taxonID	type	description	language	source
6A3BBE243EC9550B9814DF9A24D5867C.taxon	materials_examined	Additional material examined. MADAGASCAR: Antsiranana Prov. [Diana Region]: 8 ♂, 4 ♀, 7 nymphs (CAS 9072436 [BLF 11400]), Reserve Analamerana, 12 ° 44 ′ 48 ″ S 49 ° 29 ′ 41 ″ E, 5 December 2004, B. L. Fisher; 1 ♂ (CAS 9008191 [BLF 2972]), Reserve Speciale de l'Ankarana, 12 ° 54 ′ 32 ″ S 49 ° 06 ′ 35 ″ E, 10 - 16 February 2001, Fisher, Griswold et al.; 1 ♀ (CAS 9071838 [BLF 2858]), Reserve Speciale de l'Ankarana; 12 ° 54 ′ 32 ″ S 49 ° 06 ′ 35 ″ E, 10 - 16 February 2001, Fisher, Griswold et al.; Antsiranana Prov. [Sava Region]: 1 nymph (CAS 9072439 [BLF 9656]), Foret de Binara, 13 ° 15 ′ 48 ″ S 49 ° 36 ′ 12 ″ E, 3 December 2003, B. L. Fisher.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6A3BBE243EC9550B9814DF9A24D5867C.taxon	diagnosis	Diagnosis. Smallest body sizes of all species of Antsirananaella gen. nov. (e. g., body length 1.80 / 2.17 ♂ / ♀ versus 1.94 - 2.17 / 2.21 - 2.47 ♂ / ♀; carapace length 0.55 / 0.65 ♂ / ♀ versus 0.58 - 0.69 / 0.70 - 0.77 ♂ / ♀). Unique cheotaxy: sb not on the same sagittal level as est but slightly more distal (on the same sagittal level in A. marlae sp. nov. and A. leniae sp. nov.); it between eb and et closer to eb than to et (equidistant in A. leniae sp. nov.); et in the distal third between dt and eb on the fixed finger (midway in between them in A. lorenzorum sp. nov. and A. leniae sp. nov.).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6A3BBE243EC9550B9814DF9A24D5867C.taxon	etymology	Etymology. This species is named after my former Latin teacher, Gerhard Faulstich, who might be one of the wisest men ML has ever known.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6A3BBE243EC9550B9814DF9A24D5867C.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Fig. 8 A): 1.36 - 1.55 (♂), 1.43 - 1.50 (♀) times longer than broad. - Pedipalps (Fig. 8 C): Trichobothrial pattern: sb on the movable finger distinctly distal to est on the fixed finger; it between eb and et on the fixed finger closer to eb than to et; et situated in the distal third between dt and eb on the fixed finger. Chelal fixed finger with 12 teeth in the OR, 16 in the MR and 15 in the IR; movable finger with 13 teeth in the OR, 13 in the MR and 16 in the IR. - Dimensions (mm): Holotype ♂: Body length 1.98; abdomen length 1.35, width (without pleura) 1.16, 1.25 (with pleura); carapace length 0.62, width 0.42; Pedipalp: trochanter 0.25; femur length 0.57, width 0.35; patella 0.46; chela (without pedicel) 0.57; hand length 0.13, width 0.15; movable finger 0.44; Leg I: trochanter 0.11, femur 0.20, patella 0.16, tibia 0.18, tarsus 0.27; Leg IV: trochanter 0.18, femur 0.12, patella 0.33, tibia 0.29, tarsus 0.32. Allotype ♀: Body length 2.28; abdomen length 1.53, width (without pleura) 1.34, 1.45 (with pleura); carapace length 0.73, width 0.51; Pedipalp: trochanter 0.32; femur length 0.71, width 0.38; patella 0.56; chela 0.65; hand length 0.14, width 0.18; movable finger 0.50; Leg I: trochanter 0.13; femur 0.25; patella 0.22; tibia 0.20; tarsus 0.33; Leg IV: trochanter 0.23; femur 0.14; patella 0.36; tibia 0.36; tarsus 0.36.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6A3BBE243EC9550B9814DF9A24D5867C.taxon	distribution	Distribution. Known from the type locality (BLF 3012) and four additional locations (BLF 11400, BLF 2858, BLF 2972 and BLF 9656) in the Diana and Sava Regions (formerly Antsiranana Province).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
4CB49A14AAEC5D9BB788A9B6131D22B4.taxon	materials_examined	Additional material examined. MADAGASCAR: Antsiranana Province [Sava Region]: 10 ♂, 8 ♀, 15 nymphs (CAS 9072435 [BLF 9556]), Foret de Binara, 13 ° 15 ′ 18 ″ S 49 ° 37 ′ 00 ″ E, 1 December 2003, B. L. Fisher; 1 ♂, 6 ♀, 8 nymphs (CAS 9072455 [BLF 9872]), Foret de Bekaraoka, 13 ° 10 ′ 00 ″ S 49 ° 42 ′ 36 ″ E, 7 December 2003, B. L. Fisher.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
4CB49A14AAEC5D9BB788A9B6131D22B4.taxon	diagnosis	Diagnosis. Very similar to A. lorenzorum sp. nov. but has the largest female body size of Antsirananaella gen. nov. (e. g., body length 2.47 (♀) versus 2.17 (♀) in A. faulstichi sp. nov., 2.21 (♀) in A. marlae sp. nov., 2.33 (♀) in A. lorenzorum sp. nov.; carapace length 0.77 (♀) versus 0.65 (♀) in A. faulstichi sp. nov., 0.70 (♀) in A. lorenzorum sp. nov., 0.71 (♀) in A. marlae sp. nov.); most slender chelal hand of females in Antsirananaella gen. nov. (0.85 (♀) x longer than wide versus 0.71 - 0.81 in other species of Antsirananaella gen. nov.). Difference in trichobothrial pattern: sb on the same sagittal level as est (more distal in A. lorenzorum sp. nov. and A. faulstichi sp. nov.); it between eb and et equidistant to both of them (closer to eb than to et in all other species of Antsirananaella gen. nov.); et midway between dt and eb on fixed finger (in the distal third between them in A. marlae sp. nov. and A. faulstichi sp. nov.).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
4CB49A14AAEC5D9BB788A9B6131D22B4.taxon	etymology	Etymology. This species is a patronym for DH's beloved daughter.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
4CB49A14AAEC5D9BB788A9B6131D22B4.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Fig. 9 A): 1.36 - 1.51 (♂), 1.33 - 1.45 (♀) times longer than broad. - Pedipalps (Fig. 9 C): Trichobothrial pattern: sb on the movable finger and est on the fixed finger on the same sagittal level; it between eb and et on the fixed finger with the same distance to eb and et. Chelal fixed finger with 12 teeth in the OR, 19 in the MR and 11 in the IR; movable finger with 11 teeth in the OR, 17 in the MR and 13 - 17 in the IR. - Dimensions (mm): Holotype ♂: Body length 2.21; abdomen length 1.54, width (with pleura) 1.34, 1.22 (without pleura); carapace length 0.68, width 0.45; Pedipalp: trochanter 0.27; femur length 0.63, width 0.34; patella 0.50; chela (without pedicel) 0.58; hand length (without pedicel) 0.11, width 0.15; movable finger 0.54; Leg I: trochanter 0.10; femur 0.27; patella 0.19; tibia 0.20; tarsus 0.29; Leg IV: trochanter 0.18; femur 0.13; patella 0.34; tibia 0.35; tarsus 0.38. Allotype ♀: Body length 2.46; abdomen length 1.71, width (without pleura) 1.51, (with pleura) 1.66; carapace length 0.72, width 0.54; Pedipalp: trochanter 0.36; femur length 0.79, width 0.44; patella 0.62; chela (without pedicel) 0.70; hand length 0.15, width 0.19; movable finger 0.54; Leg I: trochanter 0.14, femur 0.29, patella 0.17, tibia 0.20, tarsus 0.28; Leg IV: trochanter 0.23, femur 0.15, patella 0.41, tibia 0.48, tarsus 0.43.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
4CB49A14AAEC5D9BB788A9B6131D22B4.taxon	distribution	Distribution. Presently known from three localities: type locality (BLF 10116) and two additional localities (BLF 9556 and BLF 9872) in the Sava Region (formerly Antsiranana Province).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
01120C81DCDF5B9E9FA8E8F7D7AC73D8.taxon	diagnosis	Diagnosis. Antsirananaella gen. nov. differs from other Malagasy genera, Toliaranella gen. nov. and Mahajanganella gen. nov., by the following characters: the presence of five specialized setae on the retrolateral face of the movable chelal finger that are arranged in a row between trichobothrium b and the terminal chelal teeth (arranged in a group in Toliaranella gen. nov. and Mahajanganella gen. nov.); the presence of five terminal teeth (including one large tooth) on the fixed chelal finger and seven on the movable chelal finger (number of terminal teeth varying from 7 - 9 teeth in fixed and movable fingers in other Malagasy genera); anteromedial lobes of carapace closer to each other and slightly longer than anterolateral lobes (all four lobes equidistant in other Malagasy genera); and larger body size (1.80 - 2.17 / 2.17 - 2.47 ♂ / ♀ in Antsirananaella gen. nov. and ca. 1.56 - 1.89 / 1.96 - 2.33 ♂ / ♀ in all other genera). Antsirananaella gen. nov. differs from Cybella (Southeast Asia) by having platelets on the pleural membrane (absent in Cybella), from Iporangella (Brazil) by the presence of specialized setae on the movable chelal finger (absent in Iporangella), from Feaella (Difeaella) (South Africa) and Feaella (tropical Africa) by the presence of four anterior carapaceal lobes (two and six, respectively), from Feaella (Tetrafeaella) in continental Africa by having a less pronounced depression on the base of coxa I and on top of coxa II (distinctly more pronounced in Feaella (T.) cf. mucronata in Figs 25 - 27), and from the Australian species presently attributed to Feaella (Tetrafeaella) by having fewer coxal spines (one pair versus three or four in the Australian taxa).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
01120C81DCDF5B9E9FA8E8F7D7AC73D8.taxon	etymology	Etymology. This genus is named after the former Antsiranana Province, the area where the specimens were found. The gender is feminine.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
01120C81DCDF5B9E9FA8E8F7D7AC73D8.taxon	description	Description. The following description is based on examination of all species in Antsiranana. Typical feaellid habitus with a spherical abdomen, short and robust pedipalps, four prominent carapaceal lobes and dark reddish-brown body colour. - Carapace: (Figs 5 A, 7 A, 8 A, 9 A, 10 A): With four distinctly pointed anterior lobes; anteriolateral lobes slightly broader and wider than anteriomedial ones; anteriomedial lobes distinctly longer and closer to each other than anteriolateral ones; two pairs of eyes with lenses, equal in size, second pair partly covered by cuticula; four prominent posterior lobes (pm, pl in Figs 7 A, 8 A, 9 A, 10 A) and two longitudinal furrows more distally (af, pf in Figs 7 A, 8 A, 9 A, 10 A); with two medio-lateral mounds; and two postero-lateral processes. - Pedipalp: With a distinct conical protuberance on trochanter (Figs 4 A, B, 7 D, 8 D, 9 D, 10 D); femur broad and with one prolateral triangular process plus a retrolateral hump, patella cone-shaped. Chelal hand very small and with one large medial tooth at the base of each finger. Fixed finger with 9 trichobothria including dt, movable finger with 4 trichobothria (Figs 6 A, B, C, 7 C, 8 C, 9 C, 10 C). Movable finger with 5 specialized setae on ventral face, arranged in a transverse row between b and terminal teeth (Fig. 6 A). Chelal teeth large and retrorse, arranged in three rows on both chelal fingers. Terminal teeth situated in compact groups facing medial, four equally sized teeth and one larger tooth on fixed finger, seven equally sized teeth on movable finger (Figs 6 A, B, C, 7 C, 8 C, 9 C, 10 C). With 2 sensory setae (dt in Figs 6 B, 7 C, 8 C, 9 C, 10 C) on dorsal tip of fixed finger. - Chelicera: Palm with five long and several short setae; is and ls close; sbs proximal on fixed finger; movable finger with 1 subdistal seta (gs); es on palm close to base of movable finger; galea absent; spinneret conical; serrula exterior with 16 - 20 blades; no rallum could be found in any of the Malagasy species because of dirt; movable finger short and without teeth. - Coxal region (Fig. 5 B, C): Pedipalpal coxae with one spike laterally near base (Fig. 5 B); coxae I with a basal depression and one small coxal spine only (Fig. 5 B); coxae II with irregular shaped spines various in number framing depression of coxa I (Fig. 5 B); all coxae reaching towards midline and with coxa IV bigger in size than coxa I-III; cuticle within depression strongly granulate. - Legs (Figs 4 B, 7 B, 8 B, 9 B, 10 B): Trochanter I and II rather circular, trochanter III and IV rather elliptical; femur I and II slightly longer than patella I and II; femur III and IV shorter than patella III and IV; all tarsi long and slender, without specialized tactical trichobothria; subterminal tarsi with two curved and smooth claws; all setae acuminate; arolium much shorter than claws and with fimbriate distal margin; claws divided. - Abdomen (Fig. 4 A, B, C): Paired tergites and sternites medially divided; anal plate strongly sclerotized and with a circular raised rim in the center; pleural membrane with two rows of pleural platelets, 15 in the dorsal row, 14 in the ventral row. - Genital region (Figs 4 B, 5 B): Details like outer genital setae and inner structures not visible because of dirt on all specimens.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
01120C81DCDF5B9E9FA8E8F7D7AC73D8.taxon	distribution	Distribution. Antsirananaella gen. nov. is endemic to dry deciduous forests in Diana and Sava Regions (formerly Antsiranana Province) of northern Madagascar and separated by other Malagasy genera by more than 500 km of terrain. The climate is arid or " transition tropical " with temperatures ranging from 10.5 - 41.5 ° C. Annual precipitation ranges from 400 - 1000 mm.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
AA95E7B6BE5C5CBAA27706A6E9F84462.taxon	diagnosis	Diagnosis. Differs from all congeners by body length (1.94 / 2.33 ♂ / ♀ versus 1.80 / 2.17 ♂ / ♀ in A. leniae sp. nov., and 2.14 / 2.47 ♂ / ♀ in A. faulstichi sp. nov.); smallest ratio of pedipalpal femora (1.70 / 1.74 ♂ / ♀ versus 1.80 / 1.84 ♂ / ♀ in A. marlae sp. nov., and 1.75 / 1.79 ♂ / ♀ in A. faulstichi sp. nov.); pedipalpal chaetotaxy with sb not on same sagittal level as est but slightly more distal (on same sagittal level in A. marlae sp. nov. and A. leniae sp. nov.), it between eb and et closer to eb than to et (equidistant in A. leniae sp. nov.), et situated midway between dt and eb on fixed finger (situated in distal third between dt and eb in A. marlae sp. nov. and A. faulstichi sp. nov.).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
AA95E7B6BE5C5CBAA27706A6E9F84462.taxon	etymology	Etymology. This species is a patronym for the senior author's family that include parents Sylvia and Guenther Lorenz and brother Henrik.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
AA95E7B6BE5C5CBAA27706A6E9F84462.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Fig. 7 A): 1.38 (♂), 1.35 - 1.47 (♀) times longer than broad. - Pedipalp (Fig. 7 C): Trichobothrial pattern: esb and est in the proximal half of the retrolateral face; ib, isb and ist situated basally as a slightly curved line, isb and ib closer to each other than isb and ist; eb and it situated sub-distally and very close to each other, with it more distal and more medial than eb; et situated distally and approximately the same distance to dt than to eb; dt situated very distal in a plain pit; st situated sub-basally at the same sagittal level as esb; st and est also on the same level as ist; t distinctly closer to sb than to b; sb distinctly closer to t than to st. Chelal fixed finger with 10 teeth in the OR, 20 in the MR and 15 in the IR; movable finger with 11 teeth in the OR, 14 - 15 teeth in the MR, and 14 teeth in the IR. - Dimensions (mm): Holotype ♂: Body length 1.94; abdomen length 1.33; abdomen width 1.12 (without pleura), 1.24 (with pleura); carapace length 0.58; carapace width 0.42. Pedipalp: trochanter 0.22; femur length 0.54; femur width 0.32; patella 0.46; chela (without pedicel) 0.54; hand length (without pedicel) 0.12; width 0.14; movable finger length 0.42. Leg I: trochanter 0.11; femur 0.19; patella 0.17; tibia 0.17; tarsus 0.27. Leg IV: trochanter 0.17; femur 0.13; patella 0.28; tibia 0.29; tarsus 0.35. Allotype ♀: Same as male except body length 2.26; abdomen length 1.59, width 1.36 (without pleura), 1.49 (with pleura); carapace length 0.66, width 0.49. Pedipalp: trochanter 0.30, femur length 0.75, width 0.44, patella length 0.61, chela (without pedicel) 0.64, hand length (without pedicel) 0.14, width 0.18. Leg I: trochanter 0.14, femur 0.25, patella 0.22, tibia 0.20, tarsus 0.29. Leg IV: trochanter 0.23, femur 0.15, patella 0.36, tibia 0.42, tarsus 0.39.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
AA95E7B6BE5C5CBAA27706A6E9F84462.taxon	distribution	Distribution. Only known from the type locality (BLF 3182) in Diana Region (formerly eastern Antsiranana Province), northern Madagascar.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6041D8D13CB55BF3B4F8B9F4D52F6982.taxon	diagnosis	Diagnosis. Differs from congeners by large body size (e. g., male body length 2.17 versus 1.80 - 2.14, male carapace length 0.69 versus 0.55 - 0.66); a slender carapace (1.47 times longer than broad versus 1.36 - 1.40); medio-lateral mounds more pronounced than in A. lorenzorum sp. nov.; a slender pedipalpal femur (1.80 (♂), 1.84 (♀) times longer than broad versus 1.70 - 1.79 (♂), 1.74 - 1.80 (♀ )); pedipalpal trichobothrial pattern: sb on movable finger on same sagittal level as est on fixed finger (slightly more distal in A. lorenzorum sp. nov. and in A. faulstichi sp. nov.), et situated in the distal third between dt and eb on fixed finger (situated midway between dt and eb on fixed finger in A. lorenzorum sp. nov. and in A. leniae sp. nov.); it between eb and et situated closer to eb than to et (equidistant in A. leniae sp. nov.).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6041D8D13CB55BF3B4F8B9F4D52F6982.taxon	etymology	Etymology. The specific epithet is a patronym in honour of Marla Elisa Nibasumba, the senior author's godchild.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6041D8D13CB55BF3B4F8B9F4D52F6982.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Fig. 10 A): 1.43 - 1.49 (♂), 1.30 - 1.46 (♀) times longer than broad. - Pedipalps (Fig. 10 C): Trichobothrial pattern: sb on the movable finger and est on the fixed finger on the same sagittal level. et situated in the distal third between dt and eb on the fixed finger. Chelal fixed finger with 11 teeth in the OR, 21 teeth in the MR, and 12 teeth in the IR; movable finger with 11 teeth in the OR, 13 teeth in the MR, and 13 teeth in the IR. - Dimensions (mm): Holotype ♂: Body length 1.81; abdomen length 1.25; abdomen width 0.99 (without pleura), 1.11 (with pleura); carapace length 0.55; carapace width 0.37. Pedipalp: trochanter 0.24; femur length 0.54; femur width 0.29; patella 0.43; chela (without pedicel) 0.49; hand length (without pedicel) 0.11; width 0.13; movable finger length 0.40. Leg I: trochanter 0.10; femur 0.13; patella 0.16; tibia 0.14; tarsus 0.25. Leg IV: trochanter 0.16; femur 0.11; patella 0.29; tibia 0.28; tarsus 0.31. Allotype ♀: Body length 2.30; abdomen length 1.60; abdomen width 1.21 (without pleura), 1.44 (with pleura); carapace length 0.67; carapace width 0.46. Pedipalp: trochanter 0.31; femur length 0.68; femur width 0.39; patella length 0.54; chela (without pedicel) 0.60; hand length (without pedicel) 0.14; hand width 0.18; movable finger length 0.48. Leg I: trochanter 0.11; femur 0.24; patella 0.22; tibia 0.15; tarsus 0.25. Leg IV: trochanter 0.21; femur 0.14; patella 0.35; tibia 0.41; tarsus 0.39.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6041D8D13CB55BF3B4F8B9F4D52F6982.taxon	distribution	Distribution. Only known from the type locality (BLF 9974) in Sava Region (formerly eastern Antsiranana Province) of northern Madagascar.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
9E11F2A8D44356C696C65F799A9950A3.taxon	materials_examined	Additional material examined. MADAGASCAR: Province de Toliara [Menabe Region]: 2 ♂, 1 ♀, 4 nymphs (CAS 9071535 [BLF 4432]), Parc National de Tsingi de Bemaraha, 19 ° 42 ′ 34 ″ S 44 ° 43 ′ 5 ″ E, 16 - 20 November 2001, Fisher, Griswold et al.; 17 ♂, 7 ♀, 16 nymphs (CAS 9071781 [BLF 4726]), Toliara Province [Menabe Region]: Parc National de Kirindy Mite, 20 ° 47 ′ 43 ″ S 44 ° 08 ′ 49 ″ E, 6 - 10 December 2001, Fisher, Griswold et al.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
9E11F2A8D44356C696C65F799A9950A3.taxon	diagnosis	Diagnosis. Very similar to the type species but remarkably more granulated cuticle on the carapace, especially above the posterior pair of eyes; anterior lobes on the carapace are not pointed but rounded at the tip.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
9E11F2A8D44356C696C65F799A9950A3.taxon	etymology	Etymology. This species is named after the Mexican artist Frida Kahlo de Rivera († 1954) whose unmistakable character were her striking eyebrows, which she included in many of her self-portraits. The species is reminiscent of her because of the strongly granulated cuticle above the second pair of eyes, which resemble ' eyebrows'.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
9E11F2A8D44356C696C65F799A9950A3.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Fig. 15 A): Cuticle strongly covered with granulate especially above the second eye pair which makes it look like an additional lateral pair of mounds behind the anterior margin; anterior lobes on carapace not pointed but rounded at the tips; 1.26 - 1.42 (♂), 1.18 - 1.34 (♀) times longer than broad. - Pedipalps (Fig. 15 C, D): See trichobothrial pattern in genus description. Chelal fixed finger with 9 - 12 teeth in the OR, 14 - 15 in the MR and 8 - 12 in the IR; movable finger with 6 - 9 teeth in the OR, 9 - 11 in the MR and 9 - 10 in the IR; fixed finger with 6 equally sized and one larger terminal tooth, movable finger with 7 equally sized terminal teeth. - Dimensions (mm): Holotype ♂: Body length 1.71; abdomen length 1.15, width (with pleura) 1.01, width (without pleura) 0.90; carapace length 0.49, width 0.39. Pedipalp: trochanter 0.24; femur length 0.50, width 0.27; patella 0.34; chela (without pedicel) 0.41; hand length 0.08, width 0.12; movable finger 0.34. Leg I: trochanter 0.09; femur 0.14; patella 0.13; tibia 0.12; tarsus 0.21. Leg IV: trochanter 0.14; femur 0.10; patella 0.24; tibia 0.25; tarsus 0.23. Allotype ♀: Body length 2.22; abdomen length 1.58, width (with pleura) 1.35, width (without pleura) 1.14; carapace length 0.60, width 0.45. Pedipalp: trochanter 0.26; femur length 0.54, width 0.35; patella 0.44; chela (without pedicel) 0.49; hand length 0.10, width 0.14; movable finger 0.37. Leg I: trochanter 0.10; femur 0.17; patella 0.19; tibia 0.14; tarsus 0.22. Leg IV: trochanter 0.18; femur 0.13; patella 0.29; tibia 0.30; tarsus 0.31.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
9E11F2A8D44356C696C65F799A9950A3.taxon	distribution	Distribution. Known from the type locality (BLF 4605) and two additional localities (BLF 4432 and BLF 4726) in the Menabe Region (formerly Toliara Province) of western Madagascar.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
F670E3C29FCF5842B7FC6906D632BC55.taxon	materials_examined	Additional material examined. MADAGASCAR: Mahajanga Province [Boeny Region]: 23 ♂, 18 ♀, 29 nymphs (CAS 9008212 [BLF 3571]), Parc National d'Ankarafantsika, 16 ° 17 ′ 56 ″ S 46 ° 48 ′ 47 ″ E, 26 March- 1 April 2001, Fisher, Griswold et al.; 4 ♂, 2 ♀, 11 nymphs (CAS 9072477 [BLF 6506]), Parc National de Namoroka, 16 ° 22 ′ 36 ″ S 45 ° 19 ′ 36 ″ E, 8 - 12 November 2002, Fisher, Griswold et al.; 2 ♂, 2 nymphs (CAS 9071651 [BLF 6446]), Parc National de Namoroka, 16 ° 28 ′ 00 ″ S 45 ° 21 ′ 00 ″ E, 4 - 8 November 2002, Fisher, Griswold et al.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
F670E3C29FCF5842B7FC6906D632BC55.taxon	diagnosis	Diagnosis. Largest female body length of Mahajanganella gen. nov. (2.15 versus 2.03 - 2.08); robustest pedipalps of this new genus measured in ratios (e. g., femur 1.64 / 1.62 ♂ / ♀ times longer than broad versus 1.69 - 1.75 / 1.69 - 1.74 ♂ / ♀); e. g., femur 1.14 ♂ and ♀ times longer than tibia versus 1.27 / 1.21 - 1.27 / 1.27 ♂ / ♀; e. g., femur 1.02 / 1.06 ♂ / ♀ longer than chela versus 1.12 / 1.19 - 1.14 / 1.19 ♂ / ♀); anteriomedial lobes distinctly pointed and moderate carapace cuticle above second eyes (in other species of Mahajanganella gen. nov. distinctly more granulate carapace especially above second eyes).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
F670E3C29FCF5842B7FC6906D632BC55.taxon	etymology	Etymology. This species is named after the divine hero Herakles (lat. Hercules) who, in Greek mythology, is known for his power and courage. He is often portrayed with a mace, which resembles the strong pedipalps of this species. It is to be treated as a noun in apposition.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
F670E3C29FCF5842B7FC6906D632BC55.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Fig. 14 A): 1.28 - 1.49 (♂), 1.21 - 1.36 (♀) times longer than broad. - Pedipalp (Fig. 14 C, D): Femora very robust 1.50 - 1.68 (♂), 1.56 - 1.69 (♀) times longer than broad. See trichobothrial pattern in genus description. Chelal fixed finger with 10 teeth in the OR, 15 in the MR and 11 in the IR; movable finger with 9 teeth in the OR, 12 in the MR and 8 in the IR. Terminal teeth with 7 equally sized and 1 larger tooth on the fixed finger, and 8 equally sized teeth on the movable finger. - Dimensions (mm): Holotype ♂: Body length 1.74; abdomen length 1.17, width (without pleura) 0.95, (with pleura) 1.08; carapace length 0.50, width 0.38. Pedipalp: trochanter 0.25; femur length 0.47, width 0.28; patella 0.39; chela (without pedicel) 0.43; hand length 0.10, width 0.13; movable finger 0.35. Leg I: trochanter 0.08; femur 0.17; patella 0.17; tibia 0.12; tarsus 0.22. Leg IV: trochanter 0.14; femur 0.12; patella 0.23; tibia 0.21; tarsus 0.22. Allotype ♀: Body length 2.19; abdomen length 1.57, width (without pleura) 1.25, (with pleura) 1.41; carapace length 0.55, width 0.45. Pedipalp: trochanter 0.25; femur length 0.52, width 0.32; patella 0.47; chela (without pedicel) 0.48; hand length 0.10, width 0.15; movable finger 0.42. Leg I: trochanter 0.09; femur 0.22; patella 0.16; tibia 0.17; tarsus 0.30. Leg IV: trochanter 0.16; femur 0.14; patella 0.34; tibia 0.33; tarsus 0.30.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
F670E3C29FCF5842B7FC6906D632BC55.taxon	distribution	Distribution. Presently known only from the type locality (BLF 3599) and from three additional locations (BLF 3571, BLF 6506 and BLF 6446) in the Boeny Region (formerly Mahajanga Province).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
BA67104534735E24B141557B9823E2F6.taxon	diagnosis	Diagnosis. Mahajanganella gen. nov. is morphologically unique by having the following characters, which differ from other feaelloid members in Madagascar: 5 specialized setae on the retrolateral face of the movable chelal finger arranged in a group between trichobothrium b and terminal teeth (arranged in a row in Antsirananaella gen. nov.); smaller overall size than Antsirananaella gen. nov.; stronger granulate cuticle than Antsirananaella gen. nov.; all four anterior lobes of carapace with the same distance to each other (smaller distance between anteriomedial lobes than to anteriolateral ones in Antsirananaella gen. nov.). Like Antsirananaella gen. nov. and Toliaranella gen. nov., it differs from Cybella by having platelets on the pleural membrane (absent in Cybella), from Iporangella by the presence of specialized setae on the movable chelal finger (absent in Iporangella), from Feaella (Difeaella) and Feaella (Feaella) by the presence of four anterior carapaceal lobes (two and six, respectively), from Feaella (Tetrafeaella) in continental Africa by having a less pronounced depression on the base of coxa I and on top of coxa II (distinctly more pronounced in Feaella (T.) cf. mucronata), and from the Australian species presently attributed to Feaella (Tetrafeaella) by having fewer coxal spines (one pair versus three or four in the Australian taxa).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
BA67104534735E24B141557B9823E2F6.taxon	etymology	Etymology. The genus is named after the former Mahajanga Province, where many of these feaellids occur. The gender is feminine.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
BA67104534735E24B141557B9823E2F6.taxon	description	Description. The following description is based on holotype and allotype of M. heraclis sp. nov. - Carapace (Figs 12 A, 14 A, 15 A, 16 A): Moderate granulate and all four anterior lobes with same distance to each other. - Pedipalp (Figs 11 A, B, 13 A, B, C, 14 C, D, 15 C, D, 16 C, D): Trichobothrial pattern: esb between st and ist on the sagittal level with ist slightly more distal and st slightly more proximal than esb; t distinctly closer to sb than to b; sb distinctly closer to t than to st; it on the fixed finger between eb and et distinctly closer to eb than to et. Movable finger with five specialized setae on the ventral face, arranged in a group and slightly more distally situated than b (Fig. 13 A). Terminal teeth varying in number, both in fixed and the movable fingers each with 7 - 8 terminal teeth (one large tooth included on fixed finger). - Chelicera: Most specimens were too dirty to get secure information about characters. Coxal region (Fig. 12 B, C), legs (Figs 11 B, 14 B, 15 B, 16 B), abdomen (Fig. 11 A, B, C) and genital region (Fig. 12 B) with the same characters as Antsirananaella gen. nov.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
BA67104534735E24B141557B9823E2F6.taxon	distribution	Distribution. Specimens of Mahajanganella gen. nov. have been found in dry localities (dry tropical forests) in the Boeny and Menabe Regions (formerly Mahajanga and Toliara Provinces) along the western coast of Madagascar.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
91C8C7D43B7A56BDA76F3C11C0E278F2.taxon	diagnosis	Diagnosis. Very similar to Mahajanganella heraclis sp. nov. but smaller female body size (body length 2.03 versus 2.15 in M. heraclis sp. nov.), and most slender carapace (1.47 (♂) / 1.41 (♀) x longer than broad versus 1.35 (♂) / 1.29 (♀) in M. heraclis); most slender femora of Mahajanganella gen. nov. (1.57 / 1.74 ♂ / ♀ x longer than broad versus 1.64 / 1.62 ♂ / ♀ in M. heraclis sp. nov.).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
91C8C7D43B7A56BDA76F3C11C0E278F2.taxon	etymology	Etymology. This species is a patronym honoring Arnold Schwarzenegger, a famous former bodybuilder from Austria, known as an actor in the movie " Terminator ", former governor of California and now supporting conservation programs.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
91C8C7D43B7A56BDA76F3C11C0E278F2.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Fig. 16 A): 1.47 (♂), 1.38 - 1.43 (♀) times longer than broad. - Pedipalps (Fig. 16 C, D): see trichobothrial pattern in genus description. Chelal fixed finger with 8 teeth in the OR, 13 - 17 in the MR and 9 - 12 in the IR; movable finger with 8 - 9 teeth in the OR, 11 - 13 in the MR and 9 - 12 in the IR; terminal teeth build a group of 6 equally sized and 1 larger tooth on the fixed finger and 8 equally sized teeth on the movable finger. - Dimensions (mm): Holotype ♂: Body length 1.93; abdomen length 1.34; width (with pleura) 1.17; (without pleura) 1.07; carapace length 0.56; width 0.38. Pedipalp: trochanter 0.23; femur length 0.56; width 0.32; patella 0.44; chela (without pedicel) 0.47; hand length 0.09; width 0.13; movable finger 0.36. Leg I: trochanter 0.09; femur 0.21; patella 0.14; tibia 0.15; tarsus 0.26. Leg IV: trochanter 0.15; femur 0.13; patella 0.29; tibia 0.24; tarsus 0.31. Allotype ♀: Body length 2.04; abdomen length 1.44; width (with pleura) 1.30; (without pleura) 1.18; carapace length 0.60; width 0.42. Pedipalp: trochanter 0.27; femur length 0.56; width 0.34; patella 0.48; chela (without pedicel) 0.51; hand length 0.10; width 0.15; movable finger 0.39. Leg I: trochanter 0.09; femur 0.21; patella 0.20; tibia 0.13; tarsus 0.28. Leg IV: trochanter 0.16; femur 0.13; patella 0.34; tibia 0.31; tarsus 0.33.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
91C8C7D43B7A56BDA76F3C11C0E278F2.taxon	distribution	Distribution. Known only from the type locality (BLF 6812) in the Boeny Region (formerly Mahajanga Province) of western Madagascar.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
3E8350C76DFC51EFA098A086FA0FF2B2.taxon	materials_examined	Additional material examined. MADAGASCAR: Toliara Province [Anosy Region]: 1 ♂, 1 nymph (CAS 9071750 [BLF 5314]), Reserve Prive Berenty, Foret de Bealoka, 24 ° 57 ′ 25 ″ S 46 ° 16 ′ 17 ″ E, 3 - 8 February 2002, B. L. Fisher, Griswold et al. 76; ♂, 55 ♀,> 100 nymphs (CAS 9071589 [BLF 5426]), Reserve Prive Berenty, Foret de Malaza, Mandrare River, 25 ° 00 ′ 28 ″ S 46 ° 18 ′ 22 ″ E, 6 February 2002, Fisher, Griswold et al.; 1 ♂, 2 ♀, 2 nymphs (CAS 9071578 [BLF 5278]), Foret de Mahavelo, Isantoria River, 24 ° 45 ′ 13 ″ S 46 ° 09 ′ 5 ″ E, 31 January 2002, Fisher, Griswold et al.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
3E8350C76DFC51EFA098A086FA0FF2B2.taxon	diagnosis	Diagnosis. Most similar to T. griswoldi sp. nov., but females with most slender femora of this new genus (1.71 x longer than broad versus 1.62 - 1.69) and different body size.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
3E8350C76DFC51EFA098A086FA0FF2B2.taxon	etymology	Etymology. This species is a patronym honoring Brian Fisher who collected many Malagasy feaellids.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
3E8350C76DFC51EFA098A086FA0FF2B2.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Fig. 21 A): 1.25 - 1.39 (♂), 1.27 - 1.39 (♀) times longer than broad. - Pedipalps (Fig. 21 C, D): See trichobothrial pattern in description of T. griswoldi sp. nov. Chelal fixed finger with 10 teeth in the OR, 13 - 16 in the MR and 10 - 13 teeth in the IR; movable finger with 8 - 10 teeth in the OR, 9 - 12 in the MR and 8 - 11 teeth in the IR. Fixed finger with 6 - 7 equally sized and one large terminal tooth, movable finger with 7 terminal teeth equal in size. - Dimensions (mm): Holotype ♂: Body length 1.56; abdomen length 1.10, width (with pleura) 1.00, width (without pleura) 0.85; carapace length 0.44, width 0.34. Pedipalp: trochanter 0.24; femur length 0.43, width 0.27; patella 0.36; chela (without pedicel) 0.39; hand length 0.09, width 0.12; movable finger 0.32. Leg I: trochanter 0.07; femur 0.16; patella 0.14; tibia; 0.13; tarsus 0.22. Leg IV: trochanter 0.15; femur 0.10; patella 0.25; tibia 0.26; tarsus 0.26. Allotype ♀: Body length 2.15; abdomen length 1.49, width (with pleura) 1.35, width (without pleura) 1.13; carapace length 0.57, width 0.44. Pedipalp: trochanter 0.24; femur length 0.60, width 0.33; patella 0.45; chela (without pedicel) 0.50; hand length 0.11, width 0.15; movable finger 0.41. Leg I: trochanter 0.10; femur 0.20; patella 0.16; tibia 0.15; tarsus 0.28. Leg IV: trochanter 0.19; femur 0.13; patella 0.30; tibia 0.31; tarsus 0.35.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
3E8350C76DFC51EFA098A086FA0FF2B2.taxon	distribution	Distribution. Known from three localities along the Mandrare River in the Anosy Region (formerly Toliara Province) in southern Madagascar.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
8415D05F4BBF5412AE4BF1FE8C7DC4CB.taxon	materials_examined	Additional material examined. MADAGASCAR: Toliara Province [Anosy Region]: 4 ♂, 4 ♀, 9 nymphs (CAS 9071509 [BLF 4984]), Parc National d'Andohahela, Foret d'Ambohibory, 24 ° 55 ′ 48 ″ S 46 ° 38 ′ 44 ″ E, 16 - 20 January 2002, Fisher, Griswold et al.; 1 ♂, 1 ♀ (CAS 9071516 [BLF 4815]), Parc National d'Andohahela, Foret de Manantalinjo, 24 ° 49 ′ 1 ″ S 46 ° 36 ′ 36 ″ E, 12 - 16 January 2002, Fisher, Griswold et al.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
8415D05F4BBF5412AE4BF1FE8C7DC4CB.taxon	diagnosis	Diagnosis. Differs from T. pumila sp. nov. in different cheotaxy: est on fixed finger situated between sb and t of the movable finger on sagittal level (est on same sagittal level with sb in T. pumila sp. nov.); most compact carapace of the southern feaellid members (1.27 / 1.23 ♂ / ♀ times longer than broad versus 1.31 - 1.37 / 1.31 - 1.39 ♂ / ♀).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
8415D05F4BBF5412AE4BF1FE8C7DC4CB.taxon	etymology	Etymology. This species is a patronym honoring Charles Griswold who collected many Malagasy feaellids.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
8415D05F4BBF5412AE4BF1FE8C7DC4CB.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Fig. 22 A): Very compact, heavily granulate, 1.20 - 1.31 (♂), 1.18 - 1.31 (♀) times longer than broad. - Pedipalps (Fig. 22 C, D): Similar trichobothrial pattern as T. pumila sp. nov. but the following difference: est on fixed finger situated between sb and t of the movable finger on sagittal level. Chelal fixed finger with 9 - 10 chelal teeth in the OR, 12 - 14 in the MR and 9 - 11 in the IR; movable finger with 6 - 8 in the OR, 7 - 11 in the MR and 7 - 8 in the IR. Fixed finger with 6 - 7 terminal teeth equal in size and one large one, movable finger with 7 - 8 equally sized terminal teeth. - Dimensions (mm): Holotype ♂: Body length 1.62; abdomen length 1.13, width (with pleura) 1.05, width (without pleura) 0.91; carapace length 0.46, width 0.35. Pedipalp: trochanter 0.22; femur length 0.49, width 0.27; patella 0.35; chela (without pedicel) 0.35; hand length 0.10, width 0.11; movable finger 0.26. Leg I: trochanter 0.07; femur 0.14; patella 0.13; tibia 0.11; tarsus 0.21. Leg IV: trochanter 0.13; femur 0.10; patella 0.21; tibia 0.20; tarsus 0.26. Allotype ♀: Body length 2.04; abdomen length 1.47, width (with pleura) 1.33, width (without pleura) 1.11; carapace length 0.53, width 0.45. Pedipalp: trochanter 0.25; femur length 0.59, width 0.35; patella 0.46; chela (without pedicel) 0.44; hand length 0.10, width 1.14; movable finger 0.35. Leg I: trochanter 0.11; femur 0.16; patella 0.14; tibia 0.16; tarsus 0.25. Leg IV: trochanter 0.18; femur 0.14; patella 0.27; tibia 0.31; tarsus 0.30.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
8415D05F4BBF5412AE4BF1FE8C7DC4CB.taxon	distribution	Distribution. Currently known from the type locality (BLF 4810) and two additional localities of (BLF 4984 and BLF 4815) in the Anosy Region (former Toliara Province) in southern Madagascar.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
78D6D54E21FC5C548F4EDE39CB81241B.taxon	diagnosis	Diagnosis. Toliaranella gen. nov. differs from all other Malagasy feaellids by the presence of a perforated organ above the coxal spines and smallest body size of any Malagasy groups. Like Antsirananaella gen. nov. and Mahajanganella gen. nov., it differs from Cybella by having platelets on the pleural membrane (absent in Cybella), from Iporangella by the presence of specialized setae on the movable chelal finger (absent in Iporangella), from Feaella (Difeaella) and Feaella (Feaella) by the presence of four anterior carapaceal lobes (two and six, respectively), from Feaella (Tetrafeaella) in continental Africa by having a less pronounced depression on the base of coxa I and on top of coxa II (distinctly more pronounced in Feaella (T.) cf. mucronata) and from the Australian species presently attributed to Feaella (Tetrafeaella) by having fewer coxal spines (one pair versus three or four in the Australian taxa).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
78D6D54E21FC5C548F4EDE39CB81241B.taxon	etymology	Etymology. This genus is named after the former Toliara Province, where all specimens were collected. The gender is feminine.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
78D6D54E21FC5C548F4EDE39CB81241B.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Figs 18 A, 20 A, 21 A, 22 A, 23 A, 24 A): Strongly granulate, all four anterior lobes with same distance to each other and rounded at tips. - Pedipalps (Figs 17 A, B, 19 A-C, 20 C, D, 21 C, D, 22 C, D, 23 C, D, 24 C, D): Terminal teeth with 7 - 8 teeth on both the fixed and the movable fingers (including 1 large tooth on fixed finger). - Coxal region (Figs 17 B, 18 B, C): Pedipalpal coxae each with one spike laterally near base; coxae I with basal depression, each side with one small coxal spine; slightly more cranial of coxal spine one novel organ on each side: rose-shaped, not granulate but perforated organ (Fig. 18 B, C) filling the base of the depression; coxae II with irregular shaped spines framing depression of coxae I.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
78D6D54E21FC5C548F4EDE39CB81241B.taxon	distribution	Distribution. Members of this genus are localized to the southern part of Madagascar in the Androy, Anosy and Atsimo-Andrefana Regions (formerly Toliara Province) where the main landscape is covered by arid spiny bush vegetation.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
D1B333573D415F8D9B2C63C3C60FD850.taxon	materials_examined	Additional material examined. MADAGASCAR: Toliara Province [Atsimo-Andrefana Region]: 3 nymphs (CAS 9071531 [MGF 004]), Antafoky, 23 ° 29 ′ 16 ″ S 44 ° 04 ′ 39 ″ E, 9 February 2002, Frontier Project; 3 ♂, 1 ♀, 6 nymphs (CAS 9071523 [MGF 020]), Sept Lacs, 23 ° 31 ′ 42 ″ S 44 ° 09 ′ 20 ″ E, 8 March 2002, Frontier Project; 5 ♂, 3 ♀, 6 nymphs (CAS 9071770 [MGF 028]), Sept Lacs, 23 ° 31 ′ 29 ″ S 44 ° 09 ′ 33 ″ E, 10 March 2002; Frontier Project; 3 ♂, 3 ♀, 10 nymphs (CAS 9071522 [MGF 030]), Manderano, 23 ° 31 ′ 38 ″ S 44 ° 05 ′ 15 ″ E, 10 - 14 March 2002, Frontier Wilderness Project; 6 ♂, 6 ♀, 8 nymphs (CAS 9071753 [MGF 033]), Manderano, 23 ° 31 ′ 27 ″ S 44 ° 05 ′ 34 ″ E, 29 May- 1 June 2002, Frontier Wilderness Project; 3 ♂, 4 ♀, 5 nymphs (CAS 9072461 [MGF 034]), Manderano, 23 ° 31 ′ 24 ″ S 44 ° 05 ′ 40 ″ E, 8 - 12 June 2002, Frontier Wilderness Project; 1 ♂, 2 nymphs (CAS 9071731 [MGF 037]), Sept Lacs, 23 ° 31 ′ 15 ″ S 44 ° 8 ′ 35 ″ E, 20 - 24 August 2002, Frontier Wilderness Project; 3 nymphs (CAS 9071780 [MGF 040]), 1 nymph (CAS 9072465 [MGF 043]), Fiherenana, 23 ° 10 ′ 37 ″ S 43 ° 57 ′ 39 ″ E, 21 - 24 October 2002, Frontier Wilderness Project; 2 ♂, 2 ♀, 2 nymphs (CAS 9071745 [MGF 050]), Fiherenana, 23 ° 14 ′ 07 ″ S 43 ° 52 ′ 15 ″ E, 1 - 4 December 2002, Frontier Wilderness Project; 2 ♂, 1 ♀, 3 nymphs (CAS 9071774 [MGF 056]), Ranobe, 23 ° 02 ′ 03 ″ S 43 ° 36 ′ 43 ″ E, 5 - 9 February 2003, Frontier Wilderness Project; 2 ♂, 1 nymph (CAS 9071517 [MGF 061]), Lake Ranobe, 23 ° 02 ′ 23 ″ S 43 ° 36 ′ 39 ″ E, 17 - 21 February 2003, Frontier Wilderness Project; 3 ♂, 4 ♀, 1 nymph (CAS 9072418 [MGF 064 (14120 (1 ))]), Lake Ranobe, 23 ° 02 ′ 21 ″ S 43 ° 36 ′ 42 ″ E, 25 - 28 April 2003, Frontier Wilderness Project; 1 nymph (CAS 9071722 [MGF 070]), Lake Ranobe, 23 ° 02 ′ 44 ″ S 43 ° 36 ′ 58 ″ E, 17 - 21 May 2003, Frontier Wilderness Project; 1 ♀, 1 nymph (CAS 9071644 [MGF 074]), Fiherenana, 23 ° 13.351 ′ S 43 ° 52.853 ′ E, 5 - 7 August 2003, Frontier Wilderness Project; 1 nymph (CAS 9071649 [MGF 077]), Fiherenana, 23 ° 10.619 ′ S 43 ° 57.685 ′ E, 18 - 19 August 2003, Frontier Wilderness Project; 1 ♂, 1 ♀ (CAS 9072464 [MGF 092]), Ranobe, 23 ° 02.667 ′ S 43 ° 36.917 ′ E, 26 - 29 January 2004, Frontier Wilderness Project; 21 ♂, 8 ♀, 47 nymphs (CAS 9071597 [BLF 5850]), Foret de Mite, 23 ° 31 ′ 27 ″ S 44 ° 07 ′ 17 ″ E, 27 February- 3 March 2002, Fisher, Griswold et al.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
D1B333573D415F8D9B2C63C3C60FD850.taxon	diagnosis	Diagnosis. Very similar to T. pumila sp. nov. but larger (e. g., body length 1.76 / 2.23 ♂ / ♀ versus 1.56 / 1.96 ♂ / ♀ in T. pumila sp. nov.; femur length 0.49 / 0.61 ♂ / ♀ versus 0.44 / 0.53 ♂ / ♀ in T. pumila sp. nov.); more slender femora (1.66 / 1.69 ♂ / ♀ times longer than broad versus 1.63 / 1.63 ♂ / ♀ in T. pumila sp. nov.).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
D1B333573D415F8D9B2C63C3C60FD850.taxon	etymology	Etymology. This species is a patronym honoring Volker Mahnert who worked on African pseudoscorpions.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
D1B333573D415F8D9B2C63C3C60FD850.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Fig. 23 A): 1.33 - 1.42 (♂), 1.32 - 1.37 (♀) times longer than broad. - Pedipalps (Fig. 23 C, D): See trichobothrial pattern in description of T. pumila sp. nov.; Chelal fixed finger with 10 - 11 chelal teeth in the OR, 12 - 16 in the MR and 8 - 14 in the IR; movable finger with 6 - 8 in the OR, 9 - 11 in the MR and 8 - 10 teeth in the IR; 7 equally sized and one larger terminal tooth on fixed finger and 8 equally sized terminal teeth on movable finger. - Dimensions (mm): Holotype ♂: Body length 1.77; abdomen length 1.21, width (with pleura) 1.13, width (without pleura) 0.99; carapace length 0.52, width 0.39. Pedipalp: trochanter 0.26; femur length 0.49, width 0.30; patella 0.41; chela (without pedicel) 0.44; hand length 0.10, width 0.13; movable finger 0.36. Leg I: trochanter 0.08; femur 0.17; patella 0.15; tibia 0.15; tarsus 0.24. Leg IV: trochanter 0.15; femur 0.12; patella 0.26; tibia 0.23; tarsus 0.30. Allotype ♀: Body length 2.34; abdomen length 1.70, width (with pleura) 1.13, width (without pleura) 1.22; carapace length 0.59, width 0.43. Pedipalp: trochanter 0.26; femur length 0.61, width 0.37; patella 0.47; chela (without pedicel) 0.48; hand length 0.12, width 0.15; movable finger 0.39. Leg I: trochanter 0.10; femur 0.17; patella 0.20, tibia 0.18; tarsus 0.27. Leg IV: trochanter 0.18; femur 0.14; patella 0.31; tibia 0.32; tarsus 0.34.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
D1B333573D415F8D9B2C63C3C60FD850.taxon	distribution	Distribution. Presently known from 17 localities in the Atsimo-Andrefana Region (formerly Toliara Province): type locality (MGF 001) in southwestern Madagascar close to the Onilahy River delta and additional localities in the radius of 9 km upstream along the right side of the Onilahy River and around the Fiherenana River (MGF 004, MGF 020, MGF 028, MGF 030, MGF 033, MGF 034, MGF 037, MGF 043, MGF 040, MGF 050, MGF 056, MGF 061, MGF 064, MGF 070, MGF 074, MGF 077, MGF 092).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6921D09A8C8E59929071493DBF45E65B.taxon	materials_examined	Additional material examined. MADAGASCAR: Toliara Province [Androy Region]: 8 ♂, 14 ♀, 5 nymphs (CAS 9071596 [BLF 5500]), Reserve Speciale de Cap Sainte Marie, 25 ° 34 ′ 54 ″ S 45 ° 10 ′ 6 ″ E, 11 - 15 February 2002, Fisher, Griswold et al.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6921D09A8C8E59929071493DBF45E65B.taxon	diagnosis	Diagnosis. Largest body size of the southern Malagasy feaellids (e. g., body length 1.85 / 2.33 ♂ / ♀ versus 1.56 - 1.80 / 1.96 - 2.23 ♂ / ♀; carapace length 0.53 / 0.64 ♂ / ♀ versus 0.45 - 0.51 / 0.52 - 0.58 ♂ / ♀; femur length 0.55 / 0.67 ♂ / ♀ versus 0.44 - 0.53 / 0.53 - 0.61 ♂ / ♀)	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6921D09A8C8E59929071493DBF45E65B.taxon	etymology	Etymology. This species is named after the type locality, which is the southernmost point of Madagascar.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6921D09A8C8E59929071493DBF45E65B.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Fig. 24 A): 1.25 - 1.38 (♂), 2.64 (♀) x longer than broad. - Pedipalps (Fig. 24 C, D): See trichobothrial pattern in description of T. pumila sp. nov. Chelal fixed finger with 9 - 11 teeth in the OR, 13 - 14 in the MR and 10 in the IR; movable finger with 8 - 9 teeth in the OR, 11 - 14 in the MR and 9 teeth in the IR; terminal teeth: 7 equally sized and one large one on the fixed finger, 7 teeth on the movable finger equal in size. - Dimensions (mm): Holotype ♂: Body length 1.82; abdomen length 1.29, width (with pleura) 1.18, width (without pleura) 1.04; carapace length 0.50, width 0.40. Pedipalp: trochanter 0.24; femur length 0.59, width 0.34; patella 0.44; chela (without pedicel) 0.43; hand length 0.10, width 0.13; movable finger 0.34. Leg I: trochanter 0.10; femur 0.13; patella 0.17; tibia 0.17; tarsus 0.22. Leg IV: trochanter 0.16; femur 0.12; patella 0.26; tibia 0.24; tarsus 0.30. Allotype ♀: Body length 2.33; abdomen length 1.69, width (with pleura) 1.50, width (without pleura) 1.30; carapace length 0.64, width 0.46. Pedipalp: trochanter 0.27; femur length 0.67, width 0.40; patella 0.52; chela (without pedicel) 0.50; hand length 0.11, width 0.16; movable finger 0.41. Leg I: trochanter 0.11; femur 0.20; patella 0.21; tibia 0.15; tarsus 0.29. Leg IV: trochanter 0.18; femur 0.16; patella 0.35; tibia 0.34; tarsus 0.34.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
6921D09A8C8E59929071493DBF45E65B.taxon	distribution	Distribution. Known from the two southernmost localities (type locality and BLF 5500) among all Malagasy feaellids in the Androy Region (formerly Toliara Province).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
1693A8FBD17A5E48A8C7B7419AF4BB22.taxon	materials_examined	Additional material examined. MADAGASCAR: Toliara Province [Atsimo-Andrefana Region]: 25 ♂, 10 ♀, 39 nymphs (CAS 9071598 [BLF 6070]), 6 ♂, 1 ♀, 5 nymphs (CAS 9071743 [BLF 6068]), Foret de Beroboka, 22 ° 13 ′ 59 ″ S 43 ° 21 ′ 59 ″ E, 12 - 16 March 2002, Fisher, Griswold et al.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
1693A8FBD17A5E48A8C7B7419AF4BB22.taxon	diagnosis	Diagnosis. Smallest body sizes of Toliaranella species and of all Malagasy feaellids (e. g., body length 1.56 / 1.96 ♂ / ♀ versus 1.60 - 1.85 / 2.04 - 2.33 ♂ / ♀; Pedipalps: femur length 0.44 / 0.53 ♂ / ♀ versus 0.45 - 0.55 / 0.56 - 0.67 ♂ / ♀); most robust male pedipalpal femur of Toliaranella (femur 1.63 x longer than broad versus 1.65 - 1.72); cheotaxy: est on fixed finger on same sagittal level as sb on movable finger (est more distal than sb in T. griswoldi sp. nov. and T. fisheri sp. nov.).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
1693A8FBD17A5E48A8C7B7419AF4BB22.taxon	etymology	Etymology. This species is named after the latin ' pumilus, - a, - um ' which means " dwarf ". The specific epithet references the small size of all specimens.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
1693A8FBD17A5E48A8C7B7419AF4BB22.taxon	description	Description. The following description is based on holotype and allotype. - Carapace (Fig. 20 A): 1.29 - 1.44 (♂), 1.24 - 1.47 (♀) times longer than broad. - Pedipalp (Fig. 20 C, D): Femora very robust (1.55 - 1.69 (♂), 1.43 - 1.80 (♀) times longer than broad; trichobothrial pattern: same as in Mahajanganella gen. nov., except: est on fixed finger on same sagittal level as sb on movable finger. Chelal fixed finger with 9 - 10 teeth in the OR, 12 - 15 in the MR and 8 - 12 in the IR; movable finger with 6 - 10 teeth in the OR, 9 - 11 in the MR and 9 - 10 in the IR; fixed finger with 6 equally sized and one larger terminal tooth, movable finger with 7 equally sized terminal teeth. - Chelicera, legs, abdomen and genital region: With same characters as all members before. - Coxal region: See genus description. - Dimensions (mm): Holotype ♂: Body length 1.47; abdomen length 1.02, width (with pleura) 0.93, width (without pleura) 0.86; carapace length 0.45, width 0.32. Pedipalp: trochanter 0.21; femur length 0.45, width 0.27; patella 0.35; chela (without pedicel) 0.38; hand length 0.08, width 0.11; movable finger 0.30. Leg I: trochanter 0.08; femur 0.12; patella 0.15; tibia 0.11; tarsus 0.22. Leg IV: trochanter: 0.13; femur 0.11; patella 0.23; tibia 0.21; tarsus 0.26. Allotype ♀: Body length 2.08; abdomen length 1.51, width (with pleura) 1.39, width (without pleura) 1.15; carapace length 0.55, width 0.40. Pedipalp: trochanter 0.26; femur length 0.58, width 0.36; patella 0.42; chela (without pedicel) 0.43; hand length 0.11, width 0.13; movable finger 0.35. Leg I: trochanter 0.09; femur 0.18; patella 0.19; tibia 0.15; tarsus 0.26. Leg IV: trochanter 0.15; femur 0.13; patella 0.27; tibia 0.28; tarsus 0.33.	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
1693A8FBD17A5E48A8C7B7419AF4BB22.taxon	distribution	Distribution. Presently known from the type locality (BLF 9556) and one additional locality (BLF 6070) in the Atsimo-Andrefana Region (formerly Toliara Province).	en	Lorenz, Michelle, Loria, Stephanie F., Harvey, Mark S., Harms, Danilo (2022): The Hercules pseudoscorpions from Madagascar: A systematic study of Feaellidae (Pseudoscorpiones: Feaelloidea) highlights regional endemism and diversity in one of the " hottest " biodiversity hotspots. Arthropod Systematics & amp; Phylogeny 80: 649-691, DOI: http://dx.doi.org/10.3897/asp.80.e90570, URL: http://dx.doi.org/10.3897/asp.80.e90570
