identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
1A0D16767638FF94FF080C71FD20F86D.text	1A0D16767638FF94FF080C71FD20F86D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calotes medogensis Zhao & Li 1984	<div><p>Calotes medogensis Zhao &amp; Li, 1984</p> <p>Material examined: WII-ADR1211, a female (Figure 3A,B)collected from Ramsing(28°39ˊ22.71˝ N, 94°58ˊ46.22˝ E, Elevation: 601 m asl.), Mouling National Park, Upper Siang, Arunachal Pradesh on 28 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.97951&amp;materialsCitation.latitude=28.656307" title="Search Plazi for locations around (long 94.97951/lat 28.656307)">October</a>, 2021 (Figure 1); WII-ADR452, a female (Figure 3 C, D) collected from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.97951&amp;materialsCitation.latitude=28.656307" title="Search Plazi for locations around (long 94.97951/lat 28.656307)">Poba Reserve Forest</a>, Arunachal Pradesh, on 30 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.97951&amp;materialsCitation.latitude=28.656307" title="Search Plazi for locations around (long 94.97951/lat 28.656307)">September</a>, 2018 (Figure 1).</p> <p>Diagnosis: A moderate-sized agamid lizard, SVL 76–95.9 mm (n= 3), TailL 212–271 mm (n= 3). Nuchal and dorsal crests continuous, consisting of erect compressed scales, distinctly pronounced at nuchal crests than the dorsal crests, smaller posteriorly. Two series of supratympanic spines present. Body laterally compressed, scales at the trunk heterogeneous in size, larger than ventrals, larger at the mid-body towards dorsal side, dorsally and posteriorly oriented, 40–42 dorsal scales, 53–58 rows of scales at midbody, 88–91 ventrals. Mental smaller than chin shields, first pair of chin shields are narrow and elongated, separated from each other by two small scales, the gular scales at the mid-line are distinctly smaller than the adjacent gular scales away from the mid-line. Dorsally and ventrally green in colour.</p> <p>Morphological description based on newly collected materials: Morphometric and meristic characters are given in the Table 2. Moderate-sized lizard, SVL 82.8–95.9 mm (n= 2), head distinct from neck (Figure 3), moderately large, longer than width (HeadW/HeadL= 0.67–0.68), slightly concave in front, snout rounded in dorsal view, longer than orbital diameter (OrbD/SnEye= 0.65–0.74), loreal slightly concave, supraciliary and canthal ridge sharp, nostril elliptical, nasal large, orbit three times larger than tympanum, pupil rounded, tympanum oval, rostral wider than mental, dorsal snout and head scales heterogenous in shape and size (Figure 6 A, D), scales at interorbital region, parietal region and occipital region have spinules forming irregular keel, scales at the loreal region, below and behind the orbit keeled, heterogenous in shape and size, scales surrounding the eye are granular, feebly conical, scales surrounding tympanum are smaller than that of mandibular region and nape, supralabials nearly rectangular, elongated, smooth, rostral wide and narrow, interparietal scale with a distinct pit, two series of ridge on temporal region above tympanum (Figure 6 C, F), upper row is consists of 9–10 scales, continuous with supraciliary scale to neck, posterior two or three scales of the upper row more pronounced or spinous, the lower row just above tympanum consisting of 9–10 scales and separated by two to three scales from tympanum, scales posteriorly more pronounced forming spine. Mental subtriangular or nearly hexagonal, smaller than chin shields, infralabials smooth, narrow and elongated, first pair of chin shields are narrow and elongated, separated by two small scales, posterior chin shields are separated from infralabials by a series of narrow scales, scales between the chin shields are smaller than the rest of the gular scales, irregular in shape and size (Figure 7 B, E), feebly keeled, gular scales posteriorly pointed, keeled, subimbricate, gular scales (except the middle scales) larger than that of the belly, the gular scales at the mid-line are distinctly smaller than the adjacent gular scales away from the mid-line (Figure 7 C, F). Scales on neck are similar shape, posteriorly pointed; upwardly and posteriorly directed, size increases dorsally and posteriorly, 8–9 nuchal spine, size increases posteriorly (Figure 6 C, F), length of the longest nuchal spine was more than twice the width of it (LongN/WidthN = 2.19–2.71, n= 2). Body laterally compressed; triangular in cross section. Dorsal crest reduced to mere ridge posteriorly. Dorsal and lateral scales of the trunk larger than ventrals, heterogenous in size; larger at the mid-body towards dorsal side, feebly keeled; pointed, dorsally and posteriorly oriented, 40–42 dorsal scales, 53–58 rows of scales around trunk at midbody, ventral scales smaller than that of dorsal and lateral, strongly keeled; heterogenous in size, pointed, 88–91 ventrals, scales at the axilla, groin and insertion of hind limb are smaller, a fold consisting of very small scales present in front of the shoulder. Limbs moderate and thin. Forefoot length nearly equal to upper arm length (UparmL/ForefL= 0.99–1.03). Crus length and thigh length almost equal (CrusL/UplegL= 1.02–1.09). All scales on limbs are keeled, smaller at the insertion, scales on ventral side of thigh are smaller than that of crus. Digits slender; with strong and curved claw, lamellae entire, bicarinate (more than two keels at the base). Tail long, two to three times of SVL, rounded, gradually tapering posteriorly, dorsal scales slightly smaller than that of ventral, scales at the base near cloaca smaller than the rest, ventral scales strongly keeled, forming continuous ridge.</p> <p>Colouration in preservative: WII-ADR1211, head, anterior part of back, forelimb, crus and foot dorsally bluish grey, large black patch from axilla to middle of the trunk, longer on left side, lower half of the trunk yellowish green, vertebral scales bluish grey all along the body, thigh scales dorsally green with black edge, almost anterior one third of the tail bluish grey, posteriorly greyish brown, three series of whitish scales on back arranged in “˄” shape. Fold on shoulder black. Head laterally bluish grey. Anterior gular scales reddish, rest gular scales intermixed with grey and yellowish green, chest scales yellowish, anterior part of belly intermixed with grey and yellowish scales, posterior part of the belly light yellow, hind limbs ventrally light yellow, forelimbs greyish, tail at the base and some extent of anterior part greenish yellow, posteriorly turn into grey. Small light brown patch on elbow and knee, a large light brown patch on heel to the base of 5 th toe. WII-ADR452, dorsally bluish grey, slightly dark patches on head, neck, above axilla and thigh, a large black patch from axilla to groin covering dorsolateral and lateral side of the trunk on both sides, ventrally entire body grey. Light cream coloured patch on elbow, knee and heel, larger on heel.</p> <p>Colouration in life: Dorsal, lateral and ventral side of head green (Figure 9), dark stripes present on orbit radiating from eye, sometimes faint. Dorsum and tail dorsally green, series of discontinuous white scales on back arranged in “˄” shape. A dark brown fold/pit on shoulder, ventrally belly and tail light green, light brown patch on elbow, knee and heel.</p> <p>Morphological comparison: Calotes medogensis closely similar to Calotes jerdoni, but it can be differentiated from the latter by more pronounced nuchal spines, i.e. nuchal spines distinctly longer than that of the dorsum (vs. nuchal spines not distinctly longer than those of the dorsum), length of the longest nuchal spine more than twice the width (vs. length of the longest nuchal spine less than twice the width), scales on trunk oriented obliquely upward (vs. scales on trunk oriented backward) (Figure 8), posterior 2–3 scales of the upper row of ridge above tympanum are spinous (vs. all the scales are similar), mental smaller than the first two pairs of chin shields (vs. mental larger than the chin shields) (Figure 7), the first two pairs of chin shield are longer than wide (vs. chin shields are as long as wide) (Figure 7), first pair of chin shield are separated by two small scales below mental (vs. first pair of chin shield are separated by small single scale below mental) (Figure 7), the scales between the chin shields are irregular in shape (vs. scales between the chin shields are almost similar shaped), the gular scales at the mid-line are distinctly smaller than the adjacent gular scales away from the mid-line (vs. gular scales at the mid-line are nearly equal to the adjacent gular scales away from the mid-line) (Figure 7), number ventrals is more in Calotes medogensis (88–91) than that of Calotes jerdoni (77–78). Calotes medogensis can be differentiated from Calotes maria by the presence of dark brown fold on shoulder (vs. absent) (Smith 1935), black stripes on orbit originating from eyes present (vs. absent) (Günther 1870).</p> <p>Natural history: We encountered the individuals on bushes, commonly observed roosting on shrubs and ferns along roadside and forest edges at night. We observed individuals changing body colour from green to brown (Figure 9B).</p> <p>Distribution: Calotes medogensis was known only from Yarang village, Beibeng Township of Medog County, China (Che et al. 2020). We recorded the species in Poba reserve forest, Jengging, Tuting, Yubuk, Ramsing in Arunachal Pradesh. Earlier records of Calotes jerdoni from Komsing, Yembung, Balek (Annandale 1912), Nameri National Park, Pake Tiger Reserve, Mouling National Park and Namdapha National Park (Pawar &amp; Birand 2001), Pasighat (Sanyal &amp; Gayen 2006), Zoological Survey of India campus, Itanagar (Sinha et al. 2021) could be referred to Calotes medogensis based on the proximity to the current records. Also, a record of Calotes jerdoni from Bhutan by Wangyal (2011) needs further verification.</p> </div>	https://treatment.plazi.org/id/1A0D16767638FF94FF080C71FD20F86D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Boruah, Bitupan;Narayanan, Surya;Deepak, V.;Das, Abhijit	Boruah, Bitupan, Narayanan, Surya, Deepak, V., Das, Abhijit (2022): Morphological and molecular differences in two closely related Calotes Cuvier 1817 (Squamata: Agamidae: Draconinae) with the first record of Calotes medogensis Zhao & Li, 1984 from India. Zootaxa 5219 (5): 433-455, DOI: https://doi.org/10.11646/zootaxa.5219.5.3
1A0D16767631FF96FF080BCDFD5FFB75.text	1A0D16767631FF96FF080BCDFD5FFB75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calotes jerdoni Gunther 1870	<div><p>Calotes jerdoni Günther, 1870</p> <p>Material examined: WII-ADR999, a male (Figure 5 A, B) collected from Malki forest (25°33ˊ24.4˝ N, 91°53ˊ33.9˝ E, Elevation: 1589 m asl.), Shillong, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.99358&amp;materialsCitation.latitude=25.64546" title="Search Plazi for locations around (long 93.99358/lat 25.64546)">East Khasi Hills</a>, Meghalaya, on 31 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.99358&amp;materialsCitation.latitude=25.64546" title="Search Plazi for locations around (long 93.99358/lat 25.64546)">July</a>, 2016 (Figure 1); WII-ADR966, a female (Figure 5 C, D) collected from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.99358&amp;materialsCitation.latitude=25.64546" title="Search Plazi for locations around (long 93.99358/lat 25.64546)">Dzuleke</a> (25°38ˊ43.66˝ N, 93°59ˊ36.92˝ E, Elevation: 1987 m asl.), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.99358&amp;materialsCitation.latitude=25.64546" title="Search Plazi for locations around (long 93.99358/lat 25.64546)">Kohima district</a>, Nagaland, on 8 <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=93.99358&amp;materialsCitation.latitude=25.64546" title="Search Plazi for locations around (long 93.99358/lat 25.64546)">August</a>, 2021 (Figure 1). Type series as mentioned in the Appendix 2.</p> <p>As Calotes jerdoni is a poorly known species, designating a lectotype for the species would provide taxonomic stability. Therefore, we designate the best preserved type specimen (BMNH 1946.8.11.54) among the syntypes as lectotype of C. jerdoni.</p> <p>Diagnosis: A moderate-sized agamid lizard, SVL 63.2 mm – 76.7 in male (n= 4) and SVL 74.1 mm – 102.2 mm in female (n= 4), TailL 196 mm – 240 mm in male (n= 4) and 204 mm – 300 mm in female (n= 4). Nuchal and dorsal crests continuous, consisting of erect compressed scales, smaller posteriorly. Two series of supratympanic spines present. Body laterally compressed, scales at the trunk heterogeneous in size, larger than ventrals, larger at the midbody towards dorsal side, oriented backward, 34–41 dorsal scales, 46–58 rows of scales around midbody, 68–78 ventrals. Mental larger than the chin shields, first pair of chin shields are of almost equal length and width; separated from each other by a small hexagonal scale, the gular scales at the mid-line are nearly equal to the adjacent gular scales away from the mid-line. Dorsally and ventrally green in colour.</p> <p>Description of the lectotype (BMNH 1946.8.11.54) (Figure 4): an adult male collected from Khasya (Khasi hills, Meghalaya), India by T. C. Jerdon.</p> <p>Specimen is in good condition. Moderate-sized lizard, SVL 76.6 mm, head distinct from neck, longer than width (HeadW/HeadL= 0.66), snout longer than orbital diameter (OrbD/SnEye= 0.71), loreal slightly concave, supraciliary and canthal ridge sharp, orbit three times larger than tympanum, supralabials 10 and 11 on right and left side respectively, infralabials 11 and 10 right and left side respectively. Body laterally compressed, 37 dorsal scales, 46 rows of scales around trunk at midbody, 71 ventrals, strongly keeled, a fold consisting of very small scales present in front of the shoulder. Tail rounded, TailL 213 mm, gradually tapering posteriorly, dorsal scales slightly smaller than that of ventral.</p> <p>Colouration in preservative: head dorsally and laterally light brown, a light bluish patch on left lateral side of the head below eye, gular region light brown with a light bluish patch on left side, trunk dorsally and laterally light bluish, belly whitish, tail dorsally and ventrally light brown, a white dorso-lateral line edged with black on trunk starting from neck to base of tail, a short white streak on elbow and knee.</p> <p>Morphological description of the newly collected materials: Morphometric and meristic characters are given in the Table 2. Moderate-sized lizard, SVL 63.2 mm (male) and 93 mm (female), head distinct from neck (Figure 5), longer than width (HeadW/HeadL= 0.64–0.68, n= 2), slightly concave in front, snout rounded in dorsal view, longer than orbital diameter (OrbD/SnEye= 0.72–0.79), loreal slightly concave, supraciliary and canthal ridge sharp, nostril elliptical, nasal large, orbit nearly three times larger than tympanum, pupil rounded, tympanum nearly rounded, rostral wider than mental, dorsal snout and head scales heterogenous in shape and size (Figure 6 G, J), scales at interorbital region, parietal region and occipital region with irregular ridge, scales at the loreal region, below and behind the orbit weakly keeled, heterogenous in shape and size, scales surrounding the eye are granular, feebly conical, scales surrounding tympanum are smaller than that of mandibular region and nape, supralabials nearly rectangular, elongated and smooth, a series of small elongated smooth scales above supralabials, rostral wide and narrow, interparietal scale with a distinct pit, two rows of elevated scales above tympanum, consisting of 8–9 scales, upper row starting from supraciliary to junction head and neck (Figure 6 I, L), the lower row just above the tympanum separated by two scales. Mental subtriangular, larger than the chin shields, infralabials smooth, narrow and elongated, first pair of chin shields are of almost equal length and width; separated by a small hexagonal scale, posterior chin shields are separated from infralabials by a series of narrow scales, size of the gular scales increases posteriorly, weakly keeled, scales between the chin shields are irregular in size and nearly of similar shape (Figure 7 C, D), subimbricate. Scales on neck are similar shape, posteriorly pointed, oriented backward, size increases dorsally and posteriorly, 9 nuchal spines, size increases posteriorly (Figure 6 I, L), length of the longest nuchal spine was less than twice the width (LongN/WidthN = 1.29–1.93, n= 8). Body laterally compressed, triangular in cross section. Dorsal crests reduced to a mere ridge on posterior body. Dorsal and lateral scales on trunk larger than ventrals, heterogenous in size, larger at the middle towards dorsal side, feebly keeled, pointed, more posteriorly oriented than dorsally, 39–41 dorsal scales, 54–58 rows of scales around trunk at midbody, ventral scales smaller than that of dorsal and lateral, strongly keeled; heterogenous in size, pointed, 77–78 ventrals, scales at the axilla, groin and insertion of hind limb are smaller, a fold consisting of very small scales present in front of the shoulder. Limbs moderate, thin, all scales keeled, smaller at the insertion, scales on ventral side of thigh are smaller than that of crus. Digits slender with strong and curved claw, lamellae entire, bicarinate. Tail rounded, gradually tapering posteriorly, dorsal scales slightly smaller than that of ventral, scales below cloaca smaller and size increases posteriorly, ventral scales strongly keeled, forming continuous ridge.</p> <p>Colouration in preservative: WII-ADR999 dorsally and ventrally uniform dark grey, tympanic shield and orbit whitish. WII-ADR966 dorsally anterior head to the level of interparietal grey; posterior part whitish, loreal region, behind the orbit to angle of jaw grey; tympanic region, mandibular region, below the level of nostril, both lips creamy white, dorsal, lateral and ventral of neck including nuchal spines creamy white to below axilla, trunk, tail and hindlimbs dark grey, forelimbs creamy white, ventrally head and chest whitish, rest of the belly, hind limbs and tail grey, forelimbs and crus ventrally creamy white. No light coloured patch on elbow, knee and heel visible.</p> <p>Colouration in life: Dorsal, lateral and ventral side of head green (Figure 10), dark stripes on orbit radiating from eye; sometimes slightly faint, tympanic shield light green. Trunk uniform green, on each side light brown lateral stripe starting from neck to tail may be present (Figure 10B). Tail dorsally green with light brown patches of irregular shape. White scales may be present on lateral side of trunk and forming discontinuous stripe on ventrolateral side. A dark brown black fold on shoulder. Ventrally pale uniform green. Light reddish-brown patch on elbow, knee and heel.</p> <p>Morphological comparison: Calotes jerdoni closely resembles with Calotes medogensis in external appearance. Morphological differences between these two species are mentioned above under Calotes jerdoni. It differs from Calotes maria by the presence of dark brown fold covered with small scales on the shoulder (vs. dark brown fold on shoulder absent), ridges above the tympanum not spinous (vs. spinous), nuchal spines less developed (vs. nuchal spines more pronounced) (Smith 1935).</p> <p>Natural history: We recorded the individuals of Calotes jerdoni on shrubs along road and forest edge. Frequently observed roosting on shrubs at night.</p> <p>Distribution: Calotes jerdoni is widely reported from northeast India to China. We recorded the species Malki forest, Shillong, Meghalaya and Dzuleke, Nagaland. Earlier reported from Khasi Hills, Meghalaya (Gunther 1870), Barail Wildlife Sanctuary (Das et al. 2009), Tinsukia and Dibrugarh district of Assam (Sengupta et al. 2019), Ukhrul in the state of Manipur, Kohima, Nagaland (Das et al. 2009), Chin Hills and Maymyo in Myanmar, Western Yunan (Teng-yueh), China (Smith 1935). Earlier reports from north of Brahmaputra River including Arunachal Pradesh and Bhutan could be referred to Calotes medogensis.</p> </div>	https://treatment.plazi.org/id/1A0D16767631FF96FF080BCDFD5FFB75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Boruah, Bitupan;Narayanan, Surya;Deepak, V.;Das, Abhijit	Boruah, Bitupan, Narayanan, Surya, Deepak, V., Das, Abhijit (2022): Morphological and molecular differences in two closely related Calotes Cuvier 1817 (Squamata: Agamidae: Draconinae) with the first record of Calotes medogensis Zhao & Li, 1984 from India. Zootaxa 5219 (5): 433-455, DOI: https://doi.org/10.11646/zootaxa.5219.5.3
1A0D1676762BFF8DFF080EB1FA2BF87C.text	1A0D1676762BFF8DFF080EB1FA2BF87C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calotes Cuvier 1817	<div><p>Identification keys to Calotes of northeast India:</p> <p>A. Postocular spine present</p> <p>Nuchal crests well developed.......................................................................................................................................................................... C. emma</p> <p>Nuchal crests weakly developed......................................................................... 2</p> <p>2A. 49–52 midbody scale rows.................................................................... C. zolaiking</p> <p>2.B. 42–46 midbody scale rows....................................................................... C. paulus</p> <p>B. Postocular spine absent</p> <p>A pale stripe present below eye from snout to neck................................................... C. geissleri</p> <p>Pale stripe absent below eye............................................................................. 3</p> <p>3A. Parallel rows of compressed scales above tympanum absent............................................ C. irawadi</p> <p>3B. Parallel rows of compressed scales above tympanum present.................................................... 4</p> <p>4A. Black fold on shoulder absent..................................................................... C. maria</p> <p>4B. Black fold on shoulder present........................................................................... 5</p> <p>5A. Scales on trunk obliquely and upwardly oriented................................................. C. medogensis</p> <p>5B. Scales on trunk backwardly oriented............................................................... C. jerdoni</p></div> 	https://treatment.plazi.org/id/1A0D1676762BFF8DFF080EB1FA2BF87C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Boruah, Bitupan;Narayanan, Surya;Deepak, V.;Das, Abhijit	Boruah, Bitupan, Narayanan, Surya, Deepak, V., Das, Abhijit (2022): Morphological and molecular differences in two closely related Calotes Cuvier 1817 (Squamata: Agamidae: Draconinae) with the first record of Calotes medogensis Zhao & Li, 1984 from India. Zootaxa 5219 (5): 433-455, DOI: https://doi.org/10.11646/zootaxa.5219.5.3
