taxonID	type	description	language	source
03ED87A7FF86FFEC6B02FB71FD05FE66.taxon	description	http: // zoobank. org / urn: lsid: zoobank. org: act: 5 E 86 BBCC-CDC 4 - 4314 - 9217 - 5 E 964 E 0 B 89 FB Description Female (Figs. 1 – 3) – Body weakly sclerotized. Length of idiosoma 125 (120 – 125), width 77 (72 – 78). Gnathosoma – Gnathosomal capsule, excluding palps, almost round, its length 25 (23 – 25), width 24 (23 – 25). Dorsally with two pairs of smooth subequal cheliceral setae (cha, chb). Setae cha 12 (11 – 12) blunt-tipped; chb 11 (10 – 11) pointed. Dorsal median apodeme weakly developed. Postpalpal setae (pp) rod-like with tiny distal projections, situated posterolaterally to setae cha. Venter of gnathosoma with one pair of smooth, pointed subcapitular setae m 13 (11 – 13). Palps freely articulated to gnathosomal capsule, with smooth setae dFe and dGe dorsally, setae dGe 11 (10 – 11) pointed, about two times longer than weakly blunt-tipped dFe 4 (4 – 5). Palps ventrally with solenidion almost three times longer than accessory setigenous structure (ass). Palps terminated with a welldeveloped tibial claw. Palp tibiotarsus with tiny eupathid-like distal seta and small lateral seta l ”. Cheliceral stylets strong, curved. Pharynx poorly visible, with weak lateral projections. Idiosomal dorsum (Figs. 1 A, 3 A) – All dorsal sclerites with clearly visible numerous round and very small puncta. Prodorsal shield with three pairs of setae (v 1, v 2, sc 2) and one pair of clavate, barbed trichobothria sc 1 with rounded apex. All dorsal setae blunt-tipped; tips of setae h 2 clubshaped. Setae v 1, v 2, and sc 2 smooth, other dorsal setae weakly barbed. Only cupules ia on tergite D clearly visible; other cupules not evident. Posterior margins of tergites C, D, and EF with several distinct tooth-like projections. Lengths of dorsal setae: v 1 13 (13 – 14), v 2 7 (6 – 7), sc 2 20 (20 – 22), c 1 15 (14 – 15), c 2 20 (19 – 21), d 13 (12 – 14), e 13 (12 – 13), f 13 (13 – 14), h 1 13 (12 – 14), h 2 37 (37 – 38). Distances between setae: v 1 – v 1 17 (17), v 2 – v 2 26 (25 – 26), sc 2 – sc 2 27 (27), c 1 – c 1 23 (22 – 23), c 1 – c 2 18 (18 – 19), d – d 32 (28 – 32), e – e 41 (39 – 41), f – f 29 (28 – 29), h 1 – h 1 12 (11 – 12), h 1 – h 2 7 (7). Idiosomal venter (Figs 1 B, 3 B) – All ventral plates smooth. All ventral setae weakly blunttipped; setae ps weakly barbed, other ventral setae smooth. Apodemes 1 (ap 1) and apodemes 2 (ap 2) well developed; ap 1 joined with poorly visible prosternal apodeme (appr), sejugal apodeme represented by pair of small sclerites located posterolaterad setae 2 c; apodemes 3 (ap 3) and 4 (ap 4) well developed. Poststernal apodeme absent. Coxal fields I-IV each with three pairs of setae. Lengths of ventral setae – 1 a 6 (6), 1 b 6 (6), 1 c 6 (6), 2 a 10 (8 – 10), 2 b 6 (5 – 6), 2 c 9 (8 – 9), 3 a 8 (7 – 8), 3 b 8 (7 – 8), 3 c 10 (9 – 11), 4 a 8 (7 – 8), 4 b 9 (9 – 10), 4 c 9 (9), ag 8 (8 – 9), g 1 3 (3), g 2 3 (3), ps 8 (8 – 9). Legs (Fig. 2) – Leg I slightly longer than subequal legs II-IV. Leg I (Fig. 2 A). Setal formula: 0 – 4 – 2 – 6 (2) – 11 (2). Tarsus with two small claws and semioval empodium. All leg setae smooth. Setae l’ of femur, l’ and v’ of genu, k and v’ of tibia blunt-tipped; other leg setae (except eupathidia p’, p ”, ft’, ft ”) pointed; setae (u) and (pv) of tarsus not modified. Trochanter dorsally with four short toothlike projections. Tarsus I with ventrodistal membranous flange. Lengths of solenidia ω 1 7 (6 – 7), ω 2 3 (3), φ 1 8 (8), φ 2 5 (4 – 5); solenidion ω 1 digitiform; solenidia φ 2 and ω 2 baculiform; solenidion φ 1 clavate. Leg II (Fig. 2 B). Setal formula: 0 – 2 – 1 – 4 (1) – 6 (1). Tarsal claws simple, hooked; empodium large, extending beyond tips of claws. Solenidion ω 5 (4 – 5) digitiform; solenidion φ 3 (3) weakly clavate. Trochanter dorsolaterally with two tooth-like projections. Seta l’ of genu weakly barbed, other setae smooth; all setae pointed. Leg III (Fig. 2 C). Setal formula: 0 – 1 – 1 – 4 – 5. Claws and empodium of same shape as on tarsus II. Setae d of femur blunt-tipped, other leg setae pointed; all setae smooth. Trochanter ventrally with short lobe. Leg IV (Fig. 2 D). Setal formula: 0 – 1 – 1 – 4 – 5. Claws and empodium of same shape as on tarsus II. Setae d of femur blunt-tipped, other leg setae pointed; all setae smooth. Trochanter ventrally with short lobe. Male – unknown	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): Two new species of Pavania (Acari: Heterostigmata: Dolichocybidae) associated with scarab beetles (Coleoptera: Scarabaeidae) from Tanzania and Madagascar. Persian Journal of Acarology 11 (4): 545-558, DOI: https://doi.org/10.22073/pja.v11i4.76531
03ED87A7FF86FFEC6B02FB71FD05FE66.taxon	type_taxon	Type material Female holotype, slide No. ZISP T-Dol- 003, Tanzania, Arusha, 1975 (the original label includes only a code [1777]; the Arusha locality was inferred from the labels of the beetles collected apparently in the same locality), on H. neptunus; paratypes: 50 females, same data. Type deposition The holotype and 10 paratypes are deposited in the collection of the Zoological Institute of RAS, Saint Petersburg, Russia; five female paratypes are deposited in the acarological collection of Jalal Afshar Zoological Museum, Department of Plant Protection, Faculty of Agriculture, University of Tehran, Karaj, Iran (JAZM); other paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): Two new species of Pavania (Acari: Heterostigmata: Dolichocybidae) associated with scarab beetles (Coleoptera: Scarabaeidae) from Tanzania and Madagascar. Persian Journal of Acarology 11 (4): 545-558, DOI: https://doi.org/10.22073/pja.v11i4.76531
03ED87A7FF86FFEC6B02FB71FD05FE66.taxon	diagnosis	Differential diagnosis The new species is most similar to Pavania africana Khaustov and Frolov, 2018 and P. carabidophila Khaustov, 2005 by the similar length of idiosomal setae and absence of modified setae on legs. The new species differs from P. africana in having seta c 2 blunt-tipped (pointed in P. africana), longer genital setae (3) (1 – 2 in P. africana), shorter setae ps (8 – 9) (14 in P. africana), in having four tooth-like projections on trochanter I (three projections in P. africana), and solenidion φ 2 with rounded tip (φ 2 with attenuate tip in P. africana). The new species differs from P. carabidophila in having setae c 2 blunt-tipped (pointed in P. carabidophila), setae h 2 with clubshaped tips (pointed in P. carabidophila), by much shorter solenidion ω 1 (6 – 7) (11 – 12 in P. carabidophila), and in having setae cha and chb subequal (cha distinctly longer than chb in P. carabidophila).	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): Two new species of Pavania (Acari: Heterostigmata: Dolichocybidae) associated with scarab beetles (Coleoptera: Scarabaeidae) from Tanzania and Madagascar. Persian Journal of Acarology 11 (4): 545-558, DOI: https://doi.org/10.22073/pja.v11i4.76531
03ED87A7FF86FFEC6B02FB71FD05FE66.taxon	etymology	Etymology The name of the new species is a combination of two words: Greek neos meaning new and africana, the name of closely related species.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): Two new species of Pavania (Acari: Heterostigmata: Dolichocybidae) associated with scarab beetles (Coleoptera: Scarabaeidae) from Tanzania and Madagascar. Persian Journal of Acarology 11 (4): 545-558, DOI: https://doi.org/10.22073/pja.v11i4.76531
03ED87A7FF82FFE168E9FD84FBF6FB93.taxon	description	http: // zoobank. org / urn: lsid: zoobank. org: act: CFDAAEAE- 9088 - 4266 - 9812 - EE 93 CB 9 AE 306 Description Female (Figs. 4 – 6) – Body weakly sclerotized. Length of idiosoma 120 (115 – 120), width 78 (75 – 80). Gnathosoma – Gnathosomal capsule, excluding palps, almost round, its length 24 (23 – 24), width 27 (27 – 28). Dorsally with two pairs of blunt-tipped cheliceral setae (cha, chb). Setae cha 15 (15 – 16) weakly barbed; chb 10 (10 – 11) smooth. Dorsal median apodeme weakly developed. Postpalpal setae (pp) rod-like with tiny distal projections, situated posterolaterally to setae cha. Venter of gnathosoma with one pair of smooth, pointed subcapitular setae m 12 (12 – 13). Palps with smooth setae dFe and dGe dorsally, setae dGe 11 (10 – 11) pointed, about two times longer than weakly blunttipped dFe 4 (4 – 5). Palps ventrally with solenidion almost two times longer than accessory setigenous structure. Palps terminated with well-developed tibial claw. Palp tibiotarsus with tiny eupathid-like distal seta and small lateral seta l ”. Cheliceral stylets strong, curved. Pharynx poorly visible, with weak lateral projections. Idiosomal dorsum (Figs. 4 A, 6 A) – Tergites D, EF and H with round and very small puncta, similar puncta also poorly visible on prodorsal shield; tergite C smooth. Prodorsal shield with three pairs of setae (v 1, v 2, sc 2) and one pair of clavate, barbed trichobothria sc 1 with rounded apex. Setae sc 2 and c 1 pointed, other dorsal setae blunt-tipped; tips of setae h 2 club-shaped. Setae v 1, v 2, sc 2 and c 1 smooth, other dorsal setae weakly barbed. Cupules on tergites not evident. Posterior margins of tergites C, D, and EF with several very small tooth-like projections. Lengths of dorsal setae: v 1 17 (17 – 18), v 2 6 (6), sc 2 41 (39 – 43), c 1 30 (29 – 31), c 2 18 (18 – 19), d 14 (15 – 16), e 9 (9 – 10), f 16 (17 – 19), h 1 11 (10 – 11), h 2 52 (52 – 54). Distances between setae: v 1 – v 1 20 (21 – 23), v 2 – v 2 31 (29 – 32), sc 2 – sc 2 34 (33 – 35), c 1 – c 1 26 (26 – 28), c 1 – c 2 20 (19 – 22), d – d 53 (50 – 53), e – e 42 (39 – 43), f – f 32 (31 – 34), h 1 – h 1 9 (8 – 10), h 1 – h 2 8 (8). Idiosomal venter (Figs. 4 B, 6 B) – All ventral plates smooth. Setae 2 c pointed, other ventral setae weakly blunt-tipped; all ventral setae smooth. Apodemes 1 (ap 1) and apodemes 2 (ap 2) well developed; ap 1 joined with poorly visible prosternal apodeme (appr), sejugal apodeme represented by pair of small sclerites located posterolaterad setae 2 c; apodemes 3 (ap 3) and 4 (ap 4) well developed. Poststernal apodeme absent. Coxal fields I-IV each with three pairs of setae. Lengths of ventral setae: 1 a 7 (6 – 7), 1 b 7 (6 – 7), 1 c 6 (6 – 7), 2 a 11 (10 – 11), 2 b 7 (7), 2 c 20 (18 – 20), 3 a 10 (8 – 10), 3 b 9 (8 – 10), 3 c 10 (9 – 10), 4 a 9 (9), 4 b 9 (9 – 10), 4 c 9 (8 – 9), ag 9 (8 – 9), g 1 5 (5), g 2 5 (5), ps 7 (7 – 8). Legs (Fig. 5) – All legs subequal in length. Setation of legs I-IV as in P. neoafricana sp. nov. Leg I (Fig. 5 A). Tarsus with two small claws and semi-oval empodium. At least seta d of femur weakly barbed, other setae smooth. Setae l’ of femur, l’ and v’ of genu, k, v’ of tibia, and (u) of tarsus blunt-tipped; other leg setae (except eupathidia p’, p ”, ft’, ft ”) pointed; setae (u) and (pv) of tarsus not modified. Trochanter dorsally with one short tooth-like projection. Tarsus I with large ventrodistal membranous flange. Lengths of solenidia ω 1 6 (6), ω 2 3 (3), φ 1 7 (7), φ 2 4 (4 – 5); solenidion ω 1 digitiform; solenidia φ 2 and ω 2 baculiform; solenidion φ 1 clavate. Leg II (Fig. 5 B). Tarsal claws simple, hooked; empodium large, extending beyond tips of claws. Solenidion ω 5 (4 – 5) digitiform; solenidion φ 3 (1 – 3) weakly clavate. Trochanter dorsolaterally with two tooth-like projections. All leg setae smooth; seta l’ of tibia incrassate, spiniform and slightly curved; setae tc ”, u’ and (pv) weakly blunt-tipped; other setae pointed. Leg III (Fig. 5 C). Claws and empodium of same shape as on tarsus II. All setae smooth; setae d of femur and (tc) of tarsus weakly blunt-tipped, other leg setae pointed. Trochanter ventrally with short lobe. Leg IV (Fig. 2 D). Claws and empodium of same shape as on tarsus II. All setae smooth; setae d of femur, pl ” and (tc) of tarsus weakly blunt-tipped, other leg setae pointed. Trochanter ventrally with short lobe. Male – unknown.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): Two new species of Pavania (Acari: Heterostigmata: Dolichocybidae) associated with scarab beetles (Coleoptera: Scarabaeidae) from Tanzania and Madagascar. Persian Journal of Acarology 11 (4): 545-558, DOI: https://doi.org/10.22073/pja.v11i4.76531
03ED87A7FF82FFE168E9FD84FBF6FB93.taxon	type_taxon	Type material Female holotype, slide No. ZISP T-Dol- 004, Madagascar, Ankaratra Andraraty, 19 ° 21 ' 20.78 " S, 47 ° 18 ' 18.71 " E, on beetle Helictipleurus quadripunctatus collected in cow dung, 23.02.2022, A. V. Frolov leg; paratypes: 48 females, same data. Type deposition The holotype and 10 paratypes are deposited in the collection of the Zoological Institute of RAS, Saint Petersburg, Russia; five female paratypes are deposited in the acarological collection of Jalal Afshar Zoological Museum, Department of Plant Protection, Faculty of Agriculture, University of Tehran, Karaj, Iran (JAZM); other paratypes are deposited in the collection of the Tyumen State University Museum of Zoology, Tyumen, Russia.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): Two new species of Pavania (Acari: Heterostigmata: Dolichocybidae) associated with scarab beetles (Coleoptera: Scarabaeidae) from Tanzania and Madagascar. Persian Journal of Acarology 11 (4): 545-558, DOI: https://doi.org/10.22073/pja.v11i4.76531
03ED87A7FF82FFE168E9FD84FBF6FB93.taxon	diagnosis	Differential diagnosis The new species is most similar to Pavania riparia Sevastianov, 1980 and P. copridis Khaustov and Frolov, 2020 a by the similar length of idiosomal setae and setae 2 c pointed and about twice longer than 2 a. The new species differs from both species in having seta l’ on tibia II spiniform (setiform in P. riparia and P. copridis) and in having only one tooth-like projection on trochanter I (four in P. riparia, five in P. copridis).	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): Two new species of Pavania (Acari: Heterostigmata: Dolichocybidae) associated with scarab beetles (Coleoptera: Scarabaeidae) from Tanzania and Madagascar. Persian Journal of Acarology 11 (4): 545-558, DOI: https://doi.org/10.22073/pja.v11i4.76531
03ED87A7FF82FFE168E9FD84FBF6FB93.taxon	etymology	Etymology The name of the new species refers to its geographical distribution.	en	Khaustov, Alexander A., Frolov, Andrey V. (2022): Two new species of Pavania (Acari: Heterostigmata: Dolichocybidae) associated with scarab beetles (Coleoptera: Scarabaeidae) from Tanzania and Madagascar. Persian Journal of Acarology 11 (4): 545-558, DOI: https://doi.org/10.22073/pja.v11i4.76531
