taxonID	type	description	language	source
3252E5707209BA7EBDC4B1AAFEE1FD07.taxon	description	D i s t r i b u t i o n: Known from Tanzania and Kenya, new record for Yemen and the Arabian Peninsula.	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E5707209BA7EBDC4B28AFD7EFA27.taxon	type_taxon	Type species: Epomiator yemeniticus, new species	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E5707209BA7EBDC4B28AFD7EFA27.taxon	diagnosis	Diagnosis: Body length 5 mm. Flagellum slender and filiform, slightly widened beyond middle. Temple strongly narrowed behind eye. Ocelli large. Clypeus flat, very wide, with straight apical margin. Genal carina reaching hypostomal carina far from mandibular base. Mandible with two teeth; lower tooth much smaller than upper one, not bent inwards. Epomia very long and almost reaching hind margin of pronotum. Notaulus indistinct. Mesoscutum punctate and granulate, dull. Mesopleuron densely punctate; speculum almost smooth. Juxtacoxal carina present. Scutellum slightly convex, with lateral carina. Propodeum with clearly differentiated basal horizontal and apical descending parts, similar to Microlongichneumon (see Heinrich 1968: 1234, fig. 219), but anterior transverse carina (costula) present in frontal half of area superomedia. Area basalis without median tubercle. Propodeal spiracle oval. Hind coxa without scopa. Claws simple. Areolet of fore wing strongly narrowed frontally. Vein 1 cu-a interstitial. Metasoma clearly oxypygous. Postpetiolus almost entirely smooth and shining. Gastrocoelus weak. Thyridium small, 0.5 × as wide as interval between thyridia. Ovipositor sheath slightly reaching behind metasomal apex. Male unknown so far.	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E5707209BA7EBDC4B28AFD7EFA27.taxon	discussion	Remark: This new genus belongs to the tribe Ichneumonini, subtribe Cratichneumonina as defined by HEINRICH (1968). In the key for the African genera of Cratichneumonina provided by HEINRICH (1968: 792 - 798), this genus runs to couple 45: Microlongichneumon HEINRICH. It resembles that genus in most morphological features, especially the structure of the mandible and propodeum, but differs by its smaller size, very wide clypeus with straight apical margin, and white marks on apical tergites. Especially, the epomia is unusually long and reaches almost the hind border of pronotum (fig. 8).	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E5707209BA7DBDC4B5EAFEA4FB70.taxon	description	D e s c r i p t i o n: ♀. Body length 5 mm. Flagellum slender and filiform, with 26 flagellomeres, 1 st flagellomere 4.0 × and 2 nd flagellomere 3.2 × longer than wide, combined length 0.69 × eye length, preapical flagellomer slightly longer than wide. Temple strongly narrowed behind eye. Distance between lateral ocellus and compound eye 0.9 × ocellar diameter. Frons with scattered punctures and shining; face and clypeus densely punctured, smooth and shining between punctures. Clypeus 3 × wider than long, flat, with straight apical margin. Malar space 0.8 × as long as width of mandibular base. Gena narrow, c. 0.5 × as wide as eye in lateral view. Genal carina reaching hypostomal carina far from mandibular base. Mandible slender, with almost parallel sides; upper mandibular tooth much longer than lower tooth. Side of pronotum with superficial punctures, shining. Epomia very long and almost reaching hind margin of pronotum (fig. 12). Notaulus indistinctly impressed. Mesoscutum with punctures, granulate and dull. Mesopleuron densely punctate, partly with longitudinal striation in ventral part; speculum large, almost smooth and strongly shining. Sternaulus impressed in frontal third of mesopleuron. Metapleuron punctate; juxtacoxal carina present. Scutellum slightly convex, about as long as wide, with strong lateral carina almost to apex, with very scattered punctures, strongly shining. Propodeum with clearly differentiated basal horizontal and apical vertical parts, similar to Microlongichneumon, but anterior transverse carina (costula) present in frontal half of area superomedia. Propodeal spiracle c. 2 × longer than wide. Mid and hind coxae densely punctate; hind coxa without scopa. Hind femur 4.4 × longer than wide, densely punctate. Claws simple. Areolet of fore wing strongly narrowed frontally, frontal distance between vein 2 rs-m and 3 rs-m 1.5 × their width; vein 2 m-cu reaching middle of areolet. Vein 1 cu-a interstitial. Metasoma oxypygous. Postpetiolus slightly widened to apex; median field not clearly separated, almost completely smooth and shining. 2 nd tergite 0.9 × as long as wide. Gastrocoelus not distinctly impressed. Thyridium small, 0.5 × as wide as interval between thyridia. 2 nd and 3 rd tergites densely punctate and granulate, dull; 3 rd tergite ± shining apically. 4 th and 5 th tergites with very superficial punctures, smooth and shining. Ovipositor sheath slightly reaching behind metasomal apex. Color: yellowish-brown. Stripes or rings on flagellomeres 8 - 13, stripes on frontal orbit, on vertex and on upper outer orbit ivory. Petiolus yellowish brown. Postpetiolus and following tergites dark brown, 6 th and 7 th tergites dorsally with large ivory marks. Legs reddish brown, all femora yellowish red in basal 0.1. Wings hyaline; pterostigma yellowish. ♁ unknown.	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E570720ABA7DBDC4B501FE2EFA7F.taxon	description	R e m a r k: 2 nd tergite reddish, 3 rd tergite diffusely reddish and yellowish; the specimen is otherwise typical. D i s t r i b u t i o n: A transpalaearctic species, not recorded from Yemen and the Arabian Peninsula before.	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E570720BBA7CBDC4B0F9FDA2FEE8.taxon	materials_examined	M a t e r i a l e x a m i n e d: Yemen (5962) 12 km NW Manakhah, Mal. trap, 1 ♁ 3. VII. - 21. VIII. 2001, leg. A. v. Harten [Leiden].	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E570720BBA7CBDC4B0F9FDA2FEE8.taxon	distribution	D i s t r i b u t i o n: Widespread in Western, Central, and Eastern Africa, new record for Yemen and the Arabian Peninsula.	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E570720BBA79BDC4B1B9FC23F993.taxon	description	D e s c r i p t i o n: ♁. Body length 11 - 12 mm. Flagellum slender, with 36 - 38 flagellomeres, 1 st flagellomere 2.1 × and 2 nd flagellomere 2.0 × longer than wide, combined length 0.50 - 0.52 × eye length, all flagellomeres longer than wide. Tyloids broadly oval, present on flagellomeres 9 - 21 / 24. Temple strongly narrowed behind eye. Ocelli large, distance between lateral ocellus and compound eye 1.0 × ocellar diameter. Frons partly punctate, shining. Face and clypeus coarsely and sparsely punctate, smooth. Malar space 0.2 × as long as width of mandibular base. Gena narrow, in profile 0.7 × as wide as eye. Mesosoma covered with short whitish setae dorsally and longer setae ventrally. Notaulus distinctly impressed in frontal 0.25 of mesoscutum. Mesoscutum with coarse but scattered punctures, shining. Mesopleuron and metapleuron coarsely and densely punctate; speculum smooth. Scutellum moderately elevated, about as long as wide, with lateral carina in basal 0.2 - 0.5. Propodeum with almost complete carination, but areae basalis and superomedia confluent. Area dentipara distinctly slanting downwards toward apex of propodeum (as in Coelichneumon). Hind femur 4.0 × longer than wide. Claws simple. Areolet pentagonal, frontal distance between veins 2 rs-m and 3 rs-m 2 - 3 × their width; vein 2 m-cu reaching areolet distad its middle. Vein 1 cu-a postfurcal. Nervellus of hind wing reclivous, intercepted in its apical 0.8. Postpetiolus strongly widened, median field not clearly separated, with scattered punctures and very fine and superficial aciculation. Gastrocoelus triangularly impressed. Thyridium oblique, about 0.8 × as wide as interval between thryidia. 2 nd tergite about as long as apically wide. 2 nd to 4 th tergites coarsely and densely rugulose-punctate; 5 th tergite with superficial punctures. Apical margin of hypopygium triangular. Color: Flagellum black, reddish ventrally. Head black, with ± dark reddish tint. Mandible except teeth, palps, labrum, clypeus, face, whole orbits (with two narrow interruptions on vertex and malar space) ivory. Mesosoma reddish; collar, subtegular ridge and postscutellum yellowish. 1 st tergite reddish; metasoma otherwise dark reddish-brown to black, sometimes with a bluish tint. Posterolateral edges of 1 st to 3 rd tergites with ivory triangular marks which are sometimes confluent in middle. 5 th tergite with ivory apical stripe; 6 th and 7 th tergites almost entirely ivory. 2 nd to 4 th sternites ivory apically. Legs blackish; fore and mid legs including coxae and trochanters ± ivory frontally; hind coxa ± red. Pterostigma black. R e m a r k: This new subspecies resembles the nominate species morphologically, it differs mainly by its reduced ivory coloration of mesosoma. Since the associated ♀♀ are unknown yet, this proposed subspecies of Lissosculpta tangana (KRIECHBAUMER) might represent a separated but closely related species. D i s t r i b u t i o n: The nominate subspecies is widely distributed in Central and Southern Africa (Ivory Coast, Guinea, Zimbabwe, Tanzania, and South Africa), the species has not been recorded from Yemen or the Arabian Peninsula before.	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E570720FBA78BDC4B0F8FE51FE48.taxon	description	D i s t r i b u t i o n: Known from Sudan, Ethiopia, and Uganda, new record for Yemen and the Arabian Peninsula.	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E570720FBA78BDC4B1D8FDEBFD50.taxon	description	D i s t r i b u t i o n: Widespread in Africa, known from Ethiopia and Eritrea, new record for Yemen and the Arabian Peninsula.	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E570720FBA76BDC4B321FC07FB70.taxon	description	D e s c r i p t i o n: ♀. Body length 9 - 10 mm. Flagellum slender, with 36 - 38 flagellomeres; 1 st flagellomere 2.0 - 2.2 × and 2 nd flagellomere 1.7 - 2.0 × longer than wide, combined length 0.45 - 0.52 × length of compound eye; flagellomeres behind middle slightly attenuated, widest flagellomeres almost square. Temple strongly narrowed behind eye. Ocelli large, distance between lateral ocellus and compound eye 0.8 × ocellar diameter. Frons coarsely rugose-punctate. Face punctate, partly rugose. Clypeus moderately convex, with scattered punctures, strongly shining. Malar space 1.0 × as long as width of mandibular base. Genal carina reaching hypostomal carina far from mandibular base. Mesosoma covered with short whitish setae. Notaulus indistinctly impressed at frontal margin of mesoscutum. Mesoscutum with coarse punctures, shining. Mesopleuron and metapleuron coarsely punctate, partly with longitudinal rugosity. Scutellum flat, about as long as wide, without lateral carina. Propodeum rounded in profile, but with differentiated horizontal and vertical areas. Area superomedia hexagonal, as wide as or slightly wider than long; anterior transverse carina (costula) frontal to its middle. Area petiolaris with longitudinal carina. Mid and hind coxae with scattered punctures in apical halves, strongly shining. Hind coxa without scopa. Hind femura 3.8 × longer than wide, almost smooth and without punctures in ventral half. Areolet pentagonal, frontal distance between veins 2 rsm and 3 rs-m 3 - 4 × their width; vein 2 m-cu reaching areolet distad to its middle. Vein 1 cu-a postfurcal by 0.3 - 0.4 × its length. Nervellus of hind wing reclivous, intercepted in its apical 0.8. Metasoma semi-amblypygous (fig. 1). Lateral field of postpetiolus punctate, wider than median field, the latter finely aciculate. Gastrocoelus triangularly impressed, with longitudinal ridges. Thyridium slightly impressed, 0.8 × as wide as the interval between thyridia. 2 nd and 3 rd tergites coarsely and densely punctate, polished between punctures. 2 nd tergite longitudinally rugose-striate centrally and 3 rd tergite only basally. 4 th tergite with very scattered punctures, 5 th tergite almost impunctate. Ovipositor sheath not reaching behind metasomal apex. Color: black. Stripes or rings on flagellomeres 8 / 9 - 13 / 14, frontal orbit, usually facial orbit, collar and hind edge or upper border of pronotum, usually spots on tegula and subtegular ridge, scutellum ±, and usually fore and mid coxae frontally yellow. Metasoma red, 5 th or 6 th to 7 th tergites ± darkened or black; 6 th and 7 th tergites with large ivory median spots (rarely 5 th tergite with very small ivory spot medially). Coxae and trochanters black; legs otherwise reddish; hind tarsus ± black. Pterostigma brownish, paler at base. ♁. Body length 10 - 11 mm. Flagellum with 36 - 38 flagellomeres; 1 st flagellomere 3.2 × and 2 nd flagellomere 2.0 × longer than wide, combined length 0.55 × eye lengt. Linear tyloids on flagellomeres 8 - 16 / 18, extending over entire length of flagellomeres. Malar space 0.5 × as long as width of mandibular base. Area superomedia hexagonal, slightly longer than wide. Hind femur 3.9 - 4.0 × longer than wide. Lateral field of postpetiolus about as wide as median field. 2 nd to 6 th tergites with coarse, rather dense punctures, strongly shining between punctures. 2 nd and 3 rd tergites with longitudinal rugose-striation medially. Hypopygium triangular, hind margin pointed medially. Structure otherwise as in ♀. Color: black. Distal flagellomeres pale brownish ventrally. Mandible except teeth, palps, clypeus and face except median longitudinal black stripe, scape ventrally, frontal orbit, collar and upper border of pronotum, tegula and subtegular ridge, two paramedian spots on mesoscutum, entire scutellum, fore and mid coxae and trochanters frontally, and sometimes dorsal spot on hind coxa yellowish. Metasoma red, 6 th tergite apically and 7 th tergite mainly blackish; 5 th tergite with narrow ivory stripe apically; 6 th tergite with a large triangular ivory mark and 7 th tergite largely ivory. Coxae and trochanters black; legs otherwise reddish, hind femur in apical 0.1 and hind tibia at apical 0.3 infuscate; hind tarsus ± black. Pterostigma brownish, paler at base. R e m a r k: This species is closely related to Spilothyrateles tibialis (CAMERON, 1906) from South Africa and differs slightly by the color pattern, especially in the ♁ (e. g. yellow mesoscutal spots and ivory spot on 6 th tergite absent in Spilothyrateles tibialis). Probably, the new species is an Arabian subspecies of S. tibialis, but more material from the Afrotropical area is necessary to clarify the relationship of these species.	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E5707201BA75BDC4B501FD70FBBF.taxon	materials_examined	M a t e r i a l e x a m i n e d: Holotype: (♀) Yemen (5554) Ar Rujun 15. I. - 9. IV. 2001, Mal. trap, A. v. Harten, RMNH` 02 [Leiden]. Paratype: (♀) Yemen (6007) Suq Bani Mansour, Mal. trap, 28. VIII. - 14. XI. 2001, A. v. Harten RMNH` 02 [coll. RIEDEL].	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E5707201BA75BDC4B501FD70FBBF.taxon	description	D e s c r i p t i o n: ♀. Body length 6 mm. Flagellum slender, with 29 flagellomeres; 1 st flagellomere 3.8 × and 2 nd flagellomere 3.0 × longer than wide, combined length 0.75 × eye length; flagellomeres behind middle slightly attenuated, widest flagellomeres almost square. Temple very strongly narrowed behind eye. Ocelli small, distance between lateral ocellus and compound eye 1.3 × ocellar diameter. Face, gena and clypeus with fine punctures, almost smooth, central face finely granulate. Clypeus with medio-apical impression. Malar space 0.8 × as long as width of mandibular base. Mandible with two teeth, lower tooth much smaller than upper tooth. Gena rather narrow, in profile 0.65 - 0.7 × as wide as eye. Genal carina reaching hypostomal carina far from the mandibular base. Mesosoma covered with short whitish setae. Notaulus impressed in frontal 0.3 of mesoscutum. Mesoscutum dull, with very fine superficial punctures and distinct granulation. Mesopleuron and metapleuron punctate and with fine longitudinal rugae. Scutellum moderately convex, about as long as wide, with scattered punctures, smooth, with strong and complete lateral carina. Propodeum with defined basal horizontal and apical descending parts. Coxae and femora with fine and superficial punctures. Hind coxa with scopa. Hind femur 4.4 × longer than wide. Claws simple. Areolet almost pointed frontally; frontal distance between veins 2 rs-m and 3 rs-m 1 × their width, vein 2 m-cu reaching the middle of areolet. Vein 1 cu-a interstitial. Nervellus of hind wing reclivous, intercepted in its apical 0.7. Metasoma strongly oxypygous (fig. 2). Gastrocoelus indistinctly impressed. Thyridium very wide, 2.0 × times wider than the interval between thyridia. 2 nd and 3 rd tergites densely granulate, with very superficial punctures, dull. 4 th and following tergites ± shining. Ovipositor sheath reaching far behind metasomal apex, about as long as 2 nd hind tarsomere (fig. 2). Color: reddish-brown. Flagellum brownish, flagellomeres 7 - 13 with ivory bands. Face and clypeus reddish-brown, gena brownish. Wide facial orbit, frontal orbit, vertex and upper outer orbit ivory. 2 nd and 3 rd tergites dark reddish-brown; 4 th and 5 th tergites dark brown. 5 th tergite apically, and 6 th and 7 th tergites almost entirely ivory. Fore and mid legs reddish brown, hind leg brownish. Pterostigma yellowish. ♁ unknown.	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
3252E5707201BA75BDC4B501FD70FBBF.taxon	discussion	R e m a r k: Due to its small size, this new species resembles the genus Parvaoplus HEINRICH, but the metasoma is clearly oxypygous, and the spiracle of propodeum elongate. In the key of the African members of Stenaoplus (HEINRICH 1968: 1111 - 1115) this new species runs to couplet 13, but differs from Stenaoplus impressus HEINRICH by its much smaller size and the different ivory coloration.	en	Riedel, Matthias (2022): Contribution to the knowledge of the Ichneumoninae (Hymenoptera, Ichneumonidae) of Yemen. Linzer biologische Beiträge 54 (1): 275-286, DOI: http://doi.org/10.5281/zenodo.7507448
