taxonID	type	description	language	source
039B8785FFB0FF84FF38FD20FC05FC94.taxon	type_taxon	TS: Phalaena arcuella Clerck, 1759. TL: Europe.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB0FF86FF38FCA0FE73FD40.taxon	description	(Figs 1, 4)	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB0FF86FF38FCA0FE73FD40.taxon	materials_examined	Type material. Holotype: 1 ♂, Korea, Hoengseong-Gun, Hwaheung-ri, 07. VI. 2018 (Y. B. Cha, & J. K. Ko), genitalia slide No. INU- 11284. Paratypes (10 ♂): Korea: 1 ♂, Chuncheon-Dam, 15. VI. 1995 (H. K. Lee & M. S. Go), genitalia slide No. INU- 11278; 1 ♂, Chuncheon, Kajeong-ri, 10. VI. 1999 (J. W. Jung & S. H. Won), genitalia slide No. INU- 11279; 1 ♂, Prov. Kangwon, Kajeong- ri, 10. VI. 1999 (J. C. Sohn), genitalia slide No. INU- 11280; 1 ♂, Mt. Palbong, 8. VI. 2004 (Bae et al.), genitalia slide No. INU- 11181; 1 ♂, Wando-Gun, Hwaheung-ri, 01. VI. 2015 (B. S. Park, S. M. Na, D. J. Lee), genitalia slide No. INU- 11183; 2 ♂, Kangwon, Pyung chang, 6. VI. 1996 (J. S. Lee), genitalia slide Nos. INU- 11293, INU- 11294; 1 ♂, Wando-Gun, Hwaheung-ri, 31. V. 2015 (B. S. Park, S. M. Na, D. J. Lee), genitalia slide No. INU- 11295; 2 ♂, Seoul, Mt. Inwangsan, 29. V. 1998 (S. H. Oh), genitalia slide Nos. INU- 11296, INU- 11297.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB0FF86FF38FCA0FE73FD40.taxon	diagnosis	Diagnosis. Superficially, the new species is very similar to O. captiosana (Falkovitsh, 1960) and O. subtilana (Falkovitsh, 1959). It can be distinguished from the latters by the curved angle of the preterminal fascia in the forewing that is situated near the tornus, whereas in the other two related species the preterminal fascia in forewing is situated near the termen. If compare the male genitalia of the new species with O. captiosana, the uncus of the new species is setose, bifurcated, and narrower. The new species possess a small spinous patch present at the middle of the sacculus. Also, the phallus of the new species is longer, than that of O. captiosana; cornutus is absent in the new species.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB0FF86FF38FCA0FE73FD40.taxon	description	Description. Male (Figs 1 a, 1 b). Head: Frons ivory; vertex ivory mixed with dark gray scales. Antenna filiform, covered with dark brown scales. Ocellus present. Labial palpus slightly upcurved and covered by ivory scales with a dark brown line on the outer surface, segment 2 with a dark brown mark, segment 3 exposed at the tip, erect, mixed with dark gray scales. Thorax: Patagium and tegula with yellowish-orange scales. Forewing length 15 – 18 mm, forewing elongatesubtriangular, broad; costa slightly curved outwards; apex pointed; termen convexly rounded; ground color orangeyellow, costal strigulae weak; basal area with three metallic silver, longitudinal streaks, second streak joined at a nearly right angle from the costal streak, costal streak oblique, extending slightly inward; subbasal fascia with a slightly curved metallic silver cross line, nearer to the median fascia, reticulated; median reticulated fascia with four metallic silver markings in the center of dark ovoid patch consisting of irregular dots and lines; preterminal fascia curved inward, near tornus; terminal fascia less curved than preterminal fascia, metallic silver; cilia ivory and pale dark gray. Hindwing elongate-ovate, pale dark brown, with a darker apical area; apex rather acute; cilia ivory, subbasal line dark brown. Abdomen: ivory and dark brown. Male genitalia (Figs. 4). Uncus narrow, apex bifurcate and with spines on top, setaceous. Socius drooped and setaceous. Gnathos sclerotized partly. Tuba analis shorter than uncus and membranous. Tegumen long-trapezoidal. Valva flattened, broad, not folded at middle. Cucullus narrower than sacculus, apex rounded and setaceous, spined, along ventral margin, in a row on basal costa. Sacculus with a patch of short, small spines on middle of sacculus; short and stout spines grouped on angular part. Phallus about as long as cucullus, split apically into a pair of mittensshaped processes. Female. Unknown.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB0FF86FF38FCA0FE73FD40.taxon	distribution	Distribution. Korea (GG, GW, JN).	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB0FF86FF38FCA0FE73FD40.taxon	biology_ecology	Host plant. Unknown	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB0FF86FF38FCA0FE73FD40.taxon	discussion	Remarks. This species is very similar to O. captiosana (Falkovitsh, 1960) and O. subtilana (Falkovitsh, 1959), and the three forms a tight species group.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB0FF86FF38FCA0FE73FD40.taxon	etymology	Etymology. This specific name is derived from the Latin, “ oculatus ” meaning “ eye-shaped ”, referring to ocellus markings on the forewing.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB2FF81FF38FD0CFCE7FED9.taxon	description	(Figs 2, 5)	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB2FF81FF38FD0CFCE7FED9.taxon	diagnosis	Diagnosis. This species is similar to O. subtilana (Falkovitsh) in appearance, but can be separated by the three metallic silver markings in the ocellus of the forewing, while O. captiosana has four metallic silver markings on the forewing. In the male genitalia, this species differs from O. subtilana by its sacculus which is slightly projected beyond the middle of the ventral edge, the phallus has three groups of spines on vesica in the male genitalia of O. captiosana, while in O. subtilana the phallus has simple and without cornuti.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB2FF81FF38FD0CFCE7FED9.taxon	description	Redescription. Male (Fig. 2). Head: Frons with ivory scales; vertex mixed with dark brown scales. Antenna filiform, covered with dark brown scales. Ocellus present. Labial palpus slightly upcurved and covered by ivory scales, the apical segment with tip exposed, erect, mixed with dark brown scales. Thorax: Patagium with grayish orange scales; tegula grayish orange mixed with grayish brown scales. Forewing length 22 – 25 mm in males. Forewing broad, forewing elongate-subtriangular, costa slightly curved outwards; apex pointed; termen convexly rounded; brownish orange, the basal area with three metallic silver longitudinal streaks, costal streak oblique slightly inward; second streak is straight, not joined with costa; subbasal fascia with a slightly straight metallic silver cross line; median area reticulated with three eye-like metallic silver markings in the center of dark ovoid plate consisting of irregular dots and lines; preterminal fascia with two metallic silver bands extending slightly inward, upper band shorter than lower one and two short metallic silver bands are present near costa and tornus; cilia ivory and dark brown. Hindwing elongate-ovate, apex rather acute; dark brown, darker in the apical area; cilia ivory and dark brown. Abdomen: ivory and dark brown. Male genitalia (Fig. 5). Uncus stout, apex bifurcate, setose. Socius moderately broad, setose. Gnathos partly sclerotized. Tuba analis shorter than uncus, membranous. Tegumen long-trapezoidal. Valva elongate, not folded at middle, more or less flat. Cucullus slightly narrower than sacculus, apex rounded, setose, with spines, slightly projected beyond the middle of the ventral edge. Sacculus strongly widened beyond the middle, convex, angulated on the ventral edge, with a stout spine cluster on the medial surface and along the ventral edge distally. Phallus nearly of the same length as cucullus, sclerotized on the left side, three groups of spines on vesica are present, the central group with few small spines, and two lateral groups with two stout spines that are longer than the spines of the central group. Female. Not examined.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB2FF81FF38FD0CFCE7FED9.taxon	materials_examined	Material examined (10 ♂). Korea: 5 ♂, Mt. Hambaek, Taebaek-shi, 13. VI. 1999 (Bae et al.), genitalia slide Nos. INU- 11053, INU- 11054, INU- 11055, INU- 11056, INU- 11057; 1 ♂, Mt. Taebaek, Kongwon, 27. VI. 1996 (Lee et al.), genitalia slide No. INU- 11058; 1 ♂, Undu-ryeong, Hongcheon-gun, 20. VII. 2000 (Bae et al.), genitalia slide No. INU- 11059; 2 ♂, Mt. Taebaek, Taebaek-shi, 13. VI. 1999 (Bae et al.), genitalia slide Nos. INU- 11060, INU- 11061; 1 ♂, Temp. Baekdamsa, Mt. Seolaksan, Injeagun, 4. VII. 2002 (Bae et al.), genitalia slide No. INU- 11127.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB2FF81FF38FD0CFCE7FED9.taxon	distribution	Distribution. Korea (Park, 1983), Japan (Nasu et al. 2013), China (Hua, 2005), Russia (Falkovitsh, 1960).	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB2FF81FF38FD0CFCE7FED9.taxon	biology_ecology	Host plant. Larvae have been reported to feed on fallen leaves in Japan (Nasu et al. 2013).	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB2FF81FF38FD0CFCE7FED9.taxon	discussion	Remarks. This species was misidentified as Olethreutes arcuella by Walsingham (1900) from Korea, but has lately been corrected as O. captiosana by later Park (1983).	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB5FF81FF38FEB4FCE0F9FE.taxon	description	(Figs 3, 6)	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB5FF81FF38FEB4FCE0F9FE.taxon	description	Olethreutes subtilanus Falkovitsh: Razowski, 1999, Shilap. Revista de lepidopterología, 27 (5): 103.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB5FF81FF38FEB4FCE0F9FE.taxon	diagnosis	Diagnosis. This species is similar to O. captiosana (Falkovitsh) in appearance, but it can be distinguished by the conspicuously smaller forewing length and four metallic silver markings on the forewing, while O. captiosana has three metallic silver markings in the ocellus of the forewing. In the male genitalia, this species also differs from O. captiosana by a line of stout spines at basal 1 / 3 part of cucullus, the sacculus almost as wide as cucullus, and the distal margin of the sacculus is covered by hair-like spines.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB5FF81FF38FEB4FCE0F9FE.taxon	description	Redescription. Male (Fig. 3). Head: Frons covered with ivory scales. Vertex mixed with dark brown scales. Antenna filiform, covered with dark brown scales. Ocellus present. Labial palpus weakly upcurved, covered by ivory scales, apical segment shortly exposed and erect, mixed with dark brown scales. Thorax: Patagium with orange scales; tegula grayish orange mixed with grayish brown scales. Forewing length 12 – 15 mm in males. Forewing broad, elongate-subtriangular, costa slightly curved outwards; apex pointed; termen convexly rounded; brownish orange; the basal area with three metallic silver longitudinal streaks, costal streak slightly oblique inwards; the second streak is straight, not joined with costa; subbasal fascia with a straight metallic silver cross line; the median reticulated area with four eye-like metallic silver markings in the center of the dark ovoid plate, consisting of irregular dots and lines; preterminal fascia with two metallic silver bands extending slightly inward, upper band shorter than the lower one and one short metallic silver band near tornus; cilia ivory and dark brown. Hindwing elongate-ovate, apex rather acute; dark brown, darker apical area; cilia ivory and dark brown. Male genitalia (Fig. 6). Uncus broad, hooked, apex bifurcate and setose. Socius moderate and drooped. Gnathos partly sclerotized. Tuba analis membranous. Tegumen narrow, long-trapezoidal. Valva elongate, not folded at middle. Cucullus shallowly emarginated at basal 2 / 3, with line of stout spines at basal 1 / 3. Sacculus almost as wide as cucullus; a small patch of fine, hair-like spines is present near the distal margin of the sacculus. Phallus narrow apically, partly membranous, without cornuti. Female genitalia. not examined.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB5FF81FF38FEB4FCE0F9FE.taxon	materials_examined	Material examined. 2 ♂, Mt. Hambaek, Taebaek-shi, 13. VI. 1999 (Bae et al.), Genitalia slide Nos. INU- 11287, INU- 11290.	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
039B8785FFB5FF81FF38FEB4FCE0F9FE.taxon	distribution	Distribution. Korea (Park, 1983), Japan (Nasu et al., 2013), China (Hua, 2005), Russia (St. Petersburg, Siberia) (Falkovitsh, 1959), Germany (Segerer, 2010). Hostplants. Fallen leaves in Japan (Nasu et al., 2013).	en	Kim, Jeong-Nam, Bayarsaikhan, Ulziijargal, Lee, Tak-Gi, Cha, Yeong-Bin, Kim, Hanul, Byun, Bong-Kyu, Bae, Yang-Seop (2022): A new species of Olethreutes Hübner, 1822 (Lepidoptera, Tortricidae, Olethreutinae) from Korea. Zootaxa 5219 (5): 486-492, DOI: https://doi.org/10.11646/zootaxa.5219.5.6
