taxonID	type	description	language	source
03E1879FFF94FFE97B78FAD9FC7465DC.taxon	type_taxon	TYPE SPECIES. — Baguoidea rubra Mahmood, 1967 by original designation.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF94FFE97B78FAD9FC7465DC.taxon	distribution	DISTRIBUTION. — Mainland Asia (Sri Lanka, Myanmar and China), Philippines and Japan (?), see Remarks of B. rufa below.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF94FFE97B78FAD9FC7465DC.taxon	discussion	REMARKS This monotypic genus is tentatively regarded as distinct from Dayus. It differs only in having the basal group setae of the subgenital plate blunt-tipped (also found in some Homa species) and distal macrosetae more numerous (Fig. 3 L) and in lacking an unusual ventral apodeme of the 8 th abdominal sternite, found in Dayus. In other respects, it is similar to Dayus in its overall red colour, having the 3 rd apical cell in the forewing petiolate (Fig. 3 C), male pygofer with processes (Fig. 3 G), form of the subgenital plate and aedeagus and well developed male basal dorsal abdominal apodemes (Fig. 3 M). The genus was redescribed by Yu & Yang (2013) as there were some errors in Mahmood’s original description and figures, which also had implications on subsequent studies (see Remarks under B. rufa below). Yu & Yang (2013) also noted that Dworakowska (1973) had stated (and figured) that the facial laterofrontal sutures were absent in Baguoidea but were present in their specimens, and are also present in the specimens studied here. This apparent mistake by Dworakowska may have been due to the sutures being obscure against the red facial colour.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF94FFE879CEFDBFFC9060E2.taxon	description	(Figs 1 A-F; 3)	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF94FFE879CEFDBFFC9060E2.taxon	distribution	DISTRIBUTION. — Mainland Asia (Sri Lanka, Myanmar and China), Philippines and Japan (?) see final comments in Remarks below.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF94FFE879CEFDBFFC9060E2.taxon	materials_examined	MATERIAL EXAMINED. — Sri Lanka • 1 ♀; Peradeniya; IV. 1906; Distant Coll; NHM. Myanmar • 1 ♂; Myitta, Doherty; coll. Distant; NHM; parasitized • 1 ♂; Myitta, Doherty; coll. Distant; NHMUK 013588830. Philippines • 1 ♂; Ifugao Prov., Luzon, Banaue; 20. VII. 1980; NHM.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF94FFE879CEFDBFFC9060E2.taxon	discussion	REMARKS B. rufa was described from a single specimen from Sri Lanka with the following data (translated from the German): “ Peradeniya. This nice Cicadine (1 ♂) was captured by Dr Uzel on 2 May 1903 in the Botanical Garden on the shrub Dichopsis laevifolia Benth. [= Palaquium laevifolium (Thwaites) Engl. (Sapotaceae)] ”. As the recorded host plant in Sri Lanka is an endemic (critically endangered) species and as B. rufa is known from outside Sri Lanka it clearly feeds on other hosts. The specimens recorded from Myanmar by Distant (1908) are probably the same as examined here (Fig. 1 E, F). Baguoidea rubra was described from the holotype male (Fig. 1 C, D) and five paratypes (Fig. 1 A, B) from the Philippines with data: “ Baguio, Benguet, Baker ” (USNM). The new synonymy of B. rufa and B. rubra is based on the type figures of the former given by Dworakowska (1973), the original description of the latter and images of its holotype sent by J. Zahniser (USNM) and the specimens studied. The differences between the two species, noted by Dworakowska (1973: 49), are either errors in the original description, i. e., Mahmood’s incorrect statement of forewings “ mottled with red patches ”, which are not present in the holotype images seen (see above) or an acceptable range of species variation, i. e., position of distal aedeagal processes; while the long pygofer processes figured by Mahmood for B. rubra is also probably an error. A specimen from Myanmar examined differs slightly in the male genitalia from the Sri Lanka type of B. rufa (figured by Dworakowska 1973) and the examined Philippine specimen in having the pygofer process slightly more sinuate apically and in having the lateral fine setae adjacent to the macrosetal row shorter. The same setae are shown longer and greater in number in Qin et al. ’ s (2010) fig. 26 of the junior synonym B. yunnanensis (see reproduced figure here, Fig. 3 L). The latter species was described from a single specimen from China and distinguished from B. rufa (and B. rubra, the other junior synonym of B. rufa) by the forewing colour (which according to all specimens seen is erroneous) and differences in pygofer and subgenital plate setae and spines at the apex of the aedeagal processes, all differences which fall within the accepted range of species variation. Genitalia figures drawn by Dworakowska (1973) were presumably taken from the holotype, as the only specimen examined, and as shown by Dworakowska’s figs 7 - 9 the base of the aedeagus was damaged when dissected. However, the correct aedeagal base is shown in Fig. 3 I (lateral view) and Fig. 3 J (dorsal view) which matches the specimens examined here and which is remarkably similar to that of some Dayus species (see Fig. 4 D). It should also be noted, that the subgenital plate basal group setae are dorsal (Fig. 3 L) rather than ventral as shown in Dworakowska’s (1973) figs 3, 4 and that the abdominal apodemes described by Qin et al. (2010), and shown in their figure 27 (and reproduced here, Fig. 3 M), are dorsal, and are a feature of the genus (see generic Remarks). Finally, the references for Japan for this species by Esaki (1932, 1950), Esaki & Ito (1954) and Kato (1933 b) need to be confirmed due to the similarity of some other red marked Empoascini (see Introduction). The reference of the species from Japan (Matsumura 1934) presumably refers to Dayus takagii Dworakowska, 1971, as this species was described from material in Matsumura’s collection from Japan and also Hong Kong (see Remarks under D. takagii).	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF95FFE87B6DFA9EFA246763.taxon	type_taxon	TYPE SPECIES. — D. elongatus Mahmood, 1967, by original designation.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF95FFE87B6DFA9EFA246763.taxon	distribution	DISTRIBUTION. — Widespread in Asia and the Pacific.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF95FFE87B6DFA9EFA246763.taxon	discussion	REMARKS This genus is tentatively regarded as distinct from Baguoidea (see Remarks under that genus). It differs only in having the basal group setae of the subgenital plate not blunt-tipped and in having an unusual ventral apodeme of the 8 th abdominal sternite with a pair of short anterior lobes (Fig. 4 F), and also found here with a series of short setae at the posterior lateral corner of the 8 th sternite. There is considerable variation among species in proportion of the vertex, size and colour, for example, in Dayus formosus the vertex is distinctly longer medially than next to eye but more or less the same in D. trifurcatus Yu & Yang, 2013 while the latter is yellow and not red as in some other congeners and very much bigger (two paratypes of D. trifurcatus [♂ and ♀] examined, NHM). A key to the Chinese species of the genus was given by Yu & Yang (2013). CHECKLIST AND DISTRIBUTION OF DAYUS SPECIES D. bifurcatus Yu & Yang, 2013: 3, figs 1 - 9. China (Zhejiang). D. elongatus Mahmood, 1967: 39, plate 8, fig. 1. Malaysia (Singapore). D. euryphaessus (Kirkaldy, 1907): 68. Fiji, Australia (?). D. formosus Dworakowska & Viraktamath, 1978: 544, figs 33 - 41. India, China (Hainan, Hong Kong new record, Taiwan). D. furcatus Xu, Dietrich & Qin, 2021 a: 231, figs 10 - 17. Thailand. D. lamellatus Qin & Zhang, 2007: 48, figs 22 - 31. China (Fujian, Zhejiang). D. lii Qin & Zhang, 2007: 45, figs 1 - 12. China (Fujian). D. membranaceus Qin & Zhang, 2007: 45, figs 13 - 21. China (Fujian, Jiangxi). D. serratus Yu & Yang, 2013: 4, figs 24 - 30. China (Hainan). D. takagii Dworakowska, 1971: 501, figs 1 - 11. China (Sichuan, Hong Kong, Taiwan), Japan. D. trifurcatus Yu & Yang, 2013: 3, figs 10 - 23. China (Chongqing). D. upoluanus (Osborn, 1934): 190, fig. 15. Western Samoa.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF95FFE579D5FC1FFD066260.taxon	description	(Fig. 2 A-F)	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF95FFE579D5FC1FFD066260.taxon	distribution	DISTRIBUTION. — Fiji, Australia (?) see Remarks below.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF95FFE579D5FC1FFD066260.taxon	materials_examined	MATERIAL EXAMINED. — Fiji • 1 ♂; Labasa; R. Veitch; VII. 1921; NHM • 1 ♀; Lautoka; W. Greenwood; 6. XII. 1921; breeding on leaves of Glochidion sp.; NHM • 1 ♀; Loloti; W. Greenwood; 19. XII. 1920; NHM.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF95FFE579D5FC1FFD066260.taxon	discussion	REMARKS This species was described from an unknown number of specimens (syntypic) from Fiji with data: “ Viti Levu, Rewa (Mar. - Apr.) Navna (Feb. Muir’s No. 53) on a native tree, also on Saccarum officinarum ” (Sugarcane) (BPBM). The data label on an imaged specimen seen (BPBM) labelled both “ type ” and “ Holotype ”, is as follows: “ Fiji Is 1905 [printed] 53 [handwritten] ” (Fig. 2 A, B). An image of four other syntype series specimens (labelled “ Paratype ”) has been seen (BPBM). In its description Kirkaldy noted the following: “ Allied to C. rufa (Melichar) [= Dayus rufus] but the head is longer, pronotum shorter, legs pale, etc. Bright scarlet, vertex pale testaceous with a medio-longitudinal suffused scarlet stripe, which forks at the base of the frons and extends all over the face suffusedly (sometimes the red stripe is obsolete at the fork, the frons then being entirely pale, only the genae and the clypeus, etc., red.) ”. Dworakowska (1971: 501) noted her placement of D. euryphaessus in Dayus was based on the original description, presumably its colour, proportions of the head and thorax and venation. In the original description it was also noted that the venation was similar to Baguoidea rufa and this is confirmed by Linnavuori’s (1960 a) redescription, which reads “ third apical cell of elytra triangular and stalked ” and the material examined here. However, the species is only tentatively retained in Dayus as the male genitalia show certain differences to that genus, i. e., the pygofer lacks a caudo-dorsal lobe, the subgenital plate lacks a basal setal group and the aedeagal shaft is very short without processes, all characters also found in the similar D. upoluana, although the aedeagus and connective are fused (Fig. 2 N) and dorsal abdominal apodemes are present (Fig. 2 I), as in other congeners. Linnavuori (1960 a) identified (and figured) D. euryphaessus from 29 specimens from Fiji, Lami, as part of a collection from Fiji sent to him by BPBM (see Introduction in Linnavuori 1960 a) and not the type series, also BPBM (see above). The identification is however considered correct based on the distinctive marking of the species. It is also worth noting that Linnavuori’s figure of the aedeagus is in lateral view, but its attached connective is shown in dorsal view, while his figure of long sternal abdominal apodemes are dorsal, as in other Dayus species. Linnavuori (1960 a) also described a new subspecies (Empoasca euryphaessa rubrocincta) from the holotype and allotype (BPBM) (Fig. 2 C-F) and one paratype in his own collection, from the same locality as the nominate subspecies, Fiji, Lami. It seems strange that Linnavuori (1960 a), when referring to his new subspecies, should say “ As the nominate form … ” as he described the red marking of the two subspecies differently, confirmed by the images reproduced here (Fig. 2 A, C, E). However, his observation that the male genitalia were the same in the two subspecies, and their same type locality, suggests they are the same taxon with unaccountably different markings. In synonymising the two, Dworakowska (1971) noted the type-series of Empoasca euryphaessa rubrocincta had been studied but from personal communication a single specimen was studied. The record of the species from Australia (Kuranda) by Evans (1966: 266), could be incorrect and could be the specimen of Homa haematoptila, from Kuranda, noted below.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF98FFE57B03F918FBD1657D.taxon	distribution	DISTRIBUTION. — India, China (Hainan, Hong Kong new record), Taiwan.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF98FFE57B03F918FBD1657D.taxon	materials_examined	MATERIAL EXAMINED. — India • 2 ♀; T. Nadu; Yercaud; I. Dworakowska det.; NHM. China • 1 ♂; Hong Kong (new record); NHM.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF98FFE57B03F918FBD1657D.taxon	discussion	REMARK This species was described from the holotype male and one male, three female paratypes from India. An additional male from Taiwan was also recorded.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF98FFE579DDFE1DFB686141.taxon	distribution	DISTRIBUTION. — Japan, Korea, China, Hong Kong (?) see Remarks below.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF98FFE579DDFE1DFB686141.taxon	materials_examined	MATERIAL EXAMINED. — Japan • 1 ♀; Kyushu; Fukuoka; 10. I. 1958; NHM.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF98FFE579DDFE1DFB686141.taxon	discussion	REMARKS This species was described from the holotype male and several paratypes (male and female) from Kyushu (Japan) and two female paratypes from Hong Kong. Although no depository for the types is indicated in the original description, it states in the acknowledgements to the paper that the type series was borrowed from EIHU, and this is presumed to be Matsumura’s collection. Therefore, Matsumura’s misidentification given in the original description (see above) refers to Matsumura (1934) where Empoasca rufa Melichar is listed from Japan and not (as stated) Matsumura (1931) where it is listed from Formosa. The female paratypes from Hong Kong could equally be D. formosus examined here from a male from Hong Kong and hence the question mark under Distribution. The colour of D. takagii was not originally described except that the species resembled D. euryphaessus but with paler coloration. The female tentatively identified here as this species (based on its locality Japan), is mainly reddish in colour.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF98FFE479C5FA39FD3E66DC.taxon	description	(Fig. 2 G-N)	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF98FFE479C5FA39FD3E66DC.taxon	distribution	DISTRIBUTION. — Samoa.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF98FFE479C5FA39FD3E66DC.taxon	materials_examined	MATERIAL EXAMINED. — Paratypes • 2 ♂, 1 ♀, 1 specimen missing from mount; “ Upolu Samoa ”; “ Apia, 9 - 15 - 23 ”; “ Swezey & Wilder ”; “ Brit. Mus. 1930 - 467 ”; NHMUK 013588829 • 1 ♂; “ Savaii, Samoa ” “ Safune, v. 2.24 ”; “ rain forest; 2000 - 4000 ”; “ H. Bryan Jr ”; “ Brit. Mus. 1930 - 467 ”; NHM.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF98FFE479C5FA39FD3E66DC.taxon	discussion	REMARKS This species was described from 16 specimens of both sexes (holotype, allotype and paratypes) with original data: “ Apia, 15. IX. 1923 Swezey and Wilder ” and two specimens, “ Savaii, Safume, 2. V. 1924, Rain Forest, 2000 - 4000 ft., Bryan ”. In the introduction to Osborn’s article it states that type material collected by Swezey & Wilder and by Bryan was deposited in BPBM and duplicates in Ohio State University. However, some specimens of the former collectors are in the NHM (see Material examined), and from which the first images and male genitalia drawings of the species are given here (Fig. 2 G-N). Based on its male genitalia the species is only tentatively retained in Dayus as some diagnostic features of the genus are not present, i. e., the pygofer lacks a caudo-dorsal lobe, the subgenital plate lacks a basal setal group and the aedeagal shaft is very short without processes, all characters found in the similar D. euryphaessus (see above), although the aedeagus and connective are fused (Fig. 2 N) and dorsal abdominal apodemes are present (Fig. 2 I), as in other congeners.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF99FFE47B42FCBFFC71659C.taxon	type_taxon	TYPE SPECIES. — Homa insignis Distant, 1908, by original designation.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF99FFE47B42FCBFFC71659C.taxon	distribution	DISTRIBUTION. — Widespread in Asia and the West Pacific.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF99FFE47B42FCBFFC71659C.taxon	discussion	REMARKS	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF99FFE479D6FE7DFC7E62C0.taxon	description	(Fig. 1 L, M)	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF99FFE479D6FE7DFC7E62C0.taxon	distribution	DISTRIBUTION. — Australia.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF99FFE479D6FE7DFC7E62C0.taxon	materials_examined	MATERIAL EXAMINED. — Australia • 1 ♀; Queensland; Kuranda; F. P. Dodd; VII. 1904; Homa haematoptila det. I. Dworakowska; 1969; NHM.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF99FFE479D6FE7DFC7E62C0.taxon	discussion	REMARKS Kirkaldy’s comments preceding his description that the “ tegmina are unfortunately a little mutilated ” suggests he had one specimen, the holotype, described as female with the following data: “ Queensland, Redlynch (vii) ”. This information matches the image of the type (now without abdomen), from BPBM, given here (Fig. 1 L, M). Dworakowska (1969) placed this species as a senior synonym of H. insignis but later (Dworakowska 1994 a) treated H. insignis as a valid species but this was apparently overlooked by Xu et al. (2022) who treated it still as a senior synonym of H. insignis, without justification. In fact, as H. haematoptila is known only from the female from Australia its identity remains uncertain, although from the described forewing markings and venation it is correctly placed in Homa and matches the female specimen examined here from Australia, Kuranda and some new species of Homa seen from the Pacific (NHM). The former specimen from Kuranda could represent the record of Dayus euryphaessus from Kuranda, by Evans (1966: 266). According to Murray Fletcher (pers. com.), the Redlynch type was collected as part of the Bishop Museum’s visit to Australia by Koebele and Perkins in 1904, which started Kirkaldy’s study of the Australian fauna.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF99FFE0790EF8B8FEF86181.taxon	description	(Figs 1 G-I; 6)	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF99FFE0790EF8B8FEF86181.taxon	distribution	DISTRIBUTION. — Sri Lanka, Thailand and China (new records) and India (?), see Remarks below.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF99FFE0790EF8B8FEF86181.taxon	materials_examined	MATERIAL EXAMINED. — Lectotype. Sri Lanka • ♂ (see Remarks); Sri Lanka: Peradeniya; V. 1905; Distant coll.; NHMUK 013588831. India • 1 ♀; Mishmi Hills; Delei River; 1700 ’; 7. II. 1935; M. Steele; NHM. Thailand • 10 ♂; Nakhon Nayok Khao Yai NP; behind football field; 14 ° 24.619 ’ N, 101 ° 22.778 ’ E; 770 m a. s. l.; malaise trap; 5 - 12. VII. 2006; Pong Sandao leg. T 142; INHS, QSBG. China • 1 ♂; Yunnan, Jinghong, Wild Elephant Valley; 29. IV- 4. V. 2017, coll. Ye Xu; NWAFU.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF99FFE0790EF8B8FEF86181.taxon	discussion	REMARKS This species was described from an unknown number of specimens (syntypic) with the following data: “ Ceylon; Peradeniya (Green) ”. Mahmood (1967) redescribed the species based on “ the type ” (and therefore is deemed to have designated the lectotype by inference, see ICZN 1999, Article 74.6). Although the male genitalia of the lectotype are now missing they were figured by Mahmood (1967) and although with certain inaccuracies, Mahmood’s figures are sufficient to identify the genus (see Qin et al. 2011: 31). Identification of Homa insignis from Mahmood’s poor figures is more problematical but the Asian specimens examined here (see Fig. 6) are correctly identified considering that Mahmood’s aedeagus figure is not lateral (as originally stated) but ventral, and the basal processes (as originally described and figured), are absent and possibly mistaken for the gonoduct. The specimen from India (Mishmi Hills), being female, is only tentatively identified as this species (and hence the question mark under Distribution), but the reference to the species from Lombok (Indonesia) by Jacobi (1941: 312) cannot be considered reliable. Dworakowska (1969) placed this species as a junior synonym of H. haematoptila but later (Dworakowska 1994 a) treated H. insignis as a valid species but this was apparently overlooked by Xu et al. (2022) who treated it still as a senior synonym of H. insignis, without justification (see Remarks under H. haematoptila). Figures of the type by McAtee (1934, pl. 3, fig. 26) indicate that it was probably he that made the wing mount (Fig. 1 I) and presumably corrected the poorly preserved jugal lobe (Fig. 1 H) in his drawing of the hind wing. The latter defect was probably that referred to by Matsumura (1932) in the following way: “ After studying the somewhat defect hind wing of this species in the British Museum, the author found that the genus belongs to the group Empoascaria, the first and second veins being uniting, and at the point of the uniting a cross-vein sent straightly downwardly, the third vein simple, stronger till the cross-vein; the cross-vein of the elytron at the dorsum straight ”.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF9DFFE07B63FA79FA24655D.taxon	description	(Fig. 1 J, K)	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF9DFFE07B63FA79FA24655D.taxon	distribution	DISTRIBUTION. — Malaysia: Peninsular, Malacca, Sabah (new record).	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF9DFFE07B63FA79FA24655D.taxon	materials_examined	MATERIAL EXAMINED. — Holotype. Malaysia • ♂; Malay Peninsula: Perak; Taiping; 8. XI. 1943; R. Takahashi; NHM. Paratypes. Malaysia • 1 ♂, 4 ♀, 1 specimen with abdomen missing; Malay Peninsula: Perak, Taiping; 8. XI. 1943; R. Takahashi; NHM • 1 ♀; Malay Peninsula: Kuala Lumpur; 7. VIII. 1931; on Solanum melongena (Aubergine); NHMUK 013588832 Non-types. West Malaysia • 1 ♂, 6 ♀, 1 immature; Malacca, Alor Gajah; 28. VIII. 1989; on cocoa; NHM • 1 ♀; same data; 17. VII. 1989; NMHUK 013387440 • 1 ♂, 5 ♀; Sabah; A. R. C. Tuaran; 22. V. 1973; on castor oil; NHM.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF9DFFE07B63FA79FA24655D.taxon	discussion	REMARKS In the original description of this species part of the legend is missing but communication with the author (Dr Dworakowska) indicates that the apodemes in her fig. 61 might be dorsal or ventral. The species is newly recorded from Sabah.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF9DFFE079B3FE3DFB5E67E3.taxon	distribution	DISTRIBUTION. — Taiwan, Japan (see Remarks).	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF9DFFE079B3FE3DFB5E67E3.taxon	materials_examined	MATERIAL EXAMINED. — No material examined.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF9DFFE079B3FE3DFB5E67E3.taxon	discussion	REMARKS The identity of this species, described from Japan and Taiwan is uncertain as it is known only from a female type and males are needed for species identification. However, from the figures given by Kato (1933 a, b) and images of a type given by Xu et al. (2022) taken by Dr Masami Hayashi from the Kato family collection (Japan), the marking on the forewings indicate it is correctly placed in Homa.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF9DFFE079A3FB9FFA9260D3.taxon	description	(Fig. 5)	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF9DFFE079A3FB9FFA9260D3.taxon	distribution	DISTRIBUTION. — China.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF9DFFE079A3FB9FFA9260D3.taxon	materials_examined	MATERIAL EXAMINED. — No material examined.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
03E1879FFF9FFFE27ABDFF5DFAF5606B.taxon	discussion	Note. Two species, known only from the female, i. e., Homa haematoptilus (Kirkaldy, 1906) from Australia and Homa rubrodorsata Kato, 1933 from Taiwan, are excluded from the key pending examination of the male.	en	Webb, Michael D., Xu, Ye (2022): Review of the “ red ” Empoascini leafhoppers in the genera Baguoidea Mahmood, 1967, Dayus Mahmood, 1967 and Homa Distant, 1908 (Hemiptera, Cicadellidae, Typhlocybinae) from Asia and the West Pacific. Zoosystema 44 (22): 549-563, DOI: 10.5252/zoosystema2022v44a22
