identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03AD87F6FFC9E47FFBF4AD10207BFE6C.text	03AD87F6FFC9E47FFBF4AD10207BFE6C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quantula Baker 1941	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Quantula Baker, 1941</p>
            <p> Type species.  Nanina striata Gray, 1834</p>
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	https://treatment.plazi.org/id/03AD87F6FFC9E47FFBF4AD10207BFE6C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Deeprom, Kanchanog;Tumpeesuwan, Chanidaporn;Tumpeesuwan, Sakboworn	Deeprom, Kanchanog, Tumpeesuwan, Chanidaporn, Tumpeesuwan, Sakboworn (2022): A new species of the genus Quantula Baker, 1941 (Eupulmonata: Dyakiidae) from the southern part of north-eastern Thailand. Raffles Bulletin of Zoology 70: 519-533, DOI: 10.26107/RBZ-2022-0029
03AD87F6FFC9E473FC97AD902410F842.text	03AD87F6FFC9E473FC97AD902410F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quantula weinkauffiana (Crosse & Fischer 1863)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Quantula weinkauffiana (Crosse &amp; Fischer, 1863)</p>
            <p>(Figs. 3A, 4A, 5A–G; Tables 1 –3)</p>
            <p> Helix weinkauffiana Crosse &amp; Fischer, 1863: 350–351 ; Crosse &amp; Fischer, 1864: 326, pl. 12, fig. 7. </p>
            <p> Hemiplecta weinkauffiana – Morlet, 1889: 124; Dautzenberg &amp; Fischer, 1905: 349; Fischer, 1973: 92; Panha, 1996: 34; Panha &amp; Thanamitramanee, 1997: 2. </p>
            <p> Ariophanta (Hemiplecta) weinkauffiana – Fischer, 1891: 21. </p>
            <p> Ariophanta weinkauffiana inflata Möllendorf, 1894: 149 . </p>
            <p> Nanina (Xestina) weinkauffiana – Kobelt, 1900: 984–985, pl. 255, figs. 6–8. </p>
            <p> Nanina (Hemiplecta) weinkauffiana var. minor Fischer &amp; Dautzenberg, 1904: 394 . </p>
            <p> Dyakia weinkauffiana – Maneevong, 2000: 46–47, figs. 4–11 (A–B). </p>
            <p> Ariophanta (Cryptozona) weinkauffiana – Schileyko, 2011: 29–30. </p>
            <p> Quantula weinkauffiana – Hemmen &amp; Hemmen, 2001: 45; Nabhitabhata, 2009: 249; Jumlong, 2012: 51,72, figs. 4.10, 5.1; Jumlong et al., 2013: 67–68, 71–76, 79, fig. 2D; Worachak, 2014: 40–43, figs. 4.1–4.3; Worachak et al., 2014: 524–529, fig. 2A; Inkhavilay et al., 2019: 71, 119 figs. 32B–D, 55H; Sutcharit et al., 2020b: 21, 23, figs. 9E–F; Jirapatrasilp et al., 2021: 256–257, 261–263, 268–269. </p>
            <p> Quantula sp. 2 – Jumlong, 2012: 53, 72, figs. 4.12, 5.1; Worachak, 2014: 45–48, figs. 4.7, 4.8A–E, 4.9A–B </p>
            <p> Quantula sp. 5 – Hoompuay, 2016: 40–42, figs. 4.18–4.20 </p>
            <p> Quantula weinkauffiana inflata – BEDO, 2017: 221. </p>
            <p> Quantula weinkauffiana weinkauffiana – BEDO, 2017: 221. </p>
            <p> Type locality. Cochinchina (Southern Vietnam) . </p>
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                 Material examined.   37 shells and seven living specimens NHMSU-00035;  
                <a title="Search Plazi for locations around (long 103.37139/lat 14.768139)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.37139&amp;materialsCitation.latitude=14.768139">Khao Panom Sawai</a>
                 , Surin Province; 14°46′05.3″N 103°22′17.0″E; 14 October 2017; (Figs. 3A, 4A, 5A–E)  . 
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            <p>Diagnosis. Shell depressed-conic. Last whorl angular with sharp peripheral keel and descending in front, lip is rather thick and slightly descending (see Inkhavilay et al., 2019: syntype fig. 32B–D; Jirapatrasilp et al., 2021: Fig. 3). Amatorial organ duct shorter than amatorial organ. Gametolytic duct connected to middle part of amatorial organ.</p>
            <p>Description. Shell (n= 37) (Fig. 4A): dextral, very depressed conic, brown colour with 5–5¼ whorls, suture shallow, growth line fine. Last whorl angular with strong peripheral keel descending in front. Shell height 10.7–15.6 mm, shell width 20.6–28.1 mm. Apex blunt, peripheral keel very sharp, colour of ventral side of shell is paler than that of dorsal side. Apertural lip simple in light brown, with margin slightly thickened in adult specimens. Umbilicus narrow and deep. Body (n=5) (Fig. 3A): Living snail light orange, with two lateral dark brown stripes running from each of upper tentacle backward to pneumostome.</p>
            <p>Genitalia (n=5) (Fig. 5A–E): Penis short, cylindrical. Epiphallus shorter and slender than penis. Inner wall of penis with three very short and thin longitudinal pilasters near atrium, with other regions smooth (Fig. 5D). Vas deferens short. Penial retractor muscle absent. Gametolytic sac short with gametolytic duct connecting to middle part of amatorial organ. Amatorial organ cylindrical, gradually larger at distal portion. Inner part of this portion, without longitudinal pilaster, but, with short amatorial organ papilla, which is robust, cylindrical and not exceeding half of distance between distal end of amatorial organ and junction of gametolytic duct. Orange papilla spike present (Fig. 5B, C). Inner wall of distal part with transverse lamella or nearly smooth surface, whereas, in proximal part before junction of gametolytic duct with two longitudinal pilasters (Fig. 5C). Amatorial organ duct shorter than amatorial organ. Vagina longer than penis. Surface of inner wall of vagina undulated, proximally with thin transverse lamellae (Fig. 5E).</p>
            <p>Radula (n=5) (Fig. 5F–G): There are approximately 78 rows of teeth, each of which comprises 73–173 teeth, formula (19–62) + (17–24) + 1 + (17–24) + (19–62). Central tooth tricuspid, lanceolate, smaller than lateral teeth. Lateral teeth tricuspid, oblique lanceolate. Marginal teeth unicuspid, sword shaped.</p>
            <p> Remarks. Images published in Inkhavilay et al. (2019) included the syntype, MNHN-IM-2000-27780 (Fig. 32B), specimens from Savannakhet (Figs. 32C, 55H), and specimens from Champasak (Fig. 32D). The type specimen possesses a sharp peripheral keel and thickened peristome. These characters resemble specimens from Phanom Sawai and in Schileyko (2003: Fig. 1776A), which illustrated the drawing of a shell identified as “  Quantula striata ”. According to the shell morphology and distribution, the shell from Southern Vietnam illustrated in Schileyko (2003) should be identified as  Quantula weinkauffiana . However, “  Ariophanta weinkauffiana inflata ” and “  Nanina (Hemiplecta) weinkauffiana minor ”, which were described from localities outside north-eastern Thailand, need clarification of their taxonomic status via further study. From the result of molecular phylogeny of  Dyakiidae by Jirapatrasilp et al. (2021),  Quantula weinkauffiana individuals from Khao Panom Sawai, Surin (Thailand), Champasak (Laos), and Kampot (Cambodia) were in the same clade with high support (BI= 100%). Considering the result from this study, the intraspecific genetic distances among the  Quantula weinkauffiana populations of Laos, Cambodia, and Phanom Sawai were low, as they ranged from 0.2–1.1%, while within the Panom Sawai population, it was 0.2% (Table 2), and the  Quantula weinkauffiana clade obtained strong support (Fig. 2) (BI posterior probability of 1.00 and 86% for ML, and 89% for NJ bootstrap replicate). </p>
            <p>Distribution. Southern Vietnam, Cambodia, Southern Laos, and North-eastern Thailand (Crosse &amp; Fischer, 1863; Schileyko, 2011; Inkhavilay et al., 2019; Sutcharit et al., 2020b). In Thailand, this species is currently recorded from Khao Panom Sawai, Surin Province (Jirapatrasilp et al., 2021 and this study).</p>
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	https://treatment.plazi.org/id/03AD87F6FFC9E473FC97AD902410F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Deeprom, Kanchanog;Tumpeesuwan, Chanidaporn;Tumpeesuwan, Sakboworn	Deeprom, Kanchanog, Tumpeesuwan, Chanidaporn, Tumpeesuwan, Sakboworn (2022): A new species of the genus Quantula Baker, 1941 (Eupulmonata: Dyakiidae) from the southern part of north-eastern Thailand. Raffles Bulletin of Zoology 70: 519-533, DOI: 10.26107/RBZ-2022-0029
03AD87F6FFC5E470FC10AADB2129F8BA.text	03AD87F6FFC5E470FC10AADB2129F8BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quantula doma Deeprom & Tumpeesuwan & Tumpeesuwan 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Quantula doma ,  new species</p>
            <p>(Figs. 3B–C, 4B–C, 6A–J; Tables 1 –3)</p>
            <p>Quantula sp. – Jumlong et al., 2013: 71–72, 74–76, 78–79, fig. 3.</p>
            <p> Quantula weinkauffiana – Worachak, 2014: 40–42, figs. 4.1A–B, 4.2 A–C, 4.3 B–C. </p>
            <p> Quantula sp. 1 – Worachak, 2014: 43–45, figs. 4.4A, 4.5A–E, 4.6A–B. </p>
            <p> Quantula sp. 1 – Jirapatrasilp et al., 2021: 193 (1), 262–263.  Quantula striata – Hoompuay, 2016: 22–25, figs. 4.1–4.3.  Quantula sp. 6 – Hoompuay, 2016: 43–45, figs. 4.21–4.24. </p>
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                 Material examined.   Holotype: NHMSU-00048 (Fig. 4B) from Khao Sala,  
                <a title="Search Plazi for locations around (long 103.93767/lat 14.430695)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.93767&amp;materialsCitation.latitude=14.430695">Buachet District</a>
                 , Surin Province; 14°25′50.5″N 103°56′15.6″E; 14 October 2017; SH = 18.69 mm, SW = 28.61 mm  .  Paratypes: 47 shells (25 adults &amp; 22 juveniles), 28 living specimens; same data as holotype; NHMSU-00049.  
                <a title="Search Plazi for locations around (long 104.32594/lat 16.322863)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.32594&amp;materialsCitation.latitude=16.322863">One</a>
                 shell (NHMSU-00050) from  
                <a title="Search Plazi for locations around (long 104.32594/lat 16.322863)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.32594&amp;materialsCitation.latitude=16.322863">Phu Jor Kor</a>
                 ,  
                <a title="Search Plazi for locations around (long 104.32594/lat 16.322863)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.32594&amp;materialsCitation.latitude=16.322863">Nong Soong District</a>
                 , Mukdahan Province; (16°22′22.1″N, 104°22′32.2″E), 11 October 2017 (Fig. 4C);   
                <a title="Search Plazi for locations around (long 104.32594/lat 16.322863)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.32594&amp;materialsCitation.latitude=16.322863">Two</a>
                 adult and three juvenile shells, and nine living specimens (NHMSU-00051) same locality as NHMSU-00050;   
                <a title="Search Plazi for locations around (long 104.32594/lat 16.322863)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=104.32594&amp;materialsCitation.latitude=16.322863">One</a>
                 adult and one juvenile shells and two living specimens (NHMSU-00052); Pha Nam Yoi, Nong Phok District, Roi Et Province, Thailand; 16°19′22.3″N 104°19′33.4″E; 11 October 2017;  seven adult and three juvenile shells, and five living specimens (NHMSU-00053), same data with NHMSU-00052 . 
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            <p>Diagnosis. Last whorl angular with peripheral keel, not descending in front. Amatorial organ duct longer than amatorial organ.</p>
            <p>Etymology. The specific epithet is derived from the Latin word ‘domus’ meaning roof, referring to the fact that its shell shape is similar to a hemispherical roof.</p>
            <p> Description. Shell (n=48, 26 adult &amp; 22 juveniles from Khao Sala (Fig. 4B) and n=20, 11 adults &amp; 9 juveniles from Phu Jor Kor and Pha Nam Yoi (Fig. 4C )): Dextral, domeshaped to depressed-conic lenticular, with 6–6¼ whorls, suture shallow, growth line prominent. Last whorl angular with peripheral keel, but less sharp than  Q. wienkauffiana . Shell height 14.8–21.9 mm, shell width 25.1–35.3 mm. Apex blunt, shell light brown, ventral side paler than dorsal side. Apertural lip simple and thick, and slightly expanded at columellar side in adult specimens. Umbilicus narrow and deep. </p>
            <p>Body (n=28 from Khao Sala (Fig. 3B) and n = 11 from Phu Jor Kor and Pha Nam Yoi (Fig. 3C )): Living snail orange brown, with two lateral light brown or dark stripes running from base of each upper tentacle backward to pneumostome.</p>
            <p>Genitalia (n=13, comprising three specimens from Khao Sala (one specimen from Hoompuay, 2016) (Fig. 6A–E), three specimens from Phu Jor Kor, and seven specimens from Pha Nam Yoi (five specimens from Hoompuay, 2016) (Fig. 6F–J): Penis short, cylindrical, inner wall with robust longitudinal penial pilaster alternating with 1–2 thin longitudinal pilasters (Fig. 6D), whereas specimens from Phu Jor Kor sculptured with irregular undulate penial pilasters arranged in inclined and longitudinal rows (Fig. 7D). Epiphallus shorter than penis. Vas deferens longer than penis. Gametolytic duct connecting to basal part of amatorial organ, basally bulbous, distally with short and small duct. Tapering gametolytic sac long, robust (Fig. 6A). Amatorial organ elliptical, with inner wall smooth. Papilla of amatorial organ strong, long, reaching junction of gametolytic duct. Corneous spike present (Fig. 6B–C). Amatorial organ duct longer than amatorial organ. Vagina and free oviduct shorter than penis. Inner wall of vagina smooth, with 3–4 thin longitudinal pilasters on median part (Fig. 6E). Specimens from Phu Jor Kor has gametolytic duct connecting to middle part of amatorial organ. Gametolytic sac cylindrical, long. Amatorial organ long, cylindrical, with inner wall sculptured with irregular undulated short pilasters. Papilla of amatorial organ robust, cylindrical, distally finger-shaped, without terminal corneous spike. Vagina longer than penis, internally with four to five longitudinal pilasters. Amatorial organ duct nearly as long as, or slightly longer than amatorial organ.</p>
            <p>Radula (n=13, comprising three specimens from Khao Sala (one specimen from Hoompuay, 2016) (Fig. 7A–B) three specimens from Phu Jor Kor, and seven specimens from Pha Nam Yoi (five specimens from Hoompuay, 2016) (Fig. 7C–D): with 83–90 rows of teeth. Each row with 83–123 teeth, formula: (24–42) + (17–19) + 1 + (17–19) + (24–42) in specimens from Khao Sala, and approximately 70–82 rows of teeth, each of which comprises 91–123 teeth. Fomula: (31–43) + (14–18) + 1 + (14–18) + (31–43) in specimens from Phu Jor Kor and Pha Nam Yoi. Central and lateral teeth lanceolate, tricuspid. Marginal teeth bicuspid and gradually changing to unicuspid and sword shaped.</p>
            <p> Distribution.  Quantula doma ,  new species , is found in sandstone mountains of the southern part of north-eastern Thailand, in the Surin, Roi Et, and Mukdahan Provinces. </p>
            <p> Remarks. According to the genetic distance and phylogenetic analysis,  Quantula “sp. 1” from Khao Sala in Jirapatrasilp et al. (2021) (specimen no. H60, COI and 28s accession numbers MT803088 and MT741936) is clustered with the  Q. doma ,  new species (specimen no. KSH and KSL). Genetic distances within the Khao Sala population are around 0.5% (Table 2), and all specimens have rather strong support (Fig. 2) (BI posterior probability of 1.00 and 100% for ML, and 99% for NJ bootstrap replicate), based on which we conclude that specimen H60, KSH and KSL represent the same species as  Q. doma ,  new species . Although conchologically this new species looks similar to  Q. simonei Thach &amp; Huber 2018 from Laos (see Thach, 2018: plate 49, figs. 580, 581 top), in  Q. doma ,  new species , the last whorl is angulate with weak peripheral keel and has an open umbilicus, whereas in  Q. simonei , the last whorl is well-inflated without peripheral keel, and the umbilicus is mostly covered by the columella portion of the apertural lip. </p>
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	https://treatment.plazi.org/id/03AD87F6FFC5E470FC10AADB2129F8BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Deeprom, Kanchanog;Tumpeesuwan, Chanidaporn;Tumpeesuwan, Sakboworn	Deeprom, Kanchanog, Tumpeesuwan, Chanidaporn, Tumpeesuwan, Sakboworn (2022): A new species of the genus Quantula Baker, 1941 (Eupulmonata: Dyakiidae) from the southern part of north-eastern Thailand. Raffles Bulletin of Zoology 70: 519-533, DOI: 10.26107/RBZ-2022-0029
