identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
68B743E1F92A527E99AB9271A458A8EE.text	68B743E1F92A527E99AB9271A458A8EE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bryocamptus (Bryocamptus) abramovae Novikov & Sharafutdinova & Chertoprud 2023	<div><p>Bryocamptus (Bryocamptus) abramovae sp. nov.</p><p>Figs 10, 11, 12, 13, 14, 15, 16, 17, 18</p><p>Bryocamptus sp. 2 - Novikov et al. 2021: 271.</p><p>Bryocamptus sp. 1 - Novikov and Sharafutdinova 2022: 34.</p><p>Material.</p><p>Holotype: Russia • ♀ dissected on two slides; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.489426&amp;materialsCitation.latitude=72.373" title="Search Plazi for locations around (long 126.489426/lat 72.373)">Lena 
River Delta</a>, Samoylov Island, Ruiba Lake; 72.373003°N, 126.489429°E; depth 1-1.5 m; 23 Aug. 2019; A. Novikov leg; BP 547/1-a, BP 547/1-b  .  Allotype: Russia • ♂ dissected on one slide; collection data as for holotype; BP 547/2 .  Paratypes: 5 ♀ and 3 ♂ undissected, preserved in 4% formalin; collection data as for holotype; BP 547/4 .</p><p>Additional material.</p><p>Russia • 9 ♀♀ and 6 ♂♂ undissected; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=125.28114&amp;materialsCitation.latitude=72.51792" title="Search Plazi for locations around (long 125.28114/lat 72.51792)">Lena River Delta</a>, Jangylakh Sise Island, large nameless lake; 72.517921°N, 125.281147°E; 7 Aug. 2019; A. Novikov leg; retained in the collection of the first author  .</p><p>Russia • 2 ♀♀ undissected; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.93597&amp;materialsCitation.latitude=72.55094" title="Search Plazi for locations around (long 126.93597/lat 72.55094)">Lena River Delta</a>, Baron Island, small thermokarst lake; 72.550939°N, 126.93597°E; 8 Aug. 2019; A. Novikov leg; retained in the collection of the first author  .</p><p>Russia • 3 ♀♀ and 1 ♂ undissected; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=126.265656&amp;materialsCitation.latitude=72.46886" title="Search Plazi for locations around (long 126.265656/lat 72.46886)">Lena 
River Delta</a>, Kurungnah Sise Island, Krugloe Lake; 72.468859°N, 126.265658°E; 21 Aug. 2019; A. Novikov leg; retained in the collection of the first author</p><p>Russia • 4 ♀♀ and 2 ♂♂ undissected; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=179.56738&amp;materialsCitation.latitude=70.954445" title="Search Plazi for locations around (long 179.56738/lat 70.954445)">Vrangel Island</a>, large nameless lake; 70.954443°N, 179.567387°E; 26 Aug. 2021; A. Novichkova leg: retained in the collection of the first author  .</p><p>Description.</p><p>Female (based on holotype and paratypes). Body subcylindrical (Fig. 10A). Total body length from anterior margin of rostrum to posterior margin of caudal rami: 586  µm (n = 1). Cephalothorax (Fig. 10B, C; Appendix 1), wider as remaining somites, length 152  µm, largest width 113  µm . Naupliar eye red. Rostrum (Fig. 10D) small, fused with cephalothorax, with rounded end, with one pair of sensillae and pore located proximal to sensillae. Posterior margin of cephalothorax and all pedigerous somites smooth.</p><p>Cephalothorax (Fig. 10B, C; Appendix 1) with dumbbell-shaped dorsal window, seven pairs of pores, seven pairs of sensillae of central group (group C), eight pairs of sensillae of marginal group (group P) and 13 pairs of ungrouped sensillae (in Table 4 and in Appendix 1 marked as L). Second pedigerous somite with lateral windows, dorsal unpaired pore, lateral pair of pores and six pairs of sensillae. Third pedigerous somite with dorsal unpaired pore and six pairs of sensillae. Fourth pedigerous somite with dorsal unpaired pore and five pairs of sensillae. Fifth pedigerous somite with three pairs of sensillae.</p><p>Abdomen (Fig. 11A-C) consisting of genital-double somite, two free abdominal somites and anal somite with caudal rami. All somites except anal somite slightly wavy posterior margin, on surface with spinular rows. Genital-double somite consists of last thoracic somite and first abdominal somite; wider than long; anterior part with two pairs of sensillae, dorsal unpaired pore, ventro-lateral row of spinules; posterior part with three pairs of sensillae, pairs of ventral and lateral pores and lateral rows of spinules.</p><p>P6 (Fig. 11C) fused with somite with one pinnate and one naked setae. Genital field (Fig. 11C) short, laterally with eight-pore sieves; copulatory pore located medially, copulatory duct chitinised with two additional tubes, extending proximally to pair of labyrinthic rounded ducts and one chitinised unpaired duct.</p><p>Second and third abdominal somites as in  B. minutus . Anal somite with one pair of sensillae, ventral pair of large pores, lateral pair of pores and lateral spinules. Anal operculum semilunar, with seven short bifid spinules.</p><p>Caudal rami (Fig. 11A-D). Length/width ratio 1.6, with three ventral pores; with rows of spinules on ventral and dorsal side at base of seta VI and rows spinules at base of setae II and III. Seta I small, located near seta II. Apical seta IV (Fig. 11D) bipinnate, without "helle Stelle". Apical seta V long, bipinnate, with "helle Stelle". Seta VI with wide base (Fig. 11C). Seta VII triarticulated (Fig. 11B).</p><p>Antennule (Fig. 12A) similar to that of  Bryocamptus minutus . Differences expressed in more elongated segments, especially 3th and 4th segments; one of setae on segment 2 pinnate. Armature formula: 1-[1],2-[9],3-[5],4-[1+(1+ae)],5-[1],6-[3],7-[2],8-[5+acr].</p><p>Antenna (Fig. 12B) similar to that of  Bryocamptus minutus . Allobasis and free endopodal segment slightly more elongated. Inner spinular row on coxa with extremely long spinules. Allobasis with proximal outer spinular row, basal seta pinnate.</p><p>Labrum (Fig. 12C) similar to that of  Bryocamptus minutus, but without semicircular spinular row on inner side.</p><p>Mandible (Fig. 13A, B) similar to that of  Bryocamptus minutus . The palp is shortened.</p><p>Paragnaths (Fig. 12D) similar to that of  Bryocamptus minutus, with only three lateral groups of spinules and with a more well-defined pocket.</p><p>Maxillule (Fig. 13C) similar to that of  Bryocamptus minutus . Basis with two groups of spinules.</p><p>Maxilla (Fig. 13D) as in  Bryocamptus minutus, only with slight differences in length and armature of setae.</p><p>Maxilliped (Fig. 14A) similar to that of  Bryocamptus minutus . Differences are only in shorter syncoxa and basis.</p><p>Cuticular process between maxillipeds and P1 (Fig. 14B, C) in height approximately same as in length, with long spinules, seven spinules on each side. Spinules on posterior margin.</p><p>P1 (Fig. 14D; Table 2) similar to that of  Bryocamptus minutus . Basis without inner spinules. First exopodal segment with row of small spinules on anterior side. First endopodal segment reaching end of second exopodal segment. First and second endopodal segments with smooth inner side. Differences also noticeable in shorter exopodal and endopodal segments and larger spinules on coxa and basis.</p><p>P2 (Fig. 14E; Table 2). Praecoxa with row of spinules. Coxa with one lateral row of large spinules, two anterior rows of large spinules and four anterior rows of small spinules. Intercoxal sclerite naked. Basis with proximal pore, inner group of long spinules, rows of spinules at base of endopod and exopod; with outer spine. All endopodal and exopodal segments with outer spinules. Exopod three-segmented; first exopodal segment with outer spinulose spine, apically with frill; second segment with outer spinulose spine, inner pectinate seta, inner slender spinules and apical frill; third segment with three outer spinulose spines, two apical setae and one inner pectinate seta. Endopod three-segmented; first and second segments with inner seta; third segment with outer spinulose spine, two apical pinnate setae and one inner pectinate seta.</p><p>P3 (Fig. 15A; Table 2). Praecoxa with spinular row. Coxa with one lateral row of large spinules, two anterior rows of large spinules and four anterior rows of small spinules. Intercoxal sclerite without spinules. Basis with outer seta, proximal pore, inner group of long spinules and rows of spinules at base of endopod and exopod. Exopod three-segmented; first exopodal segment with outer spinulose spine, outer spinules, apically with frill; second segment with outer spinulose spine, outer spinules, inner pectinate seta, inner slender spinules and apical frill; third segment with three outer spinulose spines, two apical setae and two inner pectinate setae. Endopod three-segmented; first and second segments with inner seta, second segment with outer spinules; third segment with outer spinules, outer spinulose spine, two apical pinnate setae and two inner pectinate setae.</p><p>P4 (Fig. 15B; Table 2). Praecoxa with spinular row. Coxa with one lateral row of large spinules, two anterior rows of large spinules and four anterior rows of small spinules. Basis with outer seta, proximal pore, rows of spinules at base of endopod and exopod. Exopod three-segmented; first exopodal segment with outer spinulose spine, outer spinules, apically with frill; second segment with outer spinulose spine, outer spinules, inner pectinate seta, inner slender spinules and apical frill; third segment with two outer spinulose spines, two apical setae and two inner pectinate setae. Endopod two-segmented; first segment with inner pectinate seta, second segment with outer spinules, outer spinulose spine, apical spiniform spinulose seta, apical pinnate seta and two inner pectinate setae.</p><p>P5 (Fig. 15C) with separate right and left baseoendopods. Baseoendopod reaching ~ 2/3 of exopodal segment; with four pores, spinular row at base of outer seta; outer seta of basis pinnate, long. Endopodal lobe with four long bipinnate setae and one short bipinnate seta V; with small process that may be pore between setae III and IV. Exopod inner thin pinnate seta, long apical pinnate seta, naked subapical seta and two pinnate outer setae.</p><p>Male. Sexual dimorphism expressed in the antennule, P2-P6, genital segmentation and ornamentation, shape of caudal rami. Cephalothorax and thoracic somites as in female. P6 (Fig. 16B) two asymmetric flaps fused to the somite, with three naked setae. Differences from female in abdomen structure as follows (Fig. 16A, B): first abdominal somite free; first to third abdominal somites with spinular row encircling somite ventrally and laterally; anal somite with ventral spinule and without lateral spinules; caudal rami without ventral spinules; seta IV with "helle Stelle".</p><p>Antennule (Fig. 17A, B) 10-segmented, haplocer with geniculation between segments 7 and 8. Segments 1, 3, 4, 5, 6, 9, and 10 almost like in  B. minutus, but more elongated. Segment 2 with small pore on anterior side. Segment 7 with articular plate, with one filiform seta, one small caudate seta and with two modified laminar setae. Segment 8 with proximal short dentate plate and two modified laminar setae. Armature formula: 1-[1],2-[9],3-[8],4-[2],5-[6+(1+ae)],6-[2],7-[2+2 modified],8-[2 modified],9-[1],10-[7+acr].</p><p>P2 (Fig. 18A) as in female, except endopod. Endopod two-segmented. First segment with inner seta. Second segment with notch on distal outer margin, outer spinules, two apical pinnate slender setae and two inner pectinate setae.</p><p>P3 (Fig. 18B-D): praecoxa, coxa, intercoxal sclerite as in female. Basis as in female, but with larger inner process. Exopod as in female, but third segment with pore. Endopod three-segmented. First endopodal segment with strong seta. Second endopodal segment with posterior thin seta and long apophysis with double tip. Third segment with two small inner setae and two apical pinnate setae.</p><p>P4 (Fig. 18E): praecoxa, coxa, intercoxal sclerite, basis, exopod as in female. Endopod two-segmented; first segment short, unarmed; second segment with outer spinules, spinulose spine, outer apical spiniform spinulose seta and inner apical bipinnate seta.</p><p>P5 (Fig. 17C) right and left fused medially. Baseoendopod with three pairs of pores, outer spinule and outer long pinnate seta; endopodal lobe with two strong spinulose apical spines. Exopod with spinule on anterior surface, two equal length outer setae, naked outer subapical seta, long apical spinulose seta, one inner spinulose seta and one long inner pectinate seta with long setules.</p><p>Variability.</p><p>Individuals with two outer spines on the third exopodal segments of P2-P4 were found.</p><p>Etymology.</p><p>This species is named after Ekaterina Abramova, teacher and mentor of the first author.</p><p>Remarks.</p><p>The species is well distinguished from other species of the  B. minutus group by the presence of only five setae on the endopodal lobe of females P5 and by simple caudal rami with unmodified setae.</p></div>	https://treatment.plazi.org/id/68B743E1F92A527E99AB9271A458A8EE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Novikov, Aleksandr;Sharafutdinova, Dayana;Chertoprud, Elena	Novikov, Aleksandr, Sharafutdinova, Dayana, Chertoprud, Elena (2023): Two new species of Bryocamptus (Copepoda, Harpacticoida, Canthocamptidae) from the Russian Arctic and comparison with Bryocamptus minutus (Claus, 1863). ZooKeys 1138: 89-141, DOI: http://dx.doi.org/10.3897/zookeys.1138.90580, URL: http://dx.doi.org/10.3897/zookeys.1138.90580
721DB59E6F34503BB17020E96BFFC3FE.text	721DB59E6F34503BB17020E96BFFC3FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bryocamptus (Bryocamptus) minutus (Claus 1863)	<div><p>Bryocamptus (Bryocamptus) minutus (Claus, 1863)</p><p>Subspecies.</p><p>B. (B.) minutus minutus (Claus, 1863),  B. (B.) minutus schizodon ( Mrázek, 1893).</p><p>Nomen dubium.</p><p>B. (B.) minnesotensis (Herrick, 1884).</p><p>Remarks.</p><p>Bryocamptus (B.) minutus is a taxonomically rather complex species due to a rather long history of study and wide distribution. According to Article 45.6 of the International Code of Zoological Nomenclature, a number of forms of this species must be treated as separate subspecies (ICZN 1999). However, in the case of  B. (B.) minutus vejdovskyiformis Thallwitz, 1916, this is probably a form that does not have subspecies status and is either an aberrant specimen(s) or simply variability (Thallwitz 1916). Simple dentiform and bifid spinules are also found in other related species, both within the same population and in one individual. This has been described in  B. hutchinsoni Kiefer, 1929 (Wilson 1956),  B. vejdovskyi ( Mrázek, 1893) (Reed 1990) and also in  B. putoranus sp. nov. (in this article).</p><p>A number of authors noted variability in the number of outer spines on the third exopodal segment of P4, which was the reason for  Lang’s description of the forms:  B. minutus f. typica Lang, 1957 and  B. minutus f. bispinosa Lang, 1957 (Lang 1957). We suggest that these forms do not have a taxonomic rank, since such variability is common for this group of species.</p><p>Another form of  B. minutus f. simplicidentata (Willey, 1934) has been synonymized with  B. hutchinsoni based on structure of caudal rami (Wilson 1956) but although figuring mistakenly and without literature support as valid in WORMS database (Walter and Boxshall 2021).</p><p>A rather interesting finding is described from the Iberian Peninsula as  B. minutus (Caramujo and Boavida 2009). Based on the depicted limbs, it can be assumed that this is either  B. minutus schizodon or a separate species. It differs from  B. minutus minutus in the two-segmented endopod P2, short bifid spinules on the anal operculum, and slight displacement of the caudal setae to the ventral side of caudal rami. In general, these characters are already enough to distinguish a separate species.</p></div>	https://treatment.plazi.org/id/721DB59E6F34503BB17020E96BFFC3FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Novikov, Aleksandr;Sharafutdinova, Dayana;Chertoprud, Elena	Novikov, Aleksandr, Sharafutdinova, Dayana, Chertoprud, Elena (2023): Two new species of Bryocamptus (Copepoda, Harpacticoida, Canthocamptidae) from the Russian Arctic and comparison with Bryocamptus minutus (Claus, 1863). ZooKeys 1138: 89-141, DOI: http://dx.doi.org/10.3897/zookeys.1138.90580, URL: http://dx.doi.org/10.3897/zookeys.1138.90580
B4DE7B68416554BBB1A6BD3DC95C1D5B.text	B4DE7B68416554BBB1A6BD3DC95C1D5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bryocamptus (Bryocamptus) minutus subsp. minutus (Claus 1863) minutus (Claus 1863	<div><p>Bryocamptus (Bryocamptus) minutus minutus (Claus, 1863)</p><p>Figs 1, 2, 3, 4, 5, 6, 7, 8, 9</p><p>B. (B.) minutus vejdovskyiformis Thallwitz, 1916: 238. syn. nov.</p><p>Material examined.</p><p>Estonia • 2 ♀♀ dissected on three slides (BP 546/1-a, BP 546/1-b, BP 546/2); 1 ♂ on one slide (BP 546/3);   9 ♀♀ and 5 ♂♂ undissected preserved in 4% formalin (retained in the collection of the first author); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.089441&amp;materialsCitation.latitude=58.18089" title="Search Plazi for locations around (long 26.089441/lat 58.18089)">Vortsjaerv
Lake</a>; 58.180888°N, 26.089441°E; 25 Sep. 2007; E. Fefilova leg; BP 546  .</p><p>Supplementary description.</p><p>Female. Body subcylindrical. Total body length from anterior margin of rostrum to posterior margin of caudal rami: 484  µm (n = 1). Cephalothorax (Fig. 1A, B; Appendix 1) wider than remaining somites, length 151  µm, largest width 124  µm . Naupliar eye not observed. Rostrum (Fig. 1C) small, fused with cephalothorax, with squared end, with one pair of sensillae. Posterior margin of cephalothorax and all pedigerous somites smooth.</p><p>Cephalothorax (Fig. 1A, B; Appendix 1) with dumbbell-shaped dorsal window, 10 pairs of pores, seven pairs of sensillae of central group (group C), 13 pairs of sensillae of marginal group (group P) and 20 pairs of ungrouped sensillae (in Table 4 and in Appendix 1 marked as L). Second pedigerous somite with lateral windows, dorsal unpaired pore, lateral pair of pores and eight pairs of sensillae. Third pedigerous somite with dorsal unpaired pore, lateral pair of pores and eight pairs of sensillae. Fourth pedigerous somite with dorsal unpaired pore, lateral pair of pores and eight pairs of sensillae. Fifth pedigerous somite with lateral pair of pores and four pairs of sensillae.</p><p>Abdomen (Fig. 2A-C) consisting of genital-double somite, two free abdominal somites and anal somite with caudal rami. All somites except anal somite on posterior margin serrated, on surface with spinular rows. Genital-double somite consists of last thoracic somite and first abdominal somite; longer than wide; anterior part with four pairs of sensillae, dorsal unpaired pore, lateral paired pores, ventro-lateral and lateral rows of spinules; posterior part with four pairs of sensillae, pairs of ventral and lateral pores and lateral rows of spinules.</p><p>P6 (Fig. 2C) fused with somite with one pinnate and one naked setae. Genital field (Fig. 2C) long, laterally with eight-pore sieves; copulatory pore displaced to posterior part of somite, copulatory duct chitinised with two additional tubes, extending proximally to pair of labyrinthic rounded ducts and one chitinised unpaired duct.</p><p>Second abdominal somite with three pairs of sensillae, pair of lateral pores; on posterior margin with lateral row of large spinules. Third abdominal somite with pair of lateral pores, on posterior margin with lateral row of large spinules and ventral row of small spinules. Anal somite with one pair of sensillae, ventral pair of large pores, lateral pair of pores, dorsal dots near base of caudal rami and lateral spinules. Anal operculum semilunar, with eight long bifid spinules.</p><p>Caudal rami (Fig. 2A-E). Length/width ratio 1.6, with three ventral pores; with rows of spinules on ventral side at base of seta IV and rows spinules at base of setae II and III. Seta I small, located near seta II. Setae IV, V and VI displaced to ventral side of caudal ramus. Apical seta IV (Fig. 2D) unipinnate, with "helle Stelle" and massive dorsal bulb located distally "helle Stelle". Apical seta V long, bipinnate, with "helle Stelle". Seta VII triarticulated (Fig. 2B).</p><p>Antennule (Fig. 1D) 8-segmented. Segment 1 short, with one pinnate seta and two rows of spinules. Other segments with bare setae. Segment 4 with fused basally seta and aesthetasc. Distal segment with acrothek consisting of aesthetasc and two setae fused basally. Armature formula: 1-[1],2-[9],3-[5],4-[1+(1+ae)],5-[1],6-[3],7-[2],8-[5+acr].</p><p>Antenna (Fig. 3A) with allobasis. Coxa with two rows of spinules. Allobasis with two naked setae and one spinular row at base of endopodal seta. Free endopodal segment with two lateral rows of big spinules, with two spinulose spines and slender seta; distally with two rows of spinules; apically with three geniculate setae, two long spines and one small accessory seta; outermost geniculate seta fused basally to small seta. Exopod two-segmented; first segment with one pinnate seta and row of spinules; second segment with three pinnate setae.</p><p>Labrum (Fig. 4A). On outer side with row of thin setules and large proximal pore. Distal margin with lateral rows of robust spinules, rows of fused spinules into comb and three rows of small spinules. On inner side medially with four unpaired pores, three pared pores, with lateral spinular row, semicircular spinular row and groups of thin setules.</p><p>Mandible (Fig. 4B, C). Coxa with spinules proximally. Gnathobase with pars incisiva, lacinia mobilis, complex dental battery and spinulose seta; pars incisiva two-pointed; lacinia mobilis three-pointed. Dental battery (Fig. 4C) consisting of five fused blocks of small short teeth, inner of which fused at base with seta. Pars molaris sharply edged. Palp one-segmented, with medial spinular row and four apical setae.</p><p>Paragnaths (Fig. 4D) with paired lateral lobes and unpaired posterior rounded lobe. Lateral lobes wrapped in distal part forming  “pocket”; proximally with lateral pore (probably); on outer side with four groups of long spinules; on inner side with three-four rows of spinules; on anterior side with three medial rows of strong spinules and proximal row of spinules.</p><p>Maxillule (Fig. 3B). Praecoxa with two rows of slender spinules on outer edge and one row of spinules on posterior side. Praecoxal arthrite medially with two rows of spinules and one proximal pore; distally with one simple strong spine, three strong spines with pectinate end, three biarticulate spines, one proximal bipinnate seta and one thin seta with long spinules. Coxa with row of spinules, coxal endite with one weakly pinnate and one spinulose geniculate setae. Basis with two subdistal setae and three distal setae, one of which spinulose and geniculate. Endopod and exopod incorporated into basis, each represented by two naked setae.</p><p>Maxilla (Fig. 3C). Basis with several rows of spinules on outer and inner edge as figured, with two endites. Proximal endite with spinular row, one spinulose spine and two pinnate setae, distal endite with one strong pinnate seta and two thin pinnate setae. Proximal endopodal segment with two setae, outer tube pore and massive distal claw. Distal endopodal segment with three naked setae, one of which proximal and small.</p><p>Maxilliped (Fig. 5A) subchelate. Syncoxa elongated with several rows of spinules as figured, distally with one pinnate seta. Basis with two rows of large spinules on anterior and posterior sides and three outer rows of small spinules. Endopod on posterior side with one seta, on anterior side with small protuberance, probably tube pore. Endopodal claw elongated, with row of small spinules.</p><p>Cuticular process between maxillipeds and P1 (Fig. 4E, F) in height approximately same as in length, with long spinules, ten spinules on each side. Spinules encircle from anterior-lateral margin to posterior margin.</p><p>P1 (Fig. 5B; Table 1) with three-segmented rami. Praecoxa with outer spinular row. Coxa rectangular, with seven spinular rows, four of which consisting of little spinules. Intercoxal sclerite wide, with one paired spinular rows. Basis with proximal pore, medial row of small spinules, rows of spinules at base of endopod and exopod, row of spinules at base of inner seta, inner row of spinules; with inner and outer strong spines. All endopodal and exopodal segments with outer spinules. First exopodal segment with one outer spinulose spine; second segment with inner pectinate seta and outer spinulose spine; third exopodal segment with two outer spinulose spines and two apical slender geniculate setae. Endopod longer than exopod. First endopodal segment reaching middle of second exopodal segment, with inner pectinate seta and inner spinular row; second endopodal segments with one inner pectinate seta, third segment with outer spinulose spine, apical long geniculate seta and inner small seta.</p><p>P2 (Fig. 6A; Table 1). Praecoxa with row of spinules. Coxa with one lateral row of large spinules and five rows of spinules on anterior side. Intercoxal sclerite with two large spinules. Basis with proximal pore, rows of spinules at base of endopod and exopod; with outer spine. All endopodal and exopodal segments with outer spinules. Exopod three-segmented; first exopodal segment with outer naked spine, apically with frill; second segment with outer naked spine, inner pectinate seta, inner slender spinules and apical frill; third segment with pore, three outer spinulose spines, two apical setae and one inner pectinate seta. Endopod three-segmented; first and second segments with inner seta; third segment with outer spinulose spine, two apical pinnate setae and one inner pectinate seta.</p><p>P3 (Fig. 6B; Table 1). Praecoxa with spinular row. Coxa with one lateral row of large spinules and five rows of spinules on anterior side. Intercoxal sclerite without spinules. Basis with outer seta, proximal pore, rows of spinules at base of endopod and exopod. Exopod three-segmented; first exopodal segment with outer naked spine, outer spinules, apically with frill; second segment with outer naked spine, outer spinules, inner pectinate seta, inner slender spinules and apical frill; third segment with pore, three outer spinulose spines, two apical setae and two inner pectinate setae. Endopod three-segmented; first and second segments with inner seta, second segment with outer spinules; third segment with outer spinules, outer spinulose spine, two apical pinnate setae and two inner pectinate setae.</p><p>P4 (Fig. 6C; Table 1). Praecoxa with spinular row. Coxa with one lateral row of large spinules and five rows of spinules on anterior side. Intercoxal sclerite without spinules. Basis with outer seta, proximal pore, rows of spinules at base of endopod and exopod. Exopod three-segmented; first exopodal segment with outer naked spine, outer spinules, apically with frill; second segment with outer naked spine, outer spinules, inner pectinate seta, inner slender spinules and apical frill; third segment with pore, two outer spinulose spines, two apical setae and two inner pectinate setae. Endopod two-segmented; first segment with inner seta, second segment with outer spinules, outer spinulose spine, two apical pinnate setae and two inner pectinate setae.</p><p>P5 (Fig. 5C) with separate right and left baseoendopods. Baseoendopod reaching ~ 1/2 of exopodal segment; with four pores, spinular row at base of outer seta; outer seta of basis pinnate, long. Endopodal lobe with four long bipinnate setae and two short bipinnate setae V and VI; with small process that may be pore between setae III and IV. Exopod with inner short pinnate seta, long apical pinnate seta, naked subapical seta and two pinnate outer setae.</p><p>Male. Sexual dimorphism expressed in the antennule, P2-P6, genital segmentation and ornamentation, shape of caudal rami. Cephalothorax and thoracic somites as in female. P6 (Fig. 7B) two asymmetric flaps fused to the somite, with three naked setae. Differences from female in abdomen structure as follows (Fig. 7A, B): first abdominal somite free; first to third abdominal somites with spinular row encircling somite ventrally and laterally; anal somite with ventral spinules; caudal rami with normal setae IV and V; anal operculum with nine bifid and simple spinules.</p><p>Antennule (Fig. 8A, B) 10-segmented, haplocer with geniculation between segments 7 and 8. Segment 5 with large aestetasc fused at base with long seta, with one strong caudate seta. Segment 7 with articular plate, with one filiform seta, one small caudate seta and with two modified laminar setae. Segment 8 with proximal dentate plate and two strong modified laminar setae. Segment 10 with acrothek consisting of slender aestetasc and two setae. Armature formula: 1-[1],2-[9],3-[8],4-[2],5-[6+(1+ae)],6-[2],7-[2+2 modified],8-[2 modified],9-[1],10-[7+acr].</p><p>P2 (Fig. 9A) as in female, except endopod. Endopod two-segmented. First segment with outer spinules and inner seta. Second segment with notch on distal outer margin, outer spinules, two apical pinnate slender setae and two inner pectinate setae.</p><p>P3 (Figs 8C, D, 9B): praecoxa, coxa, intercoxal sclerite as in female. Basis as in female, but with inner process. Exopod as in female, but third segment with broad slit-like pore. Endopod three-segmented. First endopodal segment with strong seta. Second endopodal segment with posterior seta and long apophysis with double tip. Third segment with two small inner setae, inner pore and two apical pinnate setae.</p><p>P4 (Fig. 9C): praecoxa, coxa, intercoxal sclerite, basis, exopod as in female. Endopod two-segmented; first segment short unarmed; second segment with outer spinules, spinulose spine, outer apical spiniform spinulose seta, inner apical bipinnate seta and inner pectinate seta.</p><p>P5 (Fig. 7C) right and left fused medially. Baseoendopod with three pairs of pores, outer spinular row and outer long pinnate seta; endopodal lobe with two strong spinulose apical spines. Exopod with spinules on anterior surface, three naked outer setae, long apical spinulose seta, one inner spinulose seta and one long inner pectinate seta with long setules.</p><p>Variability.</p><p>We found variability in the structure of the caudal rami. Some females have an inner group of long spinules (Fig. 2E).</p></div>	https://treatment.plazi.org/id/B4DE7B68416554BBB1A6BD3DC95C1D5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Novikov, Aleksandr;Sharafutdinova, Dayana;Chertoprud, Elena	Novikov, Aleksandr, Sharafutdinova, Dayana, Chertoprud, Elena (2023): Two new species of Bryocamptus (Copepoda, Harpacticoida, Canthocamptidae) from the Russian Arctic and comparison with Bryocamptus minutus (Claus, 1863). ZooKeys 1138: 89-141, DOI: http://dx.doi.org/10.3897/zookeys.1138.90580, URL: http://dx.doi.org/10.3897/zookeys.1138.90580
ADC42B81DD2652E38A4AB967ABA4BA5D.text	ADC42B81DD2652E38A4AB967ABA4BA5D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bryocamptus (Bryocamptus) Chappuis 1929	<div><p>Subgenus Bryocamptus Bryocamptus Chappuis, 1929</p><p>Remarks.</p><p>Bryocamptus is a very large genus with ~ 135 species and subspecies in four subgenera:  B. (Arcticocamptus) Chappuis, 1929,  B. ( Bryocamptus) Chappuis, 1928,  B. (Echinocamptus) Chappuis, 1929 and  B. (Rheocamptus) Borutzky, 1952. Additionally, two subgenera were earlier designated as not valid  B. (Limocamptus) Chappuis, 1929 and  B. (Pentacamptus) Wiley, 1934.</p><p>In our opinion, this is one of the genera of the family most in need of revision. The first reason is that there are no clear diagnostic characters for the entire genus. Previously, this character was the two-segment exopod A2; however, this character is plesiomorphic for the entire family  Canthocamptidae, so it may be an adequate solution to separate at least part of the subgenera into separate genera. The second reason is the blurred line between  B. (Bryocamptus) and  B. (Rheocamptus) . Borutzky (1952) in the differences between these subgenera indicates the difference in segmentation of the endopods P1-P4, which again contrasts plesiomorphic and apomorphic characters. In our opinion, an essential part of the  B. (Rheocamptus) species should in fact be transferred to the type subgenus.</p><p>Unfortunately, at the moment we do not have enough data and material to revise the subgenera, so in this work we adhere to the classification given by Dussart and Defaye (1990).</p></div>	https://treatment.plazi.org/id/ADC42B81DD2652E38A4AB967ABA4BA5D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Novikov, Aleksandr;Sharafutdinova, Dayana;Chertoprud, Elena	Novikov, Aleksandr, Sharafutdinova, Dayana, Chertoprud, Elena (2023): Two new species of Bryocamptus (Copepoda, Harpacticoida, Canthocamptidae) from the Russian Arctic and comparison with Bryocamptus minutus (Claus, 1863). ZooKeys 1138: 89-141, DOI: http://dx.doi.org/10.3897/zookeys.1138.90580, URL: http://dx.doi.org/10.3897/zookeys.1138.90580
30E8A0B801335BF594D646705779EE8E.text	30E8A0B801335BF594D646705779EE8E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bryocamptus (Bryocamptus) putoranus Novikov & Sharafutdinova & Chertoprud 2023	<div><p>Bryocamptus (Bryocamptus) putoranus sp. nov.</p><p>Figs 19, 20, 21, 22, 23, 24, 25, 26, 27</p><p>Material.</p><p>Holotype: Russia • ♀ dissected on two slides; Russia, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=94.17053&amp;materialsCitation.latitude=68.901985" title="Search Plazi for locations around (long 94.17053/lat 68.901985)">Putorana Plateau</a>, large nameless lake in the upper flow of the Neral River; 68.901987°N, 94.170533°E; depth 0.5-1 m; 4 Aug. 2021; E. Chertoprud leg; BP 548/1-a, BP 548/1-b  .  Allotype: Russia • ♂ dissected on one slide; collection data as for holotype; BP 548/2 .  Paratypes: Russia • ♀ dissected on two slides (BP 548/3-a, BP 548/3-b) and ♂ dissected on one slide (BP 548/4); Putorana Plateau, large nameless lake; 68.898348°N, 94.174442°E; depth 0.5-1 m; 4 Aug. 2021; E. Chertoprud leg.</p><p>Description.</p><p>Female (based on holotype and paratype). Body subcylindrical (Fig. 19A). Total body length from anterior margin of rostrum to posterior margin of caudal rami: 527  µm (n = 1). Cephalothorax (Fig. 19B, C; Appendix 1), wider than remaining somites, length 144  µm, largest width 112  µm . Naupliar eye not observed. Rostrum (Fig. 21A) small, fused with cephalothorax, with rounded end, with one pair of sensillae and pore located distal to sensillae. Posterior margin of cephalothorax and all pedigerous somites smooth.</p><p>Cephalothorax (Fig. 19B, C; Appendix 1) with dumbbell-shaped dorsal window, seven pairs of pores, seven pairs of sensillae of central group (group C), 13 pairs of sensillae of marginal group (group P) and 21 pairs of ungrouped sensillae (marked as L in Table 4 and in Appendix 1). Second pedigerous somite with lateral windows, dorsal unpaired pore, lateral pair of pores and eight pairs of sensillae. Third pedigerous somite with dorsal unpaired pore, lateral pair of pores and nine pairs of sensillae. Fourth pedigerous somite with dorsal unpaired pore, lateral pair of pores and eight pairs of sensillae. Fifth pedigerous somite with lateral pair of pores and four pairs of sensillae.</p><p>Abdomen (Fig. 20A-C) consisting of genital-double somite, two free abdominal somites and anal somite with caudal rami. All somites except anal somite with wavy posterior margin, on surface with spinular rows. Genital-double somite consists of last thoracic somite and first abdominal somite; wider than long; anterior part with four pairs of sensillae, dorsal unpaired pore, lateral paired pores, ventro-lateral and lateral rows of spinules; posterior part with four pairs of sensillae, pairs of ventral and lateral pores and lateral rows of spinules.</p><p>P6 (Fig. 20C) fused with somite with two pinnate setae. Genital field (Fig. 20C) long, laterally with eight-pore sieves; copulatory pore displaced to posterior part of somite, copulatory duct chitinised with two additional tubes, extending proximally to pair of labyrinthic rounded ducts and one chitinised unpaired duct.</p><p>Second, third abdominal and anal somites as in  B. minutus . Anal operculum semilunar, with seven long simple spinules. Caudal rami (Fig. 20A-D). Length/width ratio 1.5, with three ventral pores; with rows of spinules on ventral and dorsal side at base of seta IV and rows spinules at base of setae II and III. Seta I small, located near seta II. Setae IV, V and VI displaced to ventral side of caudal ramus. Apical seta IV (Fig. 20D) bipinnate, with massive bulbous base and "helle Stelle". Apical seta V long, bipinnate, with "helle Stelle". Seta VII triarticulated (Fig. 20B).</p><p>Antennule (Fig. 20B) similar to that of  Bryocamptus minutus . Differences expressed in more elongated segments, especially 3rd and 4th segments; one of setae on segment 2 pinnate. Armature formula: 1-[1],2-[9],3-[5],4-[1+(1+ae)],5-[1],6-[3],7-[2],8-[5+acr].</p><p>Antenna (Fig. 21B) similar to that of  Bryocamptus minutus . Allobasis and free endopodal segment slightly shorter. Allobasis with proximal outer spinular row, basal seta pinnate; without spinular row at base of endopodal seta.</p><p>Labrum (Fig. 22A) similar to that of  Bryocamptus minutus, but without semicircular spinular row on inner side.</p><p>Mandible (Fig. 21D, E). Coxa and gnathobase as in  Bryocamptus minutus . The palp elongated, with three apical setae.</p><p>Paragnaths (Fig. 22B) similar to that of  Bryocamptus minutus, with only two groups of spinules on anterior side and without proximal spinular row.</p><p>Maxillule (Fig. 22C) similar to that of  Bryocamptus minutus . Coxal endite without spinules; basis with group of spinules.</p><p>Maxilla (Fig. 22D) as in  Bryocamptus minutus, only with slight differences in length and armature of setae.</p><p>Maxilliped (Fig. 23A) similar to that of  Bryocamptus minutus . Differences are only in shorter syncoxa and basis.</p><p>Cuticular process between maxillipeds and P1 (Fig. 23B, C) extremely high, with long spinules, five spinules on each side. Spinules on posterior margin.</p><p>P1 (Fig. 23D) almost like in  Bryocamptus minutus . Basis with two inner groups of long spinules. First exopodal segment with inner spinules. First endopodal segment reaching end of second exopodal segment. Second endopodal segments with smooth inner side. Differences also noticeable in shorter exopodal and endopodal segments.</p><p>P2 (Fig. 24A; Table 3). Praecoxa with row of spinules. Coxa with one lateral row of large spinules, two anterior rows of large spinules and four anterior rows of small spinules. Intercoxal sclerite naked. Basis with proximal pore, rows of spinules at base of endopod and exopod; with outer spine. All endopodal and exopodal segments with outer spinules. Exopod three-segmented; first exopodal segment with outer spinulose spine, apically with frill; second segment with outer spinulose spine, inner pectinate seta, inner slender spinules and apical frill; third segment with three outer spinulose spines, two apical setae and one inner pectinate seta. Endopod two-segmented; first segment with inner seta; second segment with distinct border between ancestral segments, outer spinulose spine, two apical pinnate setae and two inner pectinate setae.</p><p>P3 (Fig. 24B; Table 3). Praecoxa with spinular row. Coxa with one lateral row of large spinules, two anterior rows of large spinules and four anterior rows of small spinules. Intercoxal sclerite without spinules. Basis with outer seta, proximal pore, and rows of spinules at base of endopod and exopod. Exopod three-segmented; first exopodal segment with outer spinulose spine, outer spinules, apically with frill; second segment with outer spinulose spine, outer spinules, inner pectinate seta, inner slender spinules and apical frill; third segment with three outer spinulose spines, two apical setae and two inner pectinate setae. Endopod two-segmented; first segment with inner seta; second segment with distinct border between ancestral segments, outer spinules, outer spinulose spine, two apical pinnate setae and three inner setae.</p><p>P4 (Fig. 24C; Table 3). Praecoxa with spinular row. Coxa with one lateral row of large spinules, two anterior rows of large spinules and four anterior rows of small spinules. Basis with outer seta, proximal pore, rows of spinules at base of exopod. Exopod three-segmented; first exopodal segment with outer spinulose spine, outer spinules, apically with frill; second segment with outer spinulose spine, outer spinules, inner pectinate seta, inner slender spinules and apical frill; third segment with three outer spinulose spines, two apical setae and two inner pectinate setae. Endopod two-segmented; first segment with inner seta, second segment with outer spinule, outer spinulose spine, apical spiniform spinulose seta, apical pinnate seta and two inner pectinate setae.</p><p>P5 (Fig. 25A) with separate right and left baseoendopods. Baseoendopod reaching ~ 1/2 of exopodal segment; with four pores, spinule at base of outer seta; outer seta of basis pinnate, long. Endopodal lobe with four long bipinnate setae and two short bipinnate setae V and VI; with small process that may be pore between setae III and IV. Exopod with inner spinule, inner strong pinnate seta, long apical pinnate seta, naked subapical seta and two pinnate outer setae.</p><p>Male. Sexual dimorphism expressed in the antennule, P2-P6, genital segmentation and ornamentation, shape of caudal rami. Cephalothorax and thoracic somites as in female. P6 (Fig. 26B) two asymmetric flaps fused to the somite, with one naked and one pinnate setae. Differences from female in abdomen structure as follows (Fig. 26A, B): first abdominal somite free; first to third abdominal somites with spinular row encircling somite ventrally and laterally; anal somite with ventral spinules; caudal rami with normal setae IV and V; anal operculum with eight simple spinules.</p><p>Antennule (Fig. 25C, D) 10-segmented, haplocer with geniculation between segments 7 and 8. Segments 1, 3, 4, 5, 6, 9, and 10 similar to those of  B. minutus, but differ in length. Segment 2 with small pore on anterior side. Segment 7 with articular plate, with one filiform seta, one small caudate seta and with two large modified laminar setae. Segment 8 with proximal long dentate plate and three modified laminar setae. Armature formula: 1-[1],2-[9],3-[8],4-[2],5-[6+(1+ae)],6-[2],7-[2+2 modified],8-[3 modified],9-[1],10-[7+acr].</p><p>P2 (Fig. 26C, D) as in female, except endopod. Endopod two-segmented. First segment with outer spinule and inner seta. Second segment with notch on distal outer margin, outer spinules, two apical pinnate slender setae and two inner pectinate setae.</p><p>P3 (Fig. 27A-C): praecoxa, coxa, intercoxal sclerite as in female. Basis as in female, but with inner process. Exopod as in female, but third segment with pore. Endopod three-segmented. First endopodal segment with strong seta. Second endopodal segment with posterior seta and long apophysis with double tip. Third segment with probably two small inner setae and two apical pinnate setae.</p><p>P4 (Fig. 27D, E): praecoxa, coxa, intercoxal sclerite, basis, exopod as in female. Endopod two-segmented; first segment short, unarmed; second segment with outer spinule, spinulose spine, outer apical spiniform spinulose seta and inner apical bipinnate seta.</p><p>P5 (Fig. 27B) right and left fused medially. Baseoendopod with three pairs of pores, outer spinule and outer long pinnate seta; endopodal lobe with two strong spinulose apical spines. Exopod with two outer spinulose setae, naked outer subapical seta, long apical spinulose seta, one inner spinulose seta and one inner pectinate seta with long setulles.</p><p>Variability.</p><p>Individuals with two outer spines on the third exopodal segment of P2 and P4 were found (Figs 26D, 27E). One female was also found with both simple and bifid spinules on the anal operculum.</p><p>Etymology.</p><p>The species is named so because it was found on the Putorana Plateau.</p><p>Remarks.</p><p>The species as a whole is similar to  B. hutchinsoni, including the structure of caudal rami; however, it differs well in two-segmented endopods P2 and P3. Another find of  B. hutchinsoni (Carter 1944) differs markedly in the structure of its caudal rami and is not similar to  B. putoranus sp. nov.</p></div>	https://treatment.plazi.org/id/30E8A0B801335BF594D646705779EE8E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Novikov, Aleksandr;Sharafutdinova, Dayana;Chertoprud, Elena	Novikov, Aleksandr, Sharafutdinova, Dayana, Chertoprud, Elena (2023): Two new species of Bryocamptus (Copepoda, Harpacticoida, Canthocamptidae) from the Russian Arctic and comparison with Bryocamptus minutus (Claus, 1863). ZooKeys 1138: 89-141, DOI: http://dx.doi.org/10.3897/zookeys.1138.90580, URL: http://dx.doi.org/10.3897/zookeys.1138.90580
