taxonID	type	description	language	source
03FD8787FFB4FFB9FF34FB62FA05F820.taxon	diagnosis	Diagnosis. Fortipalpa Kasparyan & Ruíz-Cancino can be differentiated from all other genera of Neotropical Cryptinae by the following combination of features: ventral tooth slightly longer than dorsal tooth; propodeum and mesoscutum smooth, polished and almost impunctate; transverse furrow anterior to propodeum very shallow, without striae; propodeum posteriorly to anterior transverse carina with uniform, closely spaced transverse striae; T 1 long and slender, T 1 LW 2.08 – 3.41; ovipositor moderately long, OST 1.25 – 1.70, straight, tip moderately compressed, apex of ventral valve not forming a lobe covering dorsal valve. Female. Fore wing 5.75 – 10.60 mm long. Body slender and shiny, somewhat subcylindrical. HEAD. Mandible moderately long, MLW 1.44 – 1.71, apex distinctly narrower than base, MWW 0.46 – 0.51; ventral tooth slightly longer than dorsal tooth, rarely teeth about the same length. Malar space narrow, MSM 0.25 – 0.39. Clypeus of moderate size, CHW 0.47 – 0.57, weakly convex, apex with two very small tubercles at center or without tubercles; apical margin translucent and laterally projected. Clypeal fovea almost indistinct. Antenna with 21 – 24 flagellomeres; apical flagellomeres gradually thicker than basal ones, not flattened; apical flagellomere with distinct cluster of short, stiff setae. Supra-antennal area ventrally distinctly concave, with weak to distinct median line. Occipital carina sharp, complete, meeting hypostomal carina far from mandible base; hypostomal carina distinct, after meeting occipital carina ventrally raised, forming a flange. Temple and gena in lateral view moderately wide. THORAX. Dorsal margin of pronotum regular, not swollen; epomia strong, short or moderately long, ending far from or near dorsal margin of pronotum. Mesoscutum subcircular 0.88 – 1.04 × as long as wide, polished smooth; notaulus moderately impressed, distinctly convergent, reaching 0.30 – 0.54 of mesoscutum length. Epicnemial carina reaching 0.60 – 0.80 of distance to subalar, almost straight to distinctly curved, ending at or above pronotal spiracle. Sternaulus moderately strong, complete but usually wide and shallow at posterior 0.25, weakly to strongly sinuous, distinctly carinate throughout. Median portion of postpectal carina present, but short. Posterior margin of metanotum with weak projections. Transverse furrow anterior to propodeum moderately deep, usually polished smooth. Juxtacoxal carina absent. Pleural carina absent. Fore tibia moderately swollen. On all legs, fourth tarsomere apically bilobed, with strong apical group of bristles. PROPODEUM. Anterior margin medially concave or slightly concave, laterally with teeth-like projections; anterior area outline regular, not raised, in lateral view at the same level as the rest of propodeum. Propodeal spiracle elliptic or elongate, SLW 1.74 – 2.24. Anterior transverse carina complete, distinct, medially straight to slightly arched forwards. Area behind anterior transverse carina with transverse striation. Posterior transverse carina absent. Median longitudinal carina absent. Lateral longitudinal carina absent. WINGS. Hyaline. Fore wing vein 1 - Rs + M with bulla placed near basal 0.2 – 0.3; ramellus absent; crossvein 1 m-cu basally bent, otherwise straight or more or less uniformly curved; limit between 1 m-cu and 1 - Rs + M more or less traceable or 1 m-cu fully continuous with 1 - Rs + M; crossvein 1 cu-a arising opposite of 1 M + Rs; vein 2 Cua 0.64 – 0.86 × as long as crossvein 2 cu-a; crossvein 2 m-cu with bulla placed about to the middle; areolet small, APH 0.47 – 0.86, pentagonal or subpentagonal, usually distinctly wider than high, AWH 0.53 – 0.96; crossvein 3 r-m spectral or nebulous; cross veins 2 r-m and 3 r-m parallel or weakly convergent, both veins usually about the same length, sometimes 2 r-m slightly shorter than 3 r-m; vein 3 - M usually distinctly shorter than 2 - M; vein 4 - M usually tubular. Hind wing vein M + Cu apically distinctly convex; vein Cua almost same length or distinctly longer than crossvein cu-a, HW 1 C 0.97 – 2.00; vein Cub straight to distinctly convex at apical half; vein 2 - 1 A ending at 0.13 – 0.27 of distance to wing margin. METASOMA. First metasomal tergite (T 1) rather slender, T 1 LW 2.08 – 3.41, apex distinctly wider than base, T 1 WW 1.78 – 2.22, without anterolateral tooth, very sparsely pilose; spiracle placed a little after the middle, SPR 0.52 – 0.57, weakly to strongly prominent; median dorsal carina absent; dorsolateral carina inconspicuous; ventrolateral carina absent; median depression of T 1 from very shallow to distinct. T 2 not particularly long, T 2 LW 0.91 – 1.16, distinctly triangular, T 2 WW 1.63 – 2.22, thyridium subcircular. T 2 – 8 sculpturing usually from T 2 distinctly coriarious to T 8 smooth or faintly coriarious. T 2 sparsely pilose, T 3 – 8 densely pilose; T 7 – 8 in lateral view lengthened. Ovipositor long, OST 1.25 – 1.70, overall shape straight or rarely slightly upcurved, compressed, polished smooth; dorsal valve with nodus weak but distinct, notch faint; ovipositor tip moderately pointed; ventral valve basad teeth with weak but distinct nodus, apex not projected dorsally as a lobe, its apex with 13 – 17 subvertical teeth. Male. Generally similar to the respective females. Morphological secondary sexual differences are usually more or less uniform within Cryptini, as noted by Santos & Aguiar (2013), and apply to the males of Fortipalpa as follows. General body size usually smaller than respective females. Antenna with significantly more flagellomeres than in female; white band of flagellum covering more articles; apical flagellomere apically pointed, without a distinct cluster of modified setae. Transverse sulcus at base of propodeum slightly wider than in female. Fore tibia regular, not inflated. Areolet distinctly larger than in female, APH 0.98 – 1.03. Metasoma and T 2 – 7 generally slenderer than in females (e. g. Figs 2 A vs. 3 C).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFB4FFB9FF34FB62FA05F820.taxon	discussion	Remarks. Specimens of Fortipalpa are superficially similar to and can be confused with the large New World genus Messatoporus Cushman (revised by Santos & Aguiar, 2013) with which they share the generally slender and polished body, long and slender T 1 and propodeum posteriorly with transverse striae. Fortipalpa can be readily differentiated by having the ventral tooth of mandible slightly longer than dorsal tooth (vs. much shorter than dorsal tooth, sometimes so short as to be indistinct); occipital carina always meeting hypostomal carina (vs. almost always fading out before meeting hypostomal carina) and ovipositor tip compressed (vs. depressed). Fortipalpa is also similar to Prosthoporus Porter, from which it can be readily differentiated by having the hind wing vein 2 - 1 A distinct (vs. absent in Prosthoporus); fore wing vein 2 Cua at least 0.60 × as long as crossvein 2 cu-a (vs. less than 0.30); and transverse furrow anterior to propodeum distinctly shorter than area anterior to anterior transverse carina (vs. about as long as area anterior to anterior transverse carina).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFB4FFB9FF34FB62FA05F820.taxon	biology_ecology	Biology. Hosts are unknown. Species of Fortipalpa seem to be relatively rare; over 40,000 specimens of Neotropical Cryptinae have been examined by the authors but only the 34 specimens reported herein were found. Most specimens have been collected by Malaise traps even though Moericke (yellow pan) traps have been extensively used in several localities as well (e. g., Aguiar & Santos 2010; Supeleto et al. 2020).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFB4FFB9FF34FB62FA05F820.taxon	distribution	Distribution. Widely distributed in the Neotropics from Mexico to Southern Brazil. Current species records are from Mexico, Costa Rica, Panama, Trinidad, Ecuador, and Brazil (States: BA, ES, MG, MS, MT, PA, PR, RJ, SP) (Fig. 19).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFB2FFB2FF34FC7DFC97FCC4.taxon	description	(Figs 1 – 3, 19 – 21) Description. Holotype ♀. Fore wing 8.20 (7.30 – 8.70) mm. Head (Figs 1, 2 A – B). MLW 1.58 (1.50 – 1.61). MWW 0.47 (0.46 – 0.51). MSM 0.29 (0.29 – 0.38). CHW 0.52 (0.47 – 0.54). Clypeus anterior margin centrally with two small tubercles. Antenna with 23 (23 – 24) flagellomeres. Supra-antennal area distinctly coriarious; median line distinct and straight. Thorax (Figs 1, 2 A). Epomia approximately straight. Mesoscutum length to width 0.89 (0.89 – 0.96) × as long as wide. Notaulus reaching about 0.54 (0.39 – 0.54) of mesoscutum length. Scuto-scutellar groove with distinct longitudinal wrinkles. Epicnemial carina reaching about 0.8 of distance to subalar ridge; ending at pronotal spiracle; almost straight. Sternaulus moderately strong, wide and shallow at posterior 0.25; weakly sinuous. Mesopleuron polished smooth, centro-anteriorly with transversal striation, ventral half punctate. Mesosternum medio-longitudinal suture with short transverse wrinkles. Transverse furrow anterior to propodeum entirely polished smooth. Metapleuron densely punctate; moderately pilose. Propodeum (Figs 1, 2 A – C). Area behind anterior transverse carina transversally strigate, generally weak, antero-laterally faint. Spiracle elongate, SLW 2.22 (1.81 – 2.24). Anterior transverse carina medially straight. Wings. Fore wing crossvein 1 m-cu more or less uniformly curved; limit between 1 m-cu and 1 - Rs + M more or less traceable. Vein 2 Cua 0.83 (0.72 – 0.86) × as long as crossvein 2 cu-a. APH 0.64 (0.60 – 0.70). Areolet pentagonal; about as high as wide. AWH 0.90 (0.68 – 0.90). AAW 1.04 (1.04 – 1.48). Crossvein 3 r-m spectral. Crossvein 2 r-m and 3 r-m about same length. Vein 3 - M about same length of 2 - M. HW 1 C 1.68 (1.17 – 1.68). Hind wing vein Cub almost straight, very slightly convex. Vein 2 - 1 A ending at 0.17 (0.17 – 0.26) of distance to wing margin. Metasoma (Figs 1, 2 A & D). T 1 LW 2.56 (2.08 – 2.57). T 1 WW 1.90 (1.81 – 2.09). T 1 spiracle at 0.54 (0.54 – 0.56) of its length. Median depression of T 1 drop-shaped, distinct, large. T 2 LW 0.95 (0.91 – 1.11). T 2 WW 1.88 (1.64 – 2.22). T 2 – 8 faintly coriarious. Ovipositor straight, OST 1.54 (1.43 – 1.54); ventral valve apex with 16 (14 – 16) teeth (Fig. 2 D). Color. Head pale yellow (233,221,143) with supra-antennal area and vertex black; supraclypeal area latero ventrally with dark brown spot. Orbital band complete. Mandible pale yellow, apical 0.25 dark brown. Antenna dark brown (61,51,38), white band (243,239,227) composed by fl 5 apical 0.40, fl 6 – 13, fl 13 dorsally. Pronotum black (36,31,21), dorsally and collar pale yellow. Propleuron pale yellow. Mesoscutum black, median lobe centrally with pale yellow spot. Tegula and subalar ridge pale yellow. Epicnemium dorsal 0.85 black, ventrally light orange (244,168,62). Mesopleuron light orange, dorso anteriorly with large black area, mid anteriorly with pale yellow spot; hypoepimeron and mesepimeron pale yellow. Metapleuron bright orange (192,117,36). Dorsal division of metapleuron, scutellum and postscutellum pale yellow. Propodeum dark orange (251,150,9), posteriorly with large pale yellow area. Fore leg: coxa pale yellow, basally with tiny brownish (152,109,82) spot; trochanter pale yellow; trochantellus and femur light orange; tibia and t 1 – 3 pale yellow (232,218,115); t 4 dark brown (71,51,36); t 5 dark brown (61,51,38). Mid leg: coxa, trochanter, trochantellus and femur light orange; tibia yellow (244,206,37); t 1 yellow (244,206,37); t 2 brown (124,80,54); t 3 dark brown (71,51,36); t 4 – 5 dark brown (61,51,38). Hind leg: coxa, trochanter, trochantellus and femur dark orange; tibia yellow (244,206,37); t 1 whitish, basal 0.20 yellow (244,206,37); t 2 – 4 whitish; t 5 whitish, apical 0.35 dark brown. Metasoma striped black and yellow. T 1 tricolored, basal 0.25 yellow (225,187,85), centrally black (36,29,20), posterior 0.20 bright yellow (246,230,113). T 2 – 8 black with posterior 0.20 – 0.30 bright yellow. S 2 – 3 laterally bicolored, anterior 0.55 – 0.75 dark brown, posteriorly pale yellow (233,221,143). Morphometry. See Table 1, Figs 20 – 21, and Appendix. Variation. Fore wing vein 3 - M distinctly shorter than 2 - M; mesopleuron sometimes centro-anteriorly polished smooth, without transversal striation; anterior transverse carina medially slightly arched forwards. Orbital band complete (5 of 13 specimens) or incomplete (8 / 13), briefly interrupted near 2, 3, or 4 h, or interrupted between 2 – 4 h; malar space black; scape rarely ventrally with yellowish mark; propodeum rarely posteriorly without distinct pale yellow area; clypeal groove sometimes black marked; hind coxa rarely dorsally with weak pale yellow spot; epicnemium dorsal 0.40 - 0.95 black, or entirely black; T 1 sometimes basally orange. Specimen from Camamú (BA) has propodeum posteriorly with brown marks, and T 1 black, posteriorly with whitish stripe. Male (Fig. 3). Same overall structure and color pattern as female, but with the following differences. Antenna with 31 flagellomeres (vs. 22 – 24 in female). Clypeus apico-ventrally dark brown; propodeum posteriorly without pale yellow area; scutellar carina pale yellow; T 1 dorso-anteriorly bright orange; and orbital band almost complete, briefly interrupted between 09: 00 – 09: 30 h. Specimen from Sinop, Mato Grosso, with propleuron dark brown; and mesepisternum without mid anterior pale yellow spot.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFB2FFB2FF34FC7DFC97FCC4.taxon	discussion	Comments. Structure. Most similar to F. panamensis, from which it differs by having the following features: Epomia after diverging from pronotal collar approximately straight (vs. distinctly curved); scuto-scutellar groove with distinct longitudinal wrinkles (vs. with vestigial longitudinal wrinkles); area behind anterior transverse carina transversally strigate, generally weak, antero-laterally faint (vs. transversally strigate, arched posteriorwards); anterior transverse carina medially straight (vs. slightly arched forwards); epicnemial carina ending at pronotal spiracle (vs. ending above pronotoal spiracle); hind wing vein Cub almost straight, very slightly convex (vs. distinctly convex on posterior half, otherwise straight); T 2 – 8 faintly coriarious (vs. from distinctly coriarious T 2 to smooth T 8); median depression of T 1 drop-shaped, distinct, large (vs. elongate, very shallow). Also similar to F. sacha, but differs by having the following features: OST 1.43 – 1.54 (vs. 1.70); sternaulus weakly sinuous (vs. strongly sinuous); area behind anterior transverse carina transversally strigate, generally weak, antero-laterally faint (vs. transversally strigate, arched posteriorwards); fore wing crossvein 1 m-cu more or less uniformly curved (vs. basally bent, otherwise straight); crossvein 3 r-m spectral (vs. nebulous); T 2 – 8 faintly coriarious (vs. from distinctly coriarious T 2 to smooth T 8); median depression of T 1 drop-shaped, distinct, large (vs. elongate, very shallow). Also similar to F. yucatanica, but differs by having the following features: Scuto-scutellar groove with distinct longitudinal wrinkles (vs. with vestigial longitudinal wrinkles); area behind anterior transverse carina transversally strigate, generally weak, antero-laterally faint (vs. transversally strigate, arched posteriorwards); supra-antennal area with median line distinct and straight (vs. with slight median line); notaulus reaching about 0.39 – 0.54 of mesoscutum length (vs. 0.32); epicnemial carina reaching about 0.8 of distance to subalar ridge (vs. reaching about 0.7 of distance to subalar ridge); T 2 – 8 faintly coriarious (vs. from distinctly coriarious T 2 to smooth T 8); sternaulus weakly sinuous (vs. strongly sinuous; median depression of T 1 drop-shaped, distinct, large (vs. elongate, very shallow); anterior transverse carina medially straight (vs. slightly arched forwards). Color pattern. Closest to F. yucatanica, from which it differs by having scape ventrally dark brown (vs. whitish); mid coxa entirely light orange (vs. dorsally with whitish area); and hind coxa without or with faint whitish spot dorsally (vs. with distinct whitish spot dorsally). Also similar to F. panamensis, from which it differs by having epicnemium mostly or almost entirely black (vs. entirely light orange); dorsal margin of pronotum pale yellow (vs. black); and propodeum posteriorly with pale yellow mark (vs. without pale yellow mark).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFB2FFB2FF34FC7DFC97FCC4.taxon	etymology	Etymology. From the Greek exelysis, feminine, meaning outlet, mouth of a river. In reference to the location of the holotype.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFB2FFB2FF34FC7DFC97FCC4.taxon	distribution	Distribution records. Brazil (BA, ES, MG, MS, MT, PA, PR, RJ, SP) (Fig. 19).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFB2FFB2FF34FC7DFC97FCC4.taxon	materials_examined	Material examined. Total of 22 specimens, 20 ♀♀, 2 ♁♁. Holotype: BRAZIL, 1 ♀, Espírito Santo, Cariacica, Reserva Biológica de Duas Bocas, Sede, 185 m, 20 ° 15 ′ 53.6 ″ S, 40 ° 29 ′ 12.5 ″ W, 2017 - 1 / 2 - 20 / 5, A. P. Aguiar & F. A. Supeleto, Malaise, FAS 1021,?, UFES • Paratypes: BRAZIL, 1 ♀, Bahia, Camamú, Fazenda Nova Sorte,?, 24 m, 13 ° 57 ′ 84 ″ S, 39 ° 09 ′ 37 ″ W, 2002 - 11 - 19, J. Cardoso & J. Maia, Malaise, UFES 45710,?, UFES • 2 ib, Espírito Santo, Cariacica, Reserva Biológica de Duas Bocas, Pau Amarelo, 640 m, 20 ° 17 ′ 16 ″ S, 40 ° 31 ′ 09 ″ W, 2005 - 10 - 20 / 28, A. P. Aguiar exped., ib, LAPIS 5261, 2 ib • 9 ib, 2005 - 10 - 23 / 25, ib, YPT, LAPIS 5262, 2 ib • ib, 1 ♁, Mato Grosso, Sinop, 2?, 330 m, 12 ° 31 ′ S, 55 ° 37 ′ W, 1975 - 10, M. Alvarenga,?, FAS 4635,?, USUC • ib, 1 ♀, Minas Gerais,?, Parque Estadual do Rio Doce, Área da Tereza 3, 302 m, 19 ° 41 ′ 31 ″ S, 42 ° 32 ′ 04 ″ W, 2002 - 11 - 7, J. C. R. Fontenelle, Malaise, FAS 4622,?, UFES • 5 ib, Trilha do Vinhático 3, 3 ib, 2003 - 10 / 11 - 26 / 2, 2 ib, FAS 4624, 2 ib • 5 ib, Porto Capim 3, 3 ib, 2005 - 5 - 14 / 21, 2 ib, FAS 4625, 2 ib • 5 ib, Campolina 2, 3 ib, 2006 - 2 - 15 / 22, 2 ib, FAS 4623, 2 ib • 2 ib, Paraná, Fênix, Reserva Estadual do ITCF,?, 360 m, 23 ° 54 ′ 43 ″ S, 51 ° 57 ′ 35 ″ W, 1987 - 1 - 5, PROFAUPAR exped., ib, FAS 4633, ib, DZUP • ib, 1 ♁, Pará, Óbidos,?, ib, 115 m, 1 ° 51 ′ 05 ″ S, 55 ° 29 ′ 41 ″ W, 1955 - 7, J. Brasilino,?, FAS 4636,?, USUC • ib, 1 ♀, Rio de Janeiro, Guanabara, Reprêsa Rio Grande, ib, 470 m, 22 ° 45 ′ 28 ″ S, 43 ° 56 ′ 46 ″ W, 1969 - 10, M. Alvarenga, ib, FAS 4631, 2 ib • 9 ib, 1969 - 12, 2 ib, FAS 4632, 2 ib • 2 ib, S „ o Paulo, Magda,?, ib, 507 m, 20 ° 38 ′ 41.53 ″ S, 50 ° 13 ′ 30.07 ″ W, 2009 - 5 - 20,?, ib, 2?, UFES • 3 ib, Pindorama, 2 ib, 510 m, 21 ° 11 ′ 10 ″ S, 48 ° 54 ′ 23 ″ W, 2009 - 3 - 31, 5 ib • Other specimens: BRAZIL, 1 ♀, Mato Grosso, Cáceres, 2?, 120 m, 16 ˚ 05 ′ 20.96 ″ S, 57 ˚ 40 ′ 32.23 ″ W, 1984 - 11, M. Alvarenga, 3?, USUC • 2 ib, Mato Grosso do Sul, Aquidauana, Reserva Ecológica UEMS, Floresta Estacional Decidual, 220 m, 20 ˚ 25 ′ 59 ″ S, 55 ˚ 39 ′ 20.8 ″ W, 2011 - 12 - 26 / 2012 - 1 - 11, C. Lamas exped., Malaise, 2 ib, MZUP • 6 ib, 233 m, 20 ˚ 26 ′ 03.7 ″ S, ib, 2011 - 10 / 11 - 26 / 11, 5 ib • 3 ib, Porto Murtinho, Fazenda Retiro Conceição, Trilha da Mata Bruta, 89 m, 21 ˚ 41 ′ 42.9 ″ S, 57 ˚ 45 ′ 51.2 ″ W, 2012 - 1 / 1 - 10 / 25, 5 ib • ib, 3 ♀♀, 3 ib, Trilha da Espinhadeira, 99 m, 21 ˚ 40 ′ 59.7 ″ S, 57 ˚ 46 ′ 42.5 ″ W, 2012 - 1 / 3 - 25 / 29, 5 ib.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFBEFFB1FF34FC6EFABDFDC0.taxon	description	(Figs 4 – 5, 19 – 21) Description. Holotype ♀. Fore wing 7.95 (6.45 – 7.95) mm. Head (Figs 4, 5 A – B). MLW 1.45 (1.45 – 1.53). MWW 0.51 (0.51 – 0.51). MSM 0.34 (0.34 – 0.34). CHW 0.50 (0.50 – 0.57). Clypeus anterior margin centrally without teeth. Antenna with 23 (22 – 23) flagellomeres. Supra-antennal area distinctly coriarious; with slight median line. Thorax (Figs 4, 5 A). Epomia distinctly curved. Mesoscutum length to width 0.95 (0.93 – 0.95) × as long as wide. Notaulus reaching about 0.45 (0.45 – 0.49) of mesoscutum length. Scuto-scutellar groove with vestigial longitudinal wrinkles. Epicnemial carina reaching about 0.7 of distance to subalar ridge; ending at pronotal spiracle; almost straight. Sternaulus moderately strong, wide and shallow at posterior 0.25; strongly sinuous. Mesopleuron polished smooth, ventral half punctate. Mesosternum medio-longitudinal suture with short transverse wrinkles. Transverse furrow anterior to propodeum entirely polished smooth. Metapleuron densely punctate-rugulose; moderately pilose. Propodeum (Figs 4, 5 A & C). Area behind anterior transverse carina transversally strigate, arched posteriorwards. Spiracle elongate, SLW 2.00 (1.74 – 2.00). Anterior transverse carina medially slightly arched forwards. Wings (Fig. 4). Fore wing crossvein 1 m-cu more or less uniformly curved; fully continuous with 1 - Rs + M. Vein 2 Cua 0.68 (0.68 – 0.71) × as long as crossvein 2 cu-a. APH 0.69 (0.69 – 0.69). Areolet subpentagonal; distinctly wider than high. AWH 0.71 (0.71 – 0.75). AAW 1.34 (1.11 – 1.34). Crossvein 3 r-m spectral. Crossvein 2 r-m slightly shorter than 3 r-m. Vein 3 - M distinctly shorter than 2 - M. HW 1 C 1.70 (1.70 – 2.0). Hind wing vein Cub almost straight, very slightly convex. Vein 2 - 1 A ending at 0.20 (0.20 – 0.27) of distance to wing margin. Metasoma (Figs 4, 5 A & D). T 1 LW 2.54 (2.54 – 3.19). T 1 WW 2.05 (1.93 – 2.05). T 1 spiracle at 0.56 (0.54 – 0.56) of its length. Median depression of T 1 elongate, very shallow. T 2 LW 1.08 (1.01 – 1.08). T 2 WW 1.95 (1.95 – 2.20). T 2 – 8 from distinctly coriarious T 2 to smooth T 8. Ovipositor straight, OST 1.38 (1.25 – 1.38); ventral valve apex with 16 (14 – 16) teeth (Fig. 5 D). Color. Head whitish (241,228,184) with supra-antennal area and vertex black (36,31,21). Orbital band interrupted between 1 – 4 h. Mandible whitish (241,228,184), apical 0.10 dark brown. Antenna dark brown (61,51,38), scape ventrally with large whitish area, white band (249,242,225) composed by fl 6 apical 0.40, fl 7 – 13 whitish, fl 14 basal 0.80. Pronotum black, dorsally and collar whitish. Propleuron whitish. Mesoscutum black, median lobe centrally with whitish spot. Tegula and subalar ridge whitish. Epicnemium dark brown (71,51,36). Mesopleuron bright orange, dorso anteriorly with dark brown area, postero ventrally with pale yellow (233,215,149) spot; hypoepimeron whitish. Mesepimeron pale yellow. Metapleuron bright orange. Dorsal division of metapleuron orange (247,189,29). Scutellum and postscutellum whitish. Propodeum bright orange. Fore leg: coxa and trochanter whitish; trochantellus, femur, tibia, and t 1 buff (207,173,110); t 2 – 4 brown (118,76,50); t 5 dark brown. Mid leg: coxa, trochanter, trochantellus and femur bright orange; tibia buff; t 1 buff, apical 0.45 dark brown; t 2 – 5 dark brown. Hind leg: coxa bright orange; trochanter bright orange; trochantellus and femur bright orange (194,107,36); tibia yellow (221,174,85), apical 0.15 dark brown (61,51,38); t 1 whitish (249,242,225), basal 0.30 dark brown; t 2 – 4 whitish; t 5 bicolored, basal 0.40 whitish, apical 0.60 dark brown. Metasoma, T 1 – 8 and S 2 – 6 laterally bright orange (230,141,25). Morphometry. See Table 1, Figs 20 – 21, and Appendix. Variation. Fore wing vein 3 - M a little shorter than 2 - M. Male. Unknown.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFBEFFB1FF34FC6EFABDFDC0.taxon	discussion	Comments. Structure. Most similar to F. yucatanica, but differs by having the following features: Mesopleuron polished smooth, ventral half punctate (vs. polished smooth, centro-anteriorly with transversal striation, ventral half punctate); clypeus anterior margin centrally without teeth (vs. with two small tubercles); APH 0.69 (vs. 0.54); notaulus reaching about 0.45 – 0.49 of mesoscutum length (vs. 0.32); AWH 0.71 – 0.75 (vs. 0.84); epicnemial carina ending at pronotal spiracle (vs. ending above pronotoal spiracle); epomia after diverging from pronotal collar distinctly curved (vs. approximately straight); MSM 0.34 (vs. 0.28). Also similar to F. panamensis, from which it differs by having the following features: Clypeus anterior margin centrally without teeth (vs. with two small tubercles); mesopleuron polished smooth, ventral half punctate (vs. polished smooth, centro-anteriorly with transversal striation, ventral half punctate); epicnemial carina reaching about 0.7 of distance to subalar ridge (vs. reaching about 0.8 of distance to subalar ridge); OST 1.25 – 1.38 (vs. 1.58 – 1.59); fore wing crossvein 1 m-cu fully continuous with 1 - Rs + M (vs. limit between 1 m-cu and 1 - Rs + M more or less traceable); epicnemial carina ending at pronotal spiracle (vs. ending above pronotoal spiracle); HW 1 C 1.70 – 2.00 (vs. 1.30 – 1.49); hind wing vein Cub almost straight, very slightly convex (vs. distinctly convex on posterior half, otherwise straight; supra-antennal area with slight median line (vs. median line distinct and straight). Also similar to F. exelysae, but differs by having the following features: Clypeus anterior margin centrally without teeth (vs. with two small tubercles); mesopleuron polished smooth, ventral half punctate (vs. polished smooth, centro-anteriorly with transversal striation, ventral half punctate); area behind anterior transverse carina transversally strigate, arched posteriorwards (vs. transversally strigate, generally weak, antero-laterally faint); scuto-scutellar groove with vestigial longitudinal wrinkles (vs. with distinct longitudinal wrinkles; epicnemial carina reaching about 0.7 of distance to subalar ridge (vs. reaching about 0.8 of distance to subalar ridge); median depression of T 1 elongate, very shallow (vs. drop-shaped, distinct, large); supra-antennal area with slight median line (vs. median line distinct and straight). Color pattern. Closest to F. shakira, from which it differs by having dorsal margin of pronotum whitish (vs. black); orbital band interrupted between 1 – 4 h (vs. interrupted between 6 – 7 h); scape ventrally whitish (vs. dark brown); and clypeus entirely whitish (vs. apical e lateral margins blackish). Also similar to F. exelysae, from which it differs by having tergites bright orange (vs. black, posteriorly with whitish stripes); scape ventrally whitish (vs. dark brown); hind t 1 basally yellowish (vs. blackish); propodeum entirely bright orange (vs. posteriorly with pale yellow mark); and hind tibia apically blackish (vs. yellow).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFBEFFB1FF34FC6EFABDFDC0.taxon	etymology	Etymology. The word frida is a free feminine approximation for Alfredo, from Alfredo Chaves, the type locality.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFBEFFB1FF34FC6EFABDFDC0.taxon	distribution	Distribution records. Brazil (ES) (Fig. 19).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFBEFFB1FF34FC6EFABDFDC0.taxon	materials_examined	Material examined. Two specimens, 2 ♀♀. Holotype: BRAZIL, 1 ♀, Espírito Santo, Alfredo Chaves, Picadão, Mata, 714 m, 20 ° 27 ′ 53 ″ S, 40 ° 42 ′ 35 ″ W, 2007 - 10 - 8 / 15, C. O. Azevedo exped., Malaise, FAS 4621,?, UFES • Paratype: BRAZIL, 1 ♀, Espírito Santo, Santa Teresa, Reserva Biológica Augusto Ruschi, Trilha da Cachoeira, 775 m, 19 ° 55 ′ 16.4 ″ S, 40 ° 33 ′ 13.5 ″ W, 2017 - 1 / 2 - 16 / 1, A. P. Aguiar & F. A. Supeleto, Malaise, FAS 1022,?, UFES.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFBDFFB4FF34FD65FC15FE08.taxon	description	(Figs 6 – 7, 19 – 21) Description. Holotype ♀. Fore wing 6.90 mm. Head (Figs 6, 7 A – B). MLW 1.71. MWW 0.50. MSM 0.33. CHW 0.54. Clypeus anterior margin centrally with two small tubercles. Antenna with 22 flagellomeres. Supra-antennal area polished smooth; median line distinct and straight. Thorax (Figs 6, 7 A). Epomia approximately straight. Mesoscutum length to width 0.96 × as long as wide. Notaulus reaching about 0.51 of mesoscutum length. Scuto-scutellar groove with vestigial longitudinal wrinkles. Epicnemial carina reaching about 0.6 of distance to subalar ridge; ending at pronotal spiracle; distinctly curved. Sternaulus moderately strong, wide and shallow at posterior 0.25; weakly sinuous. Mesopleuron entirely polished smooth. Mesosternum medio-longitudinal suture without transverse wrinkles. Transverse furrow anterior to propodeum entirely polished smooth. Metapleuron densely punctate; glabrate. Propodeum (Figs 6, 7 A & C). Area behind anterior transverse carina transversally strigate, arched posteriorwards. Spiracle elliptic, SLW 1.88. Anterior transverse carina medially slightly arched forwards. Wings (Fig. 6). Fore wing crossvein 1 m-cu more or less uniformly curved; limit between 1 m-cu and 1 - Rs + M more or less traceable. Vein 2 Cua 0.64 × as long as crossvein 2 cu-a. APH 0.86. Areolet subpentagonal; distinctly wider than high, AWH 0.62. Crossvein 3 r-m spectral. Crossvein 2 r-m slightly shorter than 3 r-m. Vein 3 - M distinctly shorter than 2 - M. HW 1 C 1.07. Hind wing vein Cub straight. Vein 2 - 1 A ending at 0.18 of distance to wing margin. Metasoma (Figs 6, 7 A & D). T 1 LW 2.51. T 1 WW 2.07. T 1 spiracle at 0.56 of its length. Median depression of T 1 subcircular, distinct. T 2 LW 0.97. T 2 WW 2.20. T 2 – 8 faintly coriarious. Ovipositor straight, OST 1.53; ventral valve apex with 14 teeth (Fig. 7 D). Color. Head whitish (249,242,225) with supra-antennal area and vertex black (36,31,21). Clypeus mid ventrally and dupraclypeal area latero ventrally with brown spot. Malar space black. Orbital band almost complete, briefly interrupted near 6 h. Mandible basally whitish, apical 0.65 dark brown. Antenna dark brown (62,39,22) with whitish band composed by fl 5 apical 0.70 whitish, fl 6 – 13, and fl 13 dorsally. Pronotum black, dorso anterior spot and collar whitish. Propleuron whitish. Mesoscutum black, median lobe centrally with whitish spot. Tegula and subalar ridge whitish. Epicnemium dorsally dark brown, ventrally yellow (245,197,66). Mesopleuron yellow, mid anteriorly with whitish (241,229,195) area, postero ventrally with whitish spot, dorso anteriorly with large dark brown area; hypoepimeron whitish. Mesepimeron and metapleuron yellow. Dorsal division of metapleuron whitish. Scutellum and postscutellum whitish (249,242,225). Propodeum shinning brown, posteriorly with large pale yellow (233,221,143) area. Legs bright yellow (255,210,87). Fore leg: coxa yellow (245,197,66), basally with brown (124,77,49) area; trochanter, trochantellus, femur, tibia and t 1 bright yellow; t 2 bright yellow, apical 0.45 brown (124,77,49); t 3 – 5 dark brown. Mid leg: coxa yellow (245,197,66); trochanter, trochantellus, femur and tibia bright yellow; t 1 bright yellow, apical 0.45 brown (124,77,49); t 2 brown (124,77,49); t 3 – 5 dark brown. Hind leg: coxa yellow (245,197,66); trochanter, trochantellus, femur and tibia bright yellow; t 1 bright yellow, apical 0.75 whitish; t 2 – 4 whitish; t 5 whitish, apically dark brown. Metasoma striped black and whitish (241,229,195). T 1 basal 0.2 and apical 0.25 whitish, centrally dark brown (45,29,16). T 2 – 8 bicolored, dark brown with posterior 0.35 – 0.50 whitish. S 2 – 3 laterally bicolored, anterior 0.75 dark brown (62,39,22), posteriorly whitish; S 4 anterior 0.45 dark brown, posteriorly whitish. Morphometry. See Table 1, Figs 20 – 21, and Appendix. Male. Unknown.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFBDFFB4FF34FD65FC15FE08.taxon	discussion	Comments. Structure. Most similar to F. panamensis, but differs by having the following features: Mesopleuron entirely polished smooth (vs. polished smooth, centro-anteriorly with transversal striation, ventral half punctate); epicnemial carina reaching about 0.6 of distance to subalar ridge (vs. reaching about 0.8 of distance to subalar ridge); epomia short, ending far from dorsal margin of pronotum (vs. moderately long, ending near dorsal margin of pronotum); epicnemial carina distinctly curved (vs. almost straight); HW 1 C 1.07 (vs. 1.30 – 1.49); epomia after diverging from pronotal collar approximately straight (vs. distinctly curved); fore wing vein 2 Cua 0.64 × as long as crossvein 2 cu-a (vs. 0.71 – 0.83); APH 0.86 (vs. 0.47 – 0.76); epicnemial carina ending at pronotal spiracle (vs. ending above pronotoal spiracle); metapleuron glabrate (vs. moderately pilose); T 1 LW 2.51 (vs. 2.67 – 2.85); median depression of T 1 subcircular, distinct (vs. elongate, very shallow); T 2 – 8 faintly coriarious (vs. from distinctly coriarious T 2 to smooth T 8); hind wing vein Cub straight (vs. distinctly convex on posterior half, otherwise straight); AWH 0.62 (vs. 0.71 – 0.96). Also similar to F. frida, but differs by having the following features: clypeus anterior margin centrally with two small tubercles (vs. without teeth); epomia short, ending far from dorsal margin of pronotum (vs. moderately long, ending near dorsal margin of pronotum); epicnemial carina reaching about 0.6 of distance to subalar ridge (vs. reaching about 0.7 of distance to subalar ridge); MLW 1.71 (vs. 1.45 – 1.53); HW 1 C 1.07 (vs. 1.70 – 2.00); APH 0.86 (vs. 0.69); sternaulus weakly sinuous (vs. strongly sinuous); mesopleuron entirely polished smooth (vs. polished smooth, ventral half punctate); epomia after diverging from pronotal collar approximately straight (vs. distinctly curved); metapleuron glabrate (vs. moderately pilose); limit between fore wing crossvein 1 m-cu and 1 - Rs + M more or less traceable (vs. fully continuous with 1 - Rs + M); median depression of T 1 subcircular, distinct (vs. elongate, very shallow); T 2 – 8 faintly coriarious (vs. from distinctly coriarious T 2 to smooth T 8); hind wing vein Cub straight (vs. almost straight, very slightly convex); OST 1.53 (vs. 1.25 – 1.38); AWH 0.62 (vs. 0.71 – 0.75); supra-antennal median line distinct and straight (vs. with slight median line). Also similar to F. exelysae, from which it differs by having the following features: Epicnemial carina reaching about 0.6 of distance to subalar ridge (vs. reaching about 0.8 of distance to subalar ridge); APH 0.86 (vs. 0.60 – 0.70); area behind anterior transverse carina transversally strigate, arched posteriorwards (vs. transversally strigate, generally weak, antero-laterally faint). mesopleuron entirely polished smooth (vs. polished smooth, centro-anteriorly with transversal striation, ventral half punctate); median depression of T 1 subcircular, distinct (vs. drop-shaped, distinct, large); hind wing vein Cub straight (vs. almost straight, very slightly convex); scuto-scutellar groove with vestigial longitudinal wrinkles (vs. with distinct longitudinal wrinkles); metapleuron glabrate (vs. moderately pilose); supra-antennal area polished smooth (vs. distinctly coriarious). Color pattern. Closest to F. yucatanica, from which it differs by having hind coxa dorsally without whitish spot (vs. dorsally with distinct whitish spot); propodeum brown (vs. orange); clypeus mid ventrally with dark brown spot (vs. whitish); supraclypeal area latero ventrally with small brown spot (vs. whitish); scape ventrally dark brown (vs. whitish); and ovipositor yellowish (vs. brown). Also similar to F. exelysae, from which it differs by having ovipositor yellowish (vs. brown); propodeum brown (vs. orange); and dorsal margin of pronotum anteriorly with whitish spot (vs. entirely yellow). Etymology. The specific epithet is a literal reference to the municipality where the holotype was collected. Distribution records. Costa Rica (Fig. 19). Material examined. One specimen, 1 ♀. Holotype: COSTA RICA, 1 ♀, Heredia, Santa Rosa, 2?, 1.100 m, 9 ° 58 ′ 10 ″ N, 84 ° 05 ′ 54 ″ W, 1981 - 8, M. Fischel,?, FAS 4628,?, TAMU.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFB8FFABFF34FDA5FA1EFE98.taxon	description	(Figs 8 – 9, 19 – 21) Description. Holotype ♀. Fore wing 7.30 (5.75 – 10.06) mm. Head (Figs 8, 9 A – B). MLW 1.67 (1.55 – 1.67). MWW 0.50 (0.47 – 0.50). MSM 0.33 (0.33 – 0.39). CHW 0.50 (0.49 – 0.53). Clypeus anterior margin centrally with two small tubercles. Antenna with 22 (22 – 24) flagellomeres. Supra-antennal area distinctly coriarious; median line distinct and straight. Thorax (Figs 8, 9 A). Epomia distinctly curved. Mesoscutum length to width 0.94 (0.88 – 1.04) × as long as wide. Notaulus reaching about 0.47 (0.30 – 0.47) of mesoscutum length. Scuto-scutellar groove with vestigial longitudinal wrinkles. Epicnemial carina reaching about 0.8 of distance to subalar ridge; ending above pronotoal spiracle; almost straight. Sternaulus moderately strong, wide and shallow at posterior 0.25; weakly sinuous. Mesopleuron polished smooth, centro-anteriorly with transversal striation, ventral half punctate. Mesosternum medio-longitudinal suture with short transverse wrinkles. Transverse furrow anterior to propodeum entirely polished smooth. Metapleuron densely punctate-rugulose; moderately pilose. Propodeum (Figs 8, 9 A & C). Area behind anterior transverse carina transversally strigate, arched posteriorwards. Spiracle elliptic, SLW 1.67 (1.67 – 2.13). Anterior transverse carina medially slightly arched forwards. Wings (Fig. 8). Fore wing crossvein 1 m-cu more or less uniformly curved; limit between 1 m-cu and 1 - Rs + M more or less traceable. Vein 2 Cua 0.71 (0.71 – 0.83) × as long as crossvein 2 cu-a. APH 0.71 (0.47 – 0.76). Areolet subpentagonal; slightly wider than high. AWH 0.74 (0.71 – 0.96). AAW 1.23 (0.98 – 1.23). Crossvein 3 r-m spectral. Crossvein 2 r-m and 3 r-m about same length. Vein 3 - M distinctly shorter than 2 - M. HW 1 C 1.34 (1.30 – 1.49). Hind wing vein Cub distinctly convex on posterior half, otherwise straight. Vein 2 - 1 A ending at 0.20 (0.20 – 0.26) of distance to wing margin. Metasoma (Figs 8, 9 A & D). T 1 LW 2.85 (2.67 – 2.85). T 1 WW 2.00 (1.78 – 2.12). T 1 spiracle at 0.53 (0.53 – 0.55) of its length. Median depression of T 1 elongate, very shallow. T 2 LW 0.97 (0.97 – 1.13). T 2 WW 2.19 (1.83 – 2.19). T 2 – 8 from distinctly coriarious T 2 to smooth T 8. Ovipositor straight, OST 1.59 (1.58 – 1.59); ventral valve apex with 14 (14 – 15) teeth (Fig. 9 D). Color. Head whitish (241,228,184) with supra-antennal area and vertex black (36,31,21). Supraclypeal area lateroventrally brown (112,63,29). Clypeus mid ventrally with dark brown (62,39,22) spot. Malar space black. Orbital band almost complete, briefly interrupted near 6 h. Mandible basally whitish, apical 0.60 dark brown. Antenna dark brown (45,29,16) with whitish band (241,229,195), composed by fl 5 apical 0.55, fl 6 – 12, and fl 13 basal 0.40. Pronotum black, collar and dorso anterior spot whitish (241,229,195) spot. Propleuron whitish. Mesoscutum black, median lobe centrally with whitish spot. Tegula and subalar ridge whitish. Epicnemium orange (244,168,62). Mesopleuron orange, dorso anteriorly black; hypoepimeron whitish (241,228,184), posteriorly with some yellow. Mesepimeron light orange, dorsally whitish. Metapleuron bright orange (240,142,56). Dorsal division of metapleuron light orange with some yellow. Scutellum and postscutellum whitish (241,229,195). Propodeum bright orange. Fore leg light orange (231,182,68); t 3 apical 0.45 brown (115,75,50); t 4 – 5 dark brown. Mid leg light orange (231,182,68); t 1 apical 0.20 brown (115,75,50); t 2 brown (115,75,50); t 3 – 5 dark brown. Hind leg: coxa, trochanter, trochantellus and femur bright orange; tibia light orange, apically with brown (115,75,50) spot; t 1 whitish (241,230,204), basal 0.25 light orange; t 2 – 5 whitish. Metasoma striped, dark brown (45,29,16) and whitish (241,229,195). T 1 – 6 bicolored, dark brown with posterior 0.15 – 0.30 whitish; T 7 posterior 0.5 and T 8 posterior 0.15 whitish. Sternites whitish; S 2 anterior 0.6 dark brown (62,39,22), S 3 – 4 anteriorly with dark brown spot. Morphometry. See Table 1, Figs 20 – 21, and Appendix. Variation. Antenna with 24 flagellomeres; one specimen from Colombia with epomia almost indistinct on dorsal half. Pronotum without anterolateral whitish spot. Fore wing vein 3 - M distinctly longer than 2 - M; anterior transverse carina medially slightly arched forwards. Paratypes with a narrow yellow stripe on dorsal margin of pronotum; scuto-scutellar groove with distinct longitudinal wrinkles; metapleuron almost entirely punctate; ventral valve of ovipositor with 15 teeth. Male. Unknown.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFB8FFABFF34FDA5FA1EFE98.taxon	discussion	Comments. Structure. Most similar to F. yucatanica, but differs by having the following features: OST 1.58 – 1.59 (vs. 1.37); T 1 LW 2.67 – 2.85 (vs. 2.54); MSM 0.33 – 0.39 (vs. 0.28); HW 1 C 1.30 – 1.49 (vs. 1.76); sternaulus weakly sinuous (vs. strongly sinuous); epicnemial carina reaching about 0.8 of distance to subalar ridge (vs. reaching about 0.7 of distance to subalar ridge); MLW 1.55 – 1.67 (vs. 1.44). Also similar to F. frida, from which it differs by having the following features: Clypeus anterior margin centrally with two small tubercles (vs. without teeth); mesopleuron polished smooth, centro-anteriorly with transversal striation, ventral half punctate (vs. polished smooth, ventral half punctate); OST 1.58 – 1.59 (vs. 1.25 – 1.38); HW 1 C 1.30 – 1.49 (vs. 1.70 – 2.00); epicnemial carina ending above pronotoal spiracle (vs. ending at pronotal spiracle); limit between fore wing crossvein 1 m-cu and 1 - Rs + M more or less traceable (vs. fully continuous with 1 - Rs + M); sternaulus weakly sinuous (vs. strongly sinuous; epicnemial carina reaching about 0.8 of distance to subalar ridge (vs. reaching about 0.7 of distance to subalar ridge). Also similar to F. exelysae, but differs by having the following features: Area behind anterior transverse carina transversally strigate, arched posteriorwards (vs. transversally strigate, generally weak, antero-laterally faint); scutoscutellar groove with vestigial longitudinal wrinkles (vs. with distinct longitudinal wrinkles); median depression of T 1 elongate, very shallow (vs. drop-shaped, distinct, large); T 2 – 8 from distinctly coriarious T 2 to smooth T 8 (vs. faintly coriarious). Color pattern. Closest to F. shakira, from which it differs by having tergites black, posteriorly with whitish stripes (vs. bright orange); and antenna with white band composed by fl 5 apical 0.55, fl 6 – 12, and fl 13 basal 0.40 (vs. by f 5 apical 0.20, f 6 – 10 and most of fl 11). Also similar to F. exelysae, from which it differs by having epicnemium entirely light orange (vs. mostly or almost entirely black); dorsal margin of pronotum black (vs. pale yellow); and propodeum posteriorly without pale yellow mark (vs. with pale yellow mark). Also similar to F. yucatanica, from which it differs by having propodeum posteriorly without pale yellow mark (vs. with pale yellow mark); hind coxa dorsally without whitish spot (vs. with distinct whitish spot centrally); and scape ventrally dark brown, or with yellow spot (vs. whitish). Etymology. The specific epithet refers to the country where the holotype was collected.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFB8FFABFF34FDA5FA1EFE98.taxon	distribution	Distribution records. Colombia and Panamá (Fig. 19).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFB8FFABFF34FDA5FA1EFE98.taxon	materials_examined	Material examined. Four specimens, 4 ♀♀. Holotype: PANAMA, 1 ♀, Darién,?, P. N. Darién, Cerro Pirre - Est. Rancho Frio, 80 m, 7 ° 55 ′ 40 " N, 77 ° 42 ′ 06 " W, 2000 - 11 - 7 / 10, R. Cambra & A. Santos, YPT, FAS 4629,?, USUC • Paratype: PANAMA, 1 ♀, 2?, Panama Canal Zone, Cocoli, 90 m, 8 ° 58 ′ 58 " N, 79 ° 35 ′ 50 " W, 1946 - 10, N. L. H. Krauss,?, FAS 4630,?, USNM • Other specimens: COLOMBIA, 1 ♀, Valle del Cauca,?, PNN Falladones de Cali; Alto Anchicaya,?, 650 m, 3 ˚ 26 ′ N, 76 ˚ 48 ′ W, 2001 - 4 / 5 - 24 / 8, S. Sarria, Malaise,?, M 1900, IAVH • 13 ib, M 1527, ib.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA7FFAEFF34FE52FAC0FF04.taxon	description	(Figs 10 – 11, 19 – 21) Description. Holotype ♀. Fore wing 9.10 mm. Head (Figs 10, 11 A – B). MLW 1.62. MWW 0.46. MSM 0.33. CHW 0.52. Clypeus anterior margin centrally with two small tubercles. Antenna with 23 flagellomeres. Supra-antennal area distinctly coriarious; median line distinct and straight. Thorax (Figs 10, 11 A). Epomia approximately straight. Mesoscutum length to width 0.93 × as long as wide. Notaulus reaching about 0.39 of mesoscutum length. Scuto-scutellar groove with distinct longitudinal wrinkles. Epicnemial carina reaching about 0.8 of distance to subalar ridge; ending above pronotoal spiracle; almost straight. Sternaulus strong but narrow, posterior 0.3 nearly indistinct; weakly sinuous. Mesopleuron polished smooth, ventral half punctate. Mesosternum medio-longitudinal suture without transverse wrinkles. Transverse furrow anterior to propodeum centrally with very weak longitudinal wrinkles. Metapleuron densely punctate-rugulose; moderately pilose. Propodeum (Figs 10, 11 A & D). Area behind anterior transverse carina transversally strigate, arched posteriorwards. Spiracle elongate, SLW 2.06. Anterior transverse carina medially slightly arched forwards. Wings (Figs X, Y). Fore wing crossvein 1 m-cu more or less uniformly curved; fully continuous with 1 - Rs + M. Vein 2 Cua 0.76 × as long as crossvein 2 cu-a. APH 0.67. Areolet subpentagonal; distinctly wider than high, AWH 0.71. Crossvein 3 r-m spectral. Crossvein 2 r-m and 3 r-m about same length. Vein 3 - M distinctly shorter than 2 - M. HW 1 C 0.97. Hind wing vein Cub straight. Vein 2 - 1 A ending at 0.13 of distance to wing margin. Metasoma (Figs 10, 11 A & D). T 1 LW 2.46. T 1 WW 1.98. T 1 spiracle at 0.57 of its length. Median depression of T 1 elongate, distinct. T 2 LW 0.91. T 2 WW 1.92. T 2 – 8 from distinctly coriarious T 2 to smooth T 8. Ovipositor slightly upcurved, OST 1.46; ventral valve apex with 13 teeth (Fig. 11 D). Color. Head pale yellow (231,209,132) with supra-antennal area and vertex black (36,31,21). Clypeus mid ventrally and supraclypeal area latero ventrally with black spot. Malar space black. Orbital band narrow, almost complete, briefly interrupted near 6 h. Mandible basal 0.25 pale yellow (231,209,132), otherwise black. Antenna black with short whitish band (233,215,149), composed by fl 6 apical 0.30, fl 7 – 10, and tiny dorsal spot basally on fl 11. Pronotum black, collar light yellow (228,205,113). Propleuron light orange (222,132,54). Mesoscutum black, median lobe centrally with light yellow spot. Tegula and subalar ridge light yellow. Mesosoma otherwise mostly light orange. Scutellum light yellow, anteriorly black (37,25,8), posterior apex with narrow light orange stripe; yellow area somewhat bilobed. Postscutellum light orange. Fore leg: coxa, trochanter, trochantellus and femur light orange (218,149,59); tibia light yellow; t 1 – 2 pale yellow (221,194,124); t 3 pale yellow, apical 0.30 brownish (170,125,81); t 4 brown (124,80,54); t 5 dark brown (61,51,38). Mid leg: coxa, trochanter, trochantellus and femur light orange; tibia light yellow; t 1 – 2 pale yellow, t 2 apical 0.30 brown (124,80,54); t 3 – 5 dark brown. Hind leg: coxa light orange (222,132,54); trochanter, trochantellus and femur fulvous (176,105,60); tibia bright yellow (209,160,35), apical 0.10 dark brown; t 1 bicolored, basal 0.60 dark brown, apical 0.40 whitish (243,239,227); t 2 – 4 whitish; t 5 whitish, apically dark brown. Metasoma striped, dark brown (45,29,16) and yellow (231,209,132). T 1 basal half dark orange (188,86,43), subapically black (36,31,21), posterior 0.2 pale yellow (231,209,132). T 2 – 8 bicolored, dark brown with posterior 0.20 – 0.50 pale yellow. S 2 – 4 anteriorly brown, posteriorly pale yellow. Morphometry. See Table 1, Figs 20 – 21, and Appendix. Male. Unknown.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA7FFAEFF34FE52FAC0FF04.taxon	discussion	Comments. Structure. Most similar to F. panamensis, but differs by having the following features: Scutoscutellar groove with distinct longitudinal wrinkles (vs. with vestigial longitudinal wrinkles); sternaulus strong but narrow, posterior 0.3 nearly indistinct (vs. moderately strong, wide and shallow at posterior 0.25); T 1 LW 2.46 (vs. 2.67 – 2.85); HW 1 C 0.97 (vs. 1.30 – 1.49); vein 2 - 1 A ending at 0.13 (vs. 0.20 – 0.26) of distance to wing margin; epomia after diverging from pronotal collar approximately straight (vs. distinctly curved); mesopleuron polished smooth, ventral half punctate (vs. polished smooth, centro-anteriorly with transversal striation, ventral half punctate); OST 1.46 (vs. 1.58 – 1.59); hind wing vein Cub straight (vs. distinctly convex on posterior half, otherwise straight); ovipositor ventral valve apex with 13 teeth (vs. 14 – 15); ovipositor slightly upcurved (vs. straight). Also similar to F. frida, but differs by having the following features: Clypeus anterior margin centrally with two small tubercles (vs. without teeth); sternaulus weakly sinuous (vs. strongly sinuous); scuto-scutellar groove with distinct longitudinal wrinkles (vs. with vestigial longitudinal wrinkles; sternaulus strong but narrow, posterior 0.3 nearly indistinct (vs. moderately strong, wide and shallow at posterior 0.25); HW 1 C 0.97 (vs. 1.70 – 2.00); hind wing Vein 2 - 1 A ending at 0.13 (vs. 0.20 – 0.27) of distance to wing margin; MWW 0.46 (vs. 0.51); T 2 LW 0.91 (vs. 1.01 – 1.08); notaulus reaching about 0.39 of mesoscutum length (vs. 0.45 – 0.49); mesosternum medio-longitudinal suture without transverse wrinkles (vs. with short transverse wrinkles); epicnemial carina reaching about 0.8 of distance to subalar ridge (vs. reaching about 0.7 of distance to subalar ridge); fore wing 9.10 mm (vs. 6.45 – 7.95 mm); ovipositor slightly upcurved (vs. straight). Also similar to F. exelysae, from which it differs by having the following features: Area behind anterior transverse carina transversally strigate, arched posteriorwards (vs. transversally strigate, generally weak, anterolaterally faint); sternaulus strong but narrow, posterior 0.3 nearly indistinct (vs. moderately strong, wide and shallow at posterior 0.25); HW 1 C 0.97 (vs. 1.17 – 1.68); mesopleuron polished smooth, ventral half punctate (vs. polished smooth, centro-anteriorly with transversal striation, ventral half punctate); metapleuron densely punctate-rugulose (vs. densely punctate); T 2 – 8 from distinctly coriarious T 2 to smooth T 8 (vs. faintly coriarious); vein 2 - 1 A ending at 0.13 (vs. 0.17 – 0.26) of distance to wing margin; fore wing crossvein 1 m-cu fully continuous with 1 - Rs + M (vs. limit between 1 m-cu and 1 - Rs + M more or less traceable); hind wing vein Cub straight (vs. almost straight, very slightly convex); transverse furrow anterior to propodeum centrally with very weak longitudinal wrinkles (vs. entirely polished smooth); ovipositor slightly upcurved (vs. straight). Color pattern. Closest to F. sacha, from which it differs by having pronotum collar yellow (vs. black); propleuron light orange (vs. dark brown); mandible apical 0.65 blackish (vs. apical 0.30 dark brown); and hind t 5 whitish (vs. dark brown). Also similar to F. exelysae, from which it differs by having dorsal margin of pronotum black (vs. yellow); epicnemium light orange (vs. mostly or almost entirely black); propodeum posteriorly without pale yellow mark (vs. with pale yellow mark); mandible apical 0.65 blackish (vs. apical 0.25 dark brown); hind t 1 basally blackish (vs. yellowish); and scutellum antero centrally with black spot (vs. entirely pale yellow).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA7FFAEFF34FE52FAC0FF04.taxon	etymology	Etymology. The specific epithet is a literal reference to the municipality where the holotype was collected.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA7FFAEFF34FE52FAC0FF04.taxon	distribution	Distribution records. Ecuador (Fig. 19).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA7FFAEFF34FE52FAC0FF04.taxon	materials_examined	Material examined. One specimen, 1 ♀. Holotype: ECUADOR, 1 ♀, Pichincha, Mindo, 2?, 1400 m, 0 ° 03 ′ 04.29 ″ S, 78 ° 46 ′ 41.89 ″ W, 2005 - 10, M. Cooper,?, FAS 4627, Cooper coll. BNHH [E] 2005 - 152, NHM.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA2FFACFF34FEAEFF67F896.taxon	description	(Figs 12 – 13, 19 – 21) Description. Holotype ♀. Fore wing 8.65 mm. Head (Figs 12, 13 A – B). MLW 1.54. MWW 0.49. MSM 0.25. CHW 0.50. Clypeus anterior margin centrally with two small tubercles. Antenna with 22 flagellomeres. Supra-antennal area distinctly coriarious; median line distinct and straight. Thorax (Figs 12, 13 A). Epomia approximately straight. Mesoscutum length to width 0.97 × as long as wide. Notaulus reaching about 0.54 of mesoscutum length. Scuto-scutellar groove with distinct longitudinal wrinkles. Epicnemial carina reaching about 0.8 of distance to subalar ridge; ending above pronotoal spiracle; slightly curved. Sternaulus moderately strong, wide and shallow at posterior 0.25; strongly sinuous. Mesopleuron polished smooth, centro-anteriorly with transversal striation, ventral half punctate. Mesosternum medio-longitudinal suture with short transverse wrinkles. Transverse furrow anterior to propodeum entirely polished smooth. Metapleuron densely punctate-rugulose; moderately pilose. Propodeum (Figs 12, 13 A & C). Area behind anterior transverse carina transversally strigate, arched posteriorwards. Spiracle elongate, SLW 2.21. Anterior transverse carina medially slightly arched forwards. Wings (Fig. 12). Fore wing crossvein 1 m-cu basally bent, otherwise straight; limit between 1 m-cu and 1 - Rs + M more or less traceable. Vein 2 Cua 0.79 × as long as crossvein 2 cu-a. APH 0.56. Areolet subpentagonal; distinctly wider than high, AWH 0.79. Crossvein 3 r-m nebulous. Crossvein 2 r-m and 3 r-m about same length. Vein 3 - M distinctly shorter than 2 - M. HW 1 C 1.19. Hind wing vein Cub almost straight, very slightly convex. Vein 2 - 1 A ending at 0.20 of distance to wing margin. Metasoma (Figs 12, 13 A & D). T 1 LW 2.33. T 1 WW 2.22. T 1 spiracle at 0.54 of its length. Median depression of T 1 elongate, very shallow. T 2 LW 0.98. T 2 WW 1.91. T 2 – 8 from distinctly coriarious T 2 to smooth T 8. Ovipositor straight, OST 1.70; ventral valve apex with 15 teeth (Fig. 13 D). Color. Head pale yellow (233,215,149) with supra-antennal area and vertex black (36,31,21). Clypeus mid ventrally with dark brown spot. A moderately wide dark brown stripe extending from tentorial pit to supraclypeal area. Orbital band almost complete, briefly interrupted near 6 h. Mandible pale yellow, apical 0.30 dark brown. Antenna dark brown (61,51,38), whitish stripe (241,229,195) composed by fl 5 apical 0.60, fl 6 – 12, and fl 13 dorsal 0.50. Pronotum black. Propleuron dark brown. Mesoscutum black, median lobe centrally with whitish (233,222,178) spot. Tegula whitish and subalar ridge whitish. Epicnemium with dark brown and orange areas. Mesopleuron bright orange, becoming dark anteriorly; hypoepimeron, mesepimeron, metapleuron and dorsal division of metapleuron bright orange. Scutellum mostly covered by a bilobed whitish (233,222,178) area, basally black. Postscutellum whitish. Propodeum bright orange. Fore leg: coxa bright orange; trochanter, trochantellus and femur light orange (221,166,64); tibia and t 1 – 3 light yellow (228,205,113); t 4 – 5 dark brown. Mid leg: coxa bright orange; trochanter, trochantellus and femur bright orange (225,146,42); tibia light yellow; t 1 yellowish brown (170,125,81), its apex and t 2 – 5 brown. Hind leg: coxa, trochanter and trochantellus bright orange (202,115,42); femur reddish brown (162,75,16); tibia light orange, apical 0.25 dark brown; t 1 whitish (241,229,195), basal 0.35 dark brown; t 2 – 4 whitish; t 5 dark brown. Metasoma striped, dark brown and whitish (233,222,178). T 1 basally bright orange, centrally dark brown, posterior margin with whitish stripe (233,222,178). T 2 – 8 bicolored, dark brown with posterior 0.20 – 0.30 whitish. S 2 – 4 anteriorly dark brown, posteriorly whitish. Morphometry. See Table 1, Figs 20 – 21, and Appendix. Male. Unknown.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA2FFACFF34FEAEFF67F896.taxon	discussion	Comments. Structure. Most similar to F. yucatanica, but differs by having the following features: HW 1 C 1.19 (vs. 1.76); OST 1.70 (vs. 1.37); T 2 WW 1.91 (vs. 1.75); scuto-scutellar groove with distinct longitudinal wrinkles (vs. with vestigial longitudinal wrinkles); fore wing crossvein 1 m-cu basally bent, otherwise straight (vs. more or less uniformly curved); notaulus reaching about 0.54 of mesoscutum length (vs. 0.32); fore wing crossvein 3 r-m nebulous (vs. spectral); MLW 1.54 (vs. 1.44); T 1 WW 2.22 (vs. 1.92); T 1 LW 2.33 (vs. 2.54); SLW 2.21 (vs. 1.88). Also similar to F. panamensis, but differs by having the following features: Sternaulus strongly sinuous (vs. weakly sinuous); OST 1.70 (vs. 1.58 – 1.59); HW 1 C 1.19 (vs. 1.30 – 1.49); T 1 LW 2.33 (vs. 2.67 – 2.85); fore wing crossvein 1 m-cu basally bent, otherwise straight (vs. more or less uniformly curved); MSM 0.25 (vs. 0.33 – 0.39); scuto-scutellar groove with distinct longitudinal wrinkles (vs. with vestigial longitudinal wrinkles); notaulus reaching about 0.54 (vs. 0.30 – 0.47) of mesoscutum length; fore wing crossvein 3 r-m nebulous (vs. spectral); epomia short, ending far from dorsal margin of pronotum (vs. moderately long, ending near dorsal margin of pronotum). Also similar to F. exelysae, from which it differs by having the following features: OST 1.70 (vs. 1.43 – 1.54); sternaulus strongly sinuous (vs. weakly sinuous); MSM 0.25 (vs. 0.29 – 0.38); area behind anterior transverse carina transversally strigate, arched posteriorwards (vs. transversally strigate, generally weak, antero-laterally faint); fore wing crossvein 1 m-cu basally bent, otherwise straight (vs. more or less uniformly curved); median depression of T 1 elongate, very shallow (vs. drop-shaped, distinct, large); fore wing crossvein 3 r-m nebulous (vs. spectral); metapleuron densely punctate-rugulose (vs. densely punctate); T 2 – 8 from distinctly coriarious T 2 to smooth T 8 (vs. faintly coriarious). Color pattern. Closest to F. pichincha, from which it differs by having pronotum collar black (vs. yellow); propleuron dark brown (vs. light orange); and mandible apical 0.30 dark brown (vs. apical 0.65 blackish).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA2FFACFF34FEAEFF67F896.taxon	etymology	Etymology. The specific epithet is a literal reference to the ecological reserve Sacha Lodge, where the holotype was collected.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA2FFACFF34FEAEFF67F896.taxon	distribution	Distribution records. Ecuador.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA2FFACFF34FEAEFF67F896.taxon	materials_examined	Material examined. Two specimens, 2 ♀♀. Holotype: ECUADOR, 1 ♀, Sucumbios,?, Napo River, Sacha Lodge, 270 m, 0.5 ° S, 76.5 ° W, 1994 - 9 - 20 / 30, P. Hibbs, Malaise, FAS 4626,?, CNCI • Paratype: COLOMBIA, 1 ♀, Putumayo, 2?, PNN La Paya Cabaña La Paya, 330 m, 0 ˚ 2 ′ S, 75 ˚ 12 ′ W, 2001 - 12 / 12 - 5 / 25, E. Lozano, Malaise, 2?, IAVH.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA1FFA3FF34FF73FD99FB38.taxon	description	(Figs 14 – 15, 19 – 21) Description. Holotype ♀. Fore wing 9.31 mm. Head (Figs 14, 15 A – B). MLW 1.54. MWW 0.50. MSM 0.38. CHW 0.49. Clypeus anterior margin centrally with two small tubercles. Antenna with 23 flagellomeres. Supra-antennal area distinctly coriarious; with slight median line. Thorax (Figs 14, 15 A). Epomia distinctly curved. Mesoscutum length to width 1.04 × as long as wide. Notaulus reaching about 0.30 of mesoscutum length. Scuto-scutellar groove with distinct longitudinal wrinkles. Epicnemial carina reaching about 0.8 of distance to subalar ridge; ending above pronotoal spiracle; slightly curved. Sternaulus moderately strong, wide and shallow at posterior 0.25; strongly sinuous. Mesopleuron polished smooth, ventral half punctate. Mesosternum medio-longitudinal suture with short transverse wrinkles. Transverse furrow anterior to propodeum centrally with very weak longitudinal wrinkles. Metapleuron densely punctate; moderately pilose. Propodeum (Figs 14, 15 A & C). Area behind anterior transverse carina transversally strigate, arched posteriorwards. Spiracle elongate, SLW 2.22. Anterior transverse carina medially slightly arched forwards. Wings (Fig. 14). Fore wing crossvein 1 m-cu more or less uniformly curved; limit between 1 m-cu and 1 - Rs + M more or less traceable. Vein 2 Cua 0.74 × as long as crossvein 2 cu-a. APH 0.55. Areolet subpentagonal; distinctly wider than high, AWH 0.53. Crossvein 3 r-m nebulous. Crossvein 2 r-m slightly shorter than 3 r-m. Vein 3 - M distinctly shorter than 2 - M. HWIC 1.19. Hind wing vein Cub almost straight, very slightly convex. Vein 2 - 1 A not measured. Metasoma (Figs 14, 15 A & D). TILW 2.45. TIWW 2.18. T 1 spiracle at 0.52 of its length. Median depression of T 1 elongate, very shallow. TIILW 1.16. TIIWW 1.63. T 2 – 8 faintly coriarious. Ovipositor straight, OST 1.20; ventral valve apex with 17 teeth (Fig. 15 D). Color. Head anteriorly whitish (234,207,178) with supra-antennal area and vertex black (35,33,35). Orbital band interrupted between 6 – 7 h. Mandible whitish (229,203,168), apical 0.4 dark brown (75,66,69). Malar space, apical and lateral margins of clypeus, including clypeal fovea, black. Antenna dark brown, white band composed by f 5 apical 0.20, f 6 – 10 and most of fl 11. Pronotum dark brown, collar whitish. Propleuron yellowish (187,137,60). Mesoscutum black, median lobe centrally with whitish spot. Tegula and subalar ridge whitish. Epicnemium dark ferruginous with 0.25 blackish (71,51,36). Mesopleuron bright orange, dorso anteriorly with blackish area; hypoepimeron in gradient of whitish to orange. Mesepimeron, metapleuron and dorsal division of metapleuron orange (247,189,29). Scutellum whitish except for very apex black. Postscutellum blackish. Propodeum bright orange. Fore leg almost entirely orange, lighter at trochanter and tibia; t 1 – 2 brownish yellow; t 3 – 5 dark brown. Mid leg: orange, progressively lighter towards apex, tibia yellow; t 1 brownish yellow; t 2 – 5 dark brown. Hind leg: coxa, femur and trochanter orange; tibia mostly yellow (221,174,85), apical 0.20 partially brownish (124,69,18); t 1 whitish, basal 0.25 brownish; t 2 – 4 whitish; t 5 bicolored, basal 0.60 whitish, apical 0.40 dark brown. Metasoma, T 1 – 8 and S 2 – 6 laterally bright orange (230,141,25), T 1 with narrow posterior yellow stripe. Morphometry. See Table 1, Figs 20 – 21, and Appendix. Male. Unknown.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA1FFA3FF34FF73FD99FB38.taxon	discussion	Comments. Structure. Most similar to F. panamensis, from which it differs by having the following features: Transverse furrow anterior to propodeum centrally with very weak longitudinal wrinkles (vs. entirely polished smooth); T 2 – 8 faintly coriarious (vs. from distinctly coriarious T 2 to smooth T 8); mesopleuron polished smooth, ventral half punctate (vs. polished smooth, centro-anteriorly with transversal striation, ventral half punctate); AWH 0.53 (vs. 0.71 – 0.96); ovipositor ventral valve apex with 17 teeth (vs. 14 – 15); scuto-scutellar groove with distinct longitudinal wrinkles (vs. with vestigial longitudinal wrinkles); metapleuron densely punctate (vs. densely punctate-rugulose). Also similar to F. exelysae, but differs by having the following features: Notaulus reaching about 0.30 of mesoscutum length (vs. 0.39 – 0.54); mesopleuron polished smooth, ventral half punctate (vs. polished smooth, centroanteriorly with transversal striation, ventral half punctate); AWH 0.53 (vs. 0.68 – 0.90); area behind anterior transverse carina transversally strigate, arched posteriorwards (vs. transversally strigate, striae generally weak, antero-laterally faint); sternaulus strongly sinuous (vs. weakly sinuous); anterior transverse carina medially slightly arched forwards (vs. straight); median depression of T 1 elongate, very shallow (vs. drop-shaped, distinct, large); transverse furrow anterior to propodeum centrally with very weak longitudinal wrinkles (vs. entirely polished smooth); epomia after diverging from pronotal collar distinctly curved (vs. approximately straight). Color pattern. Closest to F. frida, from which it differs by having dorsal margin of pronotum black (vs. whitish); orbital band interrupted between 6 – 7 h (vs. interrupted between 1 – 4 h); scape ventrally dark brown (vs. whitish); and clypeus with apical and lateral margins blackish (vs. entirely whitish). Also similar to F. panamensis, from which it differs by having tergites bright orange (vs. black, posteriorly with whitish stripes); and antenna with white band composed by f 5 apical 0.20, f 6 – 10 and most of fl 11 (vs. by fl 5 apical 0.55, fl 6 – 12, and fl 13 basal 0.40).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA1FFA3FF34FF73FD99FB38.taxon	etymology	Etymology. Shakira is a feminine given name derived from a homonymous Arabic verb meaning “ to be thankful ”. It is used here both as a reverence to the species and as a deference to the homonymous singer, who is from the same country where the holotype was collected.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA1FFA3FF34FF73FD99FB38.taxon	distribution	Distribution records. Colombia (Fig. 19).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFA1FFA3FF34FF73FD99FB38.taxon	materials_examined	Material examined. One specimen, 1 ♀. Holotype: COLOMBIA, 1 ♀, Valle del Cauca,?, PNN Falladones de Cali; Cgto. La Mesata,?, 1960 m, 3 ˚ 34 ′ N, 76 ˚ 40 ′ W, 2003 - 8 / 9 - 27 / 10, S. Sarria, 2?, M 4549, IAVH. Holotype condition: left antenna after scape broken and glued to specimen label; right antenna broken and fragments glued to specimen label.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFAFFFA6FF34FAF2FC4FF804.taxon	description	(Figs 16 – 21)	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFAFFFA6FF34FAF2FC4FF804.taxon	description	Redescription. ♀. Fore wing 7.10 mm. Head (Figs 17 A – C, 18). MLW 1.44. MWW 0.50. MSM 0.28. CHW 0.47. Clypeus anterior margin centrally with two small tubercles. Antenna with 21 – 23 flagellomeres. Supra-antennal area polished smooth; with slight median line. Thorax (Figs 16, 17 C, 18 A). Epomia approximately straight. Mesoscutum length to width 0.95 × as long as wide. Notaulus reaching about 0.32 of mesoscutum length. Scuto-scutellar groove with vestigial longitudinal wrinkles. Epicnemial carina reaching about 0.7 of distance to subalar ridge; ending above pronotoal spiracle; slightly curved. Sternaulus moderately strong, wide and shallow at posterior 0.25; strongly sinuous. Mesopleuron polished smooth, centro-anteriorly with transversal striation, ventral half punctate. Mesosternum medio-longitudinal suture with short transverse wrinkles. Transverse furrow anterior to propodeum entirely polished smooth. Metapleuron densely punctate-rugulose; moderately pilose. Propodeum (Figs 16, 17 C, 18 A & C). Area behind anterior transverse carina transversally strigate, arched posteriorwards. Spiracle elliptic, SLW 1.88. Anterior transverse carina medially slightly arched forwards. Wings (Fig. 17 D). Fore wing crossvein 1 m-cu more or less uniformly curved; limit between 1 m-cu and 1 - Rs + M more or less traceable. Vein 2 Cua 0.74 × as long as crossvein 2 cu-a. APH 0.54. Areolet subpentagonal; distinctly wider than high, AWH 0.84. Crossvein 3 r-m spectral. Crossvein 2 r-m and 3 r-m about same length. Vein 3 - M distinctly shorter than 2 - M. HW 1 C 1.76. Hind wing vein Cub almost straight, very slightly convex. Vein 2 - 1 A ending at 0.23 of distance to wing margin. Metasoma (Figs 16, 17 C & E, 18 A & D). T 1 LW 2.54. T 1 WW 1.92. T 1 spiracle at 0.55 of its length. Median depression of T 1 elongate, very shallow. T 2 LW 1.02. T 2 WW 1.75. T 2 – 8 from distinctly coriarious T 2 to smooth T 8. Ovipositor straight, OST 1.37; ventral valve apex with 15 – 16 teeth (Fig. 18 D). Color. Head whitish (243,239,227) with supra-antennal area and vertex black (37,32,24). Orbital band complete. Mandible whitish (243,239,227), apical 0.25 dark brown (48,31,18). Antenna black (39,24,12); scape ventral 0.80 whitish; white band composed by fl 5 apical 0.40, fl 6 – 13, and fl 14 dorsally. Pronotum black, dorsally and collar whitish. Propleuron whitish. Mesoscutum black, median lobe centrally with whitish spot. Tegula, subalar ridge and epicnemium whitish. Mesopleuron light orange, anteriorly black; hypoepimeron, mesepimeron, dorsal division of metapleuron, scutellum and postscutellum whitish. Propodeum orange, apically whitish (244,232,205). Fore leg: coxa whitish (243,239,227), trochanter whitish (241,230,204), trochantellus, femur, tibia and t 1 – 3 light orange; t 4 – 5 dark brown (48,31,19). Mid leg light orange, coxa anteriorly with whitish area; t 1 pale yellow, apical 0.45 dark brown (87,68,55); t 2 dark brown; t 3 – 5 dark brown (48,31,19). Hind leg light orange, coxa dorsally with whitish spot; t 1 whitish (244,243,240), basal 0.25 pale yellow (237,219,153); t 2 – 4 whitish; t 5 whitish, apically dark brown. Metasoma striped, dark brown and whitish (243,239,227). T 1 tricolored, basal 0.35 pale orange, centrally dark brown, posterior 0.20 whitish. T 2 – 8 bicolored, dark brown with posterior 0.10 – 0.45 whitish. S 2 – 4 anteriorly dark brown, posteriorly whitish. Morphometry. See Table 1, Figs 20 – 21, and Appendix. Variation. Fore wing vein 3 - M distinctly shorter than 2 - M, almost indistinct. Specimen from Trinidad with epicnemium almost entirely blackish (vs. yellowish in holotype). Overall, the Trinidad specimen is more extensively orange, while the specimens from Mexico are yellowish. Male. Unknown.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFAFFFA6FF34FAF2FC4FF804.taxon	discussion	Comments. Structure. Most similar to F. panamensis, but differs by having the following features: Sternaulus strongly sinuous (vs. weakly sinuous); epicnemial carina reaching about 0.7 of distance to subalar ridge (vs. reaching about 0.8 of distance to subalar ridge); T 1 LW 2.54 (vs. 2.67 – 2.85); HW 1 C 1.76 (vs. 1.30 – 1.49); MSM 0.28 (vs. 0.33 – 0.39); MLW 1.44 (vs. 1.55 – 1.67); OST 1.37 (vs. 1.58 – 1.59). Also similar to F. frida, from which it differs by having the following features: Clypeus anterior margin centrally with two small tubercles (vs. without teeth); mesopleuron polished smooth, centro-anteriorly with transversal striation, ventral half punctate (vs. polished smooth, ventral half punctate); AWH 0.84 (vs. 0.71 – 0.75); APH 0.54 (vs. 0.69); MSM 0.28 (vs. 0.34); notaulus reaching about 0.32 of mesoscutum length (vs. 0.45 – 0.49). Also similar to F. exelysae, but differs by having the following features: Sternaulus strongly sinuous (vs. weakly sinuous); scuto-scutellar groove with vestigial longitudinal wrinkles (vs. with distinct longitudinal wrinkles); area behind anterior transverse carina transversally strigate, arched posteriorwards (vs. transversally strigate, generally weak, anterolaterally faint); notaulus reaching about 0.32 of mesoscutum length (vs. 0.39 – 0.54); T 2 – 8 from distinctly coriarious T 2 to smooth T 8 (vs. faintly coriarious); median depression of T 1 elongate, very shallow (vs. drop-shaped, distinct, large); epicnemial carina reaching about 0.7 of distance to subalar ridge (vs. reaching about 0.8 of distance to subalar ridge. Color pattern. Closest to F. exelysae from which it differs by having scape ventrally whitish (vs. dark brown); mid coxa dorsally with whitish area (vs. entirely light orange); and hind coxa with distinct whitish spot dorsally (vs. without or with a faint whitish spot dorsally). Also similar to F. panamensis, from which it differs by having propodeum posteriorly with pale yellow mark (vs. without pale yellow mark); hind coxa with distinct whitish spot dorsally (vs. without whitish spot); and scape ventrally whitish (vs. dark brown, or with yellow spot).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFAFFFA6FF34FAF2FC4FF804.taxon	distribution	Distribution records. Mexico, and Trinidad (New record) (Fig. 19).	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
03FD8787FFAFFFA6FF34FAF2FC4FF804.taxon	materials_examined	Material examined. One specimen, 1 ♀. TRINIDAD, 1 ♀, 2?, Simla nr. Arima,?, 250 m, 10 ° 41 ′ 33 ″ N, 61 ° 17 ′ 22 ″ W, 1977 - 12 - 3 / 10, W. R. M. Mason, Malaise, FAS 4634,?, CNCI.	en	Supeleto, Fernanda A., Santos, Bernardo F., Aguiar, Alexandre P. (2022): Revision of Fortipalpa Kasparyan & Ruíz-Cancino, (Ichneumonidae, Cryptinae). Zootaxa 5219 (6): 501-533, DOI: https://doi.org/10.11646/zootaxa.5219.6.1
