taxonID	type	description	language	source
03F88794755CFFC308D0FE7CFBF78224.taxon	description	(Figs. 1, 2, 6, 9, 10, 13, 14, 19) urn: lsid: zoobank. org: act: C 332 DB 03 - 8 FAF- 47 AC-B 45 A- 2 AA 1675 DEFE 2	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755CFFC308D0FE7CFBF78224.taxon	materials_examined	Type material. J. Holotype. [India] Karnataka: Bengaluru, Kommasandra, 12 ° 52 ′ 22.7 ″ N 77 ° 44 ′ 46.4 ″ E, 898 mt., 26. VII. 2021, MVL, Karthik Reddy M (gen. slide no. L 00051806) - coll. INPC. Paratypes. 1 ♀, India: Karnataka: Bengaluru, Dommasandra lake, 12 ° 52 ′ 26.5 ″ N 77 ° 44 ′ 52.8 ″ E, 901 mt., 20. XI. 2021, MVL, Karthik Reddy M (gen. slide no. L 00051805); 1 J, 1 ♀, India: Karnataka: Bengaluru, Dommasandra lake, 12 ° 52 ′ 26.5 ″ N 77 ° 44 ′ 52.8 ″ E, 901 mt., 09. VIII. 2021, MVL, Karthik Reddy M; 1 ♀, India: Karnataka: Bengaluru, Thigala Chowdadenahalli, 12 ° 53 ′ 16.9 ″ N 77 ° 44 ′ 34.7 ″ E, 918 mt., 27. VII. 2021, MVL, Karthik Reddy M; 1 ♀, India: Karnataka: Bengaluru, Dommasandra lake, 12 ° 52 ′ 26.5 ″ N 77 ° 44 ′ 52.8 ″ E, 901 mt., 01. IX. 2021, MVL, Karthik Reddy M; 1 ♀, India: Karnataka: Bengaluru, Dommasandra lake, 12 ° 52 ′ 26.5 ″ N 77 ° 44 ′ 52.8 ″ E, 901 mt., 09. XI. 2021, MVL, Karthik Reddy M-coll. INPC.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755CFFC308D0FE7CFBF78224.taxon	diagnosis	Diagnosis. Acanthoclita bengaluruensis sp. nov. is closely related to A. trichograpta (Meyrick) from Australia. The forewings of A. trichograpta have a well-developed dorsal blotch but absent in A. bengaluruensis sp. nov.. In male genitalia, cucullus of A. bengaluruensis sp. nov. has a constriction in inner margin, obtusely curved outer margin, isolated spine close to margin at base, and aedeagus is long and strongly curved medially, whereas in A. trichograpta, cucullus is evenly curved, crescentic, with isolated spine more distant from margin, and weakly curved short aedeagus. The female genitalia of A. bengaluruensis sp. nov. is characterized by ductus bursae 2 / 3 length of corpus bursae, and larger signa, whereas A. trichograpta has ductus bursae less than half the length of corpus bursae, and with shorter signa.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755CFFC308D0FE7CFBF78224.taxon	description	Description. Male (Fig. 1). Wingspan: 15 - 16 mm (n = 2). Head (Fig. 6). Ocellus large, chaetosema welldeveloped. Vertex dark brownish orange, with long scales anteriorly projecting between antenna. Upper frons brownish orange, with anteriorly projecting moderately long scales; lower frons light brownish orange, with minute, upwardly appressed scales. Labial palpi moderately long (1.4 x diameter of eye); first segment light brownish orange; second segment strongly widened distally with long dark brownish orange scales along ventral margin, and apically along dorsal margin; terminal segment brownish orange, short and stout, downward pointing, concealed within the long scales from distal part of second segment. Antenna filiform, moderately long, light brown, reaching middle of forewing. Thorax. Posterior crest absent. Pronotal collar and tegulae dark brownish orange with pale brownish orange scale tips. Legs orange with dark brown tarsal rings, unmodified. Wings. Forewing brownish orange, elongate, subrectangular; male costal fold absent, costa evenly curve; apex rounded; termen straight; costal strigulae fine, yellowish white separated by fine brown marks along costa, ending with a conspicuous dark grey to brown rounded spot at apex edged by yellowish white strigulae 9 connected with terminal strigulae between R 5 and M 1; basal fascia indistinct; subbasal fascia indistinct; median fascia extending obliquely from costa to inner margin near tornus, light brown mixed with brownish orange, with dark drown spot beyond median fascia between base of M 1 and M 3; ocelloid patch with silvery grey striae between R 5 and CuA 2, surrounded by row of longitudinal, short, parallel black lines extending in a curve from apical 3 / 4 of costa to near tornus; with a dark brown subrectangular bar along termen below apical dark spot; underside light brown with yellowish grey spots on costa Hindwing greyish brown with sparsely scaled, translucent basal half, well-developed cubital pecten, and anal margin with modified scaling between CuA 2 and 1 A + 2 A, greyish brown scales; underside light brown Abdomen (Figs. 9, 10). greyish brown scales dorsally, with greyish orange anal tuft. S 2 with well-developed anterolateral processes. Coremata, S 8 a small crescentic plate, with U-shaped rib posteriorly; intersegmental sclerite associated with small tufts of long, broad, distally rounded scales laterally; T 8 with strongly sclerotized T-shaped plate. Male genitalia (Figs. 13, 14). Vinculum a weakly sclerotized band. Tegumen subtriangular, weakly sclerotized. Uncus and socii reduced. Gnathos band-like, arising at middle of tegumen. Valva long and slender, constricted ventromedially; sacculus with large basal excavation; costal process well-developed; cucullus with a constriction apicodorsally 1 / 3, upcurved apically, obtusely curved outer margin, with rows of sparsely, shorter spines pointing outward and longer spines pointing inward, ventrobasally with one conspicuously long and narrow. Juxta subtriangular; caulis a well-sclerotized curved band, moderately long; aedeagus long, slender, tapering towards apex, strongly bent medially and apically weakly sinuate; vesica with a bundle of deciduous, lanceolate cornuti, and a distal series of short cornuti. Female (Fig. 2). Wingspan: 16 - 17 mm (n = 5). As male, but slightly darker forewing colour, without modified scales; frenulum of three bristles. Female genitalia (Fig. 19). Ovipositor lobes elongate. Hind margin of S 7 rounded-concave; anterior margin with a sclerotized curved band. Ostium a small semi-circular opening beyond hind margin of S 7. Sterigma semicircular ring surrounding ostium. Ductus bursae membranous, slender, about 2 / 3 length of corpus bursae, with a weak sclerotization just below ostium; origin or ductus seminalis from posterior 1 / 3 of ductus bursae. Corpus bursae ovate, with two short horn-shaped signa, pointed apex.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755CFFC308D0FE7CFBF78224.taxon	distribution	Distribution. Kommasandra, Dommasandra, Thigala Chowdadenahalli (India: Karnataka: Bengaluru).	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755CFFC308D0FE7CFBF78224.taxon	biology_ecology	Host. Unknown.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755CFFC308D0FE7CFBF78224.taxon	etymology	Etymology. The specific name refers to the name of the type locality Bengaluru, Karnataka, India.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755CFFC308D0FE7CFBF78224.taxon	discussion	Remarks. Diakonoff (1982) took into consideration the characters of head, wing venation and genitalia of both sexes while erecting the genus Acanthoclita. Most of the Acanthoclita species are identified and characterized by various apomorphies such as forewing pattern and venation, hindwing with translucent base and modified anal region, male genitalia with fused gnathos and tegumen, valva with crescentic to falcate cucullus and isolated spines at base of cucullus, female genitalia of more derived species with complex sterigma around ostium with sclerotized, funnel-shaped posterior end of ductus bursae, and simple sterigma with a patch of modified scales on each side in less derived species of the genus which it shares with Matsumuraeses Issiki, 1957.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755EFFC408D0FBC4FE3E85D8.taxon	description	(Figs. 3, 7, 11, 15, 16) urn: lsid: zoobank. org: act: C 09 D 2 E 42 - C 164 - 49 CB- 9 DE 2 - 472 D 6 F 28 CC 84	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755EFFC408D0FBC4FE3E85D8.taxon	materials_examined	Type material. J. Holotype. [India] Karnataka: Bengaluru, Dommasandra lake, 12 ° 52 ′ 26.5 ″ N 77 ° 44 ′ 52.8 ″ E, 901 mt., 09. XI. 2021, MVL, Karthik Reddy M (gen. slide no. L 00051939) - coll. INPC.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755EFFC408D0FBC4FE3E85D8.taxon	diagnosis	Diagnosis. Grapholita constricta sp. nov. is closely related to G. saphinella Razowski & Becker. The adult of G. constricta sp. nov. is orangish brown with indistinct dorsal blotch, and ocelloid patch with black dashes, whereas G. saphinella is brownish grey with conspicuous dorsal blotch, and ocelloid patch indicated by a vertical leaden line above tornus. The male genitalia of G. constricta sp. nov. is characterized by a wide vinculum, weakly pointed hood-shaped tegumen with curved bristles at apex, valva with deep and narrow constriction reaching to 2 / 3 width, straight dorsal margin, basal excavation occupying 2 / 3 sacculus, and long and slender aedeagus weakly tapering towards apex, whereas G. saphinella has narrow vinculum, triangular tegumen sparsely hairy and tapered apically, valva constriction reaching to 1 / 2 width, slightly sinuate dorsal margin, basal excavation occupying 1 / 2 sacculus, and aedeagus broad basal half, then tapering towards apex.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755EFFC408D0FBC4FE3E85D8.taxon	description	Description. Male (Fig. 3). Wingspan: 9.3 mm (n = 1). Head (Fig. 7). Ocellus large, chaetosema distinct. Vertex light brown, with long scales anteriorly projecting between antennae. Upper frons light brown to orangish white, with moderately long forward projecting scales; lower frons light brown, with minute upwardly appressed scales. Labial palpi moderately long (1.8 x diameter of eye), ascending; first segment light brown; second segment slender, slightly widened distally, light brown; third segment slender with pointed apex, light brown. Antenna moderately long, brown, reaching middle of forewing. Thorax. Posterior crest absent. Pronotal collar and tegulae light brown to orangish brown. Legs brown, unmodified. Wings. Forewing light brown mixed with orange white, subrectangular; male costal fold absent, costa evenly curved; apex rounded; termen slightly notched below apex, almost straight; apical half of costa with well-developed costal strigulae, strigulae 1 and 2 indistinct, strigulae 3 - 6 paired and separated by dark brown, strigulae 7 - 9 singled; interfascia between strigulae extending obliquely from costa to termen; ocelloid patch with four black dashes between M 1 and CuA 2, surrounded by greyish brown scales; outer margin with terminal strigulae, yellowish white between R 5 and M 1; underside light brown with yellowish white spots on costa and outer margin between veins. Hindwing pale brown to orange, subtrapezoidal, with well-developed cubital pecten; underside light brown. Abdomen (Fig. 11). Brown. S 2 with well-developed anterolateral processes. Coremata: S 8 a small crescentic plate with a pair of short anterolateral projections; intersegmental sclerite bearing lever rods laterally, associated with pair of small tufts of long filiform scales. Male genitalia (Figs. 15, 16). Vinculum wide V-shaped band. Tegumen ovate, weakly sclerotized, hood-shaped apex with inward folded membrane bearing dense, curve, short bristles. Uncus and socii reduced. Gnathos medially fused, weakly sclerotized band. Valva elongate, ventral margin with a deep and narrow constriction beyond middle of valva length, with straightened dorsal margin; sacculus subrectangular, with large basal excavation occupying 2 / 3 sacculus, small costal process apicoventrally with moderately dense of short setae mixed with sparsely long setae; cucullus ovate, with dense bristles except dorsally. Juxta rhomboidal; caulis long; aedeagus moderately long, gradually narrow towards apex, slightly curve apical 1 / 3; vesica with two groups of deciduous spiniform cornuti near base and distal 1 / 3 of aedeagus. Female. Unknown.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755EFFC408D0FBC4FE3E85D8.taxon	distribution	Distribution. Dommasandra (India: Karnataka: Bengaluru).	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755EFFC408D0FBC4FE3E85D8.taxon	biology_ecology	Host. Unknown.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755EFFC408D0FBC4FE3E85D8.taxon	etymology	Etymology. The specific name constricta refers to deep ventral valval constriction.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F88794755EFFC408D0FBC4FE3E85D8.taxon	discussion	Remarks. The most recent comprehensive review of the genus Grapholita was given by Komai (1999). Most of the Grapholita species are recognized by short, upcurved labial palpi, absence of sexual dimorphism in hindwing, well-developed coremata, with paired tufts of filiform or lanceolate scales, intersegmental ventral sclerite usually with a pair of long levers, and with a small crescent-shaped eighth sternite with a pair of short projections laterally, and male genitalia with weakly sclerotized tegumen, absence of uncus and socius, valva constricted near middle with rounded or ovate cucullus.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F887947559FFC708D0F983FD238224.taxon	description	(Figs. 4, 5, 8, 12, 17, 18, 20) urn: lsid: zoobank. org: act: A 9 D 53626 - 7889 - 49 CF- 853 E-DAAE 14 BFA 613	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F887947559FFC708D0F983FD238224.taxon	materials_examined	Type material. J. Holotype. [India] Karnataka, Bengaluru, Chikkavaderapura, 12 ° 53 ′ 31.7 ″ N, 77 ° 44 ′ 40.3 ″ E, 914 m, 09. VIII. 2021, MVL, Karthik, Reddy, M (gen. slide no. L 00051932) - coll. INPC. Paratypes: J, India: Karnataka, Bengaluru, Chikkavaderapura, 12 ° 53 ′ 31.7 ″ N, 77 ° 44 ′ 40.3 ″ E, 914 m, 06. VII. 2021, MVL, Karthik, Reddy, M (gen. slide no. L 00051933); ♀, India: Karnataka, Bengaluru, Chikkavaderapura, 12 ° 53 ′ 31.7 ″ N, 77 ° 44 ′ 40.3 ″ E, 914 m, 09. VIII. 2021, MVL, Karthik, Reddy, M (gen. slide no. L 00051934) - coll. INPC.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F887947559FFC708D0F983FD238224.taxon	diagnosis	Diagnosis. Thaumatotibia ramamurthyi sp. nov. is closely related to T. aclyta (Turner) but it can be distinguished by the greyish brown to blackish brown forewings without a distinctly paler distal region. The male genitalia of T. ramamurthyi sp. nov. is characterized by a constriction between sacculus and cucullus, and undulating outer margin of valva, whereas T. aclyta has a smooth outer and inner margin of valva, and absence of constriction between sacculus and cucullus. The female genitalia of T. ramamurthyi sp. nov. is characterized by narrow sclerotized ring around ostium, a subtriangular lamella postvaginalis of dense microtrichia, colliculum a small sclerotized ringshaped sclerite, inception of ductus seminalis at anterior 1 / 3 of ductus bursae, and smaller signa, whereas T. aclyta has a broader ring around ostium, lamella postvaginalis represented by two ovate bands of dense microtrichia, colliculum a large ring-shaped sclerite anterior of ostium, inception of ductus seminalis at anterior 1 / 4 of ductus bursae, and larger signa.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F887947559FFC708D0F983FD238224.taxon	description	Description. Male (Fig. 4). Wingspan: 16 - 18 mm (n = 2). Head (Fig. 8). Ocellus large, chaetosema welldeveloped. Vertex reddish-brown intermixed with dark brown. Upper frons dark brown, with anteriorly projecting short scales; lower frons dark brown, with minute, upwardly appressed scales. Labial palpi porrect, moderately long (1.8 - 1.9 x diameter of eye), dark brown; first segment short; second segment distally widened with appressed scales; third segment short with blunt apex. Antenna filiform, dark brown, reaching half-length of forewing. Thorax. Posterior crest a tuft of whitish scales. Pronotal collar dark brown with few reddish-brown scales. Tegulae dark brown mixed with copperish brown basally, some scales with white to cream apices. Legs dark grey with whiteringed tarsi. Wings. Forewings subtriangular, greyish brown ground color; male costal fold absent, costa almost straight except slightly curved apical 1 / 5; rounded apex; termen oblique and straight; distal paler region not distinct; staggered series of dark and white bands oblique to apex and parallel to termen; wing variably overlaid with dark grey to blackish-brown scales except for small pale band parallel to termen; costal strigulae light brown to grey broken by dark brown streaks; white scale tuft near base of dorsum; a small white dot in middle of wing; semicircular copperish-brown pretornal mark arched with black scales and outlined with whitish scales; cilia mixed whitish, grey and brown scales; underside light brown with greyish yellow spots on costa and outer margin between veins. Hindwing: Greyish brown; cilia pale grey; underside light brown. Abdomen (Fig. 12). Long copper-brown scales dorsally and grey anal tuft. S 2 with well-developed anterolateral processes. T 8 a broadly sclerotized plate, posterior margin medially convex. Coremata: S 8 a crescentic plate, with paired tufts of scales laterally. Male genitalia (Figs. 17, 18). Vinculum a moderately wide band. Tegumen moderately sclerotized, ovate, with broadened base and rounded apex. Uncus and Socii absent. Gnathos moderately sclerotized semicircular depressions on each side with spinulose base. Valva large, elongate-oval, slightly constricted near ventromedially, with a small subtriangular knob dorsomedially; sacculus with large basal excavation with moderately dense bristles along ventral margin, costal process small; cucullus ovate, densely setose, with distinct, sparsely spines projecting outward along ventral margin. Juxta subtriangular; caulis moderately long and sclerotized, with rounded basal spinulose lobes; anellus surrounding base of aedeagus; aedeagus bulbous basal 1 / 4, remainder abruptly narrowed with parallel-sided; vesica with series of deciduous spiniform cornuti. Female (Fig. 5). Wingspan: 19.5 - 21 mm (n = 1). As male, but with darker pre-apical blotch in the distal half of forewing, and absence of long grey scales dorsally on abdomen. Female genitalia (Fig. 20). Ovipositor lobes moderate, subreniform. S 7 trapezoidal. Ostium in shallowed incision of S 7 medially. Sterigma moderately sclerotized, narrow ring around ostium connected with subtriangular lamella postvaginalis, covered with dense microtrichia. Ductus bursae about 1 / 2 length of corpus bursae, gradually widened posteriorly with a small, sclerotized ring-shaped colliculum, inception of ductus seminalis at anterior 1 / 3 of ductus bursae. Corpus bursae ovate, with a weakly sclerotized, narrow ring of spinules at bursa neck, with two moderately long sickle-shaped signa.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F887947559FFC708D0F983FD238224.taxon	distribution	Distribution. Chikkavaderapura (India: Karnataka: Bengaluru).	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F887947559FFC708D0F983FD238224.taxon	biology_ecology	Host. Unknown.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F887947559FFC708D0F983FD238224.taxon	etymology	Etymology. The species name is given after Dr. V. V. Ramamurthy for his contribution to the Indian insect taxonomy.	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
03F887947559FFC708D0F983FD238224.taxon	discussion	Remarks. Komai (1999) has revised Thaumatotibia, providing illustrations of their genitalia and early stages and characterized the genus by following apomorphies: 1, broadly sclerotized plate on tergum 8 in males with medially convex posterior margin laterally produced into curved points; 2, sterigma represented by a rectangular or ovate raised rim connected posteriorly with a pair of ovate depressions with modified scales; 3, ring of spinules around the entrance of corpus bursae (Komai, 1999).	en	Reddy, Karthik M., Shashank, P. R. (2022): Three new species of the tribe Grapholitini (Lepidoptera: Tortricidae: Olethreutinae) from India. Zootaxa 5219 (6): 534-542, DOI: https://doi.org/10.11646/zootaxa.5219.6.2
