution	Distribution. Canada: BC. United States: AZ, CA, CO, ID, MT, NM, NV, OR, UT, WA. Iratiquedius prostans is the most widespread species of the genus. It occurs along the entire western cordillera, including both sides of the continental divide, and as far south as New Mexico in the east and near the United States border with Mexico, in the west. The species is not yet known from mountainous southern Alberta but is expected there.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
A2615E202B5B5C889021A0F48FAD0B85.taxon	description	Figs 2 F, 3 A, D, 7 A, B, D, E, H, I, L, M, 9 D, 11 C (map)	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
A2615E202B5B5C889021A0F48FAD0B85.taxon	diagnosis	Diagnosis. Iratiquedius seriatus can be distinguished from all other Iratiquedius except I. uncifer by a combination of: pronotum missing microsculpture on at least parts of the pronotum; elytra with serial punctation. From I. uncifer, it can be distinguished either by the rounded apices of the ventral paired sclerites of the internal sac, or the evenly convex disc of female tergite X.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
A2615E202B5B5C889021A0F48FAD0B85.taxon	distribution	Distribution. Canada: BC. United States: CA, OR, WA.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
DCEE72BCE6D1541BAB67C8117FBEECAF.taxon	description	Figs 7 C, F, G, J, K, N, 9 E, 11 C (map)	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
DCEE72BCE6D1541BAB67C8117FBEECAF.taxon	etymology	Etymology. The species epithet refers to the diagnostic hooked apex of the ventral paired sclerites of the internal sac.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
DCEE72BCE6D1541BAB67C8117FBEECAF.taxon	diagnosis	Diagnosis. Iratiquedius uncifer can be distinguished from all other Iratiquedius except I. seriatus by a combination of: pronotum entirely without microsculpture; elytra with serial punctation (Fig. 2 F). From I. seriatus, it can be distinguished either by the sharp, hooked apices of the ventral paired sclerites of the internal sac, or the distinctly impressed disc of female tergite X.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
DCEE72BCE6D1541BAB67C8117FBEECAF.taxon	description	Description. Measurements ♂ (n = 5): HW / HL 1.07 - 1.11; PW / PL 1.01 - 1.08; EW / EL 1.17 - 1.32; ESut / PL 0.76 - 0.93; PW / HW 1.01 - 1.07; forebody length 2.2 - 2.4 mm. Measurements ♀ (n = 5): HW / HL 1.06 - 1.10; PW / PL 1.03 - 1.08; EW / EL 1.19 - 1.29; ESut / PL 0.74 - 0.87; PW / HW 1.01 - 1.06; forebody length 2.2 - 2.6 mm. Extremely similar to I. seriatus and overlapping in most characters except: specimens of I. uncifer on average smaller, more slender, most examined specimens with bright orange-red pronotum and greenish elytra; pronotum entirely without microsculpture, at most with a few fragments of lines around the punctures; median lobe in ventral view with apical ridge (Fig. 7 C); median lobe in lateral view less projected ventrad (Fig. 7 F, G); internal sac with ventral paired sclerites sinuate, broad at base, strongly converging toward narrow, hooked apex (Fig. 7 J, K); paramere with emargination smaller or absent, peg setae fields more truncate at base (Fig. 7 N); female tergite X with long oval impression on disc occupying about half of its length (Fig. 9 E).	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
DCEE72BCE6D1541BAB67C8117FBEECAF.taxon	distribution	Distribution. United States: CA. This species is currently known only from a cluster of localities in the Klamath Mountains and Coast Ranges of California.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
B2FA19DD7046543EA676EFE1D05A329F.taxon	description	Figs 2 G, H, 5 A, B, 8 A-J, 9 F-G	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
B2FA19DD7046543EA676EFE1D05A329F.taxon	diagnosis	Diagnosis. Paraquedius is easily recognized within Quediini by a combination of the dark metallic blue / green reflections on the forebody, the punctate head and punctate scutellum. Within the Nearctic Region, Paraquedius is the only genus of Quediini with the disc of the head evenly punctate, at least on the posterior half. Worldwide, Paraquedius is superficially similar to the members of ' Clade L' of Brunke et al. (2021) (the Oriental and Palaearctic Multipunctatus and Intricatus groups of Quedius (Raphirus)), which are also metallic blue or green and have extensive head punctation. However, the latter have an impunctate scutellum, larger eyes and shorter appendages. Paraquedius is also superficially similar in habitus to members of the West Palaearctic clade (part of ' Clade B', Brunke et al. 2021) that consists of Quedius riparius and its close relatives. However, in Paraquedius the head is evenly punctate on the posterior half and the scutellum is punctate.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
B2FA19DD7046543EA676EFE1D05A329F.taxon	distribution	Distribution. Paraquedius is endemic to western North America, from coastal British Columbia, along the Coast, Cascade and Sierra Nevada Ranges, and as far south as the San Bernardino Mountains of California (Smetana 1981).	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
1B5D457B4DC958EF8B149A36C97702E8.taxon	description	Figs 2 H, 5 B, 8 A-F, 9 G, 11 D (map)	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
1B5D457B4DC958EF8B149A36C97702E8.taxon	etymology	Etymology. The species epithet refers to the diagnostic pale margin of the pronotum.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
1B5D457B4DC958EF8B149A36C97702E8.taxon	diagnosis	Diagnosis. Paraquedius marginicollis is most easily distinguished from the only other species of the genus by the pale base of antennomere 1 and margins of the pronotum. For other differences see the key above.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
1B5D457B4DC958EF8B149A36C97702E8.taxon	description	Description. Measurements ♂ (n = 5): HW / HL 1.04 - 1.09; PW / PL 1.01 - 1.08; EW / EL 1.13 - 1.17; ESut / PL 0.83 - 0.88; PW / HW 0.99 - 1.07; forebody length 4.0 - 4.3 mm. Measurements ♀ (n = 5): HW / HL 1.06 - 1.10; PW / PL 1.01 - 1.08; EW / EL 1.11 - 1.17; ESut / PL 0.86 - 0.89; PW / HW 1.01 - 1.08; forebody length 4.3 - 4.5 mm. Similar to P. puncticeps and differing only in the following: antennomeres 1 and 2 with pale base (Fig. 5 B); marginal area of pronotum and at least extreme base of elytra, suture and sometimes scutellar area, paler (Fig. 2 H); maxillary palpi usually slightly paler; head and pronotum with microsculpture more distinct; frons with central glabrous area not reaching posteriad to middle of eyes (Fig. 5 B); frons with pair of impressions shallower and usually more linear, forming a border around raised central area (Fig. 5 B); elytra with punctures slightly finer, disc always strongly uneven, with metallic greenish blue reflection; abdominal tergites with central, sparsely punctate to impunctate, raised areas, in addition to usual basal impressions; whorls of pale setae on tergites appearing more distinct from surrounding setae; male with emargination of sternite VIII slightly shallower, about twice as wide as deep; tergite X more slender, with narrower apex and setae more restricted to apical area; sternite IX with apex broadly, shallowly emarginate; median lobe of aedeagus in ventral view subparallel-sided, with truncate or broadly rounded apex (Fig. 8 A, B); median lobe in lateral view with short apical area that is acute to obtusely pointed (Fig. 8 C, D); paramere varying from slightly longer to slightly shorter than median lobe, spoon-shaped to lancet-shaped (Fig. 8 E, F), peg setae forming long, oval-shaped median field (Fig. 8 E, F). Female with tergite X shorter, with apex broader and truncate (Fig. 9 G).	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
1B5D457B4DC958EF8B149A36C97702E8.taxon	distribution	Distribution. Canada: BC. United States: CA, OR, WA.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
8533B3422EDE597992A98BC381EAE009.taxon	description	Figs 2 G, 5 A, 8 G-J, 9 F, 11 D (map)	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
8533B3422EDE597992A98BC381EAE009.taxon	diagnosis	Diagnosis. Paraquedius puncticeps is most easily distinguished from the only other species of the genus by the entirely dark first antennomere and pronotum. For other differences see the key above.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
8533B3422EDE597992A98BC381EAE009.taxon	distribution	Distribution. Canada: BC. United States: WA. Thus far, P. puncticeps is known only from the Vancouver area and Vancouver Island, British Columbia, Canada, and from one male collected somewhere in Washington. More collecting in its preferred microhabitats is needed to determine the full distribution of P. puncticeps.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
727C08730C0A55D58AA27AE059DAE8B2.taxon	description	Figs 5 C, D, 8 K-Q, 9 H, I	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
727C08730C0A55D58AA27AE059DAE8B2.taxon	diagnosis	Diagnosis. Quediellus, in the restricted sense used here, can be recognized within Quediini by a combination of: head with genal and interocular punctures absent; pronotum without extra punctures between dorsal and sublateral rows, sublateral rows not extended posteriad of single large lateral puncture (but sometimes at same level); prosternum without trace of longitudinal carina; scutellum impunctate; elytra with punctures not arranged in distinct rows, spaces between with distinct meshed microsculpture (Fig. 5 D). Quediellus and Quedionuchus are the only genera of Quediini with meshed (scale-like) microsculpture on the elytra (Fig. 5 D, E), while other lineages may have granulose microsculpture composed of micropunctures or microsetae (Fig. 5 F), superficially giving a similar dull appearance at low magnification. Quediellus differs from Quedionuchus by the irregularly scattered (not rows) or evenly distributed elytral punctures (Fig. 5 D versus Fig. 5 E) and complete infraorbital ridge, running from the neck to the base of the mandibles under the eye. Quediellus shares plesiomorphic, simple head chaetotaxy (though the basal puncture is often doubled, e. g., Fig. 5 C) with members of the mostly Palaearctic subgenus Quedius (Raphirus) (sensu Brunke et al. 2021), with which it was long considered to be synonymous. However, all Quedius (Raphirus) differ by the lack of meshed microsculpture on the elytra and those with a dull reflection between the punctures (e. g., Q. cincticollis Kraatz, Q. fumatus (Stephens), the members of clade ' X 2 ' of Brunke et al. 2021 (Q. lateralis (Gravenhorst) and its relatives )) have micropunctures rather than meshes, much denser elytral punctation and most have an, at least partly, carinate prosternum.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
727C08730C0A55D58AA27AE059DAE8B2.taxon	distribution	Distribution. Quediellus is endemic to the western Nearctic, occurring along the western cordilleras from southern British Columbia to southern California on the western side of the continental divide, and known from the Rocky Mountains of Alberta, Idaho and Montana on the eastern side.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
E88883057D0355F9A33B62C5A65F7029.taxon	description	Figs 5 C, D, 8 K-Q, 9 H, I, 11 E (map), 12	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
C806BB117C435FA98122B5BFAF3C1ED3.taxon	diagnosis	Diagnosis. As in generic diagnosis.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
C806BB117C435FA98122B5BFAF3C1ED3.taxon	distribution	Distribution. Canada: AB, BC. United States: CA, ID, MT, NV, OR, WA. Broadly distributed along the western cordilleras on both the eastern and western sides of the continental divide (Fig. 11 E). At the northern edge of its western distribution, Q. debilis is entirely flightless and appears to extend eastward along the Fraser River in British Columbia to at least the Hope area but it is not clear how distantly east and west populations are separated by the drier, central interior, if at all.	en	Brunke, Adam J. (2022): A world generic revision of Quediini (Coleoptera, Staphylinidae, Staphylininae), part 1. Early diverging Nearctic lineages. ZooKeys 1134: 129-170, DOI: http://dx.doi.org/10.3897/zookeys.1134.87853, URL: http://dx.doi.org/10.3897/zookeys.1134.87853
