taxonID	type	description	language	source
03BE8788FFB7C759003B6ACFDE2956EC.taxon	discussion	Baeocera is a species­rich genus distributed virtually world­wide. Many members are quite common in moist forest litter and may be easily found if appropriate colleting techniques are used. Currently, 15 species are known to occur in Sabah (Löbl & Smetana 2021).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB7C75A03236BD5DC0B50D6.taxon	description	(Figs 1 ̅ 4)	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB7C75A03236BD5DC0B50D6.taxon	materials_examined	Type material. Holotype male, Sabah: Lahad Datu Ulu Segama forest Reserve, Danum Valley For. Centre, N 04 ° 57.9 ′ E 117 ° 48.1 ′, 200 m, 21. xi. 2005, D. Mann, E. Slade & J. Villanuava lgt., lowland mix. dipterocarp forest. West trail area 1, FIT, OUMNH 2006 ­ 051 (OUMNH).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB7C75A03236BD5DC0B50D6.taxon	description	Description. Length 1.28 mm, width 0.76 mm. Head and body nearly evenly reddishbrown, elytra hardly darker than pronotum, apical abdominal segments slightly lighter than pronotum, femora and tibiae as elytra, tarsi and antennae lighter. Head with interocular distance nearly as dorsoventral eye diameter. Head, thorax and elytra lacking microsculpture. Length / width ratios of antennomeres as follows: III 20 / 5: IV 20 / 6: V 30 / 7: VI 27 / 7: VII 33 / 8: VIII 27 / 7: IX 36 / 11: X: 37 / 15: XI 48 / 17. Pronotal and elytral punctation even, very fine. Lateral contours of pronotum and elytra continuously, very weakly arcuate. Pronotum with anterior margin bead broadly interrupted, lateral margins sinuate in lateral view, lateral margin striae impunctate. Margin of exposed part of scutellum rounded. Elytra with lateral margins nearly parallel, lateral margins in lateral view slightly sinuate, lateral margin carinae visible in dorsal view, lateral margin striae impunctate; sutural striae starting at base, parallel, deeply impressed, curved along pronotal lobe to form basal striae extended to humeral areas, broadly separated from lateral striae; discal punctation sparse, near base with puncture intervals about 8 to 10 larger than puncture diameters. Hind wing fully developed. Hypomera and mesanepisterna appearing impunctate. Mesoventrite with mesal ridge, punctate along lateral margins. Mesepimeron about 4 times as long as large and 5 times as long as interval to mesocoxa. Median part of metaventrite nearly flat, impunctate mesally, with dense, fairly coarse punctures on posterior third and laterally smooth mesal area, bearing conspicuous, long pubescence. Lateral parts of metaventrite with few scattered extremely fine punctures. Submesocoxal line parallel, margined with row of coarse elongate punctures extended posterior the inner part of mesepimeron. Submesocoxal area about 0.02 mm long. Metanepisterna large, about 0.10 mm wide, twice as long as wide, slightly convex, with inner margin broadly rounded anteriorly, nearly straight posteriad; inner suture impunctate. Protibiae straight. Mesotibiae and metatibiae slightly bent. Ventrite I lacking striae, with row of elongate basal punctures not interrupted in middle, extended laterally behind tip of metepimeron; punctation posterior basal row very fine and sparse. Male. Protarsomeres Ito III strongly widened, bearing long tenent setae; protarsomere I wider than apex of protibia. Ventrite VI with apical lobe subtriangular, about 0.04 mm long. Aedeagus (Figs 1 ̅ 4) 0.60 mm long. Median lobe and parameres strongly sclerotized, symmetrical, dorsal valves excepted. Articular process not prominent, distinct. Apical process of median lobe somewhat shorter than basal bulb, moderately inflexed, with weakly concave ventral margin, gradually narrowed apically, tip acute in lateral view, obtuse in dorsal view. Parameres robust, gradually narrowed apically in dorsal view, widened apical in lateral view, with broadly rounded apex. Internal sac with complex sclerites, lacking spinose and denticulate structures, with apical section of ejaculatory duct wide, weakly membranous, extruded in repos.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB7C75A03236BD5DC0B50D6.taxon	etymology	Etymology. The species epithet is a noun meaning " forest ̏ in Malay.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB7C75A03236BD5DC0B50D6.taxon	diagnosis	Differential diagnosis. This species is unique in having broadly interrupted pronotal bead. Therefore, it is placed in Baeocera tentatively (the mouth­parts and thorax of the unique available specimen were not dissected). In addition, its aedeagal characters are unusual, in particular by the structure of the internal sac and the asymmetrical valves of the median lobe. The internal sac lacking a flagellar guide sclerite and possessing a membranous ejaculatory duct extruded in repos is shared with the Chinese B. xichangana Löbl, 1999. It differs notably from B. xichangana by the not prominent articular process, the two asymmetrical dorsal valves, and the much shorter, robust parameres. Both species are also very distinct in external characters, in particular by their body­shape and size (B. xichangana is 2.0 ̅ 2.15 mm long).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB7C75A03236BD5DC0B50D6.taxon	discussion	Remarks. As to the Sabah species, only B. barbara Löbl, 1990 and B. doriae (Pic, 1920) share the wide metanepisterna. These two species have asymmetrical aedeagus and may be easily distinguished by their external characters. The antitheses 7 and 8 have been inversed in key to the Sabah Baeocera given by LÖBL & SMETANA (2021): the thesis 6 (Metanepisternum fused, ...) should lead to ̤ 8.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB4C75A03C76E1DDBDA53DC.taxon	materials_examined	Material examined. 1 male, Sabah, Mt. Kinabalu N. P., 1700 m, 16. vii. 02, Kurbatov & Zimina (MHNG); 1 male, Sabah, rd Kota Kinabalu ̅ Tambunan, km 52, 1600 ̅ 1800 m, 2 ̅ 5. vii. 02, Kurbatov & Zimina (MHNG).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB4C75A03C76E1DDBDA53DC.taxon	discussion	Remarks. This species is known from Sarawak and Sabah only. It is poorly represented in collections, a single specimen has been reported from each of these states.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB5C75B0323699DDC185722.taxon	materials_examined	Material examined. 1 male, Sabah, Lahad Datu Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04 ° 57 ′ E 117 ° 48 ′, 450 m, 1 ̅ 5. ix. 2005, Slade & Villanuava lgt., lowland mixed dipterocarp forest. Nature trail area 1, FIT, OUMNH (OUMNH).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB5C75B0323699DDC185722.taxon	discussion	Remarks. The species is widely distributed in Sabah and seems to be common in samples of forest floor litter (see LÖBL 2021: 28).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB5C75B03236A9FDC165620.taxon	materials_examined	Material examined. 1 male, Sabah, Lahad Datu Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04 ° 58 ′ E 117 ° 42 ′, 450 m, viii. 1998, D. Bebber lgt., lowland mixed dipterocarp forest, FIT, OUMNH 2003 ­ 050 (OUMNH).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB5C75B03236A9FDC165620.taxon	discussion	Remarks. As B. incisa, this species is widely distributed in Sabah and seems to be common in forest floor litter (see LÖBL 2021: 31).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB5C75B03236C5FDD1E50F5.taxon	materials_examined	Material examined. 1, Sabah, Lahat Datu, Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04 ° 57.9 ′ E 117 ° 48.1 ′, 200 m, 27 ̅ 28. x. 2005, primary forest, D. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, West trail 10 FIT, OUMNH 2006 ­ 051 (OUMNH); 1, with the same data but 21. X. 2005, West trail area (MHNG); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04 ° 59.738 ′ E 117 ° 50.165 ′, 250 m, iii ̅ iv. 2005, FIT no 2, primary forest, Slade & J. Villanuava lgt. lowland mixed dipterocarp forest, Yayasan Sabah Logging Concession OUMNH 2005 ­ 062 (OUMNH).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB5C75B03236C5FDD1E50F5.taxon	discussion	Remarks. The species is known only from East Malaysia (LÖBL et al. 2020). It was based on specimens found in the Gunung Mulu National Park in Sarawak and subsequently recorted from the Mount Kinabalu National Park, the vicinity of Tambunan and the Batu Punggul Resort in Sabah.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB5C75B00D76B91DD9E5162.taxon	discussion	The Asian members Bironium have been revised by LÖBL et al. (2020). Currently, three species are known to occur in Sabah: B. borneense Löbl, 1987, B. minutum (Achard, 1920), and B. pustulatum Löbl, 2011.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB5C75B002C6DC6DBFB5337.taxon	discussion	The genus is monospecific. It was established to accommodate a species possessing characters typical for the termitophilous members of the Baeoceridium group (see LESCHEN & LÖBL 2005) and thus is assumed to live on fungus gardens of termites.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB5C75C03236E83DD195704.taxon	materials_examined	Material examined. 184, Sabah, Batu Punggul Resort env., 24. vi. ̅ 1. vii. 1996, FIT (MHNG); 1, with the same data but in debris accumulates around large trees near river (MHNG); 1, Sabah, Lahat Datu, Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04 ° 57.9 ′ E 117 ° 48.1 ′, 200 m, 22 ̅ 23. x. 2005, primary forest, D. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, west trail area, FIT, OUMNH 2006 ­ 051 (OUMNH); 1, with the same data but 3 ̅ 13. xi., D. J. Mann & E. Slade lgt ,, forest natural trail area OUMNH 2065 ­ 051 (OUMNH); 1, Sabah, Lahat Datu, Ulu Segama Forest Reserve, Danum Valley Conservation Area, Borneo Rain Forest Lodge, N 05 ° 02.682 ′ E 117 ° 45.553 ′, 250 m, iii ̅ iv. 2005, E. Slade & J. Villaneuve lgt., FIT, primary dipterocarp forest, OUMNH 2065 ­ 062 (OUMNH).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB5C75C03236E83DD195704.taxon	discussion	Remarks. The currently known specimens of this species are the syntypes from east Kalimantan, Indonesia.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB2C75C00D56ABDDDB356FE.taxon	discussion	The genus is Asian in distribution, with most members found in the tropics and subtropics, remaining unknown south of the Wallace line. Currently, six species are known to occur in Sabah (LÖBL & TANG 2013): P. convexum Löbl & Tang, 2013, P. horaki Löbl & Tang, 2013, P. pubiventer Löbl & Tang, 2013, P. sparsepunctatum (Pic, 1915), P. subovatum Pic, 1920, and P. vitalisi (Achard, 1920). Anew record is given, and two additional species are described below.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB2C75C03C76BDADDDF519C.taxon	materials_examined	Material examined. 1 male, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04 ° 59.738 ′ E 117 ° 50.165 ′, 250 m, iii ̅ iv. 2005, FIT no 2, primary forest, E. Slade & J. Villanuava lgt. Lowland mixed dipterocarp forest, Yayasan Sabah Logging Concession OUMNH 2005 ­ 062 (OUMNH).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB2C75C03C76BDADDDF519C.taxon	discussion	Remarks. The species seems to be rather widely distributed. It is currently known from Indonesia (Sumatra and Kalimantan), Malaysia (West Malaysia, Sabah and Sarawak), and Thailand, though only a single Sabah record is published from the Gunung Emas in the Crocket Range (LÖBL & TANG 2013). The species may be easily distinguished by the long protibial setae in males.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB2C75F03C76D24DEE750BE.taxon	description	(Figs 5 ̅ 7)	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB2C75F03C76D24DEE750BE.taxon	materials_examined	Type material. Holotype male, SABAH: Poring Hot Springs 600 m 9. V. 1987 Burckhardt ̅ Löbl (MHNG); Paratypes: 1 female, SABAH: Crocker Ra. 1350 m, km 60 Kota Kinabalu­Tambunan 17. V. 1987 Burckhardt ̅ Löbl; female, BORNEO SABAH Tamis, Hwy AI 10 km NW Kinabalu Park Entr. 1100 m 24. V. 87 A. Smetana; 1 female, SABAH: E Mt. Kinabalu 1150 m, rte Ranau­Kota Kinabalu 24. V. 1987 Burckhardt ̅ Löbl (all in MHNG).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB2C75F03C76D24DEE750BE.taxon	description	Description. Length 1.78 ̅ 1.87 mm, width 1.20 ̅ 1.35 mm. Head and most of body dark reddish­brown, metaventrite somewhat darkened, apex of abdomen lighter, yellowish, femora and tibiae roughly as body, tarsi and antennae lighter, yellowish. Maxillary palpi with palpomere IV narrow, tapering, about 4 times as long as wide. Length ratio of antennomeres as follows: III 14: IV 25: V 25: VI 22: VII 26: VIII 26: IX 26: X: 26: XI 31; antennomere XI about 4 times as long as wide. Pronotal punctation about as fine as that on frons, with punctures very shallow and not well delimited; middle of pronotal disc with puncture intervals about 5 to 10 times larger than puncture diameters. Elytra lacking lateral impressions, not flattened or impressed apically, lacking humps; discal punctation on prevailing surface similar to pronotal punctation, conspicuous patch of rather coarse punctures present on anterolateral area. Puncture patch separated from lateral and basal margins by rather broad, very finely punctate areas. Lateral striae coarsely punctate. Hypomera impunctate, flattened along basal margin. Mesoventrite lacking mesal ridge, with distinct setiferous punctures at lateral margins of coxal process, setae long. Mesanepisternum very finely punctate, lacking microsculpture. Metaventrite with inconspicuous striate to punctulate microsculpture; median area nearly flat, lacking impressions and lacking tubercle, its punctation fine, arranged to form two sparse admesal rows joined posteriorly. Lateral parts of metaventrite very finely and sparsely punctate. Metacoxal process impunctate, inflexed. Submesocoxal area slightly shorter than half of shortest interval to metacoxa. Submesocoxal line coarsely punctate. Mesotibiae and metatibiae slightly bent, ventral spur of mesotibiae straight. Exposed ventrites and tergites with distinct punctulate microsculpture, setiferous punctures very fine and scattered. Male. Protarsus shorter than protibia, protarsomeres Ito III hardly widened, narrower than apex of tibiae, bearing short setae. Aedeagus (Figs 5 ̅ 7) 0.68 mm long, weakly sclerotized. Median lobe with basal bulb shorter than apical process, the latter hardly inflexed, with sinuate ventral margin (lateral view) and blunt tip; apical part of median lobe overlapped by two valves. Parameres widened basally, weakly bent. Internal sac lacking sclerotized pieces, without distinct flagellum, bearing apical spinose structures.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB2C75F03C76D24DEE750BE.taxon	etymology	Etymology. The species epithet is a Latin adjective meaning " irregulaȑ, referring to the elytral punctation.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB2C75F03C76D24DEE750BE.taxon	diagnosis	Differential diagnosis. This species may be readily distinguished from its congeners, P. impressipenne Löbl, 1973 and P. subglabrum Löbl, 1990 excepted, by elytra bearing each an anterolateral patch of coarse punctures. In addition, its small body­size and the structure of the internal sac are diagnostic. It may be readily distinguished from P. impressipenne and P. subglabrum by the parameres widened basally. It falls to the couplet 25 in the key to the species in LÖBL & TANG (2013), to P. feai Pic, though the latter species is 2.90 ̅ 3.20 mm long and has a black body. The aedeagal characters suggest relationships with members of the P. plagiferum group. It may be distinguished from the three species placed in that group (P. javanum Löbl & Tang, 2013, P. plagiferum Löbl, 1980, and P. spinipes Löbl & Tang, 2013) by the shape of the parameres and apex of the median lobe.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB1C75003236E1ADEE850B9.taxon	description	(Figs 8 ̅ 10)	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB1C75003236E1ADEE850B9.taxon	materials_examined	Type material. Holotype male, Malaysia, Sabah, Crocker Range, Mawar Waterfall env. 17. VI. 1996, 9 c, vegetation debris and forest floor litter accumulated along large fallen trees near river (MHNG).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB1C75003236E1ADEE850B9.taxon	description	Description. Length 1.95 mm, width 1.30 mm. Head and most of body dark reddishbrown, apex of abdomen lighter, yellowish; femora and tibiae roughly as body, tarsi and antennae light brown. Maxillary palpi with palpomere IV narrow, tapering, about 4 times as long as wide. Length ratio of antennomeres as follows: III 15: IV 22: V 26: VI 21: VII 26: VIII 26: IX 27: X: 26: XI 35; antennomere XI about 3.5 times as long as wide. Pronotal punctation very fine, similar to that on frons, with punctures very shallow and not well delimited; middle of pronotal disc with puncture intervals about 4 to 10 times larger than puncture diameters. Elytra lacking lateral impressions, not flattened or impressed apically, lacking humps; discal punctation similar to pronotal. Hypomera impunctate, impressed posteriad. Mesoventrite lacking mesal ridge and lacking distinct setiferous punctures. Mesanepisternum very finely punctate, lacking microsculpture. Metaventrite lacking microsculpture; median area nearly flat, lacking impressions and tubercle, with two irregular rows of distinct admesal punctures. Lateral parts of metaventrite very finely and sparsely punctate. Metacoxal process impunctate, horizontal. Submesocoxal area short, about as fourth shortest interval to metacoxa. Submesocoxal line finely punctate. Mesotibiae and metatibiae slightly bent, ventral spur of mesotibiae straight. Ventrite Ilacking microsculpture, with very fine and scattered punctures; following ventrites and exposed tergites with punctulate microsculpture. Male. Aedeagus (Figs 8 ̅ 10) 0.74 mm long. Median lobe with basal bulb shorter than apical process, the latter slightly inflexed, with sinuate ventral margin (lateral view), and blunt tip; apical part of median lobe lacking distinct valves. Parameres nearly evenly wide, bent in dorsal view, straight in lateral view. Internal sac with pair of strongly sclerotized denticles and mesal sclerite, lacking flagellum and lacking apical spinose structures.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB1C75003236E1ADEE850B9.taxon	etymology	Etymology. The species epithet is a Latin noun, referring to the canine­like denticles of the internal sac.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB1C75003236E1ADEE850B9.taxon	diagnosis	Differential diagnosis. The species is as P. irregulare less than 2 mm long and thus conspicuous by its small body­size. It may be easily distinguished from P. irregulare by the evenly very fine elytral punctation, the narrow submesocoxal areas and ventrite I lacking microsculpture. The aedeagal characters suggest relationship with members of the P. subovatum group (see LÖBL & TANG 2013), though the shape of the admesal sclerites of the internal sac is unique. The species falls in the key species to couplet 27, to P. schuhi Löbl & Tang, 2013 from which it differs notably by the parameres evenly narrow, except at their basis.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFB1C75003236E1ADEE850B9.taxon	discussion	Remarks. The single available specimen has its forelegs broken, probably lost when collected.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBEC75000E96E35DA5C53C6.taxon	discussion	This genus encompasses three Southeast Asian species presumably associated with termites. Currently, only one of them has been reported from Sabah. Unlike most other Scaphisomatini, these species are efficiently collected only by using flight intercept traps.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBFC7510323699DDEC75015.taxon	materials_examined	Material examined. 1, Sabah, Mt. Kinabalu Nat. Park, above Pring Hot Springs, 520 m, 15. viii. 1988, A. Smetana (MHNG); 70, Sabah, Batu Punggul Resort env., 24. vi. ̅ 1. vii. 1996, 11 c, FIT; 2, Sabah, Mt. Kinabalu Nat. Park, above Poring Hot Springs, 520 m, 15. viii. 1988, A. Smetana (MHNG); 4, Sabah, Danum Valley, 14. ̅ 16. ii. 1968, FIT, G. de Rougemont (MHNG); 1, Sabah, Lahad Datu, Ulu Segama, Danum Valley Forest Centre, N 04 ° 57.9 ′ E 117 ° 48.1 ′, 200 m, 1 ̅ 23. iii. 2005, primary forest, D. J. Mann & E. Slade lgt., lowland mixed dipterocarp forest, west trail area, first inferior FIT, OUMNH­ 2006 ­ 051 (OUMNH); 8, Sabah, Lahad Datu, Ulu Segama, Danum Valley Forest Centre, N 04 ° 57 ′ E 117 ° 48 ′, 450 m, 1 ̅ 5. xi. 2005, FIT, primary forest, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, Nature trail area 1 (OUMNH); 2, Sabah, Lahad Datu, Ulu Segama, Danum Valley Forest Centre, N 04 ° 57 ′ E 117 ° 48 ′, 450 m, 22 ̅ 23. xi. 2005, primary forest, D. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, west trail area first inferior FIT, OUMNH 2006 ­ 051 (OUMNH); 3, Sabah, Lahad Datu, Ulu Segama, Danum Valley Forest Centre, N 04 ° 57.9 ′ E 117 ° 48.1 ′, 200 m, 27 ̅ 28. x. 2005, primary forest, D. J. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, West trail area, first interior FIT, OUMNH 2006 ­ 051 (OUMNH); 6, Sabah, Lahad Datu, Ulu Segama, Danum Valley Forest Centre, N 04 ° 57.9 ′ E 117 ° 48.1 ′, 200 m, 22 ̅ 23. ix. 2005, primary forest, D. J. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, nature reserve area, first inferior FIT, OUMNH­ 2006 ­ 051 (OUMNH); 1, Sabah, Lahad Datu, Ulu Segama, Danum Valley Forest Centre, N 04 ° 57.9 ′ E 117 ° 48 ′, 200 m, 3 ̅ 13. ix. 2005, primary forest, D. J. Mann, E. Slade & J. Villanuava lgt., lowland mixed dipterocarp forest, West trail area, FIT, OUMNH 2006 ­ 05 (OUMNH); 5, Sabah, Lahad Datu, Ulu Segama, Danum Valley Conservation Area, Borneo Rain Forest Lodge, N 05 ° 57.682 ′ E 117 ° 45.553 ′, 250 m, iii ̅ iv. 2005, FIT, primary dipterocarp forest, E. Slade & J. Villanuava lgt., OUMNH 2005 ­ 062 (OUMNH); 36, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04 ° 59.738 ′ E 117 ° 50.165 ′, 250 m, iii ̅ iv. 2005, FIT no 2, selective lowland mixed logged dipterocarp forest, Yayasan Sabah Logging Concession, E. Slade & J. Villanuava lgt., OUMNH­ 2005 ­ 062 (OUMNH); 4, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 81 logging area, N 04 ° 58.660 ′ E 117 ° 53.410 ′, 250 m, iii. 2005, secondary forest, FIT, selectively logged dipterocarp forest, Yayasan Sabah Logging Concession, E. Slade & J. Villanuava lgt., OUMNH­ 2005 ­ 062 (OUMNH); 3, Sabah, Lahad Datu, Ulu Segama, Danum Valley Forest Centre, N 04 ° 58 ′ E 117 ° 42 ′, 450 m, viii. 1998, FIT, lowland mixed dipterocarp forest, D. Bebber lgt., OUMNH 2003 ­ 050 (OUMNH).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBFC7510323699DDEC75015.taxon	discussion	Remarks. This species appears to be widely distributed and has been reported from China: Yunnan; Indonesia: Lombok, Sumatra; Philippines: Palawan; Thailand; Vietnam (see LÖBL 2018). The description of Scaphisoma bugiamapi suggest its synonymy with Sapitia lombokiana, notably the illustration of the aedeagus, and the statement about striate elytra (based on a misintepretation of the elytral pubescence). Additional information provided by M. Prokofiev (personal communication) convinced the author to publish it.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBFC75103236DAEDC50539E.taxon	materials_examined	Material examined. 474, Sabah, Batu Punggul Resort env., 24. vi. ̅ 1. vii. 1996, 11 c, FIT (MHNG); 3, Sabah, Danum Valley, 14 ̅ 16. ii. 1968, G. de Rougemont (MHNG); 1, Sabah, Poring Hot Springs, 500 m, 11. v. 1987, Burckhard & Löbl (MHNG); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Danum Valley Conservation Area, Borneo Rain Forest Lodge, N 05 ° 02.682 ′ E 117 ° 45.553 ′, 250 m, iii ̅ iv. 2005, FIT, primary dipterocarp forest, E. Slade & J. Villanuava lgt. OUMNH 2055 ­ 062 (OUMNH).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBFC75103236DAEDC50539E.taxon	discussion	Remarks. Thie species appears to be widely distributed and has been reported from Indonesia: Sumatra; East Malaysia: Sabah and Thailand (see LÖBL 2018). It is very similar to S. sumatrana Löbl, 1978 while it differs notably from S. lombokina by the nearly fused mesepimera and the stoud legs.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBCC7520033699CDDC1577E.taxon	discussion	The genus comprises 19 Afrotropical, Oriental, Melanesian and Australian species. Several, described by J. Achard and M. Pick, still lack revisions. Asingle species is present in samples from Sabah.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBCC75203C76A5ADBBA56B1.taxon	materials_examined	Material examined. 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Danum Valley Forest Centre, N 04 ° 57.9 ′ E 117 ° 48.1 ′, 450 m, 22 ̅ 23. xi. 2005, primary dipterocarp forest, west trail area, first inferior FIT, D. Mann, E. Slade & J. Villanuava lgt., OUMNH 2006 ­ 051 (OUMNH); 1, Sabah, Lahad Datu, Ulu Segama Forest Reserve, Coupe 88 logging area, N 04 ° 59.738 ′ E 117 ° 50.165 ′, 250 m, iii ̅ iv. 2005, FIT no 2, E. Slade & J. Villanuava lgt., selectively logged dipterocarp forest, Yayasan Sabah Logging Concesion, OUMNH 2005 ­ 052 (MHNG).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBCC75203C76A5ADBBA56B1.taxon	discussion	Remarks. These specimens exhibit characters congruent with the redescription given by LÖBL & LESCHEN (2010). Notable are the widely interrupted basal striae of the elytra, the mesoventrite with distinct mesal ridge, the lateral parts of the metaventrite lacking coarser punctures, and the lateral parts of ventrite I, each bearing a subbasal patch of distinct punctures, in combination with the uniformly rufous, about 1.9 mm long body. The species is very similar to S. arcuatum Champion, 1927 and S. pulex Heller, 1917.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBCC75200E66C09DD0951AB.taxon	discussion	Four species of Scaphoxium are currently known to occur in Sabah, S. opacum Löbl, 2021, S. opertum Löbl, 2021, S. taylori Löbl, 1981, and one which remains undescribed because its male characters are unknown. The newly studied collections yield a fifth species described below.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBCC75303C76D17DDD75121.taxon	description	Figs 11 ̅ 13	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBCC75303C76D17DDD75121.taxon	materials_examined	Type material. Holotype male, Malaysia, Sabah, Batu Punggul Resort env., 24. VI. ̅ 1. VII. 1996, 11 c. vegetation debris and forest floor litter accumulated around large trees near river (MHNG).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBCC75303C76D17DDD75121.taxon	description	Description. Length 1.18 mm, width 0.55 mm, dorsoventral diameter 0.62 mm. Thorax, elytra and most of abdomen uniformly reddish­brown, head somewhat lighter, apical abdominal segments light, nearly yellowish, femora and tibiae roughly as thorax, tarsi lighter, antennae light brown. Length / width ratios of antennomeres as follows: III 22 / 5: IV 17 / 4: V 22 / 4: VI 23 / 4: VII 28 / 7: VIII 20 / 7: IX 28 / 11: X: 28 / 11: XI 38 / 12. Pronotal and elytral punctation similar, very fine, hardly visible at 40 times magnification. Scutellum concealed. Elytra with sutural striae parallel, shortened, starting about 0.15 mm posterior of margin of pronotal lobe, adsutural areas hardly raised. Hypomeron with scattered, very fine punctured and short oblique stria. Mesoventrite with mesal impression shallow, narrowed posteriad, lacking mesal ridge and lacking microsculpture. Margin of mesocoxal process rounded. Metaventrite lacking microsculpture, very finely punctate, convex in middle. Lateral stria oblique, nearly reaching convex, finely punctate submesocoxal line. Submesocoxal area about 0.03 mm, as long as half of the shortest interval to margin of metacoxa. Metanepisternum flat, with suture fairly narrow, punctate, somewhat rounded. Abdomen very finely punctate, median areas of ventrites with punctulate microsculpture. Male. Segments Ito III of protarsi hardly widened. Aedeagus (Figs 11 ̅ 13) 0.44 mm long. Parameres narrowed posterior of basal section, gradually weakly widened toward narrow apical section, not forming subapical lobe or process; outer margins of parameres concave and inner margins strongly sclerotized posterior of narrowed apical section. Internal sac bearing two pairs of sclerotized pieces, outer larger and more sclerotized than inner pair, expanded apically, inner pair widened proximally, with margins becoming indistinct apically; scale­like, spine­like or filamentous structures absent.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBCC75303C76D17DDD75121.taxon	etymology	Etymology. The species epithet is a Latin adjective meaning " sinuatȅ, referring to the outer margins of the parameres.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBCC75303C76D17DDD75121.taxon	diagnosis	Differential diagnosis. The body­size of this new species is significantly smaller than that of other Bornean species and the shape of its parameres is diagnostic for it. The new species is similar to S. bilobum Löbl, 2015 from Lombok. It shares with the latter the simple internal sac of the aedeagus lacking spine­like, denticulate or striate membranous structures in combination with the presence of two pairs of admesal sclerites. However, the outer pair of sclerites is much more expanded and the inner sclerites are much shorter and broader in S. sinuatum, which may be distinguished from B. bilobum also by its wide impression of the mesoventrite, finer punctation of the lateral parts of the metaventrite and shorter submesocoxal areas.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBDC75303236D21DBB853D1.taxon	discussion	Remarks. The species was based on several specimens found in North Sumatra, in fungus gardens of Odontotermes takensis Ahmad, 1965. It was not reported after its description while V. termitophila Champion, 1927 is widely distributed and has been referred to many times (see OGAWA & MAETO 2015). The colour pattern of the abdomen used as one of the diagnostic characters proves to be unreliable in the examined material.	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
03BE8788FFBDC75300C56C9BDC765199.taxon	discussion	The genus encompasses 31 species, all Afrotropical but the Asian V. termitophila (Champion, 1927) and V. kistneri (Löbl, 1979).	en	Löbl, Ivan (2022): Supplement to the knowledge of Scaphisomatini of Sabah, Malaysia (Coleoptera: Staphylinidae: Scaphidiinae). Acta Musei Moraviae, Scientiae biologicae 107 (1 - 2): 7-19, DOI: http://doi.org/10.5281/zenodo.7501705
