identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03F09661FFFC806EFE0BE7267A265192.text	03F09661FFFC806EFE0BE7267A265192.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis Burmeister 1844	<div><p>Byrsopolis Burmeister, 1844</p> <p>Type species: Byrsopoliscastanea Burmeister, 1844 [by monotypy].</p> <p>Gender. Feminine.</p> <p>Byrsopolis Burmeister, 1844: 425 (original description); Blanchard 1851: 219 (citation); Burmeister 1855: 523 (female diagnosis); Lacordaire 1856: 364 (redescription, key to genera); Castelnau and Lucas 1857: 130 (distribution, illustrations); Gemminger and Harold 1869: 1226 (checklist); Bates 1888: 290 (citation); Heyne and Taschenberg 1908: 88 (citation), 356 (illustration); Ohaus 1908: 251 (citation); 1912: 312 (citation); 1915: 75 (citation); Casey 1915: 1 (note); Ohaus 1925: 75 (citation); 1926: 225–226 (citation); Hardy 1971: 235 (status revalidation); Machatschke 1972: 2 (catalogue, distribution); Morón 1976: 19 (citation); Ratcliffe and Jameson 1989: 135 (citation); Jameson 1990: 377 (citation, key to genera); 1998: 5 (citation); Krajcik 2008: 58 (checklist); Smith (2009): 73 (citation); Soula 2010a (citation): 44; 2010b: 54 (key to species, illustration); Grossi and Vaz-de-Mello 2015: 221 (biology, key to genera); Morón and Nogueira 2016: 7 (citation); Ferreira 2016: 112 (illustration); Moore et al. 2017: 49 (citation); Ferreira et al. 2018: 4, 8–9 (checklist, distribution); Carvalho and Grossi 2018: 372 (checklist, distribution); Hielkema and Hielkema 2019: 126 (checklist, distribution); Menis and Rodrigues 2021: 3, 5 (citation); Vaz-de-Mello and Grossi 2022 (online catalogue).</p> <p>– Cotalpa (Byrsopolis) (Burmeister 1844): Ohaus 1915: 256 (status change, key to subgenera); 1918: 10 (checklist); 1934: 37 (subgenus redescription, checklist, distribution), 173 (illustration); Blackwelder 1944: 235 (checklist).</p> <p>Diagnosis. Frontoclypeal suture complete, almost linear or with curvilinear extremities; frons glabrous or with short setae; clypeus glabrous or setose; mandibles externally rounded with molar region striated, with or without projections; apex of prementum weakly to strongly emarginated; antennae with 10 antennomeres; antennal club longer than antennomeres II–VII combined, with marginal areas unconstricted. Thorax with mesosternal process not exceeding mesocoxae apex. Elytra punctate, with or without striae, striae with small punctures; apical calli distinct; hind wings with anterior margin (PC +C, ScA, ScP, RA 1+2, RA 3) glabrous, RA 3 margin very close to RA 4 + RP 1, but not reaching it; apex of Cu setose; AA 3 developed and converging to Cu. Parameres bifurcated, parallel, convergent or divergent; shape without projections.</p> <p>Redescription. Body. Shape elongated, oval, sides parallel to rounded in dorsal view; dorsal surface usually convex, ventrally flattened in lateral view; females usually bigger than males and with a more distinctly convex dorsal surface. Colour very variable, from light yellow to completely black, with metallic reflections in some species, or with legs, head, pronotum and venter metallic green. Length 20.40–37.81 mm; humeral width 9.50– 20.73 mm, elytral width 11.05–22.37 mm. Head. Clypeus rounded, trapezoidal, rectangular, semioval, or square; weakly to moderately concave, wider than long, widest at base; apex straight, truncated, emarginated, bilobed or trilobed; surface glabrous to hirsute; frontoclypeal suture clearly visible, almost straight or sinuous; clypeus and frons sometimes with isolated concavities, or a longitudinal midline formed by punctures; mandibles partially or totally covered by clypeus, glabrous or hirsute, smooth or punctate; maxillae symmetrical or asymmetrical; surface of prementum and area of insertion of palpigers excavated or not; premental apex with weak to strong emargination; labrum bilobed, with narrowly rounded to truncated lobes; male antennal club 1.5–3x longer than antennomeres II–VII combined, female antennal club smaller, about 1.2–1.5x longer than antennomeres II–VII combined. Thorax. Pronotum with weak to strong concavities at isolated punctures or oblique lines near posterior angles; anterior angles obtuse, incurved; scutellar plate subtriangular, with punctures and setae varying in size from small to large, sparse to densely distributed; mesoventral process with apex rounded to oval, not exceeding mesocoxa. Elytra. Humeri slightly concave, with small setae and small calli; elytral suture smooth to striate; disc with weak to strong striae, interstriae sparsely to densely punctate; epipleura sparsely to densely setose; hind wings with anterior margin near veins (PC+C, ScA, ScP, RA 1+2, RA 3) glabrous, vein AA 3 developed, converging to Cu; Cu setae only at apex; margin of RA 3 very close to RA 4 + RP 1, but not reaching it. Legs. Protibia with three outer teeth; spur developed, at least 2x longer than wide, conical (spine-like) or flattened; females with protarsomeres IV–V usually with longer setae than in males; apex of metatibiae with 14–32 spine-like setae. Abdomen. Tergite 8 striated or punctated, glabrous or hirsute. Aedeagus. Parameres dorsally or laterally symmetrical or not, weakly to strongly bifurcated; rounded, parallel, divergent or convergent apex, weakly to strongly angular; lateral margins of parameres weakly to strongly excavated longitudinally, forming a small to long carina; right paramere sometimes thicker or more divergent.</p> <p>Distribution. Byrsopolis has a disjunct distribution, with two species restricted to the coastal region of French Guiana (Soula 2010a), and the others exclusive to the Brazilian Cerrado biome and Atlantic Forest found in Bahia, Distrito Federal, Goiás, Mato Grosso do Sul, Mato Grosso, Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo and Paraná states, and Paraguay (Dept. San Pedro).</p> </div>	https://treatment.plazi.org/id/03F09661FFFC806EFE0BE7267A265192	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFFA8063FEC3E2CE7A3F50AF.text	03F09661FFFA8063FEC3E2CE7A3F50AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis castanea Burmeister 1844	<div><p>Byrsopolis castanea Burmeister, 1844 (Figures 1 (a–g), 5(a–h))</p> <p>Byrsopolis castanea Burmeister, 1844: 425 (original description); Burmeister 1855: 523 (female diagnosis); Lacordaire 1856: 364 (citation); Castelnau and Lucas 1857: 130 (distribution and illustration); Gemminger and Harold 1869: 1226 (checklist); Heyne and Taschenberg 1908: 88 (diagnosis), 356 (illustration); Ohaus 1908: 251 (distribution); 1912: 312 (citation); Hardy 1971: 236 (citation); Machatschke 1972: 3 (catalogue, distribution); Krajcik 2008: 58 (checklist); Grossi and Vaz-de-Mello 2015: 223 (biology, distribution); Ferreira 2016: 112 (illustration); Carvalho and Grossi 2018: 372 (checklist, distribution); Vazde-Mello and Grossi 2022 (online catalogue).</p> <p>Cotalpa (Byrsopolis) castanea (Burmeister, 1844): Ohaus 1915: 257 (citation); 1918: 10 (catalogue, distribution); 1934: 37 (citation, checklist), 173 (illustration); Blackwelder 1944: 235 (checklist).</p> <p>Type locality. Nova Friburgo, Rio de Janeiro, Brazil.</p> <p>Type material. Lectotype male, here designated. (a) ‘ Byrsopolis *’, (b) ‘castanea *./Bras. Bsk. [handwritten by Burmeister]’ (1♂ ZMHB) (Figure 1 (a,b)).</p> <p>Paralectotypes. (a) ‘ Byrsopolis *’, (b) ‘castanea *./Bras. Bsk. [handwritten by Burmeister]’ (1♀, 1♂ ZMHB) (Figure 1 (c–1)).</p> <p>Additional material. [22 specimens] Brazil, Rio de Janeiro. ‘ Brasil, Rio de Janeiro, Nova/ Friburgo, Macaé de Cima,/ i.2004, 1500 m, luz, D. &amp; I./Miller legs’ (1♀, 7♂ CERPE); same, except ‘Conquista,/ 850 m – XII.2004 /E. &amp; P. Grossi Leg’. // ‘COLEÇÃO E./ &amp; P. GROSSI’ (1♀, 2♂ EPGC); ‘ BRASIL, RJ, Teresópolis, 600 m / 01 a 31.xii.2006 /E. &amp; P. Grossi Legs’. // ‘COLEÇÃO E. &amp; P. GROSSI’ (1♂ EPGC); ‘Brasília/Cantagallo’ // ‘castanea Brm’ // ‘ Byrsopolis / castanea Burmeister /Museum f. Naturkunde/Berlin’ (2♂ ZMHB). Minas Gerais. ‘BR, MG, Alto Caparaó/PARNA Caparaó/Coleta noturna/ 05-06.xii.2016 /A. Orsetti &amp; S. Alóquio legs’ (1♂ CERPE). Espírito Santo. ‘ BRASIL: Espírito Santo. Conceição do Castelo./Ribeirão do Meio. 20° 18’/31’S: 41°14 ʹ 17”W. 880 m /Light. ii.2011. R. Falqueto’ (1♂ CEMT). No collection data. ‘ Byrsopolis /castanea Burm’ (1♀ ZMHB); ‘ ♂ ’ // ‘11,445’ // ‘Hist. – Coll. (Coleoptera)/ Nr.11445/ Byrsopolis castanea Burm. /Brasília, Riehl’ (2♂ ZMHB); ‘43,114’ // ‘ ♀ ’ // ‘Hist. – Coll. (Coleoptera)/Nr. 43,114/ Byrsopolis castanea Burm. /Brasília, Coll. Schaum’ (1♀, 1♂ ZMHB); ‘ Castanea Burm. Brésil’ // ‘Coll. Höhne’ // ‘ Byrsopolis /castanea Burmeister’ (1♂ ZMHB).</p> <p>Diagnosis. Body brown, reddish-brown to dark brown with metallic green reflections; clypeus subrectangular, anterior margin weakly emarginate; antennal club about 1.5x (female) to 2x (male) longer than antennomeres II–VII combined; scutellar plate with moderately rounded base; elytra with very distinct striae, densely punctate on the humeri; elytral apex truncated; protibial spur about 8x longer than wide; aedeagus with asymmetrical parameres, weakly to moderately divergent, weakly angulate laterally, apex of parameres weakly rounded.</p> <p>Description of the lectotype. Male. Body. Shape elongated, oval, with rounded sides; brown, reddish-brown to black colouration (Figure 1 (a–b)); frontoclypeal suture darker than clypeus and frons; body with marginal areas from dark brown to black; metallic green and reddish reflections. Length 25.89–29.07 mm. Humeral width 12.82–15.02 mm. Elytral width 15.03–16.54 mm. Head. Clypeus about 2x wider than long, subrectangular, with angles weakly rounded, anterior margin weakly emarginate, surface weakly concave near the apex; frontoclypeal suture weakly bisinuate; surface strongly rugose; punctures small to moderately large, glabrous; frons as long as clypeus, punctures small to moderately large; moderately to densely punctate; glabrous; mandibles moderately sinuous at the border, scissorial region C-shaped (Figure 5 (e)), rounded apex, small punctures, sparsely to densely punctate in the transverse median region; setae small to moderate in size near scissorial region, sparse to moderately distributed; lacinia with three teeth, equidistant; apical tooth longer and wider than basal one, basal tooth longer and wider than medium one (Figure 5 (f)); basal tooth with long setae; last maxillary palpomere subrectangular sensorial area; premental apex moderately emarginated (Figure 5 (g)); last labial palpomere about 1.5x longer than palpomere II (Figure 5 (g)); labrum moderately emarginate, moderately setose (Figure 5 (h)); antennomere III subcylindrical, antennomeres IV and V subquadrate, antennomeres VI and VII subconical; antennal club about 2x longer than antennomeres II–VII combined. Thorax. Pronotum transverse, about 1.5x wider than long (Figure 5 (a)); punctures small to moderately large, moderately to densely punctate, sparsely punctate at marginal areas, glabrous, two elongate oblique concavities at base, one on each side; scutellar plate subtriangular, sides straight, base moderately rounded; apex with punctures and setae small to moderate, disc moderately to densely punctate, marginal areas sparsely to moderately punctate. Elytra. About 1.25x longer than wide; surface glabrous, strongly striate and punctate, punctures small to large, moderately to densely punctate (Figure 5 (a)); interstriae with strong coalescent punctures; apex truncate; glabrous. Legs. Protibial spur about 8x longer than wide; mesotibiae with one carina, metatibiae with two carinae. Aedeagus. Parameres with proximal margin moderately rounded; lateral margins excavated, forming a carina from near base to apex; apexes rounded and weakly sinuate internally, weakly divergent, dorsally symmetrical, laterally and ventrally asymmetrical; left paramere longer than righr paramere (Figure 5 (b)).</p> <p>Female. Similar to male, varying in the aspects described below.</p> <p>Body. Length 29.70–37.81 mm. Humeral width 18.79–20.73 mm. Elytral width 20.50– 22.37 mm (Figures 1 (e–f), 5(c-d)). Head. Clypeus about 2x wider than long, apical angles rounded with slight concavity near the apex; punctures small to large, densely punctate; frons with small to moderate punctures, moderately to densely punctate; in last maxillary palpomere with excavated area larger; antennal club about 1.5x as long as antennomeres II–VII combined. Thorax. Pronotum transverse about 2x wider than long; small punctures. Elytra. About 1.40x longer than wide; convex; punctures small to large, moderately to densely punctate.</p> <p>Variation. Some specimens exhibit variation in colour, from reddish-brown to black, especially on marginal areas; apex of clypeus weakly to moderately emarginate; apex of parameres weakly to moderately divergent.</p> <p>Etymology. The species name castanea is derived from Latin, in reference to the brown colouration of the specimens.</p> <p>Distribution. Minas Gerais (Alto Caparaó), Espírito Santo (Conceição do Castelo), Rio de Janeiro (Nova Friburgo [Conquista, Macaé de Cima] and Teresópolis) (Figure 19).</p> <p>Remarks. This species is restricted to the Atlantic Forest of Serra do Mar and Serra dos Órgãos mountains in Rio de Janeiro state (near the cities of Nova Friburgo and Teresópolis), Serra de Conceição do Castelo in Espírito Santo state and Caparaó in Minas Gerais state. The locations of Alto Caparaó in Minas Gerais and Conceição do Castelo in Espírito Santo are new state records. Some specimens were collected in Nova Friburgo (Rio de Janeiro) using light traps.</p></div> 	https://treatment.plazi.org/id/03F09661FFFA8063FEC3E2CE7A3F50AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFF78060FEFEE3FB7A9A5057.text	03F09661FFF78060FEFEE3FB7A9A5057.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis crassa Blanchard 1851	<div><p>Byrsopolis crassa Blanchard, 1851 (Figures 2 (a–e), 6(a–g))</p> <p>Byrsopolis crassa Blanchard 1851: 219 (original description); Lacordaire 1856: 364 (citation); Castelnau and Lucas 1857: 130 (distribution, illustration); Gemminger and Harold 1869: 1226 (checklist); Heyne and Taschenberg 1908: 89 (diagnosis, distribution); Machatschke 1972: 3 (catalogue, distribution); Krajcik 2008: 58 (checklist); Carvalho and Grossi 2018: 372 (checklist, distribution); Vaz-de-Mello and Grossi 2022 (online catalogue).</p> <p>Cotalpa (Byrsopolis) crassa (Blanchard, 1851): Ohaus 1915: 256 (note); 1918: 10 (catalogue, distribution); 1934: 37 (citation and checklist); Blackwelder 1944: 235 (checklist).</p> <p>Type locality. Brazil, Mato Grosso, Cuiabá.</p> <p>Type material</p> <p>Holotype. (a) ‘6/47’ [handwritten], (b) ‘MUSEUM PARIS’, (c) ‘GOYAZ CUYABA/DE CASTELNAU 6–47’, (d) ‘ Byrsopolis /crassa, Bl’. [handwritten], (e) ‘ B. crassa /Cat. Mus./ Goyaz/M de Castelnau’ [handwritten], (f) ‘HOLOTYPE’ [red label], (g) ‘ MNHN /EC1803’ (1♂ specimen examined through images by A. Mantilleri; MNHN-EC-EC1803, MNHN 2016) (Figure 2).</p> <p>Additional material. [37 specimens] Brazil, Minas Gerais. ‘Brasil, Minas Gerais,/ Cordisburgo, Faz. Pontinha / i.1999, F.Z. Vaz-de-Melo’ (1♂ CERPE, 1♂ CEMT); ‘ Brasil, Minas Gerais, São Gonçalo do Rio Preto, Pq./Est. Rio Preto, Mirante,/ 22.ii.2011, luz, R.Oliveira &amp;/ Ferreira legs’ (1♂ CERPE); ‘ Minas Gerais’ // ‘Ohaus determ./ Byrsopolis / crassa Bl. / Klein. ♂ ’ (1♂ NHM); ‘ Fazenda/SÃO CAETANO – MG/ VALÉRIA LUSTOSA / 10/12/2006 ’ (1♂ CEMT); ‘ Minas Geraes [sic, Gerais]/ Ouropreto / II2 Magalhaes\Gomes S.G’. // ‘ Byrsopolis / crassa Blanch. /m.d. Type vergl./Paris 5.VII.1911 [handwritten by Ohaus]’ (1♂ ZMHB). Mato Grosso. ‘M. Grosso/xi.946 [handwritten]’ // ‘ 02 17/61 ’ // ‘ Areoda banksi Lap. /det M.L. Jameson 1997’ (1♂ MNRJ); ‘ UFMT/MT – RONDONÓPOLIS / 15.I.1991 / VILMA C. ELLER [handwritten]’ // ‘CEMT’ (1♂ CERPE); ‘ Brasil, Mato Grosso, Cuiabá,/ Fazenda Mutuca, 1998–1999,/ Peres Fo leg’ (1♂ CEMT); ‘ Rondonopolis /M. Grosso /1.64’ (1♂ CMN); ‘UFMT/MT – Barra do Garças / 9-V-1995 / Cláudia C. Queiroz’ // ‘UFMT/ Coleoptera /Scarabaeidae’ (1♂ CEMT). São Paulo. ‘ Brasil, São Paulo, Itirapina,/ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-47.83472&amp;materialsCitation.latitude=-22.24166" title="Search Plazi for locations around (long -47.83472/lat -22.24166)">Estação Ecológica Cerrado</a>,/ 600 m, 5.iv-5.v.2000, 22° 14 ʹ 30’S, 47°50 ʹ 05”W,/ pitfall G. Machado leg.’ (1♂ EPGC, 1♂ CERPE). Goiás. ‘ Faz. Nova Orlândia / Jataí, GO – Brasil /I.964 – Martins,/ Morgante &amp; Silva’. // ‘ Byrsopolis crassa [handwritten]’ (1♂ MZUSP); ‘ BRASIL: Goiás. Mineiros./PNEmas. 17°54 ʹ 45’S;52°59 ʹ 20”/W. 845 mosl. hum/Dung. 15-III- /2011. M.F. Souza.’ (3♂ CERPE, 2♂ CEMT); same, except ‘845 mosl. hum. Dung. 15-/ III-2011. M.F. Souza’ (1♂ CEMT); same, except ‘PNEmas. 18° 04 ʹ 14’S;52°55 ʹ 57” (1♂ CERPE, 2♂ CEMT); same, expect, ‘WORLD/SCARAB./DATABASE/ WSD00344175 ’ (1♂ MSPC); same, except ‘PNEmas 18°01 ʹ 12’S;52°57 ʹ 52”/W. 874 mosl.’ (1♂ CERPE, 1♂ CEMT); same, except ‘18°04 ʹ 14’S, 52°55 ʹ 57”/W. 845 mosl. Hum. Dung. 15-/ III-2011. M.F. Souza’ (2♂ CEMT); same, except ‘845 mosl. Hum. Dung. 15-III- /2011. M. F. Souza’ (2♂ CEMT); ‘ BRASIL: GO Colinas do Sul/Serra da Mesa 2–15.XII.1995/ 14°01 ʹ S, 48° 12 ʹ W / C. Campaner col’ (2♂ MZUSP). Distrito Federal. ‘ 18/11/17 /PNB/35/ Marina Regina Frizzas’ (1♂ CEMT). No collection data (2♂ CERPE).</p> <p>Diagnosis. Body almost entirely orange to dark orange, darker brown at marginal areas, without green reflections; clypeus semioval, slightly to moderately concave; antennal club about 1.70x longer than antennomeres II–VII combined; scutellar plate with sides straight and base weakly rounded basis; elytral apex truncated; elytra with indistinct elytral costae and humeri moderately punctate; protibial spur about 4x longer than wide; parameres asymmetrical, moderately divergent, glabrous, apex with acute angles.</p> <p>Redescription. Male. Body. Shape elongate oval; orange to brown-testaceous, without green reflections (Figures 2 (a- b), 6(a-b)); frontoclypeal suture darker than clypeus and frons; antennae yellow to orange; marginal areas darker, brown. Length 23.34–29.50 mm. Humeral width 12.60–15.60 mm. Elytral width 15.00– 17.75 mm. Head. Clypeus about 2x wider than long, semioval; apex moderately concave; frontoclypeal suture moderately bisinuate, weakly curvilinear near extremities; punctures small to moderate, surface moderately to densely punctate, glabrous; frons about 1.30 times longer than clypeus; punctures small to moderate, disc moderately to densely punctate, glabrous; external border of mandibles forming a right angle with the scissorial region, small projection at middle, or with small setae, densely punctate (Figure 6 (d)); galea with one long apical tooth, four small teeth at middle, basal tooth small; base of galea with four small setae (Figure 6 (e)); last maxillary palpomere elliptic, sensorial area as long as palpomere; apex of prementum moderately emarginate, moderately setose; last labial palpomere about 1.90x longer than palpomere II (Figure 6 (f)); labrum moderately emarginate (Figure 6 (g)); antennomere II weakly rounded, with setae; antennomeres III to V slightly conical, antennomere VI subconical and antennomere VII subrectangular; antennal club about 1.70x longer than antennomeres II–VII combined. Thorax. Pronotum transverse, about 1.70x wider than long (Figure 6 (a)); punctures small to moderate, moderately to densely punctate; marginal areas sparsely to moderately punctate, sparsely or moderately setose; small constriction near base; scutellar plate subtriangular, sides straight, base weakly rounded, apex with punctures and setae small to moderate, sparsely to moderately distributed; space between punctures forming a longitudinal midline. Elytra. About 1.10 times longer than wide; elytral surface without striae; punctures small to large, sparsely to densely punctate at disc, humeri moderately punctate; microsetae in moderate punctures; marginal areas with small to moderate punctures, sparsely to moderately punctate; interstriae without coalescent punctures, moderately punctate (Figure 6 (a)). Legs. Protibial spur about 4x longer than wide; mesotibiae with two carinae; metatibiae with one carina. Aedeagus. Phallobase slightly asymmetrical; parameres moderately elongate; basal margin bisinuate; transverse striae in middle region; apex rounded, moderately divergent, asymmetrical, lateral margins with acute angles; weak concavity extending longitudinally at middle (Figures 2 (d–e, 6(c))).</p> <p>Female. Unknown.</p> <p>Variation. Clypeal concavity weak to moderate, near apex; clypeus slightly shorter; mandibles with glabrous or hirsute apex; apex of prementum weakly to moderately emarginated; aedeagus can vary in length; parameres weakly to moderately asymmetrical, right paramere more divergent; transversal striae small and reduced; indistinct longitudinal concavity, very indistinct.</p> <p>Etymology. The species name crassa (crassus) is derived from Latin referring to its size and width, larger and thicker than the other species so far described.</p> <p>Distribution. Minas Gerais (Ouro Preto, Codisburgo and São Gonçalo do Rio Preto), Mato Grosso (Cuiabá, Diamantino and Rondonópolis), São Paulo (Itirapina), Distrito Federal (Brasília) and Goiás (Jataí, Mineiros and Colinas do Sul) (Figure 19).</p> <p>Remarks. The species is restricted to Cerrado biome areas with rocky outcrops, such as the Serra do Espinhaço complex (Minas Gerais), and open-field Cerrado biome areas in Mato Grosso state. Byrsopolis crassa has the second largest geographic distribution known in the genus, being found in four states, from the Midwest (Mato Grosso and Goiás) to the Southeast (São Paulo and Minas Gerais) of Brazil. The species has been collected in almost all seasons, showing great temporal plasticity. The records for the states of Minas Gerais (Cordisburgo and São Gonçalo do Rio Preto), Mato Grosso (Diamantino and Rondonópolis), Goiás (Jataí, Mineiros and Colinas do Sul), São Paulo (Itirapina), and Distrito Federal (Brasília) are new state records.</p> </div>	https://treatment.plazi.org/id/03F09661FFF78060FEFEE3FB7A9A5057	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFF48065FED5E393791C519F.text	03F09661FFF48065FED5E393791C519F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis quadraticeps Blanchard 1851	<div><p>Byrsopolis quadraticeps Blanchard, 1851 (Figures 3 (a–e), 7(a–g))</p> <p>Byrsopolis quadraticeps Blanchard 1851: 219 (original description); Lacordaire 1856: 364; Castelnau and Lucas 1857: 130 (distribution and illustration); Gemminger and Harold 1869: 1226 (checklist); Machatschke 1972: 3 (catalogue); Krajcik 2008: 58 (checklist); Carvalho and Grossi 2018: 372 (checklist, distribution); Menis and Rodrigues 2021: 3, 5 (citation); Vaz-de-Mello and Grossi 2022 (online catalogue).</p> <p>Cotalpa (Byrsopolis) quadraticeps (Blanchard, 1851): 219; Ohaus 1918: 10 (catalogue, distribution); Ohaus 1934: 38 (citation, checklist); Blackwelder 1944: 235 (checklist).</p> <p>Type locality. Brazil, Minas Gerais.</p> <p>Type material. Holotype male, here designated. (a) ‘MUSEUM PARIS/OEST CAPIT e/DES MINES’, (b) ‘OUEST DE/LA CAPITE DES MINES [handwritten]’, (c) ‘ Byrsopolis / quadraticeps, Bl. [handwritten]’, (d) B. quadraticeps /Cat. Mus/Brésil./M. Aug. Saint-Hilaire [handwritten]’, (e) HOLOTYPE [red label]’, (f) ‘MNHN/EC1804’ (1♂ MNHN, specimen examined through images by A. Mantilleri; MNHN-EC-EC1804, MNHN 2016) (Figure 3 (a-e)).</p> <p>Additional material. [59 specimens] Brazil, Minas Gerais. ‘ BRASIL: Minas Gerais. Buritis / Farm Querência. 03-xii-2012./ Light. C. M Oliveira’ (9♂ CEMT, 8♂ CERPE); ‘ IJACI, MG/ 15/X/ 03 / T. BERNARDO Col’ (1♂ CEMT). Mato Grosso. ‘ Rondonópolis /Mato Grosso/10.62’ (1♂ CMN); ‘ UNIVAG / Cuiabá – MT/ 13/10/2012 /Ana’ (1♂ CEMT); ‘ UNIVAG / V. Grande MT –/ 23/10/ 2004 / Pereira, Lucimar’ (1♂ CERPE); ‘ UNIVAG / Várzea Grande /MT/15/10.2004/ Vilma’ (1♂ CEMT); same, except ‘MT – V. Grande / 16.X.2004 / Danny Rotilli’ (1♂ CERPE). Goiás. ‘C. GOYAZ/ L. Bulhões / R. Spitz S. \ 10.xi.35 /a ♂ ’ // ‘ Ohaus determ’ (1♂ ZMHB); ‘ LEOPOLDO BULHOES / Est. Goyaz Dez. 37 / Dr. Nick’ (1♂ MNRJ); same (1♂ MNRJ); ‘ Brasil, Goiás, Cristalina,/ Fazenda Larga Grande /(Sorgatto), 03–08.xi.2016,/ Luz, N.C.M. Souza leg’ (2♂ CERPE); ‘ LEOPOLDO/BULHOES /Est de Goias/(Brasil)/DIRINGS\XII. Dr. Nick’ (2♂ MZUSP); ‘ Leopoldo Bulhões / Goiaz /12.937’ // ‘Coll. J. Guerin’ (1♂ IBSP); ‘ Brazil, Goias, Cristalina / xi.1989 /unknown collector’ // ‘Coll. Matthias/Seidel 2016’ // ‘ Byrsopolis quadraticeps / Blanchard, 1851 /det. M. Seidel 2017’ // ‘WORLD/SCARAB./DATABASE/[WSD00347498 - WSD00347499]’ (2♂ MSPC); same, except ‘WORLD/SCARAB./DATABASE/WSD00347500’ // ‘ ZMH 833674 ’ (1♂ ZMH). São Paulo. ‘Bresilien/Prov. S. Paulo/ Campinas / Alwine Braatz V’ (3♂ ZMHB); ‘ Pr. S. Paulo /Bras’. // ‘ ♂ ’ // ‘ Byrsopolis Bl. /quadraticeps/m.d. Type vergt./Paris 5. VII.1911 [handwritten by Ohaus]’ (1♂ ZMHB); ‘ Campinas /San Paulo/Bresilien’ // ‘quadraticeps Blanch’ (1♂ ZMHB); ‘ Bresilien /SanPaulo’ // ‘ Byrsopolis /quadraticeps Blanch’ (1♂ ZMHB); ‘ Pr. S. Paulo /Brasil’ // ‘ Byrsopolis quadraticeps /Blanch’ (1♂ ZMHB); ‘ Campin. [handwritten]’ // ‘ Byrsopolis /crassa Blch/?? Mey. [handwritten]’ // ‘ Byrsopolis /quadraticeps Blanch’ (1♂ ZMHB); ‘ Oct. 943/ Sao Paulo /P. Epitacio/Zellibor-leg/Coll. Martinez’ (1♂ CMN); ‘ Dic. 937/ Sao Paulo / Campinas /R. Spitz-leg./Coll. Martinez [handwritten]’ (1♂ CMN); ‘ Brasil _ SP Lins/ Faz Sta Clara / 19-IX-1981 / Luiz Evandro /Col: Solo’ (1♂ CEMT). Distrito Federal. ‘ Brasília DF/ Brasil XII 93 ’ // ‘ E. &amp; P. Grossi /legs [handwritten]’ (2♂ EPGC); ‘Brasil: Distrito Federal / Planaltina. Embrapa Cerrados./Área Agrícola. 15°36 ʹ 54’S / 47° 42 ʹ 51”W. 18-XI-2005./ Light / C. Oliveira’ (1♂ CERPE); same, except ‘ Fragmento Cerrado 15°36 ʹ 20’/S 47°42 ʹ 26”W. 18-xi-2005./ Light. C. Oliveira’ (1♂ CEMT); same, except ‘ Planaltina./ Embrapa Cerrados. Área Agrícola. 15°36’/20’S 47°42 ʹ 26”W’. (1♂ CEMT); same, except ‘ Planaltina./ Embrapa Cerrados. Área Agrícola. 15°35 ʹ 54’S/47°42 ʹ 26”W. 18-xi-2005. Light./ C. Oliveira’ (1♂ CEMT); ‘ DF. PNB/ Cerrado / 11.ii.15 / pitfall / Marina Regina Frizzas’ (1♂ CERPE); ‘ Brazil, Brasilia D. F. / xi.1992 /local collector’ // ‘Coll. Matthias/Seidel 2017’ // ‘ Byrsopolis /quadraticeps/Bl, 1851/det. M. Seidel 2017’ // ‘WORLD/SCARAB./DATABASE/ [WSD00347501 - WSD00347502]’ (2♂ MSPC). Paraná. ‘ Curitiba./Paraná/12.936’ // ‘Coll. J. Guerin’ (1♂ IBSP); ‘ Brasil – Paraná – Londrina / 15-IV-2002 /Col. André Manfio Maia’ (1♂ CEMT). No collection data (2♂ CERPE, ♂ 2 CEMT).</p> <p>Diagnosis. Body yellowish to yellowish brown, without metallic green reflections; clypeus subrectangular with rounded angles, apex moderately bilobed; antennal club about 2x longer than antennomeres II–VII combined; scutellar plate subtriangular, base subacute; elytral apex with right angles; elytra with indistinct striae, punctate, humeri weakly punctate; protibial spur about 4x longer than wide; parameres symmetrical, weakly convergent, moderately elongated, weakly angled internally, proximal margin bisinuate.</p> <p>Redescription. Male. Body. Oval, elongated, sides rounded; yellowish to yellowish brown, without green reflections, marginal areas orange-brown to dark-brown (Figures 3 (a-b), 7 (a-b)); clypeal border darker, reddish to dark red; frontoclypeal suture darker than frons, reddish. Length 20.40–25.40 mm. Humeral width 9.50–12.83 mm. Elytral width 11.05– 14.29 mm. Head. Clypeus about 1.60x wider than long, subrectangular, rounded angles, weakly to moderately bilobed (Figure 7 (a)); surface moderately concave; frontoclypeal suture almost straight between curvilinear extremities, rounded apical corners; punctures small to moderate, moderately to densely punctate; setae moderate to large, moderate to densely distributed; length of frons subequal to clypeus (0.95x shorter than clypeus); punctures small to moderately large, moderate to densely punctate; microsetae in moderate punctures; mandibles with small and sparse punctures and setae near scissorial region (Figure 7 (d)); galea with three equidistant teeth, equidistant, apical tooth longer and thicker than basal tooth, basal tooth longer and thicker than medial tooth, basal tooth with two long setae (Figure 7 (e)); last maxillary palpomere with oblong sensorial area; apex of prementum weakly emarginate; last labial palpomere about 2x longer than palpomere II (Figure 7 (f)); labrum strongly emarginate (Figure 7 (g)); antennomere III subcylindrical, antennomere IV subquadrate, antennomeres V–VII subconical; antennal club about 2x longer than antennomeres II–VII combined. Thorax. Pronotum transverse, about 1.5x wider than long (Figure 7 (a)); punctures small to moderately large, moderately to densely punctate, glabrous; marginal areas sparsely to moderately punctate, with sparse setae; longitudinal midline formed by absence of punctures; scutellar plate subtriangular, subacute, sides almost right, base moderately rounded, small to moderate setae, punctures small to moderately large, moderately to densely punctate. Elytra. About 1.10x longer than wide; elytral surface smooth, without striae; punctures small to moderately large, moderately to densely punctate (Figures 3 (a), 3(a)); microsetae in moderate punctures; humeri weakly punctate; interstriae without coalescent punctures, moderately distributed. Legs. Protibial spur about 4x longer than wide; mesotibiae with two carinae; metatibiae with one carina. Aedeagus. Parameres moderately elongated; basal margin bisinuate; apex rounded, symmetrical, subparallel, weakly convergent, separated by a Ushaped gap; apex with weakly sinuous internal angles; paramere with lateral margins longitudinally excavated forming a moderate carina; weak longitudinal concavity near base (Figures 3 (d,e), 3(c)).</p> <p>Female. Unknown.</p> <p>Variation. Some specimens exhibit asymmetrical maxillae, lacking apical and basal teeth in one or both galea; labial palps with different colour, one darker than another; some specimens have dark brown or black spots on elytral surface.</p> <p>Etymology. The species name quadraticeps is derived from Latin words quadrātus (= quadrate) and ceps (= head) reffering to the quadrate shape of the head.</p> <p>Distribution. Minas Gerais (Buritis and Ijaci), Goiás (Leopoldo de Bulhões and Cristalina), Distrito Federal (Brasília and Planaltina), São Paulo (São Paulo, Campinas) and Paraná (Curitiba, Londrina) (Figure 19).</p> <p>Remarks. This species was described from Minas Gerais in the original description of Blanchard (1851); however, he did not specify the city where Saint-Hilaire collected the specimens used for the description. The type was possibly collected in Serra do Espinhaço, occurring in sympatry with Byrsopolis laticollis. Byrsopolis quadraticeps was at the time of its description the only species within the genus with a bilobed and size-reduced clypeus compared to the remaining species. In the present study a second species similar to B. quadraticeps is described as Byrsopolis angelootatii sp. nov. (see below). Additionally, the records for the states of Goiás, Distrito Federal, São Paulo and Paraná are new state records. According to label data, this species is found in the Midwest, Southeast and South regions. It occurs predominantly in the Cerrado biome and can be collected with the aid of light traps.</p> </div>	https://treatment.plazi.org/id/03F09661FFF48065FED5E393791C519F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFF18078FEF2E2AB7B205779.text	03F09661FFF18078FEF2E2AB7B205779.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis laticollis Burmeister 1855	<div><p>Byrsopolis laticollis Burmeister, 1855 (Figures 4 (a–f), 8(a–f))</p> <p>Byrsopolis laticollis Burmeister, 1855: 523 (original description); Gemminger and Harold 1869: 1226 (checklist); Ohaus 1926: 226 (citation); Machatschke 1972: 2 (catalogue, distribution); Krajcik 2008: 58 (checklist); Grossi and Vaz-de-Mello 2015: 223 (biology); Ferreira et al. 2018 (checklist, distribution); Carvalho and Grossi 2018: 372 (checklist, distribution); Vaz-de-Mello and Grossi 2022 (online catalogue).</p> <p>Cotalpa (Byrsopolis) laticollis (Burmeister, 1855): 523 (original description); Ohaus 1918: 10 (catalogue, distribution); Ohaus 1934: 37 (citation, checklist); Blackwelder 1944: 235 (checklist).</p> <p>Type locality. Brazil, Minas Gerais (South of Minas), Cachoeira, Dona Vicenza.</p> <p>Type material. Lectotype male, here designated. (a) ‘laticollis N6/Bras. Br’. [handwritten by Burmeister], (b) ‘Cacho-/eira’ (1♂ MLUH, specimen examined through images by Dr Karla Schneider) (Figure 4 (a,b)).</p> <p>Paralectotype female. (a) ‘laticollis N6/Bras. Br’. [handwritten by Burmeister], (b) ‘Cacho-/ eira’. (1♀ MLUH, specimen examined through images, photos by Karla Schneider), (Figure 4 (d,e)).</p> <p>Additional material. [12 specimens] Brazil, Minas Gerais. ‘ Brasil, MG, 14 km ao S de/ Ponto dos Volantes,/ 16°52 ʹ S 41°40 ʹ W, 660 m / 12.ii.2010, P. Grossi, D./Parizotto &amp; G. Melo Legs’ (1♂ CERPE); same, except ‘MG, 15 km ao S/de Ponto dos Volantes/ 16°52 ʹ S 41°32 ʹ W, 640 m,/ 11.ii.2010, P. Grossi, D. Parizotto &amp; G. Melo Legs/Manual’ (1♂ EPGC); ‘Belo Horizonte, A. Costa leg. 1936?’ (1♂ CERPE); ‘ Brasil, MG,/Belo Horizonte,/ x.1943./A. Costa Jor./UFMG ICO/ 1302579’ // ‘B HORIZONTE/ Minas Gerais Brasil /A. Costa J’. // ‘367/x – 943’ (1♂ UFMG); ‘ Minas Gerais, v. Langsdorf/Zool. Mus. Berlin’ // ‘crassa/Blanch./Minas G. v Langsdorf [handwritten]’ (1♂ ZMHB); ‘Brasília’ // ‘ Byrsopolis / crassa Blanchard /Museum f. Naturkunde/Berlin’ (1♂ ZMHB); ‘ xi.1993 /Min. Ger./(Araguary)/Coll. R. Spitz’ // ‘Ohaus determ’ (1♂ ZMHB); ‘Barbacena/Minas ger’. // ‘ Byrsopolis / laticollis Bŭrm. /m. d. Type vergl./Halle 10.VIII.1911 ’ (1♂ ZMHB); ‘Brasília’ // ‘ Byrsopolis / laticollis Burmeister /Museum f. naturkunde/Berlin’ (1♂ ZMHB); ‘Minas/Brasília’ // ‘ Byrsopolis /castanea/Burm./Brasil’ (1♂ NHMUK); ‘11,446’ // ‘?gifh? in Nahren arso? [handwritten]’ // ‘Hist. – Coll. (Coleoptera)/Nr. 11,446/ Byrsopolis crassa Blanch. / Minas Gerais, v. Langsdorf/Zool. Mus. Berlin’ // ‘crassa/ Blanch./Minas G. v Langsdorf [handwritten]’ (1♂ ZMHB); ‘Brasília’ // ‘ Byrsopolis / crassa Blanchard /Museum f. Naturkunde/Berlin’; ‘ ♂ ’ // ‘Coleção Entomológica Adolph/Hempel – Instituto Biológico/IBSP – IB – 0.006.538’ // ‘J. F. Zikán/Virginia/S. Minas Ger./ 10.xii.1921 / 900 m \Paracatü [sic, Paracatu]’ // ‘ Byrsopolis /laticollis/Brm’ // ‘ Byrsopolis /laticollis, Burm’ (1♂ IBSP).</p> <p>Diagnosis. Uniformly coloured, from light brown to black or reddish; clypeal shape subtrapezoidal, with elevated and weakly rounded anterior margin; weakly concave; antennal club 2.1x longer than antennomeres II–VII combined; scutellar plate with broadly rounded base; elytral apex with acute angles; elytra moderately striate, humeri sparsely punctate; protibial spur about 3x longer than wide; parameres almost straight, basal margin of parameres slightly concave.</p> <p>Description of the lectotype. Male. Body. Oval elongated, rounded sides; orange-brown colour, reddish-brown to black, orange-reddish at the marginal areas (Figures 4 (a-b), 8(a)). Length 25.70–30.21 mm. Humeral width 14.00– 15.75 mm. Elytral width 15.30–17.80 mm. Head. Clypeus about 2x wider than long, shape subtrapezoidal, elevated apex, apex weakly rounded, rounded angles; weak to moderate concavity from base to apex; frontoclypeal suture with moderately rounded curvilinear extremities; punctures small to moderately large, densely punctate; glabrous; frons 1.30x longer than clypeus, punctures small to moderately large, moderately to densely punctate; glabrous; mandibles scissorial region with weak sinuosity; apex widely rounded; three setae moderately sized and sparsely distributed near the border (Figure 8 (c)); maxillary galea with apical tooth well developed, with a weak acute angle near the border, three small sized setae sparsely distributed, one moderately sized seta near border (Figure 8 (d)); last maxillary palpomere with moderately elliptical sensorial area; last labial palpomere about 2x longer than palpomere II (Figure 8 (e)); apex of prementum weakly emarginate; labrum moderately emarginate (Figure 8 (f)); antennomere III subcylindrical, antennomeres IV and V subquadrate, antennomeres VI and VII subconical; antennal club about 2.10x longer than antennomeres II–VII combined (Figure 8 (a)). Thorax. Pronotum transverse, about 1.60x wider than long (Figures 4 (a), 4(a)); punctures small to moderately large, moderately to densely punctate; border with punctures and setae sparsely distributed; scutellar plate subtriangular, base moderately rounded; glabrous; punctures small to moderately large, sparsely to moderately distributed, sparse at marginal areas. Elytra. About 1.10x longer than wide; moderately striate; punctures small to large, moderately to densely distributed; humeral punctures with small to moderate and sparsely punctate; elytral apex with acute angles; interstriae with punctures moderate to large, moderately coalescent (Figures 4 (a), 4(a)). Legs. Protibial spur 3x longer than wide; mesotibiae and metatibiae with one carina. Aedeagus. Parameres with rounded apex, with weakly sinuous internal angles; almost straight laterally, weakly divergent; asymmetrical; lateral margin of left paramere weakly excavate longitudinally, with a small carina; basal margin weakly rounded to straight; surface between base of parameres slightly concave only at apex; glabrous (Figure 8 (b)).</p> <p>Variation. The colouration in some specimens is reddish, black or reddish-brown; clypeus weakly rounded or almost straight, with same frontoclypeal suture and clypeal elevation.</p> <p>Female. Similar to male but with the following differences: antennal club shorter, 1.5x longer than antennomeres II–VII combined; thorax and abdomen less hirsute; scutellar plate narrower; elytra without strong coalescent punctures, less striate (Figure 4 (d,e)); legs longer.</p> <p>Etymology. The species name laticollis is derived from Latin, lati [lātus] (= large, broad) and collis (= neck), reffering to the broad pronotum in this species.</p> <p>Distribution. Minas Gerais (Ouro Preto [Itatiaia, district of the municipality] Barbacena, Agudos [Ohaus 1934], Belo Horizonte, Ponte dos Volantes, Paracatu and Araguari) (Figure 19).</p> <p>Remarks. Two localities recorded in the literature in the state of Minas Gerais have undergone name changes. Those are Cachoeira, now Cachoeira do Campo in the district of Ouro Preto, and Dona Vicenza (or Fazenda Dona Vicenza), north of Queluzito, on the other side of Serra do Deus-te-livre, today known as Itatiaia (personal communication, Vaz-de-Melo 2017). Furthermore, Ohaus (1934) recorded the species from Agudos, probably the ‘Serra de Agudos’, between the states of Minas Gerais and São Paulo. He stated that Burmeister collected in the morning a couple in copula but we found no trace of the female specimen. The new records presented here are from Minas Gerais (Barbacena, Belo Horizonte, Ponto dos Volantes, Paracatu and Araguari).</p></div> 	https://treatment.plazi.org/id/03F09661FFF18078FEF2E2AB7B205779	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFEC807DFEB5E4B47B5152D5.text	03F09661FFEC807DFEB5E4B47B5152D5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis cribricollis Ohaus (MZUSP 1912	<div><p>Byrsopolis cribricollis Ohaus, 1912 (Figure 9 (a–f))</p> <p>Byrsopolis cribricollis Ohaus, 1912: 312 (original description); Machatschke 1972: 3 (catalogue, distribution); Krajcik 2008: 58 (checklist); Carvalho and Grossi 2018: 372 (checklist, distribution); Vaz-de-Mello and Grossi 2022 (online catalogue).</p> <p>Cotalpa (Byrsopolis) cribricollis (Ohaus, 1912): 312; Ohaus 1918: 10 (catalogue, distribution); 1934: 38 (citation, checklist); Blackwelder 1944: 235 (checklist).</p> <p>Type locality. Brazil, São Paulo, Jacareí.</p> <p>Type material. Lectotype female, here designated. (a) ‘ Byrsopolis / cribricollis Ohs. [handscript by Ohaus]’, (b) ‘Typus!’ [red label], c) ‘S. PAULO/Jacarehy’ [printed label] (1♀ MZUSP) (Figure 9).</p> <p>Paralectotype female. (a) ‘ São Paulo / Ypiranga /xii’, (b) ‘COTYPUS’, (c) ‘DIC/ BRASIL /E o SAO PAULO/YPIRANGA/ Ohaus-leg./ Coll. Martinez’, (d) ‘ Ohaus determin. 1912/ Byrsopolis cotype cribricollis Ohs. ♀ ’, (e) ‘H. &amp; A. HOWDEN COLLECTION, ex. A. Martinez Coll’, (f) ‘ Cotalpa (Byrs.) /cribricollis ♀ / Ohaus /A. Martinez-DET. 1951’, (g) ‘CMNEN/2001-0613’ (1♀ CMN, specimen examined through images by François Génier).</p> <p>Diagnosis. Body with reddish-brown colouration, without metallic green reflections; clypeal margin and pronotum dark brown; clypeus subrectangular with rounded anterior margin and anterior angles; antennal club about 1.5x longer than antennomeres II–VII combined; scutellar plate with broadly rounded base and sides straight; apex of elytral marginal areas with acute angles; humeri densely punctate; elytra with strong striae; protibial spur about 2.8x longer than wide.</p> <p>Redescription. Female. Body. Shape elongate oval; reddish-brown; frontoclypeal suture dark brown; antenna orange to dark brown; pronotal margins reddish brown at sides and dark brown at apex and base; scutellar plate marginal areas reddish-brown to dark brown, brighter and smaller in width at apex, increasing gradually to darker and thicker at base; orange setae scattered near apex; elytra reddish-brown, with marginal areas dark brown, without metallic green reflections (Figure 9 (a,b)). Length 30.28–31.00 mm. Humeral width 15.94 mm. Elytral width 17.69–18.50 mm. Head. Clypeus about 2x wider than long; subrectangular, with apex and angles weakly rounded; concavity weak at middle, moderate at apex; frontoclypeal suture almost straight, with curvilinear extremities very straight laterally; strongly striate; densely puctate; punctures moderate large to large; glabrous; frons about 1.50x longer than clypeus; punctures small to large, moderately to densely punctate; glabrous; mandibles with rounded external border, forming an almost right angle between scissorial region and apex; scissorial region slightly concave; small punctures at middle, moderately to densely punctate; scissorial region and apex of external border with setae sparsely to moderately distributed (apex and disc) (Figure 9 (c)); maxillary galea with three teeth; apical tooth longer than basal and median teeth (of similar size); small basal tooth with a small seta; apex of prementum moderately emarginate; last labial palpomere about 0.75x shorter than palpomere II (Figure 9 (e)); labrum weakly emarginated (Figure 9 (e)); antennomere II subconical, antennomere III subcylindrical, antennomeres IV and V subquadrate, antennomere VI subconical and VII subelliptical; antennal club about 1.50x longer than antennomeres II–VII combined. Thorax. Pronotum transverse, about 1.60x wider than long; punctures small to moderate, moderately to densely punctate (Figure 9 (a)); glabrous at disc; marginal areas with small and moderate, sparsely distributed setae; strongly striate; longitudinal midline with concavity at the apex; scutellar plate subtriangular, straight sides, broadly rounded at the base; setae small and moderate, moderately to densely distributed; surface moderately striate; punctures small to moderately large, sparsely to moderately punctate on the sides and densely at disc. Elytra. About 1.20x longer than wide; strongly striate; punctures small to large, sparsely to densely distributed; humeri with small to moderate punctures, densely punctate; interstriae with large coalescent punctures, (Figure 9 (a,b)); elytral apex with acute angles; glabrous. Legs. Protibial spur about 2.8x longer than wide; mesotibiae and metatibiae with two carinae; apex of mesotibia with about 14 spine-like setae and metatibiae with about 29 spine-like setae.</p> <p>Male. Unknown.</p> <p>Etymology. The species name is derived from Latin, cribricollis = crībrum (= riddled) and collis (= neck), referring to the strongly punctate pronotum.</p> <p>Distribution. São Paulo (Jacareí and Ipiranga) (Figure 19).</p> <p>Remarks. According to Ohaus (1912), the type specimens were deposited at ‘Museum of São Paulo’ (MZSP). But with the deposition of the Ohaus collection in ZMHB the lectotype from Jacareí found its way to Berlin, where it has been ever since. In agreement with Johannes Frisch (ZMHB), the specimen was finally transferred to the MZSP. The second specimen (paralectotype) from Ipiranga was found in the collection of Antonio Martínez, and is now deposited in the Canadian Museum of Nature (CMN).</p></div> 	https://treatment.plazi.org/id/03F09661FFEC807DFEB5E4B47B5152D5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFE98073FF63E01D7AD65074.text	03F09661FFE98073FF63E01D7AD65074.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis angeloottatii Medeiros, Seidel and Grossi 2022	<div><p>Byrsopolis angeloottatii Medeiros, Seidel and Grossi sp. nov. (Figure 10 (a–g))</p> <p>Holotype. Male, dissected, labelled. (a) ‘ Brasil, Mato Grosso do Sul,/ Três Lagoas, Três Lagoas /Agroflorestal Ltda. Horto Rio/ Verde parte A – Lageado /ethanol-baited FIT cerradão/fragment, 26.x.1993,/ 20°55 ʹ 31.94’S 52° 8 ʹ 15.14” W/ Flechtmann, C.A.H. leg.’ (CERPE).</p> <p>Paratypes [4 specimens]</p> <p>Males, not dissected. Three males with same data as holotype (CERPE, CEMT, MZSP); one male labelled, ‘ Brasil, Mato Grosso do Sul,/ Selvíria, UNESP Farm –/ Bovinocultura, 10. v.1991,/ 20°22 ʹ 36.13’S / 51°24 ʹ 58.92”W hand/collected Perozim, C.R. leg.’ (MEFEIS).</p> <p>Diagnosis. Clypeus, frons and elytra reddish-brown; antennae, pronotum, scutellar plate, venter and legs dark orange, without metallic green reflections; clypeus subrectangular, weakly bilobed, moderately concave; antennal club about 2.60x longer than antennomeres II–VII combined; scutellar plate subtriangular, with sides and basis rounded; elytral apex with right angles; elytra without striae, punctate, humeri weakly punctate; protibial spur about 2.60x longer than wide; parameres symmetrical, parallel, weakly angled internally, basal margin bisinuate.</p> <p>Description of holotype. Body. Shape elongate oval; without metallic green reflections; colour reddish-brown to brown-testaceous on clypeus, frons and elytra; pronotum and scutellar plate lighter than other body areas, yellowish-brown to orange-dark (Figure 10 (a, b)); frontoclypeal suture black on sides and reddish in middle; marginal areas of clypeus black on apex and reddish-brown on base; clypeus with orange setae; antenna yellow to orange; scutellar plate dark with marginal areas; dark brown. Length 19.90 mm. Humeral width 10.00 mm. Elytral width 11.25 mm.</p> <p>Head. Clypeus about 1.80x wider than long, subrectangular, weakly bilobed, sides almost straight (Figure 10 (a)), angles and basis rounded, anterior margin straight; anterior area of clypeus moderately concave, apex elevated; frontoclypeal suture bisinuate, with weakly V-shaped sinuosity at middle, sides rounded; frons with punctures small to moderately large, moderately to densely punctate; moderate to large setae, moderately to densely distributed; frons equal to clypeus in length; punctures small to moderately large, moderately to densely punctate, setae scattered near eyes, posterior region moderately punctate; moderately sized punctures setigerous; mandibles with rounded external border, with a small tooth at incisive region; maxillary galea with six teeth (apical region without teeth, two toothlets at middle, and three teethelets at base [toothlets without moderate setae]) (Figure 10 (e)); last maxillary palpomere with elliptical sensorial area (Figure 10 (e)); apex of prementum moderately emarginate (Figure 10 (g)); last labial palpomere about 2x longer than palpomere II (Figure 10 (g)); labrum weakly emarginated at middle of apex (Figure 10 (g)); antennomere III subcylindrical, antennomere IV subquadrate, antennomeres V–VI subconical; antennal club about 2.60x longer than antennomeres II–VII combined (Figure 10 (a)). Thorax. Pronotum transverse, about 1.50x wider than long (Figure 10 (a)); punctures small to moderately large, moderately to densely punctate; microsetae in moderate punctures, sparsely to moderately distributed; marginal areas with sparse punctures; scutellar shield subtriangular, with sides and basis weakly rounded; surface covered with small to large setae; setae on apex sparsely to densely distributed; punctures small to moderately large, base sparsely and disc densely punctate. Elytra. About 1.10x longer than wide; punctures small to large, moderately to densely punctate (Figure 10 (a,b)); microsetae in moderate punctures, sparsely to moderately distributed; indistinct striae, punctate; humeri weakly punctate; interstriae with moderately sized punctures. Legs. Protibial spur about 2.60x longer than wide; mesotibiae and metatibiae with two carinae. Aedeagus. Parameres symmetrical, parallel, elongated, separated by a Ushaped gap, rounded at apex, weakly obtuse on internal angles; lateral margins weakly excavated longitudinally; basis of parameres with a short carina; basal margin bisinuate (Figure 10 (c)).</p> <p>Female. Unknown.</p> <p>Variation. Length 19.90–21.00 mm. Humeral width 10.00– 10.70 mm. Elytral width 11.25– 12.00 mm.</p> <p>Etymology. The specific epithet is a atribute to Angelo Thadeu Ottati. He is currently a Federal Agricultural Auditor with MAPA (Ministério da Agricultura, Pecuária e Abastecimento). Angelo dedicated himself for years to sorting, cataloguing, pinning and recording specimens trapped in light and ethanol-baited flight traps at the company known then as Três Lagoas Agroflorestal (today Suzano Papel e Celulose), in Três Lagoas, state of Mato Grosso do Sul, Brazil. The type locality (Três Lagoas and Selvíria, Mato Grosso do Sul state) (Figure 19) has a predominant Cerrado vegetation, but there are also Atlantic Forest strips alternating with Cerrado close to the Paraná River.</p> <p>Remarks. All specimens are known only from the type locality. Byrsopolis angellottati sp. nov. resembles B. quadraticeps, but differs from this species by: the reddish colouration, clypeus more setose, premental apex moderately emarginate, labrum weakly emarginate; scutellar plate more rounded on sides; shorter protibial spur; metatibia with two carinae. Most specimens were collected with black light traps.</p> </div>	https://treatment.plazi.org/id/03F09661FFE98073FF63E01D7AD65074	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFE78071FF1BE3BE7AFF519F.text	03F09661FFE78071FF1BE3BE7AFF519F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis schmidti Medeiros, Seidel and Grossi 2022	<div><p>Byrsopolis schmidti Medeiros, Seidel and Grossi, sp. nov. (Figure 11 (a–g))</p> <p>Holotype. Male, dissected, labelled: (a) ‘ Brasil, BA, Barreiras,/ Serra Geral de Goiás,/ 15. xii. 2002, 160 km W,/ P.A. Schmidt leg’, (b) ‘COLEÇÃO E./&amp; P. GROSSI’ (1♂ CERPE).</p> <p>Paratypes [5 specimens]. Males, dissected, labelled. (a) ‘ Brasil, BA, Barreiras,/Serra Geral de Goiás,/ 15.xii. 2002, 160 km W,/P.A. Schmidt leg’ (1♂ CEMT); (b) ‘11/06/ Barreiras BA/ Monsanto’ (2♂ CERPE, 1♂ CEMT, and 1♂ ESALQ /USP).</p> <p>Diagnosis. Head, pronotum, abdomen and legs metallic green, with orange elytra presenting black spots on margins and interstriae; clypeus subrectangular, weakly trilobed; antennal club about 2.6x longer than antennomeres II–VII combined; scutellar plate subtriangular, with sides and basis weakly rounded; elytral apex with right angles; elytra with indistinct striae, smooth, humeri weakly punctate; protibial spur about 3.5x longer than wide; aedeagus symmetrical, parallel, with weak sinuous internal angles, basal margin weakly sinuate.</p> <p>Description of holotype. Body. Oval, elongated; colour metallic green; with elytra orange; without metallic green reflections, with marginal areas dark red to dark brown; clypeus with orange setae; antennomeres II–VII from orange colour to brown, antennomeres VIII–X dark brown; pronotal setae yellowish (Figure 11 (a,b)). Length 21.10 mm. Humeral width 9.50 mm. Elytral width 11.60 mm. Head. Clypeus about 1.60x wider than long; subrectangular, rounded angles, weakly trilobed apex; weakly concave from base to sides, moderately concave at apex (Figure 11 (a)); frontoclypeal suture with curvilinear extremities and moderately rounded laterally; punctures small to moderate, densely punctate; setae small to moderate, moderately to densely distributed; frons about 0.80x shorter than clypeus; punctures small to moderately large, moderately to densely punctate; glabrous; scissorial region of mandibles with three projections (Figure 11 (d)); small to moderate punctures, disc and apex sparsely to densely punctate; scissorial region with moderate setae, sparsely to moderately distributed; maxillary galea with three long, equidistant, teeth in dorsal view (Figure 11 (e)); two small punctures near basal tooth; moderately sized setae on middle and basal teeth; last maxillary palpomere with oblong sensorial area; premental apex weakly emarginate; last labial palpomere about 2x longer than palpomere II (Figure 11 (f)); labrum moderately emarginated, densely setose (Figure 11 (g)); antennomere III subcylindrical, antennomeres IV–V subrectangular, antennomeres VI–VII subconical; antennal club about 2.6X longer than antennomeres II–VII combined. Thorax. Pronotum transverse, about 1.60x wider than long; punctures small to moderate, moderately to densely punctate; glabrous (Figure 11 (a)); marginal areas with small punctures, sparsely to moderately punctate, moderate to large setae, moderately to densely distributed, setae at base densely distributed; longitudinal midline without punctures; scutellar plate subtriangular, sides and base weakly rounded; punctures small to moderately large, moderately to densely punctate; apex and disc with small to large setae, moderate to densely distributed (Figure 11 (a)). Elytra. About 1.10x longer than wide; punctures small to large in size, moderately to densely punctate; humeri with small to moderate setae, sparsely distributed near scutellar plate; small to moderate setae on marginal areas up to elytral suture, moderately to densely distributed; elytral apex with right angles; distinct striae, smooth surface (Figure 11 (a,b)), humeri weakly punctate. Legs. Protibial spur about 3.5x longer than wide; mesotibiae and metatibiae with two carinae. Aedeagus. Parameres symmetrical, parallel, separated by a U-shaped gap, moderately elongate, rounded at apex, with weakly sinuous internal angles; lateral surface of parameres weakly excavated longitudinally, forming a small carina; middle weakly striate; with weak longitudinal concavity; basal margin moderately sinuate (Figure 11 (c)).</p> <p>Female. Unknown.</p> <p>Variation. Length 21.10–24.65 mm. Humeral width 9.50–12.25 mm. Elytral width 11.60– 13.40 mm. One of the paratypes exhibits a greenish colour in apical marginal areas and elytral suture; clypeal apex can be weakly to moderately trisinuate.</p> <p>Etymology. The specific epithet is a atribute to Paulo Almeida Schmidt, a Brazilian agronomist who collected the first specimens of the species.</p> <p>Distribution. This species is known from the type locality only, situated next to Serra Geral de Goiás, Goiás state, in the extreme west of the municipality of Barreiras, Bahia state (Figure 19). The type locality consists of Cerrado area, and hosts the only species of the genus exclusively from Northeast Brazil.</p> <p>Remarks. Byrsopolis schmidti sp. nov. is very different from all other species because of its almost totally metallic green colouration, a weakly trilobed clypeal apex, and antennal club about 3x longer than antennomeres II–VII combined. Moreover the elytra are orange with black elytral suture and epipleuron. The addition of this species increases the taxonomic and morphological limits of the genus.</p> </div>	https://treatment.plazi.org/id/03F09661FFE78071FF1BE3BE7AFF519F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFE58076FF69E2AB7B61501D.text	03F09661FFE58076FF69E2AB7B61501D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis burmeisteri Medeiros, Seidel and Grossi 2022	<div><p>Byrsopolis burmeisteri Medeiros, Seidel and Grossi, sp. nov. (Figure 12 (a–h))</p> <p>Holotype. Male, dissected, labelled (a) ‘ S. Paulo / Cantareira / L. Guimarães [handwritten]/ XI.940’, (b) ‘MZUSP’ (1♂ MZUSP).</p> <p>Paratypes [13 specimens]. Males and females, dissected (only males), labelled: (a) ‘Coleção Entomológica Adolph/Hempel – Instituto Biológico/IBSP – IB – 0.006.537’, (b) ‘Cantareira./S. Paulo./11.936’, (c) ‘Coll. J. Guerin./S. Paulo. Brasil./3995’, (d) ‘ Cotalpa cribicolis Ohs. [sic, cribricollis]’ (1♀ IBSP); (a) ‘Coleção Entomológica Adolph/Hempel – Instituto Biológico/IBSP – IB – 0.006.536’, (b) ‘Cantareira./S. Paulo./12.936’, (c) ‘Coll. J. Guerin./S. Paulo. Brasil./2039’, (d) ‘ Cotalpa / castanea Burm. /comp. M. P. 1936’ (1♂ IBSP); (a) ‘Coleção Entomológica Adolph/Hempel – Instituto Biológico/IBSP – IB – 0.006.535’, (b) ‘Cantareira./ S. Paulo./12.935’ (c) ‘Coll. J. Guerin./S. Paulo. Brasil./3597’ (1♂ IBSP); (a) ‘Brasil – S. Paulo/Est. S. Paulo’, (b) ‘ Byrsopolis /castanea/Burm. [handwritten]’, (c) ‘MNRJ’ (1♀ MNRJ); (a) ‘ BRASIL / Cantareira/Bairro S. Paulo/Dirings\ XII.1939 ’ (b) ‘Birsopolis castanea Burm. [sic, Byrsopolis, handwritten]/Dr. Nick determ’. (c) ‘MZUSP’ (1♂ MZUSP); (a) ‘Cantareira/I.53. [handwritten]/ Est. S; Paulo/Coll. H. Zellibor’ (1♂ CEMT); (a) ‘Cantareira 20.I.54 [date handwritten]/Est. S. Paulo/Coll. H. Zellibor’ (1♂ CERPE); (a) ‘S. Paŭlo [sic, Paulo]/Campinas [handwritten]’ (b) ‘ Byrsopolis / castanea Burmeister /Museum f. Naturkunde/ Berlin’ (1♀ ZMHB); (a) ‘Cantareira/S. Paulo 31.xii.39/Coll. Zellibor-Hauff’, (b) ‘H. &amp; A. HOWDEN/Collection/ex. A. Martinez coll’, (c) ‘ Cotalpa /castanea/Burm./A. Martinez-DET 1948’ (specimen examined through images), (1♂ CMN); same, except (a) ‘Cantareira/S. Paulo 1.I.41/Coll.H.Zellibor’ (specimen examined through images), (1♂ CMN); same, except (a) ‘10.xii.41’ (specimen examined through images), (1♂ CMN); São Paulo. ‘S. Paŭlo [sic, Paulo]/Mogy das Crŭzes [sic, Cruzes]/C. P. Vianna S’ (1♂ ZMHB); (a) ‘ Sao Paulo, Brasil./Serra da Cantareira/ 1000 m 20.11.52/leg.G.H.Nick’ (b) ‘WORLD/SCARAB./DATABASE/WSD00347503’ (1♂ ZSM).</p> <p>Diagnosis. Body brown-reddish to dark brown with metallic green reflections; elytra orange, dark orange to dark brown borders; clypeus subrectangular, with anterior margin weakly emarginate, almost straight; antennal club about 1.2x (female) to 1.7x (male) longer than antennomeres II–VII combined; scutellar plate with broadly rounded basis; elytral apex with right angles; elytra with strong striae, humeri moderately punctate; protibial spur about 4x longer than wide; parameres symmetrical, strongly divergent and angulose, proximal margin weakly bisinuate.</p> <p>Description of holotype. Body. Oval, elongated; reddish-brown to dark brown; metallic green reflections on legs, marginal areas of pronotum and pygidial disc; frontoclypeal suture dark brown to black; antennae orangish to dark brown; elytra orange, dark orange to dark brown; interstriae with dark brown to black punctures (Figure 12 (a,b)). Length 25.85 mm. Humeral width 12.82 mm. Elytral width 15.03. Head. Clypeus about 1.80x wider than long; subrectangular, moderately rounded at angles, apex weakly emarginate, almost straight (Figure 12 (a)); clypeal longitudinal midline moderately concave, weak concavity at sides and apex; frontoclypeal suture with weak curvilinear extremities, almost straight; sinuosities weakly rounded at apex, rounded corners; surface of clypeus moderately striate; base with small to moderate punctures, moderately to densely punctate; glabrous; frons about 0.90x shorter than clypeus; punctures moderate to large, densely punctate; glabrous; two concavities near frontoclypeal suture; mandibles with weak concavity at middle of scissorial region, rounded apex; setae near marginal areas of scissorial region, small to moderate, moderately to densely distributed; punctures small, sparsely to moderately punctate near scissorial region (Figure 12 (e)); maxillary galea with three teeth, middle and basal teeth closest; apical tooth longer and larger than basal tooth, basal tooth longer and larger than median tooth; basal tooth with elongated setae (Figure 12 (f)); last maxillary palpomere with elliptic sensorial area; apex of prementum with strong emargination; last labial palpomere about 2.50x longer than palpomere II (Figure 12 (g)); labrum moderately emarginate, sparsely setose (Figure 12 (h)); antennomere III subcylindrical, IV–V subquadrate, VI–VII subconical; antennal club about 1.70x antennomeres II–VII combined. Thorax. Pronotum transverse, about 1.60x wider than long; punctures small to moderately large, moderately to densely punctate; glabrous; two small lateral concavities near posterior margin; marginal punctures small to moderate, sparsely to densely punctate; setae at marginal areas near to humeri small and moderately distributed; punctures forming a longitudinal midline; scutellar plate subtriangular, base broadly rounded (Figure 12 (a)); punctures small to moderately large, sparsely to moderately puctate at base and densely punctate at apex and disc surface; scutellar plate with apical setae small to moderate, moderately to densely punctate. Elytra. About 1.10x longer than wide; strongly striate; punctures small to large, moderately to densely punctate; glabrous (Figure 12 (a,b)); humeri with punctures moderately distributed; border with few setae; interstriae with large, coalescent punctures. Legs. Protibial spur about 4x longer than wide; mesotibiae and metatibiae with two carinae. Aedeagus. Parameres strongly sinuous laterally, strongly divergent (Figure 12 (c)); symmetrical dorsally, asymmetrical laterally and ventrally; right paramere with thicker base; lateral margins weakly excavated longitudinally, forming a small carina; basal margin weakly bisinuate.</p> <p>Description of paratype female. Similar to holotype, differing in the following aspects.</p> <p>Body. Oval, elongate, to weakly spheroidal, more robust than male (Figure 12 (d)). Head. Antennal club about 1.20x longer than antennomeres II–VII combined. Thorax. Pronotum with two concavities at base and near to humeri; scutellar plate with rounded base (Figure 12 (d)), moderately punctate; border sparsely to moderately punctate; metallic green or red reflections.</p> <p>Variation. Males. Length 25.85–29.07 mm. Humeral width 12.82–15.00 mm. Elytral width 15.03–16.35 mm. Females. Length 30.78–37.81 mm. Humeral width 17.16–20.73 mm. Elytral width 19.90–22.37 mm. Some males show distinct colouration with pronotum the same colour as elytra, which can be caused by natural wear with the storage time.</p> <p>Some specimens exhibit metallic green reflections on some parts of the body such as the ventral side, pygidial disc and pronotal concavity.</p> <p>Etymology. The specific epithet is a atribute to German entomologist Carl Hermann Conrad Burmeister (1807–1892), who described the genus Byrsopolis and two species within it.</p> <p>Distribution. São Paulo (São Paulo and Mogi das Cruzes). The distribution ranges where the species occur are located in the rock formations of the Serra do Mar and Mantiqueira (São Paulo state), which constitutes the denominated Planalto Atlântico with vegetation typical of the Atlantic Forest and high elevations (Figure 19).</p></div> 	https://treatment.plazi.org/id/03F09661FFE58076FF69E2AB7B61501D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFE28074FF65E3297957568A.text	03F09661FFE28074FF65E3297957568A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis vazdemelloi Medeiros, Seidel and Grossi 2022	<div><p>Byrsopolis vazdemelloi Medeiros, Seidel and Grossi, sp. nov. (Figure 13 (a–g))</p> <p>Holotype. Male, dissected, labelled. (a) ‘ BRASIL: Mato Grosso./Porto Estrela. Est. Eco/ Serra das Araras. Trilha /Andorinhas. Pitfallfaec/ XI-2013.RVNunes(leg.)’ (1♂ CERPE).</p> <p>Paratypes [15 specimens].. Males, labelled: Mato Grosso. (a) ‘ BRASIL: Mato Grosso./ Porto Estrela. Est. Eco / Serra das Araras. Trilha / Andorinhas. Pitfallfaec / XI-2013. RVNunes(leg.)’ (1♂ CERPE, 2♂ CEMT); same, except (a) ‘ Porto Estrela, EE Serra das/ Araras -sede, 15°39 ʹ 09’S, 57° 12 ʹ 52”W, 18–19.xi.2017,/light, R.J.P. Machado &amp;/R. Nunes’ (1♂ CEMT); same, except (a) ‘ Araras – heliporto, 15°39 ʹ 19’/S, 57°12 ʹ 50”W, 20.xi.2017,/light, RJP Machado’ (2♂ CEMT); (a) ‘ Brasil, Mato Grosso, Barra do/ Bugres, CE. AS., Alojamento,/ 20.xi.1987, C.A. Caetano leg’ (1♂ CEMT); (a) ‘BR – MT – B. BUGRES/CE. SA/ALOJAMENTO/20-XI-87/WILSON A. PINTO’ (1♂ CEMT); (a) ‘ Brasil, MT, Barra do Bu /gres, Est. Ecol. Serra / das Araras 8.XII/1988 C.A. Caetano’ (1♂ CERPE, 1♂ CEMT). Mato Grosso do Sul. (a) ‘ Ordem: Coleoptera / Família: Scarabaeidae’, (b) ‘ Stefanes, R /Dourados 10,XII,2000’, (c) ‘ Byrsopolis sp./P. Grossi det. 2009’./ (1♂ CERPE). Paraguay. (a) ‘ PARAGUAY: Dept. San Pedro /off Hwy. No. 3, E. of Santa Rosa /RIO VERDE, YAGUARETE FOREST/ 22-XII-1995, J. D. Beierl lgt. BL/ S 23° 47’ 44 ’, W 56° 03’ 59”, 225 m / Moist – Warm Temperature Forest’ (1♂ WWPC); (a) same, (b) ‘ BYRSOPOLIS /SP./ Det. M.L. Jameson 2019’, (c) ‘WORLD/SCARAB./ DATABASE/WSD00347555’ (1♂ RCPC); (a) same, (b) ‘ Coll. Matthias / Seidel 2021’, (c) ‘WORLD/SCARAB./ DATABASE/WSD00347556’ (1♂ MSPC) ’. No collection data (1♂ MZUSP).</p> <p>Diagnosis. Body almost totally orange to reddish brown, with metallic green reflections ventrally; clypeus parabolic, wider than long; antennal club about 2.2x longer than antennomeres II–VII combined; scutellarplate subtriangular, moderately rounded at basis; elytral posterior margins with right angles; elytra with strong striae, humeri moderately punctate; protibial spur about 4x longer than wide; parameres symmetrical, weakly divergent, with or without small, thin, spine-like setae, sparsely to moderately distributed.</p> <p>Description of holotype. Body. Oval, elongate; orange to reddish-brown, frontoclypeal suture dark reddish brown to black; orange setae on apex of scutellar plate (Figure 13 (a,b)); metallic green reflections ventrally. Length 23.30 mm. Humeral width 12.60 mm. Elytral width 15.00 mm. Head. Clypeus about 2.20x wider than long; parabolic;moderately concave near sides and frontoclypeal suture, base weakly concave; frontoclypeal suture with curvilinear extremities weakly straight laterally, with obtuse angles; punctures small to moderately large, moderately to densely punctate; microsetae at basal region, between frons and clypeus; frons about 1.20x longer than clypeus; moderate concavity posteriorly the frontoclypeal suture; punctures small to moderately large in size, moderately to densely punctate; glabrous; mandibles with external border forming a straight angle with scissorial region; scissorial region strongly elongated transversely, narrow longitudinally; glabrous (Figure 13 (d)); with two punctures small and sparse; galea with five teeth, apical and basal tooth moderate in size (Figure 13 (e)), middle region with three small teeth, setae extending over small teeth; last maxillary palpomere with moderate elliptic sensorial area; apex of prementum strongly emarginate; last labial palpomere about 1.80x longer than palpomere II (Figure 13 (f)); labrum strongly emarginate, moderately setose (Figure 13 (g)); antennomere III subcylindrical, antennomeres IV–V subquadrate, antennomeres VI–VII subconical; antennal club about 2.2x longer than antennomeres II–VII combined. Thorax. Pronotum transverse, about 1.60x wider than long; punctures small to moderately large, moderately to densely punctate (Figure 13 (a)); setae near humeri; punctures forming a longitudinal midline; glabrous; scutellar plate subtriangular, base moderately rounded; setae small to moderate (Figure 13 (a)); punctures small to moderately large, sparsely to moderately punctate at base and densely punctate at apex. Elytra. About 1.10x longer than wide; punctures small to large, moderately to densely punctate (Figure 13 (a,b)); microsetae in small and moderate punctures. Legs. Protibial spur about 4x longer than wide; mesotibiae and metatibiae with one carina. Aedeagus. Parameres weakly divergent, almost parallel, asymmetrical, V-shaped gap angle between parameres (Figure 13 (c)); basal margin with three small longitudinal lines (one at middle and one at each side); longitudinal concavity at middle; glabrous; slightly asymmetrical phallobase; right paramere more divergent than left.</p> <p>Female. Unknown.</p> <p>Variation. Length 23.30–28.00 mm, humeral width 12.60–13.50 mm, elytral width 15.00– 16.50 mm.</p> <p>Etymology. The specific epithet is a atribute to Dr Fernando Zagury Vaz-de-Mello (UFMT), in honour of his contributions to Coleoptera research, as well as for providing part of the type series of this species and others.</p> <p>Distribution. The regions where it is found in Brazil are in the two municipalities of ‘Barra do Bugres’ and ‘Porto Estrela’; it is also found in the ‘Departamiento de San Pedro’ in Paraguay (Figure 19).</p> <p>Remarks. This species resembles Byrsopolis crassa, but can be differentiated from it through many characters such as: the reduced body size and reddish colour; concavities posterior to the frontoclypeal suture; narrow mandibles, smooth, with scissorial region elongated transversely. Furthermore, some specimens exhibit setae on the aedeagus. Byrsopolis crassa have large body size (view redescription), colour orange to browntestacious; apex of clypeus moderately concave; wide mandibles, elongate longitudinally; aedeagus glabrous.</p> </div>	https://treatment.plazi.org/id/03F09661FFE28074FF65E3297957568A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFE0804AFF0AE5C47B8256A7.text	03F09661FFE0804AFF0AE5C47B8256A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis ohausi Medeiros, Seidel and Grossi 2022	<div><p>Byrsopolis ohausi Medeiros, Seidel and Grossi, sp. nov. (Figure 14 (a–f))</p> <p>Holotype. Male, dissected, labelled. (a) ‘ Byrsopolis Grão Mogol / Brasil. Minas Gerais, Grão / Mogol, 16.i.2004, J. De Nadai /&amp; N. dos Anjos legs’ (1♂ CERPE).</p> <p>Paratype. Male, dissected, labelled. (a) ‘ Grão Mogol-MG – Brasil / 15.xii.2002/N. Anjos col’, (b) ‘ B. laticollis?/ Burmeister, 1855 /P. Grossi det. 2015’ [handwritten] (1♂ CERPE).</p> <p>Diagnosis. Body yellowish-brown to reddish-brown, with metallic green reflections; clypeus parabolic; antennal club about 1.5 longer than antennomeres II–VII combined; scutellar plate subtriangular, with broadly rounded basis; elytral posterior margins with divergent angles; elytra strongly striate and humeri moderately punctate; protibial spur about 4.2x longer than wide; parameres symmetrical, weakly divergent; basal margin bilobed.</p> <p>Description of holotype. Body. Oval, elongate; yellowish-brown to reddish-brown (Figure 14 (a)), with metallic green reflections; frontoclypeal suture dark reddish-brown to black; yellow, orange or reddish-brown setae on scutellar plate. Length 23.10 mm. Humeral width 12.60 mm. Elytral width 13.15 mm. Head. Clypeus about 1.75x wider than long; parabolic; weakly concave only at marginal area; frontoclypeal suture with curvilinear extremities moderately rounded and almost straight laterally, with obtuse angles; punctures small to moderate in size, moderately to densely punctate; frons about 1.80x longer than clypeus; weak concavity on disc; punctures small to moderately large, moderately to densely punctate; glabrous; mandibles with moderately rounded external border, scissorial region weakly rounded, with two setae, one near apex and one near intern border (Figure 14 (c)); galea with four teeth, one long apical tooth, one median tooth, and one basal tooth with three small toothlets, each with a single seta, anterior tooth with two setae (Figure 14 (d)); last maxillary palpomere with large, elliptic sensorial area, not exceeding middle of palpomere; apex of prementum moderately emarginate; last labial palpomere about 2.4x longer than palpomere II (Figure 14 (e)); labrum weakly emarginate, moderately setose (Figure 14 (f)); antennomere III subcylindrical, antennomeres IV–V subquadrate, antennomeres VI–VII subconical; antennal club about 1.5x longer than antennomeres II–VII combined. Thorax. Pronotum transverse, about 1.70x wider than long (Figure 14 (a)); punctures small to moderate, moderately to densely punctate; humeri with small setae sparsely distributed; one transverse line formed by a concavity on each side near marginal areas; scutellar plate subtriangular, base broadly rounded (Figure 14 (a)); small setae on the apex, sparsely to moderately distributed; punctures small to moderately large, plate sparsely punctate at base, moderately punctate at apex. Elytra. About 1.10x longer than wide; strongly striate; punctures small to large, moderately to densely punctate; interstriae with strong coalescent punctures (Figure 14 (a)); elytral apex with divergent angles; humeri with small setae in moderate punctures. Legs. Protibial spur about 4.2x longer than wide; mesotibiae and metatibiae with one carina. Aedeagus. Parameres rounded, weakly sinuous, weakly divergent, asymmetrical, acute angle between parameres; parameres wide and flattened (Figure 14 (b)); lateral margins moderately excavated, longer on left paramere, forming a carina; proximal margin sinuated; longitudinal midline concave; glabrous.</p> <p>Female. Unkown.</p> <p>Variation. Length 24.10 mm. Humeral width 13.50 mm. Elytral width 15.00 mm.</p> <p>Etymology. The specific epithet is a atribute to the great German entomologist Friedrich Ohaus (1864–1946), also called the ‘Father of Rutelinae’, who established the modern systematics of Rutelinae that is still largely in use today.</p> <p>Distribution. This species is only found in the ‘Serra do Espinhaço’, north of Minas Gerais, in the Grão Mogol city (Figure 19).</p> <p>Remarks. This species is very similar to Byrsopolis laticollis and B. blanchardi sp. nov., but differs from these two by the characters of the clypeus being 1.75x wider than long, elytral apexes divergent, aedeagus with longitudinal midline formed by a concavity, and wide, flat parameres. Byrsopolis laticollis have clypeus 2.1x wider than long; elytral apexes with acute angles;aedeagus with rounded and thick parameres. Byrsopolis blanchardi sp. nov. have clypeus 2.2x wider than long; elytral apexes truncate; aedeagus with rounded and thin parameres.</p> </div>	https://treatment.plazi.org/id/03F09661FFE0804AFF0AE5C47B8256A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFDE8048FF11E5E37A3B5753.text	03F09661FFDE8048FF11E5E37A3B5753.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis blanchardi Medeiros, Seidel and Grossi 2022	<div><p>Byrsopolis blanchardi Medeiros, Seidel and Grossi, sp. nov. (Figure 15 (a–f))</p> <p>Holotype. Male, dissected, labelled. Bahia. (a) ‘Águas Vermelhas ixi.1992, E.J. Grossi leg’ (1♂ EPGC).</p> <p>Paratypes [2 specimens]. Males, dissected, labelled. Minas Gerais. (a) ‘ Brasil, Minas Gerais, Divisa / Alegre. entroncamento/BR116-MG000. 13.xii.2012./ Pto.08. manual dia. E.&amp; P./ Grossi legs’ (1♂ CERPE); (a) ‘ Brasil, Bahia / Encruzilhada /BR 116/ Divisa xi./1970. O. Roppa col’ (1♂ MZFS).</p> <p>Diagnosis. Body with uniform colouration; from reddish-brown to black, with metallic green reflections ventrally; clypeus trapezoidal, with broadly rounded angles; antennal club about 1.75x longer than antennomeres II–VII combined; scutellar plate subtriangular, moderately rounded at basis, laterally straight; elytral apex truncate; elytra strongly striate, humeri moderately punctate; protibial spur about 4.4x longer than wide; parameres symmetrical, thin, weakly divergent.</p> <p>Description of holotype. Body. Oval rounded; reddish-brown to black, frontoclypeal suture dark reddish-brown to black (Figure 15 (a)); metallic green reflections ventrally. Length 23.30 mm. Humeral width 12.60 mm. Elytral width 15.00 mm.</p> <p>Head. Clypeus about 2.2x wider than long; subtrapezoidal, broadly rounded angles; weakly concave; frontoclypeal suture with curvilinear extremities weakly rounded, almost straight laterally, with obtuse angles; punctures small to moderately large, moderately to densely punctate; frons about 1.1x longer than clypeus; punctures small to moderately large, base sparsely to moderately punctate, apex densely punctate, glabrous; mandibles with a weak projection at middle, with three small sparse setae, weakly punctate (Figure 15 (c)); galea with two teeth, one apical tooth, basal tooth small, truncate, with small setae (Figure 15 (d)); last maxillary palpomere with moderate elliptic sensorial area; apex of prementum weakly emarginate; last labial palpomere about 2.20x longer than palpomere II (Figure 15 (e)); labrum moderately emarginate, moderately setose (Figure 15 (f)); antennomere III subcylindrical, antennomeres IV–V subquadrate, antennomeres VI–VII subconical; antennal club about 1.75x longer than antennomeres II–VII combined. Thorax. Pronotum transverse, about 1.7x wider than long (Figure 15 (a)); punctures small to moderately large, moderately punctate, base densely punctate; longitudinal midline with punctures; glabrous; scutellar plate subtriangular, base moderately rounded, laterally straight; glabrous (Figure 15 (a)); punctures small, base moderately punctate, apex sparsely to moderately punctate, base densely punctate. Elytra. About 1.1x longer than wide; strongly striated; humeri with small to moderate punctures, moderately punctate; elytral punctures small to large, moderately to densely punctate; interstriae near elytral suture with strong coalescent punctures (Figure 15 (a)); elytral apex truncate; humeri with microsetae in moderate punctures. Legs. Protibial spur about 4.4x longer than wide; mesotibiae and metatibiae with two carinae. Aedeagus. Parameres weakly sinuous, weakly divergent, narrow, asymmetrical, thin, apex rounded; rounded gap between parameres (Figure 15 (b)); lateral paramere margins moderately excavated, longer on left paramere, forming a carina; proximal margin weakly bisinuate; longitudinal midline weakly concave; glabrous.</p> <p>Female. Unkown.</p> <p>Variation. Length 23.30–26.00 mm. Humeral width 12.60–13.50 mm. Elytral width 15.00– 16.50 mm.</p> <p>Etymology. The specific epithet is a atribute to the great French zoologist and entomologist Charles Émile Blanchard (1819–1900), whose significant contributions also include the description of one Byrsopolis species.</p> <p>Distribution. This species occurs in the south of Bahia state, in the municipality of Encruzilhada and in the north of Minas Gerais state, in the cities of Divisa Alegre and Águas Vermelhas (Figure 19), considered a transition zone, with vegetations and faunas of Atlantic Forest, Caatinga and Cerado biomes, called Mata Seca.</p> <p>Remarks. The species resembles Byrsopolis laticollis, and differs by the frons and clypeus subequal in size; scutellar shield with punctures sparsely distributed at the base and densely distributed at the apex; elytral apex truncate; parameres slender. Byrsopolis laticollis have frons 1.3x longer than clypeus; scutellar shield sparsely to moderately distributed at the marginal areas</p> </div>	https://treatment.plazi.org/id/03F09661FFDE8048FF11E5E37A3B5753	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFDC8048FEC9E48F7A66548A.text	03F09661FFDC8048FEC9E48F7A66548A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis aenescens , Ohaus 1926	<div><p>Byrsopolis aenescens Ohaus, 1926 (Figures 16 (a–d), 17(a–h))</p> <p>Byrsopolis aenescens Ohaus, 1926, p. 225 –226 (original description); Machatschke 1972: 3 (catalogue, distribution); Krajcik 2008: 58 (checklist); Soula 2010a: 44 (redescription, distribution); Soula 2010b: 54 (key to species, illustration); Carvalho and Grossi 2018: 372 (checklist, distribution); Hielkema and Hielkema 2019: 126 (checklist, distribution).</p> <p>Cotalpa (Byrsopolis) aenescens (Ohaus, 1926): 225–226; Ohaus 1934: 37 (citation, checklist); Blackwelder 1944: 235 (checklist).</p> </div>	https://treatment.plazi.org/id/03F09661FFDC8048FEC9E48F7A66548A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFDB804CFF28E5C77C3C53A2.text	03F09661FFDB804CFF28E5C77C3C53A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis chassaini Soula 2010	<div><p>Byrsopolis chassaini Soula, 2010a new junior subjective synonym</p> <p>Byrsopolis chassaini, 2010a: 46 (original description); Carvalho and Grossi 2018: 372 (checklist, distribution); Hielkema and Hielkema 2019: 127 (checklist, distribution).</p> <p>Type locality. Nouveau Chantier, Saint-Laurent-du-Maroni, French Guiana.</p> <p>Type material. Byrsopolis aenescens Ohaus, 1926: Lectotype male, dissected, here designated. (a) ‘ HOLOTYPUS / Byrsopolis / aenescens Ohaus 1926 /labelled by MHHUB 2010’ [red label], (b) ‘Oyapok Oberlf./Pied Saüt, III.1918 /’, (c) ‘ GUYANE FRANÇ SE/Nouveau Chantier/ COLLECTION LE MOULT’, (d) ‘Type’ [red label], (e) ‘ Byrsopolis /aenescens/Ohs’. [handwritten by Soula], (1♂ ZMHB) (Figure 16 (a-b, d)); Byrsopolis chassaini Soula, 2010a: Holotype male, dissected. (a) ‘ Holotype / Byrsopolis / chassaini S./2010 Soula’ [red label, handwritten by Soula], (b) ‘Mussé des Confluences/MHNL/47030019’, (c) ‘Cacao/Guyane Fç./M Soula det 19’ [handwritten by Soula], (1♂ CCECL, specimen examined through images) (Figure 16 (e–h)); (1♂).</p> <p>Additional material. [15 specimens] (a) ‘ Guiana Francesa, Route de/St. George, Pk 125, 07./iii.2008, P.L. J.G./ Giúglaris Leg’ (1♂ CERPE) (Figure 16 (c)); ‘ FRENCH GUIANA: Saül, Point / de Vue, Belvédère de Saul-/ Montagne Pelée, Saül, Colline /en lisière forêt et bourg de/ Saül, N 3°37 ʹ 22 ’ W 53°12 ʹ 58” ’ // ‘ 09.iii.2017, PVB:/automatic light trap/(blue)’ // ‘ Byrsopolis aenescens / Ohaus, 1926 /det. M. Seidel 2021’ // ‘WORLD/SCARAB./ DATABASE/ WSD00347508’ (1♂ MSPC); same, except ‘ 19.iv.2017, PVB:/automatic light trap/(blue)’ // ‘WORLD/SCARAB./ DATABASE/WSD00347509’ (1♂ MSPC); same, except ‘ 23.iii.2017, PVB:/ automatic light trap/(blue)’ // ‘WORLD/SCARAB./ DATABASE/[WSD00347510- WSD00347512]’ (3♂ MSPC); same, except ‘ 17.iii.2017, PVB:/automatic light trap/(blue)’ // ‘WORLD/SCARAB./ DATABASE/WSD00347513]’ (1♂ MSPC); same, except ‘ 06.iv.2017, PVB:/ automatic light trap/(blue)’ // ‘WORLD/SCARAB./ DATABASE/[WSD00347517- WSD00347518]’ (2♂ MSPC); same, except ‘ 03.iii.2017, PVB:/automatic light trap/(blue)’ // ‘WORLD/SCARAB./ DATABASE/WSD00347519’ (1♂ MSPC); ‘ FRENCH GUIANA: Piton/ <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-52.319973&amp;materialsCitation.latitude=3.73225" title="Search Plazi for locations around (long -52.319973/lat 3.73225)">Rocheux de l’Armontabo</a>, Saint/Georges, Savane-roche,/sommet de l’inselberg (353 m),/ N 3°43 ʹ 56,1 ’ W 52°19 ʹ 11,9” ’ // 11.ii.2018,/L.T: light trap’ // ‘ Byrsopolis aenescens / Ohaus 1926 /det. M. Seidel 2021’ // ‘WORLD/SCARAB./ DATABASE/WSD00347514’ (1♂ MSPC); same, except ‘ 16.ii.2018,/L.T: light trap’ // ‘WORLD/SCARAB./ DATABASE/WSD00347515’ (1♂ MSPC); same, except ‘ 16.ii.2018, PVP:/automatic light trap/(pink)’ // ‘WORLD/ SCARAB./ DATABASE/WSD00347516’ (1♂ MSPC). No collection data.</p> <p>Diagnosis. Body testacious-brown to reddish-brown, without metallic green reflections; clypeus subrectangular; antennal club about 1.97x longer than antennomeres II–VII combined; scutellar plate subtriangular, base weakly rounded; elytral apex with divergent angles; elytra with indistinct striae, humeri moderately punctate; protibial spur about 3.5x longer than wide; parameres symmetrical, weakly divergent; sides of parameres almost linear, basal margin with strong small emarginations.</p> <p>Description. Male. Body. Oval, elongate; testaceous-brown to reddish-brown, only elytra and antennomeres testaceous-brown to dark orange (Figures 16 (a), 17(a–c)), frontoclypeal suture black; reddish-brown setae on scutellar shield; without metallic green reflections. Lectotype. 27.00 mm. Elytral width 15.50 mm. Head. Clypeus about 2x wider than long; semioval; weakly concave only at marginal areas; frontoclypeal suture with weak curvilinear extremities; punctures small to moderate, moderately to densely punctate; frons about 1.50x longer than clypeus; punctures small to large, moderately punctate on base, densely punctate on apex; glabrous; mandibles with moderately rounded external border, scissorial region weakly rounded, with few and sparse setae on disc, base of scissorial region with short and dense setae, small, tooth-like projection on mesal internal border (Figure 17 (e)); galea with four teeth, one long apical tooth, two median small toothlets, and one small basal tooth with a single seta (Figure 17 (f)); apex of prementum moderately emarginate, with triangular lobes; last labial palpomere about 2.87x longer than palpomere II (Figure 17 (g)); labrum weakly emarginate, moderately setose (Figure 17 (h)); antennomere III subcylindrical, antennomeres IV–V subquadrate, antennomeres VI– VII subconical; antennal club about 1.97x longer than antennomeres II–VII combined. Thorax. Pronotum transverse, about 1.67x wider than long (Figures 16 (a), 1616(a)); punctures small to moderate, moderately punctate at disc, densely punctate at marginal areas; one transverse line at apex formed by a small concavity on each side near anterior marginal areas; scutellar plate subtriangular, base weakly rounded (Figure 17 (a)); small setae on apex, moderately distributed; punctures small to moderate, sparsely punctate at apex, moderately punctate at disc and base. Elytra. About 1.15x longer than wide; punctures small to moderately large, moderately punctate at disc and marginal areas; interstriae indistinct (Figure 17 (a)); elytral apex with divergent angles; humeri with microsetae in moderate punctures, sparsely distributed. Legs. Protibial spur about 3.5x longer than wide; mesotibiae with one carina and metatibiae with two carinae. Aedeagus. Parameres apex rounded, weakly sinuous, moderately divergent, symmetrical; acute angle between parameres; parameres wide (Figures 16 (b-c), 17(d)); lateral margins without excavation or carina; proximal margin with strong small emarginations; longitudinal midline concave at apex; glabrous.</p> <p>Female. Unkown.</p> <p>Variation. Length 23.10–28 mm. Humeral width 10.50–13.50 mm. Elytral width 13.15– 17.00 mm.</p> <p>Etymology. The specific epithet in latin is ‘aenescens’, which means bronze-coloured.</p> <p>Distribution. Byrsopolis aenescens can be found only in French Guiana in the cities of Saint-Laurent-du-Maroni, Cacao, Saül, Saint-Georges.</p> <p>Remarks. Byrsopolis aenescens differ from B. ferrandi in the following: apex of prementum with moderate emargination (strong emargination); labrum weakly emarginated (moderately emarginated); protibial spur 3.5x longer than wide (2.80x longer); mesotibia with one carina (two carinae); elytral apex with divergent angles (right angles).</p> </div>	https://treatment.plazi.org/id/03F09661FFDB804CFF28E5C77C3C53A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
03F09661FFD88047FEFCE0EC7CA85378.text	03F09661FFD88047FEFCE0EC7CA85378.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Byrsopolis ferrandi Soula 2010	<div><p>Byrsopolis ferrandi Soula, 2010a (Figures 16 (i–j), 18(a–h))</p> <p>Byrsopolis ferrandi Soula 2010a: 45 (original description); Carvalho and Grossi 2018: 372 (checklist, distribution); Hielkema and Hielkema 2019: 127 (checklist, distribution); Vaz-deMello and Grossi 2022 (online catalogue).</p> <p>Type locality. Bélizon, French Guiana.</p> <p>Type material. Holotype male, dissected. (a) ‘ Holotype / Byrsopolis / ferrandi S./2010 Soula’ [red label, handscript by Soula], (b) ‘Mussé des Confluences/MHNL/47030014’, (c) ‘Piste de/Belizon/I.94 G.F./col. SOULA’ [handwritten by Soula], (1♂ CCECL, specimen examined through images) (Figures 16 (i–m).</p> <p>Additional material. [4 specimens] ‘ FRENCH GUIANA: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-53.084164&amp;materialsCitation.latitude=3.0219445" title="Search Plazi for locations around (long -53.084164/lat 3.0219445)">Mont Itoupé</a> /(sommet 830 m), Maripa-Soula,/sommet (830 m), N 03°01 ʹ 19 ’, W 53°05 ʹ 03” // ‘L. T: light trap,/ 13.i.2016 ’ // ‘ Byrsopolis ferrandi /Soula, 2010/det. M. Seidel 2021’ // ‘ WORLD/SCARAB./DATABASE/ WSD00347520’ (1♂ MSPC); same, except ‘L. T: light trap,/ 16.i.2016 ’ // ‘WORLD/SCARAB./ DATABASE/WSD00347521’ (1♂ MSPC); same, except ‘WORLD/SCARAB./ DATABASE/ WSD003475213’ (1♂ MSPC); same, except ‘ 16.i.2016, PVP:/automatic light trap/(pink)’ // ‘WORLD/SCARAB./ DATABASE/WSD00347522’ (1♂ MSPC).</p> <p>Diagnosis. Body light yellow to reddish-brown, without metallic green reflections; clypeus subrectangular with weakly rounded apex; antennal club about 1.70x longer than antennomeres II–VII combined; scutellar shield subtriangular, base broadly rounded; elytral apex with right angles; elytra almost smooth, humeri weakly punctate; protibial spur about 2x longer than wide; parameres symmetrical, moderately divergent; parameres with weak sinuosities, proximal margin without strong emarginations.</p> <p>Description of holotype. Body. Shape elongate oval; light yellow on elytra to reddishbrown (Figures 16 (i), 18(a-c)); without green metallic reflections; frontoclypeal suture reddish-brown to black; yellow to orange setae on scutellar shield. Lectotype. Length 27.00–28.00mm. Humeral width 17.00–22.00. Elytral width 14.00–17.00 mm. Head. Clypeus about twice wider than long; subrectangular with weakly rounded apex; weakly concave only at marginal areas; punctures small to moderately large, densely punctate; frontoclypeal suture with curvilinear extremities weakly rounded, with obtuse angles; frons about 1.41x longer than clypeus; punctures small to moderately large, sparsely punctate at base, densely punctate at apex; glabrous; mandibles with apex weakly rounded external border, without setae, scissorial region weakly rounded, with one small tooth at middle (Figure 18 (e)); galea with one moderate apical tooth, three small setae, a single seta at middle, two setae at anterior part (Figure 18 (f)); apex prementum strongly emarginated; last labial palpomere about 2.18x longer than palpomere II (Figure 18 (g)); labrum moderately emarginate, moderately setose (Figure 18 (h)); antennomere III subcylindrical, IV–V subquadrate, antennomeres VI–VII subconical; antennal club about 1.70x longer than antennomeres II–VII combined. Thorax. Pronotum transverse, about 1.73x wider than long (Figure 18 (a)); punctures small to moderate, moderately punctate at disc, densely punctate at marginal areas; a small, anterior concavity on each side near marginal areas; scutellar plate subtriangular, base moderately rounded (Figure 18 (a)); small setae on apex, sparsely distributed; punctures small to moderate, sparsely punctate at marginal areas, moderately punctate at disc surface. Elytra. About 1.12x longer than wide; punctures small to moderate, moderately punctate at disc, densely punctate at marginal areas; interstriae indistinct (Figure 18 (a)); elytral apex with right angles; humeri with microsetae, moderately distributed. Legs. Protibial spur about 2.80x longer than wide; mesotibiae and metatibiae with one carina. Aedeagus. Parameres with apex rounded, weakly sinuous, moderately divergent, symmetrical; acute angle between parameres; parameres wide and flat (Figures 16 (j–l), 18(d)); lateral margins excavated at base forming a small carina; basal margin with weak sinuosities, basal margin without strong emarginations; longitudinal midline concave at apex; glabrous.</p> <p>Female. Unkown.</p> <p>Variation. Humeral width 12.50–13.00 mm. Elytral width 16.00–17.00 mm.</p> <p>Etymology. The specific epithet is a atribute to Michel Ferrand, given by Soula.</p> <p>Distribution. Byrsopolis ferrandi was described from Bélizon (Cayenne, French Guiana).</p> <p>Remarks. Byrsopolis ferrandi and B. aenescens can be found in French Guiana, but there are localities within cities bordering Brazil – for example, Oiapoque.</p> <p>Key to the genera of Areodina (modified from Grossi and Vaz-de-Mello (2015) after Jameson (1990))</p> <p>10(4). Frontoclypeal suture almost straight or sinuous; clypeus broadest at base, apex rounded, trapezoidal, rectangular, semioval or squared; surface weakly to moderately concave; hind wings with anterior margin near RA1+2, RA3 and apical hinge glabrous................................................................................................ Byrsopolis Burmeister 10’. Frontoclypeal suture rounded or arched; clypeus broadest at middle, apex quadrate; surface moderately to strongly concave; hind wings with anterior margin near RA1+2, RA3, and apical hinge setose................ Moronius Grossi and Vaz-de-Mello</p> <p>Key of the South American Areodina</p> <p>1 Mesoventral process moderate to long (exceeding the apex of mesocoxae); parameres strongly asymmetric.............................................................................................................. 2</p> <p>- Mesoventral process short (not exceeding the apex of mesocoxae); parameres symmetric or weakly to moderately asymmetric............................................................................ 3</p> <p>2 Apex of clypeus weakly rounded (males); parameres with hook-like projection...................................................................................................................................... Areoda MacLeay, 1819</p> <p>- Apex of clypeus trilobed (males); parameres without hook-like projection.................................................................................................................................... Oplognathus MacLeay, 1819</p> <p>3 Clypeus weakly to moderately concave; parameres weakly asymmetrical....................................................................................................................................... Byrsopolis Burmeister 1844</p> <p>- Clypeus strongly concave; parameres symmetrical........................................................................................................................................................ Moronius Grossi and Vaz-de-Mello 2015.</p> <p>Identification key for adults of the Byrsopolis species Burmeister</p> <p>1. Species from Cerrado and Atlantic Forest (castanea, crassa and quadraticeps groups)...................................................................................................................................................... 2</p> <p>- Species from Amazon (unknown females) [aenescens group]......................................... 12</p> <p>2. Elytral surface rugostriate, with weak to strong striae, deep punctures (males and females) [castanea group]................................................................................................................. 3</p> <p>- Elytral surface punctatostriate to shallowly punctate (unknown females) [crassa and quadraticeps groups]............................................................................................................................ 8</p> <p>3. Clypeus subrectangular with apex emargination; general colour usually dark brown......................................................................................................................................................... 4</p> <p>- Clypeus subrectangular, subtrapezoidal, parabolic, without apex emargination; general colour usually light brown or metallic green.................................................................. 5</p> <p>4. Scutellar plate with sparse to moderate setation (males and females); parameres weakly to moderately divergent; parameres with basal margin rounded.................................................................................................................... Byrsopolis castanea Burmeister 1844</p> <p>- Scutellar plate with moderate to dense setation (males and females); parameres strongly divergent; parameres with basal margin weakly bisinuate.................................................................. Byrsopolis burmeisteri Medeiros, Seidel and Grossi, 2020 sp. nov.</p> <p>5. Elytral suture with strong striae and coalescent punctures; antennal club less than 2x longer than antennomeres II–VII combined............................................................................. 6</p> <p>- Elytral suture with indistinct striae and coalescent punctures; antennal club about 2x longer than antennomeres II–VII combined.. Byrsopolis laticollis Burmeister 1855</p> <p>6. Frontoclypeal suture bisinuate; pronotum and scutellar shield weakly punctate (males known)..................................................................................................................................................... 7</p> <p>- Frontoclypeal suture almost straight; pronotum and scutellar shield strongly punctate (males unknown).............................................................. Byrsopolis cribricollis Ohaus 1912</p> <p>7. Clypeus about 1.75x wider than long; labrum weakly emarginate; elytral apex with divergent angles; wide parameres (females unknown)..................................................................................................... Byrsopolis ohausi Medeiros, Seidel and Grossi, 2020 sp. nov.</p> <p>- Clypeus about 2.2x wider than long; labrum moderately emarginat;; elytral apex truncate; narrow parameres (females unknown)........................................................................................................ Byrsopolis blanchardi Medeiros, Seidel and Grossi, 2020 sp. nov.</p> <p>8. Clypeus subrectangular, weakly bilobed or trilobed, hirsute; parameres separated by a U-shaped gap, symmetrical, parallel to slightly convergent [quadraticeps group] 9</p> <p>- Clypeus semioval, rounded, sparsely setose or glabrous; parameres separated by a Vshaped gap, dorsally symmetrical to slightly asymmetrical, parameres moderately divergent [crassa group].................................................................................................................. 11</p> <p>9. Metallic green on most parts of the body; clypeus weakly trilobed; antennal club about 3x longer than antennomeres II–VII combined (females unknown)........................................................... Byrsopolis schmidti Medeiros, Seidel and Grossi, 2020 sp. nov.</p> <p>- Yellowish to reddish brown on almost the whole body; clypeus weakly to moderately bilobed; antennal club about 2.60x longer than antennomeres II–VII combined.. 10</p> <p>10. Frontoclypeal suture almost straight between curvilinear extremities; pronotum glabrous; parameres with lateral margins moderately excavated longitudinally, forming a moderate carina at base (females unknown)......................................................................................................................................................... Byrsopolis quadraticeps Blanchard 1851</p> <p>- Frontoclypeal suture with a slight V-shaped concavity between curvilinear extremities; pronotum with short setae, sparsely to moderately distributed; parameres with lateral margins weakly excavated longitudinally, forming a short carina at the base of parameres. (females unknown)...................................................................................................................................... Byrsopolis angeloottatii Medeiros, Seidel and Grossi, 2020 sp. nov.</p> <p>11. Mandibles glabrous or hirsute in the angle formed by external border and scissorial region, wide longitudinally; labrum moderately emarginated; parameres with transverse striae in middle region. (females unknown) Byrsopolis crassa Blanchard 1851</p> <p>Mandibles glabrous, without projections; narrow longitudinally; labrum strongly emarginated; parameres without transverse striae in middle region (females unknown)........ Byrsopolis vazdemelloi Medeiros, Seideland Grossi, 2020 sp. nov.</p> <p>12. Apex prementum with triangular lobes; labrum moderately emarginate; scutellar shield with apex weakly rounded; elytral apexes with divergent angles............................................................................................................................ Byrsopolis aenescens Ohaus 1926</p> <p>- Apex of prementum with rounded lobes; labrum weakly emarginate; scutellar shield with apex broadly rounded; elytral apexes with right angles........................................................................................................................................................ Byrsopolis ferrandi Soula 2010a</p></div> 	https://treatment.plazi.org/id/03F09661FFD88047FEFCE0EC7CA85378	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Medeiros, Rone A. F.;Seidel, Matthias;Grossi, Paschoal C.	Medeiros, Rone A. F., Seidel, Matthias, Grossi, Paschoal C. (2022): Revision of the genus Byrsopolis Burmeister, 1844 (Coleoptera: Melolonthidae: Rutelinae: Rutelini), with the description of six new species endemic to Brazil and Paraguay. Journal of Natural History (J. Nat. Hist.) 56 (29 - 32): 1315-1364, DOI: 10.1080/00222933.2022.2115950, URL: http://dx.doi.org/10.1080/00222933.2022.2115950
