taxonID	type	description	language	source
039687CBB4450A42906CF97C8324F929.taxon	materials_examined	Type species: Tellina fluminalis O. F. Müller, 1774.	en	Bespalaya, Yulia V., Kropotin, Alexander V., Kondakov, Alexander V., Aksenova, Olga V., Gofarov, Mikhail Yu., Kim, Sang Ki, Lee, Jin Hee, Travina, Oksana V., Vikhrev, Ilya V., Vinarski, Maxim V., Bolotov, Ivan N. (2023): A taxonomic reassessment of native and invasive species of Corbicula clams (Bivalvia: Cyrenidae) from the Russian Far East and Korea. Zoological Journal of the Linnean Society 197 (1): 104-126, DOI: 10.1093/zoolinnean/zlac078, URL: http://dx.doi.org/10.1093/zoolinnean/zlac078
039687CBB4450A419081F8E08369F883.taxon	description	TAXONOMY OF NATIVE AND INVASIVE CLAMS 115	en	Bespalaya, Yulia V., Kropotin, Alexander V., Kondakov, Alexander V., Aksenova, Olga V., Gofarov, Mikhail Yu., Kim, Sang Ki, Lee, Jin Hee, Travina, Oksana V., Vikhrev, Ilya V., Vinarski, Maxim V., Bolotov, Ivan N. (2023): A taxonomic reassessment of native and invasive species of Corbicula clams (Bivalvia: Cyrenidae) from the Russian Far East and Korea. Zoological Journal of the Linnean Society 197 (1): 104-126, DOI: 10.1093/zoolinnean/zlac078, URL: http://dx.doi.org/10.1093/zoolinnean/zlac078
039687CBB4450A419081F8E08369F883.taxon	materials_examined	Type material: Syntypes ZMB 32.198 (two pairs of valves ex coll. Gundlach; ‘ Corbicula japonica Prime, Japan, Gulick, Prime’); syntypes ZMB 170.411 (two pairs of valves ex coll. Dunker; ‘ Corbula japonica, von Mr Prime selbst erhalten’) inspected during this work. According to Glaubrecht et al. (2007), three other specimens (presumable syntypes) are kept in the Museum of Comparative Zoology, Harvard University, Cambridge, MA, USA (not seen). Type locality: ‘ Japonia (fide Gulick) collect. Wheatley, Jay, Browne, Cooper, Lyc. Hist. Nat., Prime and Lea’ = Japan (Glaubrecht et al., 2007). Material examined: Far East of Russia: Artemovka, Kiparisovka, Razdolnaya, Partizanskaya and Kievka rivers, in total 194 specimens; IX. 2020, Aksenova O. V., Bespalaya Yu. V., Kropotin A. V., Travina O. V., Vinarski M. V. leg. [lot nos RMBH Corb 54 and RMBH Corb 113 – 119], 43 specimens from the Razdolnaya and Artemovka rivers are kept in LMBI (accession numbers 21 - 043 and 21 - 045). Description: Shell relatively large, oval – triangular (Fig. 5). The umbo is centrally located and prominent. External shell colour varies from olive or greenish brown to black. The exterior exhibits concentric sculpture with regular ribs. Internal shell coloration matte white, with pale violet or dark purple marks in the area of the umbo. Hinge of the heterodont dentition type, with three cardinal teeth on each valve and two crenulated lateral teeth. Shell sculpture of young individuals characterized by well-marked concentric ridges and pronounced transverse stripes; shell coloration is lighter than in adults. Nacre varies from pure purple to purple with wide white marks. Siphons conical, both narrow. Inhalant siphon with a row of long papillae and with an additional row of short papillae. Black pigment is concentrated in rings internally at the base of both siphons and in the edge of the mantle forming the siphons. The outer surface of the siphons is white; on the inside, the siphons are bright orange. Larger papillae of the inhalant siphon, with dark rings. Distribution: East Asia: the Japan Sea (including some lakes and rivers near the sea), Korea, southern Sakhalin, southern Kurile Islands and the lower reaches of the Amur River (Glaubrecht et al., 2003; Yamada et al., 2014; Vinarski & Kantor, 2016).	en	Bespalaya, Yulia V., Kropotin, Alexander V., Kondakov, Alexander V., Aksenova, Olga V., Gofarov, Mikhail Yu., Kim, Sang Ki, Lee, Jin Hee, Travina, Oksana V., Vikhrev, Ilya V., Vinarski, Maxim V., Bolotov, Ivan N. (2023): A taxonomic reassessment of native and invasive species of Corbicula clams (Bivalvia: Cyrenidae) from the Russian Far East and Korea. Zoological Journal of the Linnean Society 197 (1): 104-126, DOI: 10.1093/zoolinnean/zlac078, URL: http://dx.doi.org/10.1093/zoolinnean/zlac078
039687CBB4410A4593C3F94F8360FC20.taxon	materials_examined	Type material: The lectotype [a pair of valves; ‘ Corbicula elatior Marts., Naktonggang’; designated by Glaubrecht et al., 2007], ZMB 55.624; paralectotypes ZMB 38.439 (ten pairs of valves, one right valve; ‘ Corbicula elatior jung, Naktonggang, mittl. Arm Korea, Gottsche’); paralectotypes ZMB 38.429 (six pairs of valves, one right valve; ‘ Corbicula No. 1, N’aktonggang, Gottsche’); paralectotypes ZMB 38.430 b (three pairs of valves, one tiny right valve; ‘ Corbicula no. 2, Imjingang bei Imjin 20 / 9 Gottsche’); paralectotype ZMB 55.626 (a pair of valves; specimen depicted in Fig. 6 as ‘ C. elatior? ’) (Glaubrecht et al., 2007). Type locality: ‘ Mittlerer Arm des Naktonggang, Hauptfluss der Provinz Kyöngsangdo, zwischen Tongnai und Kimhai. Imjingang bei Imjin, Provinz Kyöngkwido, Korea’; South Korea: restricted to Naktong River near Kimhae, Kyongsang-namdo (NakdongRivernearGimhae, Gyeongsangnam Prov., 35 ° 11 ′ 57 ″ N, 128 ° 54 ′ 00 ″ E). Locality of paralectotypes: Prov. Kyonggi-do: Imjin River near Imjin (Glaubrecht et al., 2007). Material examined: The type series. Our samples: Far East of Russia: lower reaches of the Lower Amur River near Nikolaevsk-on-Amur City, 53 ° 7 ′ 12.948 ″ N, 140 ° 50 ′ 47.646 E ″, 17 July 2014, 28 specimens, Bolotov I. N. & Vikhrev I. V. leg. (lot nos RMBH Corb 15); Nizhnetambovskoe settlement, 16 September 2020, 50.952778 N, 138.204444 E, 89 specimens, Aksenova O. V., Kropotin A. V., Travina O. V. & Vinarski M. V. leg.; 61 specimens are stored in RMBH; 28 specimens from the same locality are kept in LMBI under accession number 21 - 044; Nizhnetambovskoe settlement, 7 July 2021, 50.921111 N, 138.1775 E, 23 specimens, Aksenova O. V., Kropotin A. V., Travina O. V. & Khrebtova I. S. leg.; one specimen from Razdolnaya River is kept in ZIN collection, accession number 3. Description: The shell is large, triangular, and the ventral margin is rounded (Fig. 6). The umbo issignificantly prominent and slightly offset. Sculpture is represented by well-marked deep concentric ridges; the width of the ribs and distance between them are variable in the same specimen. External shell coloration varies from dark brown, greenish brown to yellow – brown, sometimes with light or dark concentric bands. Internal shell coloration varies from matte white with pale violet marks to light purple. Hinge has a heterodont dentition type, with three cardinal teeth on each valve and two crenulated lateral teeth. Siphons conical, both narrow, with a row of short papillae. Black pigment is concentrated in rings internally at the base of both siphons and in the edge of the mantle forming the siphons. The outer surface of the siphons is white; on the inside, the siphons are bright orange. The papillae of both siphons are without dark rings. Corbicula elatior is dioecious. Differential diagnosis: This species could be distinguished from the sympatric Corbicula leana, C. f l u m i n e a a n d C. j a p o n i c a b y t h e f o l l o w i n g combination of characters: larger size, significantly prominent umbo, well-marked deep concentric ridges, external shell with light or dark concentric bands, the siphons without dark pigmentation and with short papillae. Distribution: East Asia: Amur River and the Korean Peninsula (Graf & Cummings, 2021). The records of this species in the South Primorye Region (Razdolnaya River) and China (Itunghe River) (Zatravkin & Bogatov, 1987) need to be checked. Comments: Considering that the samples from the Lower Amur River are genetically identical to the samples from the Seomjin River (South Korea) collected by Park & Kim (2003), we also analysed the original descriptions of some nominal species described from South Korea: C. producta, C. elatior, C. colorata and C. papyracea. Presumably, C. producta is a junior synonym of C. fluminea (for details, see Discussion). According to their original descriptions (Heude, 1887; von Martens, 1905), C. colorata and C. papyracea are small clams with a flattened umbo. At the same time, the specimens of Corbicula from the Lower Amur River are distinguished by a relatively large shell and prominent umbo. We were unable to find any significant conchological differences between our samples of Corbicula from the Lower Amur and the type specimens of nominal species described by Bogatov & Starobogatov (1994) (Figs 6, 7). The original morphological description of C. elatior published by von Martens (1905) (Fig. 7 E) corresponds well to the morphological features of our samples from the Lower Amur River. Hence, we assume that C. elatior represents a species endemic to the Far East of Russia and the Korean Peninsula. The nominal species C. amurensis, C. neƲelskoyi and C. sirotskii are considered here junior synonyms of C. elatior.	en	Bespalaya, Yulia V., Kropotin, Alexander V., Kondakov, Alexander V., Aksenova, Olga V., Gofarov, Mikhail Yu., Kim, Sang Ki, Lee, Jin Hee, Travina, Oksana V., Vikhrev, Ilya V., Vinarski, Maxim V., Bolotov, Ivan N. (2023): A taxonomic reassessment of native and invasive species of Corbicula clams (Bivalvia: Cyrenidae) from the Russian Far East and Korea. Zoological Journal of the Linnean Society 197 (1): 104-126, DOI: 10.1093/zoolinnean/zlac078, URL: http://dx.doi.org/10.1093/zoolinnean/zlac078
