taxonID	type	description	language	source
2E5EF5096358FFD2B7BFFF58FD8FFA61.taxon	diagnosis	Ceratozamia reesii can be distinguished from other species in the genus by having 4 – 12 leaves per crown, light green color in emerging leaves, petioles armed with thin prickles, leaflets coriaceous, elliptic to lanceolate or oblanceolate, 11 – 23 pairs. Megastrobilus solitary erect, brown, peduncle 4.2 – 6 cm long. Seed 2 – 2.8 cm long.	en	Vovides, Andrew P., Pérez-Farrera, Miguel Angel, Salinasrodríguez, María Magdalena, Galicia, Sonia, Díaz-Jiménez, Pedro, Calonje, Michael, Gutiérrez-Ortega, José Said (2022): Morphological and anatomical analyses clarify the species definition of Ceratozamia latifolia Miq. (Zamiaceae) and lead to the description of a new species: Ceratozamia reesii. Phytotaxa 575 (3): 224-252, DOI: 10.11646/phytotaxa.575.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.575.3.2
2E5EF5096358FFD2B7BFFF58FD8FFA61.taxon	materials_examined	Holotype. MEXICO, San Luis Potosí: Xilitla, Camino a San Pedro Huiztquilico, 790 m a. s. l., 31 July 2021, M. A. Pérez-Farrera, P. Díaz, & M. M. Salinas 4017 ♀ (HEM). Isotype: XAL.	en	Vovides, Andrew P., Pérez-Farrera, Miguel Angel, Salinasrodríguez, María Magdalena, Galicia, Sonia, Díaz-Jiménez, Pedro, Calonje, Michael, Gutiérrez-Ortega, José Said (2022): Morphological and anatomical analyses clarify the species definition of Ceratozamia latifolia Miq. (Zamiaceae) and lead to the description of a new species: Ceratozamia reesii. Phytotaxa 575 (3): 224-252, DOI: 10.11646/phytotaxa.575.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.575.3.2
2E5EF5096358FFD2B7BFFF58FD8FFA61.taxon	description	Plant rupicolous, unbranching, stem short, cylindrical, erect, covered with persistent leaf bases, 12 – 33 cm tall, 10.5 – 17 cm in diameter. Cataphylls persistent, brown and densely tomentose at emergence, triangular apex acuminate. Leaves pinnate, 4 – 12 per crown, forming a semi-open crown, erect, ascending, olive green, 69.5 – 201 cm long, 39.6 – 71 cm wide, light green color at emergence, turning olive green at maturity. Petiole terete, 34 – 67.8 cm long, armed with thin prickles. Rachis green, terete, 34.5 – 138 cm long, erect, with sparse prickles diminishing into the distal end of the rachis. Leaflets 11 – 23 pairs, linear, long oblanceolate or oblanceolate, coriaceous, alternate to sub-alternate in the basal part of the leaf, subopposite in the median part, opposite in the apical part, subfalcate into apical part of leaflet, margin entire, apex acute to acuminate, asymmetric; base broad attenuate, articulation yellow adaxially, light green abaxially, 0.4 – 1.3 cm wide, veins 28 – 37, parallel, inconspicuous; median leaflets 19.8 – 35.5 cm long, 1.5 – 5.1 cm wide, spaced 1.2 – 7.2 cm between leaflets. Microstrobilus solitary, conical, erect, light brown when mature, 33 – 35 cm long, 4.8 – 5 cm diameter, peduncle densely tomentose, light brown, 14 – 15 cm long, 1.5 – 1.8 cm diameter. Microsporophyll cuneiform, 10.11 – 12.59 mm long, 4.69 – 6.25 mm wide, distal face bicornate, with light brown color at the central portion, sporangia zone on abaxial surface 4.10 – 7.65 mm long, microsporangia grouped in 3 – 4 per sorus. Megastrobilus solitary, cylindrical, erect, 16 – 16.3 cm long, 6.5 – 8 cm diameter; apex mucronate, brown pubescent at mature, peduncle short, brown, tomentose, 4.2 – 6 cm long, 1.06 – 1.56 cm in diameter. Megasporophylls peltate, bicornate, 3.7 – 4.2 cm wide, 1.5 – 2.1 cm tall, distal face pubescent, when mature, distal face presents a light brown to brown color at the central portion, which fades to brown-reddish towards the margins. Seed ovoid, sarcotesta cream when immature, sclerotesta beige when mature, 1.45 – 1.67 cm diameter, 2.0 – 2.28 cm long, with micropylar ridges.	en	Vovides, Andrew P., Pérez-Farrera, Miguel Angel, Salinasrodríguez, María Magdalena, Galicia, Sonia, Díaz-Jiménez, Pedro, Calonje, Michael, Gutiérrez-Ortega, José Said (2022): Morphological and anatomical analyses clarify the species definition of Ceratozamia latifolia Miq. (Zamiaceae) and lead to the description of a new species: Ceratozamia reesii. Phytotaxa 575 (3): 224-252, DOI: 10.11646/phytotaxa.575.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.575.3.2
2E5EF5096358FFD2B7BFFF58FD8FFA61.taxon	materials_examined	Specimens examined: MEXICO. San Luis Potosí: Xilitla, Las Pozas, Jardín Escultórico Edward James, 665 m, 24 April 2018, M. A. Pérez-Farrera 4021 (HEM); Xilitla, El Cañón, 580 m, 30 July 2021, M. A. Pérez-Farrera, P. Díaz, & M. M. Salinas 4016 (HEM); Xilitla, Cerro de la Silleta, 1300 m, 20 July 2021, M. M. Salinas s. n. (QMEX); Cultivated at Jardín Botánico Francisco Javier Clavijero (Veracruz), accession 2001 - 110 A, originally collected by S. Avendaño 5304 at Cerro del Muerto, Xilitla, 25 August 2014, A. P. Vovides 1522 (XAL).	en	Vovides, Andrew P., Pérez-Farrera, Miguel Angel, Salinasrodríguez, María Magdalena, Galicia, Sonia, Díaz-Jiménez, Pedro, Calonje, Michael, Gutiérrez-Ortega, José Said (2022): Morphological and anatomical analyses clarify the species definition of Ceratozamia latifolia Miq. (Zamiaceae) and lead to the description of a new species: Ceratozamia reesii. Phytotaxa 575 (3): 224-252, DOI: 10.11646/phytotaxa.575.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.575.3.2
2E5EF5096358FFD2B7BFFF58FD8FFA61.taxon	etymology	Etymology: The specific epithet was chosen to honor the memory of John David Rees (1932 – 2021), an American botanist and a mentor on cycads to the principal author. His guidance and knowledge on Mexican cycads were invaluable to the later creation of the Mexican National Cycad Collection of the Francisco Javier Clavijero Botanic Garden of INECOL, Xalapa, Veracruz.	en	Vovides, Andrew P., Pérez-Farrera, Miguel Angel, Salinasrodríguez, María Magdalena, Galicia, Sonia, Díaz-Jiménez, Pedro, Calonje, Michael, Gutiérrez-Ortega, José Said (2022): Morphological and anatomical analyses clarify the species definition of Ceratozamia latifolia Miq. (Zamiaceae) and lead to the description of a new species: Ceratozamia reesii. Phytotaxa 575 (3): 224-252, DOI: 10.11646/phytotaxa.575.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.575.3.2
2E5EF5096358FFD1B7BFF9A4FBB9FC7C.taxon	materials_examined	Neotype. MEXICO, San Luis Potosí: Route 70, 46 k West of [Ciudad] Valles, 20 Jul 1983, Stevenson 565 E (Neotype: NY).	en	Vovides, Andrew P., Pérez-Farrera, Miguel Angel, Salinasrodríguez, María Magdalena, Galicia, Sonia, Díaz-Jiménez, Pedro, Calonje, Michael, Gutiérrez-Ortega, José Said (2022): Morphological and anatomical analyses clarify the species definition of Ceratozamia latifolia Miq. (Zamiaceae) and lead to the description of a new species: Ceratozamia reesii. Phytotaxa 575 (3): 224-252, DOI: 10.11646/phytotaxa.575.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.575.3.2
2E5EF5096358FFD1B7BFF9A4FBB9FC7C.taxon	materials_examined	MEXICO, San Luis Potosí: Municipio of Ciudad del Maíz, at Ejido las Abritas, 850 m, 1974, J. Rees 1613 (MO, XAL).	en	Vovides, Andrew P., Pérez-Farrera, Miguel Angel, Salinasrodríguez, María Magdalena, Galicia, Sonia, Díaz-Jiménez, Pedro, Calonje, Michael, Gutiérrez-Ortega, José Said (2022): Morphological and anatomical analyses clarify the species definition of Ceratozamia latifolia Miq. (Zamiaceae) and lead to the description of a new species: Ceratozamia reesii. Phytotaxa 575 (3): 224-252, DOI: 10.11646/phytotaxa.575.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.575.3.2
2E5EF5096358FFD1B7BFF9A4FBB9FC7C.taxon	description	Plant rupicolous, unbranching. Stem short semihypogeous, ovoid-cylindrical, erect, sometimes, covered with persistent leaf bases, 8 – 10 cm tall, 10 – 12 cm in diameter. Cataphylls persistent, brown and densely tomentose at emergence, triangular apex acuminate. Leaves pinnate, 2 – 6 per crown, forming a open crown, erect, ascending, olive green, 60 – 219 cm long, 19.8 – 56.2 cm wide, reddish-brown color at emergence, turning olive green at maturity. Petiole, terete, greenish-light brown, 14.5 – 62 cm long, unarmed to rarely armed with thin, tiny and spread prickles. Rachis greenishbrown, terete, 42 – 161 cm long, unarmed or rarely with thin, tiny and sparse prickles diminishing into the distal end of the rachis. Leaflets 7 – 20 pairs, oblong or long oblanceolate, papyraceous, subopposite to alternate in the basal part of the leaf, opposite to subopposite in the median part, opposite in the apical part, margin entire, apex acute to acuminate, asymmetric; base broad attenuate, articulation green or brown adaxially, light green abaxially 0.33 – 1.09 cm wide, veins 32 – 46, parallel, inconspicuous; median leaflets 9.9 – 28.1 cm long, 2.5 – 4.9 cm wide, spaced 1.9 – 7.5 cm between leaflets. Microstrobilus solitary, conical, erect, green to brown, tomentose, 17 – 18 cm long, 2.2 – 2.6 cm diameter, peduncle brown, densely tomentose, 8 – 10 cm long, 0.8 – 1.2 cm diameter. Microsporophyll cuneiform, 8.10 – 10.70 mm long, 4.36 – 5.79 mm wide, distal face bicornate, with brown color at the central portion, sporangia zone on abaxial surface 4.58 – 7.82 mm long, microsporangia grouped in 3 – 4 per sorus. Megastrobilus solitary, cylindrical, erect, 8 – 13.5 cm long, 5.7 – 5.9 cm diameter; apex mucronate, light brown pubescent at mature, peduncle long, brown to orange, tomentose, 6.5 – 10 cm long, 0.78 – 1.25 cm in diameter. Megasporophylls peltate, bicornate, 2.3 – 3.9 cm wide, 1.4 – 1.8 cm tall, distal face pubescent, when mature, distal face presents a brown color at the central portion, which fades to green to brown towards the margins. Seed ovoid, sarcotesta cream when immature, sclerotesta beige when mature, 1.12 – 1.53 cm diameter, 1.21 – 1.68 cm long, with micropylar ridges.	en	Vovides, Andrew P., Pérez-Farrera, Miguel Angel, Salinasrodríguez, María Magdalena, Galicia, Sonia, Díaz-Jiménez, Pedro, Calonje, Michael, Gutiérrez-Ortega, José Said (2022): Morphological and anatomical analyses clarify the species definition of Ceratozamia latifolia Miq. (Zamiaceae) and lead to the description of a new species: Ceratozamia reesii. Phytotaxa 575 (3): 224-252, DOI: 10.11646/phytotaxa.575.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.575.3.2
2E5EF5096358FFD1B7BFF9A4FBB9FC7C.taxon	materials_examined	Specimens examined: MEXICO, San Luis Potosí: Ciudad de Maíz, 1100 m, 25 Apr 2001, S. Avendaño & M. A. Pérez Farrera 5320 (MEXU); 850 m, 2 August 2021, M. A. Pérez-Farrera et al. 4026 (HEM); 750 m, 13 Oct 1968, Puig 3979 (P); 850 m, 7 Nov 1974, J. Rees 1613 (MO, XAL); 1100 m, 5 Feb 1984, S. Sabato et al. 2340 (MEXU, MO); El Naranjo, 800 m, 13 Jun 2008; 1611 (XAL); 910 m, 20 Jul. 1983, D. W. Stevenson et al. 567 (MEXU); 895 m, 14 Jan 2001, T. Walters 2001 - 10 (MEXU); Rayón, 1100 m, 30 Jun 1962, F. Medellín Leal 1330 (MEXU); M. A. PérezFarrera et al. 4027 (HEM); 912 m, 5 Aug. 2003 Andrew P. Vovides 1466 (XAL); Tamasopo, 1000 m, 24 April 2001, S. Avendaño & M. A. Pérez-Farrera 5282 (MEXU); 900 m, 24 May 1981, P. A. Fryxell & W. R. Anderson 3586 (MEXU); 2 Jun 1968, Medellín Leal s. n. (MEXU accession 171861); 180 m, 01 August 2021, M. A. Pérez-Farrera 4023 (HEM); 20 July 1983, D. W. Stevenson et al. 565 A; 565 B; 565 C; 565 D; 565 G; 567 K (MEXU); 700 m, 15 Aug 2003, A. P. Vovides et al. 1465 (MEXU, XAL); 895 m, 13 Jan 2001, T. Walters et al. 2001 - 8 (XAL).	en	Vovides, Andrew P., Pérez-Farrera, Miguel Angel, Salinasrodríguez, María Magdalena, Galicia, Sonia, Díaz-Jiménez, Pedro, Calonje, Michael, Gutiérrez-Ortega, José Said (2022): Morphological and anatomical analyses clarify the species definition of Ceratozamia latifolia Miq. (Zamiaceae) and lead to the description of a new species: Ceratozamia reesii. Phytotaxa 575 (3): 224-252, DOI: 10.11646/phytotaxa.575.3.2, URL: http://dx.doi.org/10.11646/phytotaxa.575.3.2
