taxonID	type	description	language	source
03A95E063E19FFB8FD90053F03C2B8BF.taxon	materials_examined	Type material Holotype: YPM IZ 003366. CR (not examined). Type locality: Flatts Inlet, Bermuda. Material examined MARTINIQUE • 1 ♂; Grande Anse d’Arlets; 14 ° 29.9 ʹ N, 61 ° 05.4 ʹ W; depth 28 m; 7 Sep. 2016; st. AB 157; MNHN-IU- 2016 - 849 / MK 411162 • 1 ♂; Grande Anse d’Arlets; 14 ° 30.5 ʹ N, 61 ° 06.1 ʹ W; depth 20 – 23 m; 6 Sep. 2016; st. AB 152; MNHN-IU- 2016 - 851 / MK 411164 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 888 / MK 411197 • 1 juv.; Baie de Fort-de-France; 14 ° 32.6 ʹ N, 61 ° 04.7 ʹ W; depth 0 – 1 m; 6 Sep. 2016; st. AM 002; MNHN-IU- 2016 - 85 / MK 411170 • 1 ♂ ov.; same collection data as for preceding; MNHN-IU- 2016 - 1193 • 1 ♂; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 16 m; 18 Sep. 2016; st. AB 189; MNHN-IU- 2016 - 881 / MK 411191 • 1 ♂; Ste-Marie, 14 ° 46.3 ʹ N, 60 ° 57.5 ʹ W; depth 20 m; 20 Sep. 2016; st. AB 260; MNHN-IU- 2016 - 1062 / MK 411040 • 1 juv.; Les Anses-d’Arlet; 14 ° 29.7 ʹ N, 61 ° 05.4 ʹ W; depth 19 m; 7 Sep. 2016; st. AB 155; MNHN-IU- 2016 - 1114 / MK 411053 • 1 ♀ gr.; E of Le Vauclin; 14 ° 34.7 ʹ N, 60 ° 45.9 ʹ W; depth 22 m; 16 Sep. 2016; st. AB 181; MNHN-IU- 2016 - 1180 / MK 411073 • 1 ♀ gr.; Le Robert; 14 ° 41.1 ʹ N, 60 ° 51.4 ʹ W; depth 10 m; 19 Sep. 2016; st. AB 195; MNHN- IU- 2016 - 1182 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1330 / MK 411104 • 2 ♂♂, 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 1408 • 1 ♀; Trois Rivières; 14 ° 27.7 ʹ N, 60 ° 57.8 ʹ W; depth 4 – 5 m; 25 Sep. 2016; st. AB 354; MNHN-IU- 2016 - 1276 / MK 411095 • 1 ♀ gr.; same collection data as for preceding; MNHN-IU- 2016 - 1380 / MK 411108 • 1 ♂ ov.; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2016 - 1379 / MK 411107 • 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 1384.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E19FFB8FD90053F03C2B8BF.taxon	discussion	Remarks This species was distinguished from Achelia sawayai Marcus, 1940 based on the key in Müller & Krapp (2009), relying on the number of palp articles (7 in A. gracilis, 8 in A. sawayai) and the number of dorsolateral tubercles on coxae 1 on the first and second legs of males (4 in A. gracilis, 3 in A. sawayai). Achelia gracilis was previously recorded from Martinique by Bourdillon (1955) on the hulls of scows in Fort-de-France port. Madibenthos sampling reveals that this species is present on both the Atlantic and Caribbean coasts of Martinique. Achelia gracilis is a common species in the Caribbean, almost as common as its close congener, Achelia sawayai (Child 1996 a). DNA barcoding data (Sabroux et al. 2019 b) suggest that this species name hides two potential cryptic species (intracluster p-distance = 0 – 0.010; intercluster p-distance = 0.105 – 0.112), here named group 1 (MNHN-IU- 2016 - 849, - 851, - 857, 881, - 888, - 1114, - 1180, - 1276, - 1379 and - 1380) and group 2 (MNHN-IU- 2016 - 1062 and - 1330) (Table 2 and Appendix).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E19FFB8FD90053F03C2B8BF.taxon	distribution	Distribution Tropical West Atlantic (Bermuda, Caribbean and Brazil). Depth range 0 – 18 m. One record at 157 m is regarded as a likely contamination by shallow fouling on the hull of a ship. We extend this depth-range to 28 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E17FFBAFD9E03270192BBC2.taxon	materials_examined	Type material Holotype not designated (see Lucena & Christoffersen 2018 a). Type locality: Baía de Santos, São Paulo, Brazil. Material examined MARTINIQUE • 1 ♂; Canal de Ste Lucie; 14 ° 22.7 ʹ N, 60 ° 51.3 ʹ W; depth 65 m; 11 Sep. 2016; st. AD 223; MNHN-IU- 2016 - 850 / MK 411163 • 1 ♂; Pointe du Diamant; 14 ° 27.5 ʹ N, 61 ° 02.9 ʹ W; depth 3 m; 15 Sep. 2016; st. AB 126; MNHN-IU- 2016 - 859 / MK 411172 • 1 ♀ gr.; same collection data as for preceding; MNHN-IU- 2016 - 1037 • 1 ♀ gr.; same collection data as for preceding; MNHN-IU- 2016 - 1329 / MK 411103 • 1 ♂; Baie du Robert; 14 ° 42 ʹ N, 60 ° 53.8 ʹ W; depth 2 m; 24 Sep. 2016; st. AB 452; MNHN- IU- 2016 - 884 / MK 411193 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 885 / MK 411194 • 1 ♀ gr.; same collection data as for preceding; MNHN-IU- 2016 - 1073 / MK 411044 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1378 • 1 ♂; Canal de Ste Lucie; 14 ° 22.7 ʹ N, 60 ° 51.3 ʹ W; depth 65 m; 11 Sep. 2016; st. AD 223; MNHN-IU- 2016 - 886 / MK 411195 • 1 ♀ gr.; N of Presqu’Île de la Caravelle; 14 ° 52.9 ʹ N, 60 ° 50.6 ʹ W; depth 61 m; 19 Sep. 2016; st. AD 249; MNHN-IU- 2016 - 1083 • 1 ♀ gr.; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 29 m; 18 Sep. 2016; st. AS 253; MNHN- IU- 2016 - 1108 / MK 411051 • 1 ♀ gr.; Les Anses-d’Arlet; 14 ° 28.5 ʹ N, 61 ° 05.1 ʹ W; depth 29 m; 26 Sep. 2016; st. AD 271; MNHN-IU- 2016 - 1136 / MK 411060 • 1 ♂; Trois Rivières; 14 ° 27.7 ʹ N, 60 ° 57.8 ʹ W; depth 4 – 5 m; 25 Sep. 2016; st. AB 354; MNHN-IU- 2016 - 1172 • 1 ♀; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2016 - 1249 • 1 ♀ gr.; same collection data as for preceding; MNHN-IU- 2016 - 1385 • 1 ♂, 1 ♀ gr.; N of Presqu’Île de la Caravelle; 14 ° 52.5 ʹ N, 60 ° 51.1 ʹ W; depth 58 m; 19 Sep. 2016; st. AD 248; 1 ♀ gr.; MNHN-IU- 2016 - 1297 • 1 ♀ gr.; same collection data as for preceding; MNHN-IU- 2016 - 1331 / MK 411105 • Le François; 14 ° 38.5 ʹ N, 60 ° 46.3 ʹ W; depth 63 m; 15 Sep. 2016; st. AD 234; MNHN-IU- 2016 - 1314 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1332 / MK 411106 • 1 ♂ ov.; Pointe Baham; 14 ° 25 ʹ N, 60 ° 50.4 ʹ W; depth 0 – 2 m; 17 Sep. 2016; st. AM 021; MNHN-IU- 2016 - 1381 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1383 / MK 411109.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E17FFBAFD9E03270192BBC2.taxon	discussion	Remarks DNA barcoding (Sabroux et al. 2019) suggests that A. sawayai is an umbrella-name including three potential cryptic species (intracluster p-distance = 0 – 0.013; intercluster p-distance = 0.133 – 0.191) here designated as group 1 (MNHN-IU- 2016 - 859 and - 1329), group 2 (MNHN-IU- 2016 - 850, - 886, - 1108, - 1136, - 1331 and - 1332) and group 3 (MNHN-IU- 2016 - 884, - 885, - 1073 and - 1383) (Table 2, Appendix). Achelia sawayai was previously recorded from Martinique by Müller (1990 a) at Cap Chevalier (South- East Martinique), and the present sampling reveals that this species is found on both the Atlantic and Caribbean coasts. The three possible cryptic species here detected in Martinique suggest that A. sawayai may represent a species complex including many more species. Indeed, it is a common species of the Tropical Western Atlantic, and its distribution extends to the Eastern Atlantic, the Mediterranean, the Indian Ocean and the Pacific.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E17FFBAFD9E03270192BBC2.taxon	distribution	Distribution Apparently a circumtropical species, mostly found in the West Atlantic (US Georgia, Caribbean, Gulf of Mexico and Brazil), occasionally recorded in the East Atlantic (Gulf of Guinea, Mauritania, Cap Verde), West Mediterranean, Indian Ocean (Papua-New Guinea, Indonesia, Pakistan, Madagascar) and Pacific Ocean (French Polynesia, Fiji). Depth range 0 – 115 m. One record at 157 m is regarded as a likely contamination by shallow fouling on the hull of a ship.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E14FFBAFDA406CD0577BA32.taxon	type_taxon	Type species Ammothea (Ammothella) rugulosa Verrill, 1900, by original designation.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E14FFBBFDE2079E0079B8EB.taxon	materials_examined	Type material Syntypes (?) (see Stock 1955; Dunlop et al. 2007): ZMB _ Pyc _ 46 (not examined). Type locality: Santa Lucia, Naples Bay, Italy. Material examined MARTINIQUE • 1 juv.; Canal de Ste Lucie; 14 ° 23.9 ʹ N, 60 ° 52.6 ʹ W; depth 0 – 2 m; 12 Sep. 2016; st. AM 012; MNHN-IU- 2016 - 1278 / MK 411096 • 1 juv.; same collection data as for preceding; MNHN- IU- 2016 - 1310 / MK 411098 • 1 ♂ ov.; same collection data as for preceding; MNHN-IU- 2016 - 1573 • 1 ♀; Baie de Sans-Souci; 14 ° 34.7 ʹ N, 60 ° 50.7 ʹ W; depth 0 – 1 m; 9 Oct. 2016; st. AM 325; MNHN- IU- 2016 - 1279 • 1 ♀; Le Prêcheur; 14 ° 47.2 ʹ N, 61 ° 13.1 ʹ W; depth 10 – 17 m; 3 Oct. 2016; st. AR 383; MNHN-IU- 2017 - 220.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E14FFBBFDE2079E0079B8EB.taxon	discussion	Remarks This species was already recorded in Martinique by Bourdillon (1955) from the Petite Anse du Diamant, and was sampled during Madibenthos from both the Atlantic and Caribbean coasts. It is commonly sampled in the Atlantic, and available illustrations (e. g., Dohrn 1881; Stock 1955; Child 1982 a; Müller & Krapp 2009) show significant morphological variations. Stock (1955) considered that a slender form and a stout form co-exist, which he suspected to correspond to two adult stages separated by a moult, though this theory was proven invalid by observations of Child (1982 a) on Belize material. Madibenthos material corresponds to the stout form and strongly resembles the drawing of Child (1992 b); this stout form is the one represented by the type material according to the drawings in Stock (1955) (although the type status of this material is not totally clear). The slender form mentioned by Stock (1955) may correspond to one of the hereafter newly described species, Ammothella dirbergi sp. nov. or A. krappi sp. nov., or to A. aff. krappi, most likely the first given Stock’s drawing. Species delimitation based on CO 1 data support the distinction of these three species from A. appendiculata (Sabroux et al. 2019 b). The p-distances between A. appendiculata and these species are very high and vary between 0.172 and 0.197 (see Appendix). Another harsh debate regarding this species is the possible synonymy of Ammothella appendiculata and A. rugulosa. Lucena et al. (2019) suggested that the two species can be differenciated by the length of their appendages.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E14FFBBFDE2079E0079B8EB.taxon	distribution	Distribution Rather cosmopolitan, with records in the West and East Mediterranean, West Atlantic (Florida, Gulf of Mexico, Caribbean, French Guiana, Brazil) and Indo-Pacific (Pakistan, Indonesia, Hong-Kong, New Caledonia). Depth range 0 – 76 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E12FFBFFD9603270057BA18.taxon	description	urn: lsid: zoobank. org: act: 7 CCAA 126 - D 880 - 4 F 80 - B 6 D 7 - A 7 CB 0 C 532 E 6 C Fig. 2	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E12FFBFFD9603270057BA18.taxon	materials_examined	Material examined Holotype MARTINIQUE • ♂; Pointe du Diamant; 14 ° 27.5 ʹ N, 61 ° 02.9 ʹ W; depth 3 m; 15 Sep. 2016; st. AB 126; MNHN-IU- 2016 - 833 / MK 411148. Paratypes MARTINIQUE • 3 ♂♂; 2 ♀♀, 3 ♀♀ gr.; Case-Pilote; 14 ° 38.3 ʹ N, 61 ° 08.4 ʹ W; depth 12 m; 8 Sep. 2016; st. AB 159; MNHN-IU- 2016 - 554 • 1 ♂, 3 ♀♀, 2 ♀♀ gr., 1 juv.; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2016 - 577 • 1 ♂ ov., 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 1252 • 3 ♀♀, 1 ♀ gr., 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 1253 • 1 ♀ gr.; Fond Boucher; 14 ° 39.3 ʹ N, 61 ° 09.6 ʹ W; depth 37 – 40 m; 1 Oct. 2016; st. AD 283; MNHN-IU- 2016 - 1054 • 1 juv.; same collection data as for preceding; MNHN- IU- 2016 - 1224 • 2 ♂♂, 1 juv.; Baie de St-Pierre; 14 ° 45.1 ʹ N, 61 ° 11 ʹ W; depth 17 m; 4 Oct. 2016; st. AB 388; MNHN-IU- 2016 - 1058 • 2 juvs; same collection data as for preceding; MNHN-IU- 2016 - 1308 • 1 ♂; Ste-Marie; 14 ° 46.3 ʹ N, 60 ° 57.5 ʹ W; depth 20 m; 20 Sep. 2016; st. AB 260; MNHN-IU- 2016 - 1066 • 1 ♀; Presqu’Île de la Caravelle; 14 ° 47.5 ʹ N, 60 ° 57.4 ʹ W; depth 14 m; 4 Oct. 2016; st. AB 562; MNHN- IU- 2016 - 1091 / MK 411047 • 1 ♀, 1 juv.; Baie du Robert; 14 ° 40.3 ʹ N, 60 ° 52.8 ʹ W; depth 1 – 8 m; 25 Sep. 2016; st. AR 455; MNHN-IU- 2016 - 1095 • 1 ♀ gr., 1 juv.; same collection data as for preceding; MNHN- IU- 2016 - 1275 • 2 ♀♀ gr., 1 juv.; Anse Couleuvre; 14 ° 50.4 ʹ N, 61 ° 13.4 ʹ W; depth 7 m; 1 Oct. 2016; st. AB 463; MNHN-IU- 2016 - 1120 • 1 ♂ ov.; same collection data as for preceding; MNHN-IU- 2016 - 1399 • 1 juv.; Baie de Fort-de-France; 14 ° 36.1 ʹ N, 61 ° 04 ʹ W; depth 0 – 1 m; 29 Sep. 2016; st. AM 034; MNHN-IU- 2016 - 1153 • 2 ♂♂, 1 ♂ ov.; Rocher du Diamant; 14 ° 26.5 ʹ N, 61 ° 02.4 ʹ W; depth 24 m; 14 Sep. 2016; st. AB 173; MNHN-IU- 2016 - 1158 • 1 ♀; Bellefontaine; 14 ° 39.7 ʹ N, 61 ° 09.6 ʹ W; depth 9 m; 30 Sep. 2016; st. AS 373; MNHN-IU- 2016 - 1174 / MK 411070 • 1 ♂; E of Le Robert; 14 ° 40.1 ʹ N, 60 ° 51.1 ʹ W; depth 16 m; 18 Sep. 2016; st. AS 255; MNHN-IU- 2016 - 1176 • 1 ♀; same collection data as for preceding; MNHN-IU- 2016 - 1315 • 1 juv.; Baie de St-Pierre; 14 ° 44.5 ʹ N, 61 ° 10.8 ʹ W; depth 44 – 47 m; 8 Oct. 2016; st. AS 572; MNHN-IU- 2016 - 1208 • 1 ♀; Grande Anse du Diamant; 14 ° 27.9 ʹ N, 61 ° 01.4 ʹ W; depth 17 m; 26 Sep. 2016; st. AB 358; MNHN-IU- 2016 - 1210 • 1 ♂; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.2 ʹ W; depth 12 m; 21 Sep. 2016; st. AS 092; MNHN-IU- 2016 - 1226 • 1 ♂; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 00.1 ʹ W; depth 12 m; 26 Sep. 2016; st. AB 360; MNHN-IU- 2016 - 1238 • 1 ♀; same collection data as for preceding; MNHN-IU- 2016 - 1239 • 1 ♂; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 29 m; 18 Sep. 2016; st. AS 253; MNHN-IU- 2016 - 1260 / MK 411088 • 1 ♀ gr.; Le Robert; 14 ° 41 ʹ N, 60 ° 49.4 ʹ W; depth 23 m; 24 Sep. 2016; st. AB 401; MNHN-IU- 2016 - 1266 / MK 411092 • 1 juv.; Anse Noire; 14 ° 32 ʹ N, 61 ° 05.3 ʹ W; depth 2 – 8 m; 6 Sep. 2016; st. AR 100; MNHN-IU- 2016 - 1286 • 1 ♂ ov.; Presqu’Île de la Caravelle; 14 ° 46.5 ʹ N, 60 ° 51.5 ʹ W; depth 15 m; 22 Sep. 2016; st. AB 350; MNHN-IU- 2016 - 1303 • 1 ♀ gr.; Le Prêcheur; 14 ° 47.2 ʹ N, 61 ° 13.1 ʹ W; depth 10 – 17 m; 3 Oct. 2016; st. AR 383; MNHN-IU- 2016 - 1306.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E12FFBFFD9603270057BA18.taxon	etymology	Etymology Latinized name, 2 nd decl. (masc.), genitive singular. This species is named after Guillaume Dirberg (MNHN), who collected many specimens of this species by sight, which is remarkable when considering the inconspicuousity of these animals.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E12FFBFFD9603270057BA18.taxon	description	Description (holotype, ♂, MNHN-IU- 2016 - 833) BODY. Trunk completely segmented, cuticle smooth. No dorsomedian ornamentation. Ocular tubercle long, about 5.5 times as long as base width, distal part broad and ovoid carrying pigmented eyes, pointy tip and 2 small pointy lateral sense organs (see Lehmann et al. 2017; Brenneis 2022). Lateral processes about twice as long as wide, well separated by about half of their own diameter, unornamented. PROBOScIS. Pyriform, movable, longer than chelifore or trunk, with blunt tip. ABDOMEN. Long, not extending beyond coxa 2, distal part inflated, then tapering at anus. Dorsally with club-shaped setae and simple setae medially and distally. Abdomen insertion articled on trunk. CHELIFORE. 3 - articled. Scape 2 - articled, 1 st scape article about half as long as 2 nd. 2 nd article long, trumpet-shaped, carrying simple and club-shaped setae. Chela reduced as rounded buds, with distal seta on outer side. PALP. 9 - articled, carrying many setae mostly on last 5 articles. 1 st article shortest, 2 nd slightly shorter than 4 th. 3 rd article about as long as wide. 4 th article longest, about as long as 2 nd, about 6 times as long as wide, with scarce setae. 5 th article about 2.5 times as long as wide. 6 th article shorter than 5 th and twice as long as 8 th, about 4.5 times as long as wide. 7 th to 9 th articles shorter than 6 th, 9 th slightly longer. OvIGER. 10 - articled, with scarce setae. 1 st article as long as wide. 2 nd about 5 times as long as wide, 3 rd about 3 times as long as wide. 4 th article longest, slightly curved, about ⅙ longer than 2 nd. 5 th article slightly curved, about ⅔ of length of 4 th article. 7 th article longest within striglis, 10 th shortest. Strigilis spines compound, formula 3: 2: 1: 2. LEGS. Slender, carrying several short, simple and club-shaped setae, some of them in bouquet. Coxa 1 about 1.5 times as long as wide, with 2 dorsodistal club-shaped setae and often 2 tiny normal setae laterally. Coxa 2 subequal to coxae 1 and 3 together. Coxa 3 about twice as long as wide. Femur 6 times as long as wide; cement gland opening standing on distal margin on dorsal surface, as long tube about ⅓ of femur length. Tibia 1 longest. Tibia 2 slenderer, slightly shorter than tibia 1 but longer than femur. Tarsus short, trapezoid. Propodus about ⅓ as long as tibia 2, with 3 large basal spines, and smaller spines and setae on distal half of sole; main claw about half as long as propodus; auxiliary claws present, thinner, about same size as main claw. MEASUREMENTS (mm). Trunk 0.54; abdomen 0.62; proboscis 0.70; chelifore 0.47; coxa 1 0.14; coxa 2 0.40; coxa 3 0.30; femur 0.84; tibia 1 1.03; tibia 2 0.92; tarsus 0.05; propodus 0.34; main claw 0.17. Sexual dimorphism Oviger articles more elongated in males, legs inflated in mature females. Individual variability Number of ovigeral spines variable, 1 to 4 on 1 st strigilis article, 1 or 2 on 2 nd strigilis article, 1 or 2 on 3 rd strigilis article. Lateral processes variable in length. Club-shaped setae sometimes slightly cleft, as on coxa 1 of left 2 nd leg of holotype.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E12FFBFFD9603270057BA18.taxon	discussion	Remarks This species has a strong affinity with Ammothella indica Stock, 1954, which is a common species in the Indo-Pacific, but never recorded in the Atlantic or Caribbean. The two species belong to the same group of Ammothella with club-shaped spines, and the lengths of the ocular tubercle, of leg and chelifore articles, and of the cement gland tube in males, are about the same. The relative lengths of palp articles are also similar between the two species. There are however some differences in the representations given by Stock (1954 b) and Nakamura (1987). The three main morphological differences between A. indica and A. dirbergi sp. nov. are the following: (i) lateral sense organs are inconspicuous in A. indica; (ii) the abdomen of A. dirbergi does not present a diadem of setae as in A. indica; and (iii) the abdomen of A. dirbergi is not as curved as in A. indica. This species can be differentiated from most other described species of Ammothella based on the long ocular tubercle and the club-shaped spines on the legs and abdomen. Species sharing these features are Ammothella fistella Lee & Arango, 2003, A. gertrudae Müller & Krapp, 2009, A. panamensis Child, 2004, A. prolixa Child, 1990 and A. setacea (Helfer, 1938). Ammothella dirbergi sp. nov. differs from A. fistella in lacking tubular spines on the lateral processes and cephalon, and by its longer ocular tubercle and leg articles; it differs from A. gertrudae in lacking long tubercles on the lateral processes; it differs from A. panamensis in lacking club-shaped setae on the lateral processes, has longer tibiae 2, and has shorter lateral processes; it differs from A. prolixa in having a shorter distance between the lateral processes and no dorsal ornamentation of the lateral processes; and it differs from A. setacea in lacking long tubercles on the lateral processes and in having a shorter ocular tubercle and a completely segmented trunk (Arnaud & Child 1988; Child 1990, 2004; Lee & Arango 2003; Müller & Krapp 2009). Finally, Ammothella krappi sp. nov. is a close relative of A. dirbergi sp. nov. Ammothella dirbergi differs in having a shorter ocular tubercle (less than 6 times as long as wide in A. dibergi, more than 7 times in A. krappi) and a shorter cement gland tube (⅓ as long as the femur in A. dirbergi, half as long in A. krappi). The two species can be distinguished based on CO 1 data (Sabroux et al. 2019): while in A. dirbergi intraspecific p-distances are between 0 and 0.012, the p-distances between the types of A. dirbergi and A. krappi are between 0.123 and 0.134, and those between A. dirbergi and A. aff. krappi are between 0.089 and 0.097.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E12FFBFFD9603270057BA18.taxon	distribution	Distribution Only known from Martinique. Ammothella dirbergi was sampled on the Atlantic and Caribbean coasts. Depth range 0.5 – 47 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E11FFA0FDB107AC03AFBC8D.taxon	materials_examined	Type material Holotype: NL ZMA. PYC. P. 1062 (not examined). Type locality: southeast of Oyster Pond, St Martin. Material examined MARTINIQUE • 1 ♀ gr.; Grande Anse du Diamant; 14 ° 28.5 ʹ N, 61 ° 01.1 ʹ W; depth 7 m; 9 Sep. 2016; st. AD 216; MNHN-IU- 2016 - 829 / MK 411146 • 1 ♀ gr.; Canal de Ste Lucie; 14 ° 23.9 ʹ N, 60 ° 52.6 ʹ W; depth 0 – 2 m; 12 Sep. 2016; st. AM 012; MNHN-IU- 2016 - 830 / MK 411147 • 1 ♀ gr.; Baie de Fort-de-France; 14 ° 33 ʹ N, 61 ° 03.7 ʹ W; depth 2 m; 7 Sep. 2016; st. AB 104; MNHN-IU- 2016 - 831 • 1 juv.; Pointe du Vauclin; 14 ° 33.6 ʹ N, 60 ° 49.7 ʹ W; depth 2 m; 12 Sep. 2016; st. AD 230; MNHN-IU- 2016 - 878 / MK 411188 • 1 ♀ gr.; same collection data as for preceding; MNHN-IU- 2016 - 555 • 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 1285 • 1 ♂ ov., 1 ♀ gr.; Ste-Luce; 14 ° 27.3 ʹ N, 60 ° 55.5 ʹ W; depth 15 m; 10 Sep. 2016; st. AB 062; MNHN-IU- 2016 - 556 • 1 ♂, 1 ♂ ov., 5 ♀♀ gr.; Baie de Sans-Souci; 14 ° 34.7 ʹ N, 60 ° 50.7 ʹ W; depth 0 – 1 m; 9 Oct. 2016; st. AM 325; MNHN-IU- 2016 - 1098 • 1 juv.; Banc du Diamant; 14 ° 26.4 ʹ N, 61 ° 01.7 ʹ W; depth 18 m; 15 Sep. 2016; st. AB 177; MNHN-IU- 2016 - 1105 • 1 ♀ gr.; Schoelcher; 14 ° 37.7 ʹ N, 61 ° 07.2 ʹ W; depth 4 m; 27 Sep. 2016; st. AB 460; MNHN-IU- 2016 - 1154 • 1 ♂, 1 ♀; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 00.1 ʹ W; depth 12 m; 26 Sep. 2016; st. AB 360; MNHN- IU- 2016 - 1217 • 1 ♀; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 02.8 ʹ W; depth 0 – 1 m; 27 Sep. 2016; st. AM 033; MNHN-IU- 2016 - 1464.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E11FFA0FDB107AC03AFBC8D.taxon	discussion	Remarks Ammothella exornata was recorded from Martinique by Müller (1990 a) from Ilet Cabrits, Petite Anse Macabou and Baie de Tartane. We found it on both the Atlantic and Caribbean coasts.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E11FFA0FDB107AC03AFBC8D.taxon	distribution	Distribution Exclusively in the Caribbean area. Depth range 0 – 10 m. We extend this depth range to 18 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0EFFA3FD8906080048BD8C.taxon	description	urn: lsid: zoobank. org: act: 016 C 5 DC 5 - 7 B 37 - 4 A 2 F-B 4 CC- 3176173 A 3 C 65 Fig. 3	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0EFFA3FD8906080048BD8C.taxon	materials_examined	Material examined Holotype MARTINIQUE • ♂; N of Le Lorrain, 14 ° 55.1 ʹ N, 61 ° 02.4 ʹ W; depth 63 m, 21 Sep. 2016; st. AD 257, MNHN-IU- 2016 - 1132 / MK 411059. Paratypes MARTINIQUE • 2 ♂♂, 4 ♀♀ gr.; Le Prêcheur; 14 ° 49.1 ʹ N, 61 ° 13.8 ʹ W; depth 20 – 25 m; 8 Oct. 2016; st. AS 576; MNHN-IU- 2016 - 1085 • 1 ♀ gr.; SW of Vétiver; 14 ° 37.6 ʹ N, 61 ° 08.6 ʹ W; depth 66 m; 7 Oct. 2016; st. AD 612; MNHN-IU- 2016 - 1113 / MK 411052 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1290 • 1 ♂; Trois Rivières; 14 ° 27.5 ʹ N, 60 ° 58.2 ʹ W; depth 17 m; 9 Oct. 2016; st. AB 578; MNHN-IU- 2016 - 1168 / MK 411069 • 1 ♂; Macouba; 14 ° 55.1 ʹ N, 61 ° 09 ʹ W; depth 80 m; 27 Sep. 2016; st. AD 274; MNHN-IU- 2016 - 1243 • 1 ♂; Trois Rivières; 14 ° 27.5 ʹ N, 60 ° 58.2 ʹ W; depth 17 – 19 m; 9 Oct. 2016; st. AS 579; MNHN-IU- 2016 - 1281 • 1 ♀, 1 ♀ gr.; Le Prêcheur; 14 ° 48.5 ʹ N, 61 ° 13.8 ʹ W; depth 0 – 56 m; 8 Oct. 2016; st. AS 574; MNHN-IU- 2016 - 1287 • 1 ♂, 1 ♀ gr., 1 juv.; Baie de St-Pierre; 14 ° 44.5 ʹ N, 61 ° 10.8 ʹ W; depth 44 – 47 m; 8 Oct. 2016; st. AS 572; MNHN-IU- 2019 - 3395.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0EFFA3FD8906080048BD8C.taxon	etymology	Etymology Latinized name, 2 nd decl. (masc.), genitive singular. This species is named after Franz Krapp, who passed away in early 2022. Through this dedication, we wish to express not only our recognition of his important contribution to pycnogonid taxonomy, but also in memory of the great help he was to the first author.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0EFFA3FD8906080048BD8C.taxon	description	Description (holotype, ♂, MNHN-IU- 2016 - 1132) BODY. Trunk completely segmented; cuticle smooth. No dorsomedian ornamentation. Ocular tubercle long, about 8.5 times as long as base width, distal part broad and round, carrying pigmented eyes, blunt tip and 2 small, rounded lateral sense organs. Lateral processes about 3 times as long as base width, well separated by more than their own diameter, sometimes carrying setae. PROBOScIS. Pyriform with slender elongated base, movable, longer than chelifore or trunk, with blunt tip. ABDOMEN. Long, segmented at base, not extending beyond coxa 2, distal part inflated, then tapering at anus. Dorsally with club-shaped setae and with simple setae medially and distally. CHELIFORE. 3 - articled. Scape 2 - articled, 1 st scape article about half as long as 2 nd. 2 nd article long, trumpet-shaped, carrying simple and club-shaped setae. Chela reduced as rounded buds, with finger and distal seta on outer side. PALP. 9 - articled, carrying many setae mostly on last 5 articles. 1 st article shortest, 2 nd article slightly shorter than 4 th. 3 rd article about as long as wide. 4 th article longest, about as long as 2 nd, about 6 times as long as wide, with scarce setae. 5 th article about 2.5 times as long as wide. 6 th article shorter than 5 th and twice as long as 8 th, about 4.5 times as long as wide. 7 th to 9 th articles shorter than 6 th, 9 th longest among them. OvIGER. 10 - articled, with scarce setae. 1 st article about as long as wide, 2 nd about 7 times as long as wide, 3 rd about 3 times. 4 th article as long as 2 nd but slenderer, slightly curved. 5 th article about ⅔ of length of 4 th article. 7 th article longest within striglis, about as long as 6 th, 10 th shortest. Strigilis spines compound, formula 2: 1: 1: 2. LEGS. Slender, carrying several short, simple and club-shaped setae, some of them in bouquet. Coxa 1 about 5 / 4 as long as wide, with two dorsodistal club-shaped setae and often two tiny normal setae laterally. Coxa 2 longer than coxae 1 and 3 together. Coxa 3 about 3 times as long as wide. Femur about 8 times as long as wide, cement gland opening standing on distal margin on dorsal surface, as very long tube about half as long as femur. Tibia 1 longest. Tibia 2 slenderer, slightly shorter than tibia 1 but longer than femur. Tarsus short, trapezoid. Propodus about 0.4 times as long as tibia 2, with 3 large basal spines, and smaller spines and setae on distal half of sole; main claw about half as long as propodus; auxiliary claws present, thinner, more than ⅘ as long as main claw. MEASUREMENTS (mm). Trunk 0.52; abdomen 0.78; proboscis 0.92; chelifore 0.56; coxa 1 0.17; coxa 2 0.53; coxa 3 0.26; femur 0.91; tibia 1 1.13; tibia 2 0.96; tarsus 0.06; propodus 0.37; main claw 0.19. Sexual dimorphism Oviger articles more elongated in males, legs inflated in mature females. Individual variability Ocular tubercle size from 7 to 9 times as long as base width. Number of ovigeral spines variable, 1 to 4 on 1 st strigilis article, and 1 or 2 on 2 nd – 3 rd strigilis articles.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0EFFA3FD8906080048BD8C.taxon	discussion	Remarks This species is distinguished from most other species of Ammothella using the same criteria as for A. dirbergi sp. nov., of which this species seems to be a close relative. Distinction from A. dirbergi is supported by CO 1 data (Sabroux et al. 2019 b): while for A. krappi sp. nov. intraspecific p-distances are between 0.011 and 0.036, the p-distances in A. dirbergi are between 0.123 and 0.134. Morphologically, A. krappi has a longer ocular tubercle than A. dirbergi (more than seven times as long as wide in A. krappi, less than six times as long as wide in A. dirbergi), the tip of the ocular tubercle is blunt rather than acute, and the lateral sense organs are smaller than in A. dirbergi. The cement gland tube is also longer in A. krappi, as well as the leg articles. The species was sampled on the Caribbean and northern coasts of Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0EFFA3FD8906080048BD8C.taxon	distribution	Distribution Only known from Martinique. Depth range 17 – 80 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0DFFA5FDA001200079BD8C.taxon	description	Fig. 4	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0DFFA5FDA001200079BD8C.taxon	materials_examined	Material examined MARTINIQUE • 1 ♂; Grande Anse d’Arlets; 14 ° 29.9 ʹ N, 61 ° 05.4 ʹ W; depth 28 m; 7 Sep. 2016; st. AB 157; MNHN-IU- 2016 - 832 • 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 553 • 2 ♂♂; same collection data as for preceding; MNHN-IU- 2016 - 567 • 1 ♀ gr.; same collection data as for preceding; MNHN-IU- 2016 - 1316 • 1 juv.; Passe du Marin; 14 ° 26.9 ʹ N, 60 ° 54 ʹ W; depth 0 – 15 m; 11 Sep. 2016; st. AS 066; MNHN-IU- 2016 - 834 / MK 411149 • 1 ♂; same collection data as for preceding; MNHN- IU- 2016 - 1147 • 1 ♂; Les Anses-d’Arlet; 14 ° 28.1 ʹ N, 61 ° 05.3 ʹ W; depth 72 m; 8 Oct. 2016; st. AD 616; MNHN-IU- 2016 - 1042 • 1 ♂; same collection data as for preceding; MNHN-IU- 2019 - 3400 • 7 ♂♂, 2 ♀♀; Rocher du Diamant; 14 ° 26.5 ʹ N, 61 ° 02.4 ʹ W; depth 26 – 32 m; 14 Sep. 2016; st. AS 071; MNHN- IU- 2016 - 1050 • 1 ♂, 1 ♀ gr.; same collection data as for preceding; MNHN-IU- 2016 - 1109 • 1 ♂; Baie du Robert; 14 ° 42 ʹ N, 60 ° 53.8 ʹ W; depth 2 m; 24 Sep. 2016; st. AB 452; MNHN-IU- 2016 - 1075 • 1 ♂; Passe du Marin; 14 ° 26.6 ʹ N, 60 ° 54.3 ʹ W; depth 15 m; 9 Sep. 2016; st. AB 058; MNHN-IU- 2016 - 1131 • 1 ♀, 1 juv.; Baie de Fort-de-France; 14 ° 33.2 ʹ N, 61 ° 01.6 ʹ W; depth 1 – 8 m; 6 Oct. 2016; st. AR 391; MNHN-IU- 2016 - 1175 • 1 juv.; SW of Vétiver; 14 ° 37.6 ʹ N, 61 ° 08.6 ʹ W; depth 66 m; 7 Oct. 2016; st. AD 612; MNHN-IU- 2017 - 221 • 1 ♂; Anse Couleuvre; 14 ° 50.4 ʹ N, 61 ° 13.4 ʹ W; depth 7 m; 1 Oct. 2016; st. AB 463; MNHN-IU- 2019 - 3393 • 1 ♂, 1 ♀ gr.; Baie de St-Pierre; 14 ° 44.5 ʹ N, 61 ° 10.8 ʹ W; depth 44 – 47 m; 8 Oct. 2016; st. AS 572; MNHN-IU- 2019 - 3394 • 1 juv.; Fond Boucher; 14 ° 39.3 ʹ N, 61 ° 09.6 ʹ W; depth 37 – 40 m; 1 Oct. 2016; st. AD 283; MNHN-IU- 2019 - 3396 • 1 ♂, 1 ♀; same collection data as for preceding; MNHN-IU- 2019 - 3397 • 2 ♂♂ ov.; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2019 - 3398 • 1 ♂; Baie du Robert; 14 ° 40.3 ʹ N, 60 ° 52.8 ʹ W; depth 1 – 8 m; 25 Sep. 2016; st. AR 455; MNHN-IU- 2019 - 3399.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0DFFA5FDA001200079BD8C.taxon	discussion	Remarks The juvenile specimen MNHN-IU- 2016 - 834 shows strong affinities to A. krappi sp. nov., even sharing with this species a taller ocular tubercle than in A. dirbergi sp. nov. However, this specimen differs slightly from the holotype of A. krappi by having a more acute ocular tubercle tip and more conspicuous lateral sense organs. This distinction from A. krappi is supported by a high molecular barcode divergence (p distance = 0.129 – 0.131). Several adults that could not be barcoded (listed above) present a similar shape of the ocular tubercle, and they are likely to belong to the same species as the juvenile, MNHN- IU- 2016 - 834. Their ocular tubercle is taller, but this is coherent with the fact that the barcoded specimen is a juvenile. However, all these specimens are extremely similar to the holotype of A. krappi, except regarding the shape of the distal region of the ocular tubercle. While we regard this group as likely to represent a new species, the similarities between A. krappi and the adults listed here as A. aff. krappi make a new species description untimely, as long as the identification of the adults as the same species as the juvenile, MNHN-IU- 2016 - 834, is not confirmed by barcoding. These specimens were sampled on both the Atlantic and Caribbean coasts.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0DFFA5FDA001200079BD8C.taxon	distribution	Distribution Only recorded in Martinique at the moment. Depth range 2 – 72 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0BFFA5FDC7012003D8B863.taxon	materials_examined	Type material Holotype: USNM 81099 (not examined). Type locality: Logger-head Key, Tortugas, southwest Florida. Material examined MARTINIQUE • 1 ♂; Presqu’Île de la Caravelle; 14 ° 46.5 ʹ N, 60 ° 51,5 ’ W; depth 15 m; 22 Sep. 2016; st. AB 350; MNHN-IU- 2016 - 1302.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0BFFA5FDC7012003D8B863.taxon	discussion	Remarks Only one specimen was collected, from which we did not succeed in amplifying the CO 1 barcode sequence due to the poor quality of the DNA extract (probably due to the advanced state of decay of the specimen, which hardly consists of more than an empty cuticle, indicating that it was probably dead upon sampling). This species is generally rare in samplings, though Child (1979) examined a rather rich collection at the Smithsonian Tropical Reseach Institute in Panama. The species was recorded in Martinique by Müller (1990 a) in Baie de Tartane, so both records of this species in Martinique are from the region of Presqu’Île de la Caravelle.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0BFFA5FDC7012003D8B863.taxon	distribution	Distribution Typically Caribbean, with one record on the Pacific shore of Mexico (Oaxaca). Depth range 0 – 5 m. We extend this depth range to 15 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0BFFA6FDAF05550079B8E4.taxon	materials_examined	Type material Holotype: USNM 231856 (not examined). Type locality: San Diego, California. Material examined MARTINIQUE • 1 ♂; Pointe du Diamant; 14 ° 27.5 ʹ N, 61 ° 02.9 ʹ W; depth 3 m; 15 Sep. 2016; st. AB 126; MNHN-IU- 2016 - 826 / MK 411145 • 1 ♀; Pointe du Vauclin; 14 ° 33.6 ʹ N, 60 ° 49.7 ʹ W; depth 2 m; 12 Sep. 2016; st. AD 230; MNHN-IU- 2016 - 827 • 2 ♂♂, 1 ♀; same collection data as for preceding; MNHN- IU- 2016 - 1141 • 1 ♂ ov.; Ste-Luce; 14 ° 27.3 ʹ N, 60 ° 55.5 ʹ W; depth 15 m; 10 Sep. 2016; st. AB 062; MNHN-IU- 2016 - 828 • 1 juv.; Pointe Michel; 14 ° 26.4 ʹ N, 60 ° 49.3 ʹ W; depth 2 m; 12 Sep. 2016; st. AB 120; MNHN-IU- 2016 - 557 • 1 ♂; Pointe Baham; 14 ° 25 ʹ N, 60 ° 50.4 ʹ W; depth 0 – 2 m; 17 Sep. 2016; st. AM 021; MNHN-IU- 2016 - 581 • 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 582 • 1 juv.; Le Prêcheur; 14 ° 47.2 ʹ N, 61 ° 13.1 ʹ W; depth 2 m; 3 Oct. 2016; st. AB 382; MNHN-IU- 2016 - 584 • 1 ♂, 2 ♂♂ ov., 1 ♀, 1 juv.; Grand’Rivière; 14 ° 52 ʹ N, 61 ° 11.7 ʹ W; depth 6 m; 3 Oct. 2016; st. AB 556; MNHN-IU- 2016 - 1070 • 1 ♂ ov., 1 ♀; Presqu’Île de la Caravelle; 14 ° 47.5 ʹ N, 60 ° 57.4 ʹ W; depth 14 m; 4 Oct. 2016; st. AB 562; MNHN-IU- 2016 - 1090 • 1 ♂, 1 ♀; Banc du Diamant; 14 ° 26.4 ʹ N, 61 ° 01.7 ʹ W; depth 18 m; 15 Sep. 2016; st. AB 177; MNHN-IU- 2016 - 1103 • 2 ♂♂, 1 ♀; Anse Couleuvre; 14 ° 50.4 ʹ N, 61 ° 13.4 ʹ W; depth 7 m; 1 Oct. 2016; st. AB 463; MNHN-IU- 2016 - 1119 • 1 ♂; Baie de Fort-de-France; 14 ° 36.1 ʹ N, 61 ° 04 ʹ W; depth 0 – 1 m; 29 Sep. 2016; st. AM 034; MNHN-IU- 2016 - 1150 • 1 juv.; Trois Rivières; 14 ° 27.5 ʹ N, 60 ° 58.2 ʹ W; depth 17 m; 9 Oct. 2016; st. AB 578; MNHN-IU- 2016 - 1169 • 1 ♂ ov., 2 ♀♀; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 00.1 ʹ W; depth 12 m; 26 Sep. 2016; st. AB 360; MNHN- IU- 2016 - 1219 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1400 • 1 ♂; Grand’Rivière; 14 ° 52 ʹ N, 61 ° 11.7 ʹ W; depth 0 – 6 m; 3 Oct. 2016; st. AS 557; MNHN-IU- 2016 - 1222 • 1 juv.; Anse des Galets; 14 ° 51.4 ʹ N, 61 ° 12.8 ʹ W; depth 0 – 10 m; 7 Oct. 2016; st. AB 567; MNHN-IU- 2016 - 1247 • 1 ♂, 1 ♂ ov., 1 juv.; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2016 - 1251 • 1 ♂; Baie de Sans-Souci; 14 ° 34.7 ʹ N, 60 ° 50.7 ʹ W; depth 0 – 1 m; 9 Oct. 2016; st. AM 325; MNHN-IU- 2016 - 1270 • 1 ♂; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.3 ʹ W; depth 4 – 10 m; 14 Sep. 2016; st. AB 123; MNHN-IU- 2016 - 1295.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0BFFA6FDAF05550079B8E4.taxon	discussion	Remarks The species is generally easily identified by the two characteristic dorsal clavate setae on the second trunk segment (Müller & Krapp 2009). It was recorded for the first time in Martinique by Müller (1990 a) at Vauclin, Petite Anse Macabou and Baie de Tartane, and was abundantly collected during Madibenthos from both the Atlantic and Caribbean coasts.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E0BFFA6FDAF05550079B8E4.taxon	distribution	Distribution Tropical Western Atlantic (Caribbean, Brazil) and East Pacific (Ecuador, Panama, California). Depth range 0 – 28 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E09FFA7FDE503FF0079BAA0.taxon	materials_examined	Type material Holotype: USNM 170494 (not examined). Type locality: San Francisquito Bay, South California. Material examined MARTINIQUE • 1 ♂; Pointe du Diamant; 14 ° 27.5 ʹ N, 61 ° 02.9 ʹ W; depth 3 m; 15 Sep. 2016; st. AB 126; MNHN-IU- 2016 - 812 / MK 411134.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E09FFA7FDE503FF0079BAA0.taxon	discussion	Remarks This species is regularly sampled in the Tropical West Atlantic, but is never abundantly collected. This is the first record of N. duodorsospinosa for Martinique, on the Caribbean coast.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E09FFA7FDE503FF0079BAA0.taxon	distribution	Distribution East Pacific (California, Galápagos, Mexico, Panama, Costa Rica), West Atlantic (US Georgia, Caribbean Sea, Gulf of Mexico). Depth range 0 – 72 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E09FFA7FDB3032602EEBECD.taxon	type_taxon	Type species Nymphopsis armatus Haswell, 1884, by original designation.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E09FFA7FD9D04130392B998.taxon	type_taxon	Type species Nymphon styligerum Miers, 1875, by monotypy.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E09FFA8FDC7052C03AFBB79.taxon	materials_examined	Type material Holotype: NL ZMA. PYC. P. 1486 (not examined). Type locality: Gustavia, Saint Barthelemy (East Caribbean). Material examined MARTINIQUE • 1 ♂; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 16 m; 18 Sep. 2016; st. AB 189; MNHN-IU- 2016 - 856 / MK 411169.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E09FFA8FDC7052C03AFBB79.taxon	discussion	Remarks Lucena et al. (2019) described a velvety region of the femur posteriorly to the cementary tube in this species, a character also observed in the single collected specimen from Martinique. The species was identified in Martinique by Müller (1990 a) in Petite Anse Macabou, and it is therefore still not known from the Caribbean coast of Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E09FFA8FDC7052C03AFBB79.taxon	distribution	Distribution Caribbean and Brazil. Depth range 0 – 15 m. We extend this depth range to 16 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E06FFA8FDB7064B0079B8BF.taxon	materials_examined	Type material Holotype: USNM 170494 (not examined). Type locality: Galeta Island reef, Caribbean Panama. Material examined MARTINIQUE • 1 ♂; Le Robert; 14 ° 41.1 ʹ N, 60 ° 51.4 ʹ W; depth 10 m; 19 Sep. 2016; st. AB 195; MNHN- IU- 2016 - 1407.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E06FFA8FDB7064B0079B8BF.taxon	discussion	Remarks This species has rarely been recorded, mostly from Panama or Belize. The species was previously recorded from West Indies from Martinique by Müller (1990 a) in Petite Anse Macabou. Therefore, the few records of Tanystylum birkelandi are all from the Atlantic coast. This species shows characteristic setae with microstetae on coxae 1 (Child 1979).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E06FFA8FDB7064B0079B8BF.taxon	distribution	Distribution Caribbean (Panama, Belize, Martinique). Depth range 0 – 15 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E07FFABFD910327006BB9AE.taxon	description	urn: lsid: zoobank. org: act: E 1915 AAA-F 3 A 8 - 496 C-AF 45 - 3 BD 9355 DEB 5 E Fig. 5 A – H nec Tanystylum hummelincki Stock, 1954 – Sabroux et al. 2019 b (pro parte): tab. 1, figs 3, 5.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E07FFABFD910327006BB9AE.taxon	materials_examined	Material examined Holotype MARTINIQUE • ♂ ov.; Baie du Robert; 14 ° 42 ʹ N, 60 ° 53.8 ʹ W; depth 2 m; 24 Sep. 2016; st. AB 452; MNHN-IU- 2016 - 1074 / MK 411045.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E07FFABFD910327006BB9AE.taxon	etymology	Etymology Latinized name, 2 nd decl. (masc.), genitive singular. The species is named after Philippe Bouchet, head of the Madibenthos Expedition.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E07FFABFD910327006BB9AE.taxon	description	Description (holotype, ♂, MNHN-IU- 2016 - 1074) BODY. Trunk completely unsegmented, discoidal; cuticle granular. No dorsomedian ornamentation. Ocular tubercle lower than abdomen, about as tall as base width, with two acute terminal tips, one large anteriorly and one smaller posteriorly, and 2 small lateral sense organs laterally, and carrying four large pigmented eyes. Cephalon with two low lateral tubercles at anterolateral margin. Low dorsal rising at base of abdomen. Lateral processes about as long as wide, jointed; 1 st lateral process with one submedian tubercle, 2 nd and 3 rd lateral processes with two dorsolateral tubercles on distal margin, 4 th lateral process with one anterior tubercle on distal margin. Lateral process tubercles carrying one short seta. PROBOScIS. Conical in dorsal view, rounded proximally and conical distally in lateral view, with blunt tip, about as long as two anterior trunk segments. ABDOMEN. Vertically oriented, higher than ocular tubercle, not reaching beyond lateral processes. Setae present on distal part. No basal segmentation. CHELIFORE. 1 - articled, composed of one elongated knob with two distal setae. PALP. 6 - articled, with setae on all articles except 1 st article. 1 st article shortest, wider than long. 2 nd article about 2.5 times as long as wide. 3 rd article shorter. 4 th article longest, about 2.5 times as long as 2 nd article. 5 th article about as long as wide, with ventral setae. 6 th article about 4 times as long as wide. OvIGER. 10 - articled with scarce setae. 1 st article as long as wide. 2 nd article about twice as long as wide. 3 rd article about 1.5 times as long as wide. 4 th article about twice as long as 3 rd. 5 th longest, curved. 6 th article 4 / 3 as long as wide. 7 th article about as long as wide, presenting one distal low spur carrying long setae. 8 th and 9 th articles subequal. 10 th article shortest. Strigilis spines simple on articles 7 th to 9 th, compound on 10 th article. Strigilis formula 2: 1: 1: 2. LEGS. Stout and nodulous, with scarce small setae. Coxa 1 shorter than wide, carrying 3 distal tubercles: anterodistal tubercle bifid, dorsomedian and distal tubercles simple, each tubercle with one short seta at tip. Coxa 2 longer than coxa 1 or 3. Coxa 3 as long as broad, with ventral setae. Femur stout, about twice as long as wide, with one elevated, dorsdistal cement gland tube. Tibia 1 and tibia 2 with large rounded tubercles and long setae on dorsal surface. Femur, tibia 1 and tibia 2 subequal. Tarsus trapezoid, short, about as long as wide, carrying one large spine and setae on ventral surface. Propodus stout and curved, with 3 large basal spines, and small spines on distal half of sole. Main claw curved, about ⅔ as long as propodus. Auxiliary claws present, about half as long as main claw. MEASUREMENTS (mm). Trunk 0.61; abdomen 0.20; proboscis 0.33; chelifore 0.12; coxa 1 0.12; coxa 2 0.26; coxa 3 0.10; femur 0.42; tibia 1 0.40; tibia 2 0.45; tarsus 0.05; propodus 0.35; main claw 0.21. Sexual dimorphism Female unknown.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E07FFABFD910327006BB9AE.taxon	discussion	Remarks The closest species to Tanystylum boucheti sp. nov. is T. isthmiacum Stock, 1955; we distinguish this species from the Stock’s species by its vertical abdomen, the presence of bifid tubercles at the dorsal margin of coxae 1 rather than rounded, and its more pronounced dorsal tubercles on the margin of the lateral processes (Müller & Krapp 2009). It may also be worth noting that the original description of T. isthmiacum from Panama mentioned 5 - articled palps, while many specimens (often regarded as belonging to the subspecies T. isthmiacum difficile Stock, 1966) are generally recorded with 6 - articled palps in the Atlantic (e. g., Fage & Stock 1966; Stock 1966 a, 1979; Müller & Krapp 2009). This may be a whole distinct species, morphologically closer to T. boucheti (although different regarding the criteria mentioned above). It is therefore possible that T. boucheti has been identified as T. isthmiacum or T. isthmiacum difficile in the past. Another similar species is Tanystylum hummelincki Stock, 1954 (Fig. 5 I – J), with which the present material was mixed in Sabroux et al. (2019 b). Indeed, the two species show similar appendages and lateral processes, and have bifid tubercles on the dorsal distal margin of the coxae. However, T. boucheti sp. nov. differs from T. hummelincki by the length of the proboscis, the height and shape of the ocular tubercle, and by the higher abdomen, which reaches beyond the ocular tubercle. The CO 1 p-distances between the T. boucheti specimen and the material in this study identified as T. hummelincki range from 0.104 to 0.106 (see Appendix). The species also strongly resembles Tanystylum evelinae Marcus, 1940, described and only recorded from Saõ Paulo State, Brazil (Marcus 1940). Tanystylum boucheti sp. nov. differs from T. evelinae by the presence of a posterior tubercle on the ocular tubercle, by the shape of the proboscis, which is conical rather than barrel-shaped, by the orientation of the abdomen, and by the size of the cement gland tube, which is longer in T. boucheti. This species can be differentiated from other Tanystylum relatives using the following combination of characters: bifid tubercles on the anterodistal margin of coxae 1, the presence of one or two dorsal tubercles on the lateral processes, the conical shape of the proboscis, and the shape of the ocular tubercle. A single specimen of the species was collected on the Atlantic coast of Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E07FFABFD910327006BB9AE.taxon	distribution	Distribution Only known from the type locality. Depth range 0 – 2 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E05FFACFDC2051B00F5B897.taxon	materials_examined	Type material Tanystylum duospinum Hilton, 1939. Holotype: USNM 81531 (not examined). Type locality: Baja California, Pacific Mexico. Tanystylum oculospinum Hilton, 1942. Holotype: USNM 81518 (not examined). Type locality: Todos Santos Bay, Baja California, Pacific Mexico. Tanystylum tubirostre Stock, 1954. Holotype: NL ZMA. PYC. P. 1520 (not examined). Type locality: Punt Vierkant, Bonaire. Tanystylum mexicanum Child, 1979. USNM 170650 (not examined). Type locality: Gulf of California, Pacific Mexico. Material examined MARTINIQUE • 1 ♀ gr.; Le Robert; 14 ° 41.1 ʹ N, 60 ° 51.4 ʹ W; depth 10 m; 19 Sep. 2016; st. AB 195; MNHN-IU- 2016 - 578 / MK 411118 • 1 juv.; Le Robert; 14 ° 43.2 ʹ N, 60 ° 50.6 ʹ W; depth 23 m; 25 Sep. 2016; st. AB 405; MNHN-IU- 2016 - 1049 / MK 411036 • 1 ♀; E of Le Robert; 14 ° 40.1 ʹ N, 60 ° 51.1 ʹ W; depth 0 – 16 m; 18 Sep. 2016; st. AS 255; MNHN-IU- 2016 - 1177 / MK 411071.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E05FFACFDC2051B00F5B897.taxon	discussion	Remarks We follow the suggestion of Bamber et al. (2022) in considering T. oculospinosum, T. tubirostrum and T. mexicanum as junior synonyms of T. duospinum. This is the first record of this species in Martinique. All records in the present material are from Atlantic coast off Le Robert.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E05FFACFDC2051B00F5B897.taxon	distribution	Distribution Circumtropical, mostly recorded in America with records in the East Pacific (California, Mexico, Panama, Galápagos, Ecuador, Peru), and West Atlantic (Bermuda, Gulf of Mexico, Caribbean). Other records from Kenya, Taiwan, Indonesia and Australia (Carpentaria Gulf). It is worth noting that the distribution range nominatively for T. duospinum is restricted to the East Pacific American coast, and that the widest distribution for the species has been recorded under the synonym T. oculospinosum (American Pacific coast from Mexico to Galápagos and Peru, and Australia). Tanystylum tubirostrum was nominatively recorded from Atlantic and Pacific American tropical waters, while T. mexicanum was nominatively recorded only from the Pacific Mexico. Depth range Intertidal to 54 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E03FFADFDCA032703A0BA72.taxon	description	Fig. 5 I – J	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E03FFADFDCA032703A0BA72.taxon	materials_examined	Type material Holotype: NL ZMA. PYC. P. 1485 (not examined). Type locality: La Pecha, Los Frailes Islands, Venezuela. Material examined MARTINIQUE • 1 ♀ gr.; Presqu’Île de la Caravelle; 14 ° 46.5 ʹ N, 60 ° 51.5 ʹ W; depth 15 m; 22 Sep. 2016; st. AB 350; MNHN-IU- 2016 - 868 / MK 411179 • 1 ♂; Ste-Marie; 14 ° 46.3 ʹ N, 60 ° 57.5 ʹ W; depth 20 m; 20 Sep. 2016; st. AB 260; MNHN-IU- 2016 - 1063 / MK 411041 • 1 ♂, 1 juv.; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2016 - 1248 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1398 / MK 411111 • 1 juv.; same collection data as for preceding; MNHN-IU- 2017 - 218 • 1 ♂; Canal de Ste Lucie; 14 ° 23.9 ʹ N, 60 ° 52.6 ʹ W; depth 0 – 2 m; 12 Sep. 2016; st. AM 012; MNHN-IU- 2016 - 1309.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E03FFADFDCA032703A0BA72.taxon	discussion	Remarks This is another rare species of Tanystylum collected for the first time in Martinique, though the most abundantly sampled here in this genus. Most specimens were collected on the Atlantic coast of Martinique, except MNHN-IU- 2016 - 1309, which was sampled near the southern cape of the Island. This is the first record of the species out of the Caribbean Sea stricto sensu.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E03FFADFDCA032703A0BA72.taxon	distribution	Distribution Caribbean (Venezuela, Colombia, Martinique). Depth range 0 – 3 m. We extend the known range to 23 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E03FF90FD90075E0048B928.taxon	description	urn: lsid: zoobank. org: act: 2 CBAF 7 F 7 - 5414 - 4617 - 80 C 9 - 809155 ADA 7 FE Fig. 6 Tanystylum sp. 3 – Sabroux et al. 2019 b: tab. 1, fig. 3.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E03FF90FD90075E0048B928.taxon	materials_examined	Material examined Holotype MARTINIQUE • ♂; Fond Boucher; 14 ° 39.3 ʹ N, 61 ° 09.6 ʹ W; depth 37 – 40 m; 1 Oct. 2016; st. AD 283; MNHN-IU- 2016 - 1055 / MK 411038. Paratype MARTINIQUE • 1 ♂; Macouba; 14 ° 55.1 ʹ N, 61 ° 09 ʹ W; depth 80 m; 24 Sep. 2016; st. AD 261; MNHN- IU- 2016 - 867 / MK 411178.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E03FF90FD90075E0048B928.taxon	etymology	Etymology Contraction of ‘ in grallis ’, in + plural accusative of the Latin gralla, - ae (1 st decl, fem.): ‘ on stilts’, referring to the long legs of this species, relatively to most instars of the genus.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E03FF90FD90075E0048B928.taxon	description	Description (holotype, ♂, MNHN-IU- 2016 - 1055) BODY. Trunk completely unsegmented, discoidal; cuticle granular. No dorsomedian ornamentation. Ocular tubercle cylindrical, higher than abdomen, about 4 / 3 as tall as wide, carrying four large pigmented eyes, one acute terminal spur anteriorly and 2 small lateral sense organs. Cephalon with 2 small lateral tubercles at anterior margin. 1 st lateral processes about 4 / 3 as long as wide, other lateral processes about as long as wide; all lateral processes jointed; 1 st lateral process without ornamentation, 2 nd, 3 rd and 4 th lateral processes with one anterior tubercle on distal margin, carrying one seta. PROBOScIS. Proximal half broad and rounded, distal half tubular, blunt at tip, bent ventrally, about as long as distance between cephalon distal margin and abdomen base. ABDOMEN. Long, diagonally oriented, inflated distally, with light inflation at base. Anus reaching beyond first coxae of 4 th legs. Abdomen carrying setae, mostly on distal inflated part. No basal segmentation. One dorsal tubercle in distal part, carrying setae. CHELIFORE. 1 - articled, composed of one small knob with one distal spine. PALP. 6 - articled, with setae mostly on last 2 distal articles. 1 st article shortest, wider than length. 2 nd article about 2.5 times as long as wide. 3 rd article as long as 1 st article. 4 th article longest, about 1.7 times as long as 2 nd article, about 3.5 times as long as wide, with cannula (gland?) on outer side. 5 th article about 0.2 times as long as wide. 6 th article about 3.5 times as long as wide. OvIGER. 10 - articled, with scarce setae. 1 st article as long as wide. 2 nd article more than 1.5 times as long as wide. 3 rd article about twice as long as wide. 4 th and 5 th articles subequal and longest, 4 th about 3.5 times as long as wide, 5 th about 4 times as long as wide. 6 th article about twice as long as wide. 7 th article about as long as wide, presenting one distal low spur carrying long setae. 8 th article about as long as wide. 9 th article about twice as long as wide. 10 th article about 1.5 times as long as wide. Strigilis spines simple, strigilis formula 2: 1: 1: 2. LEGS. Rather long for the genus, 1 st leg pair longer than others. Coxa 1 shorter than wide, with 4 tubercles mounted with seta on distal margin of 1 st and 2 nd legs: one dorsomedian, two dorsolateral and one ventrolateral on posterior side; or with 3 tubercles on 3 rd and 4 th legs: one dorsomedian, one dorsolateral and one ventrolateral on posterior side. Coxa 2 longer than coxa 1 or 3, with one large distal tubercle on anterior side and 2 or 3 distal tubercles on posterior side, all mounted with one seta at tip. Coxa 3 as long as broad, with ventral setae. Femur straight, with long dorsal spur at distal margin with one lateral cement gland tube on inner side, rounded at base and tubular distally. Tibia 1 shorter than femur, with two low dorsal knobs mounted by long setae and distal dorsomedian spur. Tibia 2 longer than tibia 1 or femur, with scarce setae, and 3 low knobs on dorsal surface mounted with long setae. Tarsus trapezoid, short, about as long as wide, carrying one large spine and setae on ventral surface. Propodus gently curved, with low heel carrying 2 large heel spines, plus one smaller spine more distally. Main claw curved, about half as long as propodus. Auxiliary claws present, about 0.6 times as long as main claw. MEASUREMENTS (mm). Trunk 0.84; abdomen 0.55; proboscis 0.80; coxa 1 0.22 (1 st leg), 0.24 (2 nd leg); coxa 2 0.46 (1 st leg), 0.36 (2 nd leg); coxa 3 0.27 (1 st leg), 0.28 (2 nd leg); femur 1.25 (1 st leg), 0.82 (2 nd leg); tibia 1 1.17 (1 st leg), 0.78 (2 nd leg); tibia 2 1.40 (1 st leg), 1.04 (2 nd leg); tarsus 0.10; propodus 0.50; main claw 0.28. Sexual dimorphism No female currently available. Individual variability On the other available specimen, the chelifores are almost not visible in dorsal view.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E03FF90FD90075E0048B928.taxon	discussion	Remarks Tanystylum ingrallis sp. nov. is readily discriminated from most other described species by the peculiarly long and slender legs and the absence of nodules on legs. Most species of Tanystylym have their propodus conspicuously longer than half the tibia 2, while here the propodus is just half as long as tibia 2 for the second, third and fourth legs, and much smaller for first legs. Only T. lamonti Staples, 2019, T. tiara Staples, 2019, T. zuydtorpi Arango, 2009 and T. paramexicanum Müller & Krapp, 2009 have long legs as well. Tanystylum lamonti, T. tiara and T. zuydtorpi are much different from T. ingrallis, due for example to the shape of their proboscis (long and barrel shape in the two former species, pyriform in the latter), and their longer chelifores. On the other hand, T. paramexicanum shares some morphological characters with this new species: the shape of the proboscis, the strong spur on the anterior margin of the ocular tubercle and the presence of 3 or 4 tubercles on coxae 1. However, the two species do not only differ by the absence of tubercles on the lateral processes, the much lower tubercles on coxae 1, the shorter size of coxae 2 and the bent ocular tubercle tip in T. paramexicanum, but also by the longer abdomen, the presence of tubercles on coxae 2 and the presence of a high tubercle on the distal margin of tibia 1 in T. ingrallis. Also, there is no specific mention of a longer first pair of legs in T. paramexicanum. Regrettably, only females are known for T. paramexicanum, so it is not possible to compare the cement gland opening of the two species. Indeed, the lateral opening of the cement gland tube in T. ingrallis is another strong diagnostic character. The morphologically closest species to T. ingrallis sp. nov. is probably T. duospinum Hilton, 1939, which presents a similar proboscis, and has the same palp structure. These two species are, however, differentiated by several conspicuous characters: (i) the abdomen of T. ingrallis extends beyond the first coxae of the fourth leg, while it does not reach the distal margin of the lateral processes in T. duospinum; (ii) in T. ingrallis, the abdomen is oriented diagonally, while it is almost vertical in T. duospinum; (iii) the anterior tubercle-ornamented coxa 1 is bifurcated in T. duospinum on the first leg (Müller & Krapp 2009), but not in T. ingrallis; (iv) the dorsodistal spur of the legs is longer in T. ingrallis compared to T. duospinum; (v) knobs dorsally positioned along the legs are smaller and more spaced in T. ingrallis compared to T. duospinum; (vi) tibia 2 is longer relative to tibia 1 in T. ingrallis compared to T. duospinum. Two specimens of the species were collected on the Caribbean and northern coasts of Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E03FF90FD90075E0048B928.taxon	distribution	Distribution Only known from Martinique. Depth range 40 – 80 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3EFF92FDBD049C0057BEBF.taxon	materials_examined	Type material Holotype: USNM 37781 (not examined, illustrated in Child 1992 b). Type locality: Long Island Sound, off Noank, Fishers Island Sound, New York, USA. Material examined MARTINIQUE • 1 ♂ ov.; Baie de Fort-de-France; 14 ° 36.1 ʹ N, 61 ° 04 ʹ W; depth 0 – 1 m; 29 Sep. 2016; st. AM 034; MNHN-IU- 2016 - 872 / MK 411183 • 1 ♂; same collection data as for preceding; MNHN- IU- 2016 - 873 / MK 411184 • 1 ♀; same collection data as for preceding; MNHN-IU- 2016 - 874 / MK 411185.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3EFF92FDBD049C0057BEBF.taxon	discussion	Remarks This species was misidentified as Tanystylum conirostre in Sabroux et al. (2019 b), as the examined specimens differed from the description of T. orbiculare given by Krapp (1973) by having slenderer legs and a straight abdomen. However, T. orbiculare can be distinguished from T. conirostre by the number of palp articles (5 rather than 4) and the proboscis shape, which is more barrel-shaped in T. orbiculare. Although this species is very common in the Atlantic, it was not yet recorded from Martinique. CO 1 p-distances to previously barcoded material (Arango & Wheeler 2007; Masta et al. 2010; respectively GenBank accession numbers DQ 390064 from Mar del Plata, Argentina, and GU 370064 from Massachusetts) are high (0.172 and 0.11 – 0.14, respectively; see Appendix). All specimens were collected at a single station in the Baie de Fort-de-France. It should be noted that this species is mostly amphi-Atlantic, but two records were provided from New South Wales by Stock (1954 b) and Clark (1963). However, Stock regarded T. orbiculare as synonymous to T. conirostre and explicitly stated that he found the material similar to the type material of Clotenia conirostris (= T. conirostre), so this record should be regarded as T. conirostre. In turn, it is possible that Clark’s material was also erroneously identified as T. orbiculare following Stock.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3EFF92FDBD049C0057BEBF.taxon	distribution	Distribution Widely recorded in the Atlantic (New England, Massachusetts, Virginia, South Carolina, Georgia, Brazil, Argentina, Senegal, Congo, Angola, Cap Verde); also found in the Caribbean, Gulf of Mexico, occidental and oriental Mediterranean Sea, as well as the Bosphorus. It was also recorded in New South Wales (Clark 1963), though this record is questionable considering the otherwise amphi-Atlantic distribution of this species. Depth range 0.5 – 58 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3CFF92FDA405660446B885.taxon	type_taxon	Type species Pallenopsis (Pallenopsis) fluminensis (Krøyer, 1844), by subsequent designation (Child 1998 a).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3CFF92FDE90031008CB994.taxon	materials_examined	Type material Holotype: SMF 1111 (not examined). Type locality: Arreçifes near Punta el Diamante, Santa Marta, Colombia. Material examined MARTINIQUE • 1 ♂; Pointe du Vauclin; 14 ° 33.6 ʹ N, 60 ° 49.7 ʹ W; depth 2 m; 12 Sep. 2016; st. AD 230; MNHN-IU- 2016 - 858 / MK 411171 • 3 ♂♂; Presqu’Île de la Caravelle; 14 ° 44.5 ʹ N, 60 ° 53.5 ʹ W; depth 5 – 12 m; 17 Sep. 2016; st. AD 241; MNHN-IU- 2016 - 1292 • 1 ♂; Ste-Luce; 14 ° 27.3 ʹ N, 60 ° 55.5 ʹ W; depth 15 m; 10 Sep. 2016; st. AB 062; MNHN-IU- 2016 - 1300 / MK 411097.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3CFF92FDE90031008CB994.taxon	discussion	Remarks This species was reported for the first time in Martinique (Petite Anse Macabou) by Müller (1990 a) as Tanystylum sp., and was formally described later by Müller & Krapp (2009) from Colombia. The characteristic tubercle near the abdomen base is quite variable in size, but it is generally lower in the studied material compared to Müller & Krapp’s drawing. This tubercle is the only conspicuous character differentiating T. tayronae from T. geminum Stock, 1954. A more diversified sampling of this species and DNA barcodes are needed to make sure the two species are not synonymous. All specimens were collected on the Atlantic and Caribbean coasts of Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3CFF92FDE90031008CB994.taxon	distribution	Distribution So far only known from the Caribbean (Colombia and Martinique). Depth range 0 – 2 m, extended to 15 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3DFF93FDDA0327005FBBC1.taxon	materials_examined	Type material Holotype: MP and EMB-IOC (see Lucena & Christoffersen 2018 a) (not examined). Type locality: between Ilha do Francês and Ilha da Armaçaõ, Florianópolis, Santa Catarina, Brazil. Material examined MARTINIQUE • 1 ♀; Le Lorrain; 14 ° 50.7 ʹ N, 61 ° 03.8 ʹ W; depth 11 – 14 m; 6 Oct. 2016; st. AS 565; MNHN-IU- 2016 - 814 / MK 411136.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3DFF93FDDA0327005FBBC1.taxon	discussion	Remarks This is the first record of this species in Martinique and in the Caribbean, which fills a gap in the distribution of this species, already recorded from the northwest Atlantic, Surinam and Brazil. The specimen was collected on the Atlantic coast.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3DFF93FDDA0327005FBBC1.taxon	distribution	Distribution West Atlantic (Brazil, Surinam, Caribbean and US Georgia). Depth range 14 – 102 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3DFF94FDC206F3005FBEE4.taxon	materials_examined	Type material Syntypes: USNM 76517 (examined). Type locality: off West Florida and Cuba, Gulf of Mexico. Material examined MARTINIQUE • 1 ♂; SW of Vétiver; 14 ° 37.5 ʹ N, 61 ° 08.3 ʹ W; depth 74 m; 2 Oct. 2016; st. AD 290; MNHN-IU- 2016 - 813 / MK 411135.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3DFF94FDC206F3005FBEE4.taxon	discussion	Remarks This is the first record of this species in Martinique. The single specimen was collected on the Caribbean coast. This species was already recorded in the Caribbean from Colombia, Cuba and Florida.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3DFF94FDC206F3005FBEE4.taxon	distribution	Distribution Tropical West Atlantic (Gulf of Mexico, Caribbean, French Guiana, Brazil). Depth range 15 – 600 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3AFF94FDD000CD0079B887.taxon	materials_examined	Type material Holotype: ZMH-A 0000719 (not examined). Type locality: Puerto Cabello, Venezuela. Material examined MARTINIQUE • 1 ♂ ov.; Baie de Fort-de-France; 14 ° 36.1 ʹ N, 61 ° 04 ʹ W; depth 0 – 1 m; 29 Sep. 2016; st. AM 034; MNHN-IU- 2016 - 853 / MK 411166 • 1 ♀ gr.; same collection data as for preceding; MNHN- IU- 2016 - 882 • 1 ♀ gr.; same collection data as for preceding; MNHN-IU- 2016 - 883 / MK 411192 • 5 spec.; same collection data as for preceding; MNHN-IU- 2016 - 574 • 4 ♂♂, 2 ♂♂ ov., 2 ♀♀, 1 ♀ gr., 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 1084 • 1 juv.; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2016 - 573 • 1 ♀; same collection data as for preceding; MNHN-IU- 2016 - 576 • 1 ♂; Le Prêcheur; 14 ° 47.2 ʹ N, 61 ° 13.1 ʹ W; depth 2 m; 3 Oct. 2016; st. AB 382; MNHN-IU- 2016 - 583.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3AFF94FDD000CD0079B887.taxon	discussion	Remarks Widely distributed in the Tropical West Atlantic, the species is here recorded for the first time in Martinique. Most of the specimens were collected at two stations, one off the Presqu’Île de la Caravalle (Atlantic coast) and the other at Baie de Fort-de-France; one record from Le Prêcheur (north Caribbean coast).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3AFF94FDD000CD0079B887.taxon	distribution	Distribution Tropical West Atlantic: Caribbean (Venezuela, Panama, Florida, Columbia, Barbados, Martinique, Bonaire, Curaçao) and Brazil (Pernambuco and Paraíba). Depth range 0 – 30 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3AFF94FDA40005023EBDF2.taxon	type_taxon	Type species Ascorhynchus abyssi Sars, 1877, by original designation.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3BFF95FDCD03270048BB3D.taxon	materials_examined	Type material Holotype: USNM 213519 (not examined). Type locality: off West Florida. Material examined MARTINIQUE • 1 ♀; E of Le Vauclin; 14 ° 34.2 ʹ N, 60 ° 44.8 ʹ W; depth 90 m; 14 Sep. 2016; st. AD 231; MNHN-IU- 2016 - 1199 • 1 ♀ gr.; same collection data as for preceding; MNHN-IU- 2016 - 1322.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3BFF95FDCD03270048BB3D.taxon	discussion	Remarks This is the first record of this species since its original description from central West Florida by Child (1992 b). The two specimens were sampled at the same station, off Le Vauclin (Atlantic coast). They are morphologically very similar to Child’s description. Unfortunately, CO 1 PCR amplifications were negative for the two DNA extracts.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3BFF95FDCD03270048BB3D.taxon	distribution	Distribution Only recorded from West Florida (Gulf of Mexico) and Martinique. Depth range 73 – 90 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3BFF98FDB6068F0048BD2B.taxon	description	urn: lsid: zoobank. org: act: AF 5026 C 5 - 94 E 5 - 4 B 01 - BBC 1 - 684199 D 91 CBE Fig. 7 Ascorhynchus sp. 5 – Sabroux et al. 2019 b: tab. 1, fig. 3.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3BFF98FDB6068F0048BD2B.taxon	materials_examined	Material examined Holotype MARTINIQUE • subadult; Le Robert; 14 ° 43.2 ʹ N, 60 ° 50.6 ʹ W; depth 23 m; 25 Sep. 2016; st. AB 405; MNHN-IU- 2016 - 1047 / MK 411035. Paratype MARTINIQUE • juv.; E of Le Robert; 14 ° 40.1 ʹ N, 60 ° 51.1 ʹ W; depth 14 m; 18 Sep. 2016; st. AB 191; MNHN-IU- 2016 - 1190 / MK 411077.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3BFF98FDB6068F0048BD2B.taxon	etymology	Etymology From Latin iguana, - ae, 1 st decl. (fem.), genitive plural. Named after the green iguanas of Fort SaintLouis, the sharp teeth of which are painfully remembered by the first author, resembling the shape of the dorsomedian trunk tubercles of this new species.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3BFF98FDB6068F0048BD2B.taxon	description	Description (holotype, MNHN-IU- 2016 - 1047) BODY. Trunk stout, smooth, completely segmented. Trunk segments 1 to 3 bearing tall dorsomedian pointing tubercles, slightly curved anteriorly. Ocular tubercle shorter than dorsomedian tubercles, about 2.5 times as tall as wide, with acute tip, bearing four pigmented eyes distally. Lateral margin of cephalon carrying pointy tubercle near ocular tubercle. Oviger base touching first lateral processes. Lateral processes about as long as wide, 4 th pair shortest, well separated by less than half their own diameter, distally bearing a dorsomedian pointing tubercle, without any seta or spine at tip. PROBOScIS. Very movable, pyriform, proximal part narrow, inflated before mid-length, tapering and blunt distally. Longer than cephalon and first trunk segment together. ABDOMEN. Horizontal, long, extending beyond coxa 2 of 4 th leg. Distal part inflated, carrying few setae. Abdomen segmented at base. CHELIFORE. 2 - articled, shorter than proboscis. Scape 1 - articled, carrying seta at mid-length and 3 additional setae at distal margin. Chela not functional, bearing one lateral seta. Finger oriented ventrally. PALP. 10 - articled, with setae mostly on ventral side of last 5 articles. 1 st article shorter than wide, carrying dorsal pointy tubercle. 2 nd article shortest, shorter than width. 3 rd article subequal to 5 th article and longer than other articles, with scarce spines. 4 th article slightly longer than wide. 5 th article with one long lateral seta on outer surface near proximal end, plus several small spines near distal end. 6 th article bent, as long as wide, with setae on ventral surface. 7 th and 8 th articles subequal, about 1.5 times as long as wide. 9 th and 10 th articles subequal, about as long as wide. OvIGER. 10 - articled, with scarce setae. 1 st article shortest, as long as wide. 2 nd article about twice as long as wide. 3 rd article 1.5 times as long as wide, glabrous. 4 th and 5 th articles subequal in length, 5 th article curved. 6 th and 7 th articles subequal. 6 th article twice as long as wide. 7 th article longest within strigilis, 8 th, 9 th and 10 th articles subequal. Strigilis with 2 rows of compound spines. Terminal claw short, with blunt tip. LEGS. Slender with scarce setae. Coxa 1 shorter than wide, dorsodistally with 2 tubercles, each surmounted with spine. Coxa 2 slightly longer than maximal width, with ventral seta. Coxa 3 as long as wide, with ventral setae. Femur and tibia 1 subequal in length. Tibia 2 longest, about ⅕ longer than tibia 1 or femur, with many long setae on dorsal surface and shorter setae on ventral surface. Tarsus mediumsized, trapezoid. Propodus straight, about 0.6 times as long as tibia 2, sole with 8 small spines. Main claw curved, about 0.4 times as long as propodus. No auxiliary claw. MEASUREMENTS (mm). Trunk 1.13; abdomen 0.39; proboscis 0.73; chelifore 0.21; coxa 1 0.14; coxa 2 0.29; coxa 3 0.18; femur 0.57; tibia 1 0.57; tibia 2 0.71; tarsus 0.09; propodus 0.38; main claw 0.15. Sexual dimorphism No female currently available. Individual variability The paratype is a juvenile, which presents developed chelae as expected, and is overall smaller in size.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3BFF98FDB6068F0048BD2B.taxon	discussion	Remarks The high, curved, conical dorsomedian tubercles of this species is a strong diagnostic character for this species. Only Ascorhynchus athernus Child, 1982 shows dorsomedian tubercles comparable in size and longer than the ocular tubercle, but it is otherwise very dissimilar to A. iguanarum sp. nov. This species is morphologically close to A. castelli (Dohrn, 1881), which can be found in the same region, from which it can be discriminated not only by the higher dorsomedian tubercles, but also by the larger tubercles on the lateral processes, the presence of small pointy tubercles on the 1 st article of the palp and at the base of palp insertion, the absence of a spur on the dorsodistal margin of tibiae 1 and the smaller distance between the lateral processes. The two specimens were collected on the Atlantic coast of Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E3BFF98FDB6068F0048BD2B.taxon	distribution	Distribution Only known from Martinique. Depth range 14 – 23 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E36FF98FDB600B40079B9E9.taxon	materials_examined	Type material Holotype: MCZ CRU- 6947 (not examined). Type locality: Sweeting’s Village, Great Abaco, Bahamas. Material examined MARTINIQUE • 1 ♀; Passe du Marin; 14 ° 27 ʹ N, 60 ° 53.8 ʹ W; depth 4 – 8 m; 9 Sep. 2016; st. AS 112; MNHN-IU- 2016 - 816 / MK 411137 • 1 ♂; Schoelcher; 14 ° 37.7 ʹ N, 61 ° 07.2 ʹ W; depth 4 m; 27 Sep. 2016; st. AB 460; MNHN-IU- 2016 - 861 / MK 411173 • 1 ♀; Pointe Baham; 14 ° 25 ʹ N, 60 ° 50.4 ʹ W; depth 0 – 2 m; 17 Sep. 2016; st. AM 021; MNHN-IU- 2016 - 862 / MK 411174 • 1 ♂ ov.; Schoelcher; 14 ° 37.7 ʹ N, 61 ° 07.2 ʹ W; depth 4 – 9 m; 27 Sep. 2016; st. AR 461; MNHN-IU- 2016 - 871 / MK 411182.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E36FF98FDB600B40079B9E9.taxon	discussion	Remarks This is the first record of this species in Martinique, where it was collected mostly on the Caribbean and southern coasts, although one specimen was found in Pointe Baham (southern Atlantic coast).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E36FF98FDB600B40079B9E9.taxon	distribution	Distribution Tropical, amphi-Atlantic (Gulf of Mexico, Caribbean, Senegal). Depth range 0 – 19 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E36FF99FD5904F20048BCFA.taxon	description	Fig. 8 Ascorhynchus sp. 4 – Sabroux et al. 2019 b: tab. 1, fig. 3.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E36FF99FD5904F20048BCFA.taxon	materials_examined	Material examined MARTINIQUE • 1 juv.; Macouba; 14 ° 55 ʹ N, 61 ° 08.9 ʹ W; depth 78 – 80 m; 24 Sep. 2016; st. AD 263; MNHN-IU- 2016 - 1039 / MK 411031.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E36FF99FD5904F20048BCFA.taxon	discussion	Remarks The only available specimen is a juvenile that is rather poorly preserved. The ovigers are broken. In these circumstances, it seems unwise to name this species, although it is probably new to science. The species differs from most other species of Ascorhynchus by the following character combination: ocular tubercle present, with pigmented eyes positioned below the ocular tubercle tip; slender dorsomedian tubercles on trunk segments; lateral processes less than twice as long as wide, separated by about their own diameter, carrying median tubercle; ocular tubercle near anterior cephalon boarder; chelifore scapes monosegmented. These characters are shared with A. okai Nakamura & Child, 1983 from Japan, A. cactoides Stock, 1954 from the Indo-Pacific (Stock 1991 b) and A. pudicus Stock, 1970 from the Eastern Atlantic and Mediterranean (Munilla and Soler-Membrives 2014). Ascorhynchus sp. 4 differs from A. okai by the longer distance between the insertions of the ovigers and palps (Nakamura & Child 1983), from A. cactoides in having the abdomen straight and from A. pudicus in having no dorsomedian tubercle on the 4 th trunk segment.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E36FF99FD5904F20048BCFA.taxon	distribution	The specimen was collected in Northern Martinique. Depth range 78 – 80 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E37FF99FDA101CD03D4BB32.taxon	type_taxon	Type species Zetes hispidus Krøyer, 1844 by monotypy.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E37FF9BFDA4068500C4BAC6.taxon	materials_examined	Type material Holotype: NHMD 110654 (not examined). Type locality: sound between St John and St James (Virgin Islands). Material examined MARTINIQUE • 1 ♂; Pointe du Diamant; 14 ° 27 ʹ N, 61 ° 04.1 ʹ W; depth 70 m; 9 Sep. 2016; st. AD 214; MNHN-IU- 2016 - 823 / MK 411142 • 1 ♂; Les Anses-d’Arlet; 14 ° 29.7 ʹ N, 61 ° 05.4 ʹ W; depth 19 m; 7 Sep. 2016; st. AB 155; MNHN-IU- 2016 - 824 / MK 411143 • 1 ♂; Canal de Ste Lucie; 14 ° 23.1 ʹ N, 60 ° 50.2 ʹ W; depth 65 m; 11 Sep. 2016; st. AD 224; MNHN-IU- 2016 - 825 / MK 411144 • 1 preadult ♀; same collection data as for preceding; MNHN-IU- 2016 - 579 • 1 ♂, 1 juv.; Canal de Ste Lucie; 14 ° 22.7 ʹ N, 60 ° 51.3 ʹ W; depth 65 m; 11 Sep. 2016; st. AD 223; MNHN-IU- 2016 - 545 • 7 ♂♂, 4 ♀♀, 1 ♀ gr., 2 juvs; Canal de Ste Lucie; 14 ° 22.7 ʹ N, 60 ° 51.6 ʹ W; depth 65 m; 11 Sep. 2016; st. AD 222; MNHN-IU- 2016 - 547 • 1 ♀; Grande Anse d’Arlets; 14 ° 29.9 ʹ N, 61 ° 05.4 ʹ W; depth 28 m; 7 Sep. 2016; st. AB 157; MNHN- IU- 2016 - 549 • 1 ♀; Vétiver; 14 ° 37.8 ʹ N, 61 ° 07.7 ʹ W; depth 25 m; 8 Sep. 2016; st. AB 161; MNHN- IU- 2016 - 550 • 6 ♂♂, 3 ♀♀; Les Anses-d’Arlet; 14 ° 28.1 ʹ N, 61 ° 05.3 ʹ W; depth 72 m; 8 Oct. 2016; st. AD 616; MNHN-IU- 2016 - 1041 • 6 ♂♂, 3 ♀♀, 4 juvs; Fond Boucher; 14 ° 39.3 ʹ N, 61 ° 09.6 ʹ W; depth 37 – 40 m; 1 Oct. 2016; st. AD 283; MNHN-IU- 2016 - 1053 • 5 juvs; same collection data as for preceding; MNHN-IU- 2016 - 1223 • 1 ♂, 1 ♀, 1 juv.; Baie de St-Pierre; 14 ° 45.1 ʹ N, 61 ° 11 ʹ W; depth 17 m; 4 Oct. 2016; st. AB 388; MNHN-IU- 2016 - 1057 • 2 ♀♀; Bellefontaine; 14 ° 40.2 ʹ N, 61 ° 10.5 ʹ W; depth 70 m; 2 Oct. 2016; st. AD 289; MNHN-IU- 2016 - 1112 • 1 ♂; SW Vétiver; 14 ° 37.6 ʹ N, 61 ° 08.3 ʹ W; depth 60 m; 30 Sep. 2016; st. AD 280; MNHN-IU- 2016 - 1134 • 1 ♂; Les Anses-d’Arlet; 14 ° 28.5 ʹ N, 61 ° 05.1 ʹ W; depth 29 m; 26 Sep. 2016; st. AD 271; MNHN-IU- 2016 - 1135 • 1 ♀ gr?; Le Carbet; 14 ° 42.3 ʹ N, 61 ° 11.7 ʹ W; depth 90 m; 3 Oct. 2016; st. AD 293; MNHN-IU- 2016 - 1146 • 1 juv.; Bellefontaine; 14 ° 40.2 ʹ N, 61 ° 10 ʹ W; depth 40 m; 2 Oct. 2016; st. AD 286; MNHN-IU- 2016 - 1192 • 1 ♀; Grande Anse d’Arlets; 14 ° 30.1 ʹ N, 61 ° 06.3 ʹ W; depth 71 m; 8 Oct. 2016; st. AD 617; MNHN-IU- 2016 - 1194 • 1 ♀; Le Prêcheur; 14 ° 49.1 ʹ N, 61 ° 13.8 ʹ W; depth 20 – 25 m; 8 Oct. 2016; st. AS 576; MNHN-IU- 2016 - 1198 • 1 ♀; Pointe du Diamant; 14 ° 26.9 ʹ N, 61 ° 04 ʹ W; depth 84 m; 8 Oct. 2016; st. AD 614; MNHN-IU- 2016 - 1212 • 1 juv.; Baie de Fortde-France; 14 ° 35.4 ʹ N, 61 ° 01.5 ʹ W; depth 8 m; 6 Oct. 2016; st. AD 602; MNHN-IU- 2016 - 1214 • 1 ♂, 3 ♀♀; Pointe de la Baleine; 14 ° 31.1 ʹ N, 61 ° 05.9 ʹ W; depth 17 – 19 m; 30 Sep. 2016; st. AB 369; MNHN- IU- 2016 - 1255 • 1 ♂; SW of Vétiver; 14 ° 37.6 ʹ N, 61 ° 08.6 ʹ W; depth 66 m; 7 Oct. 2016; st. AD 612; MNHN-IU- 2016 - 1289.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E37FF9BFDA4068500C4BAC6.taxon	discussion	Remarks A noticeable feature of this species compared to other representatives of Eurycyde is the strong bending of the abdomen about mid-length in a right angle. This feature is illustrated by Müller & Krapp (2009) and Stock (1955), although the bending seems to be only very slight in Child (1992 b). This is the first record of this species in Martinique. The species was abundantly sampled but only on the Caribbean coast. Records of E. clitellaria outside the Caribbean Sea are actually rare, with only one record of four specimens in Jalisco, Pacific Mexico (Child 1979) and one record of one specimen from North Carolina (McCloskey 1967).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E37FF9BFDA4068500C4BAC6.taxon	distribution	Distribution Caribbean, North Carolina, Pacific Mexico (Jalisco). Depth range Intertidal to 100 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E35FF9FFD7F07FA0079BCA4.taxon	description	urn: lsid: zoobank. org: act: 8 B 0 E 31 E 8 - 5 FB 3 - 47 DF- 8 BED-F 63 E 014 DA 6 B 7 Fig. 9 A – J nec Eurycyde raphiaster Loman, 1912 – Sabroux et al. 2019 a: fig. 2 d – f; 2019 b (pro parte): 1525, 1531, tab. 1, figs 3, 5.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E35FF9FFD7F07FA0079BCA4.taxon	materials_examined	Material examined Holotype MARTINIQUE • ♂; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.4 ʹ W; depth 14 m; 14 Sep. 2016; st. AB 175; MNHN-IU- 2016 - 1129 / MK 411058. Paratypes MARTINIQUE • 1 ♀; E of Le Vauclin; 14 ° 34.2 ʹ N, 60 ° 44.8 ʹ W; depth 90 m; 14 Sep. 2016; st. AD 231; MNHN-IU- 2016 - 821 • 1 ♀; same collection data as for preceding; MNHN-IU- 2016 - 1200 • 1 ♂; CasePilote; 14 ° 38.3 ʹ N, 61 ° 08.4 ʹ W; depth 12 m; 8 Sep. 2016; st. AB 159; MNHN-IU- 2016 - 820 • 2 ♂♂, 1 ♂ ov.; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.3 ʹ W; depth 19 – 21 m; 9 Sep. 2016; st. AS 057; MNHN- IU- 2016 - 540 • 1 ♂, 1 ♂ ov.; Pointe du Diamant; 14 ° 27.5 ʹ N, 61 ° 02.9 ʹ W; depth 3 m; 15 Sep. 2016; st. AB 126; MNHN-IU- 2016 - 542 • 1 ♂ ov., 1 ♀; Bellefontaine; 14 ° 39.7 ʹ N, 61 ° 09.6 ʹ W; depth 12 m; 30 Sep. 2016; st. AB 372; MNHN-IU- 2016 - 1032 • 1 ♂; Baie de Fort-de-France; 14 ° 34.3 ʹ N, 61 ° 02.7 ʹ W; depth 7 m; 7 Oct. 2016; st. AB 394; MNHN-IU- 2016 - 1045 / MK 411033 • 1 ♂; Fond Boucher; 14 ° 39.3 ʹ N, 61 ° 09.6 ʹ W; depth 37 – 40 m; 1 Oct. 2016; st. AD 283; MNHN-IU- 2016 - 1052 • 1 ♂, 2 ♀♀; 2 juvs; Baie de St-Pierre; 14 ° 45.1 ʹ N, 61 ° 11 ʹ W; depth 17 m; 4 Oct. 2016; st. AB 388; MNHN-IU- 2016 - 1056 • 2 ♂♂, 2 ♀♀; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.3 ʹ W; depth 4 – 10 m; 14 Sep. 2016; st. AB 123; MNHN-IU- 2016 - 1067 • 1 juv.; Anse Couleuvre; 14 ° 50.4 ʹ N, 61 ° 13.4 ʹ W; depth 7 m; 1 Oct. 2016; st. AB 463; MNHN-IU- 2016 - 1077 • 8 ♂♂, 3 ♀♀, 2 juvs; same collection data as for preceding; MNHN- IU- 2016 - 1121 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1145 / MK 411064 • 2 ♀♀; N of Presqu’Île de la Caravelle; 14 ° 52.9 ʹ N, 60 ° 50.6 ʹ W; depth 61 m; 19 Sep. 2016; st. AD 249; MNHN- IU- 2016 - 1082 • 1 ♂; Presqu’Île de la Caravelle; 14 ° 47.5 ʹ N, 60 ° 57.4 ʹ W; depth 14 m; 4 Oct. 2016; st. AB 562; MNHN-IU- 2016 - 1089 / MK 411046 • 4 ♂♂, 2 ♀♀; Le Lorrain; 14 ° 50.7 ʹ N, 61 ° 03.8 ʹ W; depth 11 – 14 m; 6 Oct. 2016; st. AS 565; MNHN-IU- 2016 - 1115 • 1 ♀; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2016 - 1137 / MK 411061 • 1 ♂; Fond Boucher; 14 ° 39.3 ʹ N, 61 ° 09.4 ʹ W; depth 0 – 14 m; 27 Sep. 2016; st. AS 363; MNHN-IU- 2016 - 1162 • 1 ♂, 1 ♂ ov.; Trois Rivières; 14 ° 27.5 ʹ N, 60 ° 58.2 ʹ W; depth 17 m; 9 Oct. 2016; st. AB 578; MNHN- IU- 2016 - 1166 • 2 ♂♂; Trois Rivières; 14 ° 27.7 ʹ N, 60 ° 57.8 ʹ W; depth 4 – 5 m; 25 Sep. 2016; st. AB 354; MNHN-IU- 2016 - 1170 • 1 ♂; Baie de Fort-de-France; 14 ° 33.2 ʹ N, 61 ° 01.6 ʹ W; depth 6 m; 6 Oct. 2016; st. AB 390; MNHN-IU- 2016 - 1178 / MK 411072 • 1 juv.; Le Robert; 14 ° 41.1 ʹ N, 60 ° 51.4 ʹ W; depth 10 m; 19 Sep. 2016; st. AB 195; MNHN-IU- 2016 - 1181 • 1 ♂; Le Vauclin; 14 ° 32 ʹ N, 60 ° 49 ʹ W; depth 1 m; 16 Sep. 2016; st. AB 130; MNHN-IU- 2016 - 1187 / MK 411075 • 2 ♀♀; Grande Anse du Diamant; 14 ° 27.9 ʹ N, 61 ° 01.4 ʹ W; depth 17 m; 26 Sep. 2016; st. AB 358; MNHN-IU- 2016 - 1209 • 1 ♀; Presqu’Île de la Caravelle; 14 ° 47.3 ʹ N, 60 ° 57.4 ʹ W; depth 19 m; 21 Sep. 2016; st. AB 301; MNHN-IU- 2016 - 1229 • 1 ♂, 1 juv.; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 00.1 ʹ W; depth 12 m; 26 Sep. 2016; st. AB 360; MNHN- IU- 2016 - 1237 • 2 ♀♀; Grande Anse du Diamant; 14 ° 27.9 ʹ N, 61 ° 01.2 ʹ W; depth 15 m; 15 Sep. 2016; st. AB 179; MNHN-IU- 2016 - 1241 • 1 ♀; Anse des Galets; 14 ° 51.4 ʹ N, 61 ° 12.8 ʹ W; depth 0 – 10 m; 7 Oct. 2016; st. AB 567; MNHN-IU- 2016 - 1244 / MK 411085 • 1 ♀; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 29 m; 18 Sep. 2016; st. AS 253; MNHN-IU- 2016 - 1261 / MK 411089 • 2 ♂♂, 3 ♀♀; N of Presqu’Île de la Caravelle; 14 ° 52.5 ʹ N, 60 ° 51.1 ʹ W; depth 58 m; 19 Sep. 2016; st. AD 248; MNHN- IU- 2016 - 1296 • 1 ♀; Rocher du Diamant; 14 ° 26.6 ʹ N, 61 ° 02.4 ʹ W; depth 22 m; 9 Sep. 2016; st. AB 163; MNHN-IU- 2019 - 3391 • 2 ♂♂; Rocher du Diamant; 14 ° 26.5 ʹ N, 61 ° 02.4 ʹ W; depth 24 m; 14 Sep. 2016; st. AB 173; MNHN-IU- 2019 - 3392.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E35FF9FFD7F07FA0079BCA4.taxon	etymology	Etymology Apposition of the Kalinago (Caribbean) word ‘ kaiouti ’, ‘ which has spines’, in reference to the long tubercles of coxae 1.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E35FF9FFD7F07FA0079BCA4.taxon	description	Description (holotype, ♂, MNHN-IU- 2016 - 1129) BODY. Trunk granulous, completely segmented with posterior margin flaring. No dorsomedian ornamentation. Ocular tubercle tall, carrying four pigmented eyes and seven thick setae around ocular tubercle tip. Oviger base almost touching lateral processes. Lateral processes about 1.5 times as long as wide, 4 th pair shortest, well separated by less than half their own diameter, distally bearing a dorsomedially pointing tubercle, without seta or spine at tip. PROBOScIS. Bipartite, anterior part as petiole, posterior part pyriform, about as long as body, anterior and posterior parts articulated together. ABDOMEN. Long, extending beyond coxa 1 of 4 th leg. Distal part inflated, carrying long thick setae. Base articled with trunk, movable, tip conical. CHELIFORE. 3 - articled, reaching beyond proximal petiole of proboscis. Scape 2 - articled, 1 st article longest, with distal dorsomedian thick seta. 2 nd article carrying many thick setae dorsally. Chela residual, with non-functional finger. PALP. 10 - articled, with setae mostly on ventral side of last five articles. 1 st article about as long as wide, carrying one dorsal pointy tubercle. 2 nd article shortest, shorter than wide. 3 rd article longest, with distal thick setae. 4 th article slightly longer than wide. 5 th article 0.7 times as long as 3 nd, carrying several thick setae. 6 th article about twice as long as wide, 6 th and 10 th articles subequal in length, shorter than 7 th. 7 th, 8 th and 9 tharticles subequal in length. OvIGER. 10 - articled, with scarce setae. 1 st article shortest, as short as wide. 2 nd and 3 rd articles subequal. 4 th and 5 th articles subequal in length, 5 th article curved. 6 th articles 3 times as long as broad, longer than 7 th article. 7 th article longest within strigilis. 8 th, 9 th and 10 th articles subequal. Compound spines of strigilis in 2 rows. Terminal claw shorter than 10 th article. LEGS. Slender with many thick setae. Coxa 1 shortest, about as long as wide, with 2 long dorsolateral pointy tubercles on distal margin, anterior one about as long as 1 st coxa, posterior one at least twice as long as anterior one and carrying long thick seta at base for 1 st to 3 rd legs. Coxa 2 slightly longer than broad. Coxa 3 as long as wide. Femur with cement gland tube at about mid-length on anterior side and many thick setae distally. Tibia 1 13 times as long as wide, about 1.3 times as long as femur, thinner, with many dorsal thick setae at about mid-length. Tibia 2 subequal to tibia 1. Tarsus medium-sized, trapezoid. Propodus gently curved, about half as long as tibia 2; sole carrying many small spines. Main claw curved, about ⅓ as long as propodus. No auxiliary claw. MEASUREMENTS (mm). Trunk 1.05; abdomen 0.55; proboscis 1.42; chelifore 1.05; coxa 1 0.23; coxa 2 0.27; coxa 3 0.16; femur 0.72; tibia 1 0.92; tibia 2 0.90; tarsus 0.10; propodus 0.48; main claw 0.16. Sexual dimorphism Females with shorter oviger articles; anterior dorsolateral pointy tubercle of coxa 1 often shorter than in males. Individual variability Size of pointy dorsolateral tubercles of coxa 1 variable among specimens, posterior one always longer (or rarely as long) than length of coxa 1, anterior one always shorter, posterior tubercle always conspicuously longer than anterior tubercle. In holotype specimen, posterior dorsolateral pointy tubercle of coxa 1 curved anteriorly, but may curve posteriorly in other specimens.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E35FF9FFD7F07FA0079BCA4.taxon	discussion	Remarks Eurycyde kaiouti sp. nov. can be discriminated from most other species of Eurycyde by the following combination of characters: (i) the ocular tubercle presents long thick setae (the number of these setae may vary with loss and we do not recommend using it as a criterion); (ii) the coxa 1 carries dorsaly a long thick seta on the first, second and third legs, as well as two laterodistal tubercles on all four legs; (iii) lateral processes bear one pointy tubercle dorsally. These characters are only shared with E. raphiaster Loman, 1912, to which it is very close morphologically. Comparing with the illustration in Bouvier (1917) of the type material, the two tubercles of coxae 1 in E. raphiaster are subequal in length and less long than coxa 1 (shorter in females). Instead, E. kaiouti presents a posterior coxa 1 tubercle that is much longer than the anterior one: the anterior tubercle is shorter than coxa 1, the posterior one longer, or rarely as long. This difference between E. raphiaster and E. kaiouti is recovered also in juveniles. Furthermore, we observe that, in Martinique material, the tibia 1 in E. kaiouti is shorter than in E. raphiaster, e. g., 13 times as long as wide for the E. kaiouti holotype against 15 times for E. raphiaster specimen MNHN-IU- 2016 - 818. The two species can be distinguished based on CO 1 barcodes (E. kaiouti intraspecific p-distance = 0 – 0.013; interspecific p-distance with E. raphiaster = 0.173 – 0.187; see Appendix). One specimen recorded in Sabroux et al. (2019 b), MNHN-IU- 2016 - 819, is morphologically very similar to the holotype of Eurycyde kaiouti sp. nov., but falls in a barcode cluster on its own that is very divergent from the E. kaiouti cluster (p-distance = 0.182 – 0.188, see Appendix). Because it is the only specimen showing such genetic divergence, and since morphological differences are small, we cannot discriminate it as another species relying on morphological characters. For this reason, we choose to keep this specimen as Eurycyde sp. Further material from NWTA Eurycyde should be barcoded and studied to resolve the status of this specimen.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E35FF9FFD7F07FA0079BCA4.taxon	distribution	Eurycyde kaiouti was widely collected on both Atlantic and Caribbean coasts. Distribution Only known from Martinique. Depth range 1 – 90 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E31FF80FDB006180079BBC4.taxon	description	Fig. 9 K	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E31FF80FDB006180079BBC4.taxon	materials_examined	Type material Holotype: MOM INV- 0001360 (not examined). Type locality: 4 miles southwest of Boa Vista, Cap Verde. Material examined MARTINIQUE • 1 ♂; Passe du Marin; 14 ° 26.8 ʹ N, 60 ° 54.3 ʹ W; depth 15 m; 9 Sep. 2016; st. AB 165; MNHN-IU- 2016 - 818 / MK 411139 • 1 juv.; Grande Anse d’Arlets; 14 ° 29.9 ʹ N, 61 ° 05.4 ʹ W; depth 28 m; 7 Sep. 2016; st. AB 157; MNHN-IU- 2016 - 822 / MK 411141 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 541 / MK 411114 • 6 ♂♂; same collection data as for preceding; MNHN-IU- 2016 - 544 • 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 548 • 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 1282 • 1 ♂; Ste-Luce; 14 ° 27.3 ʹ N, 60 ° 55.5 ʹ W; depth 15 m; 10 Sep. 2016; st. AB 062; MNHN-IU- 2016 - 543 • 1 ♂ ov., 1 ♀; Rocher du Diamant; 14 ° 26.6 ʹ N, 61 ° 02.4 ʹ W; depth 22 m; 9 Sep. 2016; st. AB 163; MNHN-IU- 2016 - 546 • 2 juvs; N Le Lorrain; 14 ° 55.1 ʹ N, 61 ° 02.4 ʹ W; depth 63 m; 21 Sep. 2016; st. AD 257; MNHN-IU- 2016 - 1133 • 2 ♀♀; Rocher du Diamant; 14 ° 26.5 ʹ N, 61 ° 02.4 ʹ W; depth 24 m; 14 Sep. 2016; st. AB 173; MNHN-IU- 2016 - 1156 • 1 juv.; Vétiver; 14 ° 37.8 ʹ N, 61 ° 07.7 ʹ W; depth 25 m; 8 Sep. 2016; st. AB 161; MNHN-IU- 2016 - 1189 / MK 411076 • 2 ♂♂, 1 juv.; Pointe de la Baleine; 14 ° 31.1 ʹ N, 61 ° 05.9 ʹ W; depth 17 – 19 m; 30 Sep. 2016; st. AB 369; MNHN- IU- 2016 - 1256 • 2 ♀♀; Le Prêcheur; 14 ° 47.2 ʹ N, 61 ° 13.1 ʹ W; depth 10 – 17 m; 3 Oct. 2016; st. AR 383; MNHN-IU- 2016 - 1305.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E31FF80FDB006180079BBC4.taxon	discussion	Remarks Unlike Müller & Krapp (2009), we have never observed an absence of the lateral process tubercle in females. Eurycyde raphiaster and E. kaiouti sp. nov. are very similar species that may have been mistaken in the past. After revising literature illustrations, we agree with the determinations of Müller & Krapp (2009) (at least the illustrated male), while Hedgpeth (1948) and Child (1992 b) may have illustrated E. kaiouti rather than E. raphiaster. Although the species is abundant in the Caribbean, this is its first record in Martinique. Our data suggest that it is less common in Martinique than E. clitellaria or E. kaiouti. The species was most often collected on the Caribbean coast, though a few specimens were sampled off Le Lorain and Presqu’Île de la Caravelle on the Atlantic coast.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E31FF80FDB006180079BBC4.taxon	distribution	Distribution Bahamas, Gulf of Mexico, Caribbean, Cap Verde, Guyana and Senegal. Depth range 0 – 91 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2EFF80FDA0073602E7BAFD.taxon	type_taxon	Type species Pallene brevirostris Johnston, 1837, by original designation.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2EFF82FDDF07F60048BEE9.taxon	materials_examined	Type material Pallene brevirostris Johnston, 1837. Type (s): unknown. Type locality: Berwickshire, southeast Scotland. Pallene empusa Wilson, 1878. Syntypes: USNM 37777 (not examined). Type locality: Vineyard Sound or Long Island Sound, or Noank (Connecticut, USA). Material examined MARTINIQUE • 1 ♂; Ste-Luce; 14 ° 27.3 ʹ N, 60 ° 55.5 ʹ W; depth 15 m; 10 Sep. 2016; st. AB 062; MNHN- IU- 2016 - 842 / MK 411156 • 1 ♀; Pointe du Vauclin; 14 ° 33.6 ʹ N, 60 ° 49.7 ʹ W; depth 2 m; 12 Sep. 2016; st. AD 230; MNHN-IU- 2016 - 843 / MK 411157 • 1 ♂ ov.; Baie du Robert; 14 ° 42 ʹ N, 60 ° 53.8 ʹ W; depth 2 m; 24 Sep. 2016; st. AB 452; MNHN-IU- 2016 - 869 / MK 411180 • 1 ♂ carrying larvae; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.3 ʹ W; depth 4 – 10 m; 14 Sep. 2016; st. AB 123; MNHN-IU- 2016 - 1294 • 1 ind. (sex indet.); Pointe du Diamant; 14 ° 27.5 ʹ N, 61 ° 02.9 ʹ W; depth 3 m; 15 Sep. 2016; st. AB 126; MNHN- IU- 2016 - 1312.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2EFF82FDDF07F60048BEE9.taxon	discussion	Remarks This very common species is recorded for the first time in Martinique, though in low numbers (five specimens). The species was found on both the Atlantic and Caribbean coasts.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2EFF82FDDF07F60048BEE9.taxon	distribution	Distribution Widespread in both sides of the Atlantic: North Sea (Norway, Denmark, Scotland, England and the Netherlands), West Scotland, Irish Sea, West Ireland, Scilly Islands, English Channel, Gulf of Biscay, Portugal, Mediterranean, Black Sea, Bay of Fundy, Massachusetts, Virginia, West Florida, Caribbean, French Guiana). Also recorded from Pakistan, Thailand and Indonesia. Depth range 0 – 316 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2CFF82FDDA03C3009ABA4A.taxon	materials_examined	Type material Holotype: SMF 1497 (not examined). Type locality: Bahía Cinto in Tayrona National Park, Colombia. Material examined MARTINIQUE • 1 ♀ gr.; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 16 m; 18 Sep. 2016; st. AB 189; MNHN-IU- 2016 - 880 / MK 411190.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2CFF82FDDA03C3009ABA4A.taxon	discussion	Remarks This is the first record of this species since its original description from Caribbean Colombia by Müller & Krapp (2009). A single specimen was collected off the Presqu’Île de la Caravelle, on the Atlantic coast.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2CFF82FDDA03C3009ABA4A.taxon	distribution	Distribution Caribbean Colombia and Martinique. Depth range 1 – 3.5 m, extended to 16 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2CFF83FDBA0765005FBDF4.taxon	materials_examined	Type material Callipallene brevirostris longicoxa Stock, 1955. Holotype: NHMD 110621 (not examined). Type locality: off Calf and Cow, St James Bay, St Thomas (Virgin Islands). Callipallene belizae Child, 1982. Holotype: USNM 171035 (not examined). Type locality: Carrie Bow Cay, Belize. Material examined MARTINIQUE • 1 ♂ carrying larvae; Grande Anse d’Arlets; 14 ° 29.9 ʹ N, 61 ° 05.4 ʹ W; depth 28 m; 7 Sep. 2016; st. AB 157; MNHN-IU- 2016 - 841 / MK 411155 • 1 ♂, 1 ♂ ov., 2 ♀♀, 2 juvs; same collection data as for preceding; MNHN-IU- 2016 - 1277.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2CFF83FDBA0765005FBDF4.taxon	discussion	Remarks This is the first record of this species in Martinique. All specimens were found at the same station near Grande Anse d’Arlets. Comparatively to other specimens, collected between 10 and 30 meters, the single specimen collected by Stock (1986) was found strikingly deep into the Strait of Florida (512 – 549 m).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2CFF83FDBA0765005FBDF4.taxon	distribution	Distribution Caribbean and Strait of Florida. Depth range 10 – 512 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2DFF83FD9C00C70206BCB8.taxon	type_taxon	Type species Pallenoides magnicollis Stock, 1951, by original designation.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2DFF83FD9C00C70206BCB8.taxon	description	It is worth noting for this genus that specific epithets should be accorded to masculine, following the ICZN code recommandations for genera ending in - oides.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2DFF85FDD1060C0079BD75.taxon	description	Fig. 10	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2DFF85FDD1060C0079BD75.taxon	materials_examined	Type material Pallenoides amazonicus Stock, 1975. Holotype: NL ZMA. PYC. P. 1356 (not examined). Type locality: off Pará State, Brazil. Material examined MARTINIQUE • 1 ♀; SW of Vétiver; 14 ° 37.6 ʹ N, 61 ° 08.6 ʹ W; depth 66 m; 7 Oct. 2016; st. AD 612; MNHN-IU- 2016 - 866 / MK 411177.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2DFF85FDD1060C0079BD75.taxon	discussion	Remarks This unique specimen is very close to Stock (1975 a) ’ s original description; however, we note a few discripencies that make this identification unsure, i. e., the shorter auxillary claws of the Martinique specimen (about half as long as in the holotype) and its conspicuous pointy tubercle on the dorsal surface of the chelifores. In the absence of a CO 1 barcode sequence for P. amazonicus, we refrain from describing this species as new. A single specimen was collected on the Caribbean coast, off Vévier.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2DFF85FDD1060C0079BD75.taxon	distribution	Distribution Pallenoides amazonicus is known from Brazil, with one juvenile that has been considered as a possible P. amazonicus collected in South Africa (Arnaud & Child 1988), though this record seems very doubtful. If confirmed, the present record would be the first one for the Caribbean.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2DFF85FDD1060C0079BD75.taxon	description	Depth range Pallenoides amazonicus is known to dwell between 17 and 136 m. The present specimen was sampled at 66 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2BFF85FDBB005F0057B859.taxon	materials_examined	Type material Holotype: NHMD 110703 (not examined). Type locality: between St Jan and St Thomas, Virgin Islands. Material examined MARTINIQUE • 1 juv.; Canal de Ste Lucie; 14 ° 22.6 ʹ N, 60 ° 52.7 ʹ W; depth 21 – 30 m; 11 Sep. 2016; st. AR 064; MNHN-IU- 2016 - 846 / MK 411159 • 1 juv.; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.3 ʹ W; depth 19 – 21 m; 9 Sep. 2016; st. AS 057; MNHN-IU- 2016 - 847 / MK 411160 • 1 ♂ carrying larvae; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.3 ʹ W; depth 4 – 10 m; 14 Sep. 2016; st. AB 123; MNHN-IU- 2016 - 848 / MK 411161 • 1 ♀ gr.; Pointe du Diamant; 14 ° 27.5 ʹ N, 61 ° 02.9 ʹ W; depth 3 m; 15 Sep. 2016; st. AB 126; MNHN-IU- 2016 - 551 • 2 ♂♂, 1 ♂ carrying larvae; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.2 ʹ W; depth 22 m; 22 Sep. 2016; st. AS 096; MNHN-IU- 2016 - 1044 • 1 preadult ♀; Baie de St-Pierre; 14 ° 45.1 ʹ N, 61 ° 11 ʹ W; depth 17 m; 4 Oct. 2016; st. AB 388; MNHN-IU- 2016 - 1061 • 2 juvs; Le Robert; 14 ° 41.1 ʹ N, 60 ° 51.4 ʹ W; depth 10 m; 19 Sep. 2016; st. AB 195; MNHN-IU- 2016 - 1405.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2BFF85FDBB005F0057B859.taxon	discussion	Remarks This is the first record of this species in Martinique. Specimens were mostly collected on the Caribbean coast, except for two specimens found off Le Robert (Atlantic Coast).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2BFF85FDBB005F0057B859.taxon	distribution	Distribution Caribbean, Yucatán Channel. Depth range 1.5 – 37 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E2BFF85FDBB05630205B888.taxon	type_taxon	Type species Pallene australiensis Hoek, 1881, by original designation	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E28FF86FDAB077A05E1BA81.taxon	type_taxon	Type species Nymphon grossipes O. Frabricius, 1780, by subsequent designation (Child 1998 a).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E28FF86FDD303270026BB91.taxon	materials_examined	Type material Holotype: NHMUK ZOO- 1957.11.6.2 (not examined). Type locality: off Castle Roads, Bermuda. Material examined MARTINIQUE • 1 juv.; Pointe de la Baleine; 14 ° 31.1 ʹ N, 61 ° 05.9 ʹ W; depth 24 m; 30 Sep. 2016; st. AR 413; MNHN-IU- 2016 - 1221 / MK 411083.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E28FF86FDD303270026BB91.taxon	discussion	Remarks Despite the specimen being a juvenile, it can easily be identified as Parapallene bermudensis thanks to the characteristic pectinated spines on tibia 2 and the ventrodistal extremity of the tarsus, and on the propodal sole (Child 1982 a). This is the first record of this species in Martinique, at Pointe Baleine (Caribbean coast).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E28FF86FDD303270026BB91.taxon	distribution	Distribution Caribbean (Belize, Martinique), Bermuda, Gulf of Mexico (off West Florida). Depth range 16.8 – 54 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E28FF87FDA5040B0057BA6A.taxon	materials_examined	Type material Holotype: NL ZMA. PYC. P. 1267 (not examined). Type locality: Dickinson Bay, Antigua. Material examined MARTINIQUE • 2 ♀♀; Le François; 14 ° 37 ʹ N, 60 ° 49.5 ʹ W; depth 1 m; 17 Sep. 2016; st. AB 134; MNHN-IU- 2016 - 1234 • 1 ♀ gr., 1 ind. (sex indet.); same collection data as for preceding; MNHN- IU- 2016 - 1234 • 1 ♀ gr.; Pointe du Vauclin; 14 ° 33.6 ʹ N, 60 ° 49.7 ʹ W; depth 2 m; 12 Sep. 2016; st. AD 230; MNHN-IU- 2016 - 845 • 1 ♀; Îlet Chevalier; 14 ° 25.9 ʹ N, 60 ° 49.5 ʹ W; depth 0 – 2 m; 19 Sep. 2016; st. AM 023; MNHN-IU- 2016 - 1034 • 1 ♀; Pointe Michel; 14 ° 26.4 ʹ N, 60 ° 49.3 ʹ W; depth 2 m; 12 Sep. 2016; st. AB 120; MNHN-IU- 2016 - 1038 • 1 ♂; E of Le Vauclin; 14 ° 34.7 ʹ N, 60 ° 45.9 ʹ W; depth 20 m; 16 Sep. 2016; st. AS 078; MNHN-IU- 2016 - 1186 / MK 411074 • 1 ♀; Baie du Robert; 14 ° 42 ʹ N, 60 ° 53.8 ʹ W; depth 1 – 3 m; 24 Sep. 2016; st. AR 453; MNHN-IU- 2016 - 1213 / MK 411082 • 1 ♀; Le Robert; 14 ° 41.1 ʹ N, 60 ° 51.4 ʹ W; depth 10 m; 19 Sep. 2016; st. AB 195; MNHN-IU- 2016 - 1318 • 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 1320 • 1 juv.; Pointe de la Baleine; 14 ° 31.1 ʹ N, 61 ° 05.9 ʹ W; depth 24 m; 30 Sep. 2016; st. AR 413; MNHN-IU- 2016 - 1221.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E28FF87FDA5040B0057BA6A.taxon	discussion	Remarks Like all Nymphon but one found in Madibenthos material (i. e., Nymphon timons sp. nov.), this species is part of the aequidigitatum group as defined by Child (1988 b) and for which Takahashi et al. (2012) provided a complete identification key. Species of this group share the following five characters: (i) a long fifth palp article relative to the other four; (ii) an oviger claw without denticulation, and most often (iii) bifurcate chela teeth, (iv) propodus main claw as long as or shorter than auxiliaries, and (v) one or all propodal claws bearing endal rugosities or setules. This is the first record of this species for Martinique. It was only collected on the Atlantic coast, where it was found to be rather common.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E28FF87FDA5040B0057BA6A.taxon	distribution	Distribution Tropical West Atlantic (Gulf of Mexico, Caribbean, Surinam). Depth range 0.5 – 37 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E29FF8AFD75075603C0BA25.taxon	description	urn: lsid: zoobank. org: act: 8 B 1908 A 6 - 04 E 4 - 4 B 2 E-B 142 - 8 E 6 E 9 F 0990 AD Fig. 11	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E29FF8AFD75075603C0BA25.taxon	materials_examined	Material examined Holotype MARTINIQUE • ♂; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 16 m; 18 Sep. 2016; st. AB 189; MNHN-IU- 2016 - 879 / MK 411189. Paratypes MARTINIQUE • 1 ♂; same collection data as for holotype; MNHN-IU- 2016 - 870 / MK 411181 • 1 ♂; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN- IU- 2016 - 1298 • 1 ♂; Le François; 14 ° 38.5 ʹ N, 60 ° 46.3 ʹ W; depth 63 m; 15 Sep. 2016; st. AD 234; MNHN-IU- 2016 - 1313 • 1 juv.; Pointe de la Baleine; 14 ° 31.1 ʹ N, 61 ° 05.9 ʹ W; depth 24 m; 30 Sep. 2016; st. AR 413; MNHN-IU- 2016 - 1221.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E29FF8AFD75075603C0BA25.taxon	etymology	Etymology Noun in apposition. Named after the Dorlis, also called ‘ night husband’, a malevolent, invisible character of Martinique folklore. It fits somehow with the aspect of this species, with horn-like lateral sense organs on the ocular tubercle and large setose chelae; and only males were found.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E29FF8AFD75075603C0BA25.taxon	description	Description (holotype, ♂, MNHN-IU- 2016 - 879) BODY. Species rather small and stout, trunk completely segmented, cuticle smooth. No dorsomedian tubercle. Ocular tubercle rounded, as tall as wide, positioned anteriorly to 1 st lateral processes, with 2 conspicuous pointy lateral sense organs beside tip and four pigmented eyes. Preocular neck short, shorter than wide. Oviger base jointing 1 st lateral process. Lateral processes without ornamentation, well separated by less than their own diameter. PROBOScIS. Rounded and large, reaching distal margin of chelifore scape in lateral view. ABDOMEN. Medium-sized, not reaching beyond lateral processes, carrying several setae at about midlength, directed diagonaly. No basal segmentation. CHELIFORE. 2 - articled, reaching far beyond proboscis. Scape 1 - articled, 4.5 times as long as wide, carrying setae. Chela very large, palm longer than fingers ornamented with setae. Fingers sub-equal in length, bearing simple and bifurcated teeth, 10 bifurcated and 3 simple teeth on both fingers. PALP. 5 - articled. 1 st article shortest, shorter than wide. 2 nd article longest, about 7 times as long as wide, with distal setae. 3 rd article about 0.8 times as long as 2 nd, carrying ventrodistal setae. 4 th article about 0.6 times as long as 5 th, about 3 times as long as wide, with many long ventral setae along its length. 5 th article 0.8 times as long as 3 rd, carrying many ventral setae. OvIGER. 10 - articled. 1 st article shorter than wide. 2 nd article about 1.5 times as long as wide. 3 rd article about twice as long as wide. 4 th article about 4 times as long as wide, with a few distal setae. 5 th article longest, about twice as long as 4 th, straight, strongly swollen distally, with light constriction at base. 6 th article about ⅔ as long as 4 th, with many setae along its length. 7 th and 10 th articles subequal, about ¾ of 6 th article length. Strigilis formula 12: 10: 9: 10. Strigilis spines compound, carrying many lateral teeth, proximalmost shortest and spatulated to oblong, median spines lancelolate, distalmost spines spatulated to oblong, longer than proximalmost. Intermediary position spines roughly intermediary in shape. Distalmost spines of 9 th and 10 th articles longest among strigilis articles. Terminal claw about ⅔ length of 10 th article. LEGS. Slender, setose. Coxa 1 no longer than broad. Coxa 2 about 3 times as long as maximal width, longer than coxae 1 and 3 together. Coxa 3 about as long as wide. Femur about 8 times as long as wide and more than twice as long as coxa 2, ventrally carrying one row of hardly conspicuous cement glands and cement gland pores, 44 glands counted on 3 rd leg. No cement gland tube. Tibia 1 gently curved, slender, about as long as femur. Tibia 2 even slenderer, longest, about 1.5 times length of femur or tibia 1, with many tall setae along and two ventral spines on distal margin. Tarsus medium-sized, trapezoid, one spine distally on ventral surface. Propodus straight, slender, about 3 times as long as tarsus, and ¼ of tibia 2 length. Main claw short, about ¼ of propodal length, well curved. Auxiliary claws present, well curved, as long as main claw. Both main claw and auxiliary claws with small teeth on inner surface. MEASUREMENTS (mm). Trunk 1.13; abdomen 0.22; proboscis 0.61; chelifore scape 0.61; chela palm 0.56; chela fingers 0.29; coxa 1 0.16; coxa 2 0.54; coxa 3 0.20; femur 1.24; tibia 1 1.34; tibia 2 1.82; tarsus 0.15; propodus 0.36; main claw 0.12; auxillary claw 0.12. Sexual dimorphism No female available. Individual variability Strigilis formula more or less with one compound spine per article in investigated specimens, the highest number of compound spines always on the first strigilis article. Chela teeth number variable, 13 – 16 on movable finger and 13 – 20 on immovable finger in investigated specimens.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E29FF8AFD75075603C0BA25.taxon	discussion	Remarks Nymphon dorlis sp. nov. is a new species in the aequidigitatum - group (Child 1988 b). It can be discriminated from most of its relatives by the following three criteria: (i) preocular neck short, (ii) tarsus about ⅓ as long as propodus; (iii) tibiae 2 about 1.5 times as long as femorae or tibiae 1. Additionally, this species can be distinguished from N. biformidens Stock, 1974 from Madagascar by the larger chelifore palms and shorter fingers of N. dorlis, as well as its conspicuous horn-like lateral sense organs (Stock 1974). It can be distinguished from N. aequidigitatum Flynn, 1919 from Australia by the shorter tarsi (⅓ of propodus length in N. dorlis against ½ in N. aequidigitatum) and the prominent lateral organs of N. dorlis (see Stock 1973). It differs from N. draconis Child, 1990 from the Indo-Pacific by having no conspicuous range of cement gland tubes on the ventral side of the femorae, a longer main claw relatively to the auxiliary claws, and the tarsus only about ⅓ of propodus length (against about ½ for N. draconis; Child 1990). Finally, it differs from N. megacheles Child, 1988 by the larger space between the lateral processes, the longer 5 th palp segment, the shorter main claw relatively to the propodus and the slenderer leg articles (Child 1988 a). Among Martinique species, Nymphon dorlis sp. nov. could be mistaken for N. macabou Müller, 1990. The two species differ by the propodus, which is straight in N. dorlis and curved in N. macabou; and the tibiae 2, which are slender and 1.5 times as long as the femorae, while in N. macabou tibiae 2 are only slightly longer than the femorae (Müller 1990 a).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E29FF8AFD75075603C0BA25.taxon	description	The species was only collected on the Atlantic coast. Depth range Only known from type locality. 16 – 63 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E24FF8EFD8707970048BCFB.taxon	description	urn: lsid: zoobank. org: act: 53 F 8 FAE 5 - 3380 - 479 C-A 5 AE- 8 C 6 B 30 CAC 9 A 5 Figs 12 – 13	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E24FF8EFD8707970048BCFB.taxon	materials_examined	Material examined Holotype MARTINIQUE • ♂ ov. carrying larvae; Les Anses-d’Arlet; 14 ° 29.6 ʹ N, 61 ° 05.5 ʹ W; depth 10 – 26 m; 7 Sep. 2016; st. AR 050; MNHN-IU- 2016 - 835 / MK 411150. Paratypes MARTINIQUE • 1 ♀; Canal de Ste Lucie; 14 ° 22.7 ʹ N, 60 ° 51.6 ʹ W; depth 65 m; 11 Sep. 2016; st. AD 222; MNHN-IU- 2016 - 836 / MK 411151 • 1 ♂; Passe du Marin; 14 ° 26.6 ʹ N, 60 ° 54.3 ʹ W; depth 15 m; 9 Sep. 2016; st. AB 058; MNHN-IU- 2016 - 837 / MK 411152 • 1 ind. (sex indet.); Passe du Marin; 14 ° 26.7 ʹ N, 60 ° 54 ʹ W; depth 19 – 15 m; 10 Sep. 2016; st. AB 060; MNHN-IU- 2016 - 558 • 1 ♂; Grande Anse d’Arlets; 14 ° 29.9 ʹ N, 61 ° 05.4 ʹ W; depth 28 m; 7 Sep. 2016; st. AB 157; MNHN-IU- 2016 - 561 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 562 • 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 1274 / MK 411094 • 1 juv.; same collection data as for preceding; MNHN- IU- 2016 - 1323 / MK 411099 • 1 ♂ ov. carrying larvae; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 29 m; 18 Sep. 2016; st. AS 253; MNHN-IU- 2016 - 1107 • 1 ♀; same collection data as for preceding; MNHN-IU- 2017 - 219 • 1 juv.; Vétiver; 14 ° 37,8 ’ N- 61 ° 07,7 ’ W; depth 25 m; 8 Sep. 2016; st. AB 161; MNHN-IU- 2016 - 1118 / MK 411054 • 1 ♂ ov.; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2016 - 1317 • 1 ♂; N Presqu’Île de la Caravelle; 14 ° 51.9 ʹ N, 60 ° 50.7 ʹ W; depth 57 m; 19 Sep. 2016; st. AD 247; MNHN-IU- 2017 - 3500.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E24FF8EFD8707970048BCFB.taxon	etymology	Etymology From the Latin first name Ludovicus, - i, (masc., 2 nd decl.), from which the French name Louis originates. The species is named after the Fort Saint-Louis naval base in Fort-de-France, Martinique, where the Madibenthos crew was warmly welcome and hosted for the whole expedition.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E24FF8EFD8707970048BCFB.taxon	description	Description (holotype, ♂, MNHN-IU- 2016 - 835) BODY. Medium size, slender species. Trunk completely segmented, cuticle smooth. No dorsomedian ornamentation. Ocular tubercle positioned anteriorly to 1 st lateral processes; about as tall as its diameter, rounded, slightly bent backward, with wide, slightly bifid rounded tip carrying lateral sense organs distally; four pigmented eyes. Preocular neck long, about 3 times as long as wide, markedly segmented at base. Oviger base well separated from 1 st lateral processes. Lateral processes well separated by about twice their diameter between 1 st and 2 nd, about 1.5 times between 2 nd and 3 rd, and about their diameter between 3 rd and 4 th; lateral processes without ornamentation. PROBOScIS. Robust, with distal swelling on 3 antimers, bearing a few setae around mouth. Proboscis reaching about distal part of chelifore scape in lateral view. ABDOMEN. Small, rounded with pointed anus, directed diagonally. No basal articulation. CHELIFORE. 2 - articled, long, reaching beyond proboscis. Scape 1 - articled, about 5 times as long as wide, with dorsal setae on distal part, peculiarly on distal margin. Chela large, palm ornamented with a few setae, longer than fingers. Fingers crossing at tip, carrying bifid teeth plus simple teeth at base. 16 bifurcate teeth plus 3 simple teeth on imovable finger, 14 bifurcate teeth plus 2 normal teeth on movable finger. PALP. 5 - articled. 1 st article shortest, shorter than wide, glabrous. 2 nd article longest, about 4 / 3 length of 3 rd or 5 th, more than 10 times as long as wide. 3 rd article about 7 times as long as wide, with several setae on ventrodistal margin. 4 th article about 1.5 times as long as wide, covered with ventral setae. 5 th article of about same size as 3 rd, about 6 times as long as wide, covered ventrally with setae. OvIGER. 10 - articled, slender, 1 st and 2 nd articles as long as wide, with scarce setae mostly on last 5 articles, 3 rd article less than 5 times as long as wide. 4 th article about twice as long as 3 rd. 5 th article longest, more than 1.5 times as long as 4 th, slightly curved, inflated distally. 6 th article about half as long as 4 th. 7 th article about 0.6 times as long as 6 th. 8 th, 9 th and 10 th articles shorter than 7 th, subequal. Strigilis formula 15: 12: 13: 13. Strigilis spines compound, carrying many lateral teeth. On 7 th to 9 th articles proximalmost compound spine shortest and oblong, median spines lanceolate and longer than proximal ones, distalmost ones spatulated, with most basal teeth of compound spines conspicuously broader than distal teeth. Intermediate position spines roughly intermediary in shape. On 10 th article, all spines but two distal ones round to oblong and short, two distal spines spatulated with basal teeth of spines conspicuously broader than distal teeth, distalmost spine longest. Terminal claw about ⅔ of 4 th strigilis article length. LEGS. Long and slender, with scarce setae. Coxae 1 and 3 subequal, about 1.5 times as long as wide. Coxa 2 more than twice as long as coxae 1 and 3 together. Femur about 12 times as long as median width, with one row of cement glands and cement gland pores on ventral surface, 46 counted on 4 th leg. No cement gland tube. Tibia slightly longer than femur, about 12 times as long as median width. Tibia 2 longest, about 1.5 times as long as femur and 22 times as long as median width. Tarsus medium-sized, about 3 times as long as wide, trapezoid, with one ventrodistal spine. Propodus straight, about 0.2 times as long as tibia 2 and 6 times as long as broad, no basal spine; sole with 8 spines and setae. Main claw curved, short, less than ¼ of propodal length. Auxiliary claws curved, slightly longer than main claw. Both auxiliary claws and main claw with denticles on inner surface. MEASUREMENTS (mm). Trunk 2.08; abdomen 0.16; proboscis 0.87; chelifore scape 0.98; chela palm 0.62; chela fingers 0.36; coxa 1 0.26; coxa 2 1.1; coxa 3 0.31; femur 1.97; tibia 1 2.15; tibia 2 2.83; tarsus 0.23; propodus 0.71; main claw 0.18; auxiliary claw 0.22. Sexual dimorphism Available female poorly preserved. Oviger articles shorter than in males, no distal inflation of 5 th article. Females chela palm not as wide as in males (Fig. 13). Individual variability Number of teeth on chela fingers variable, up to 18 teeth on movable finger and 16 – 19 on imovable finger in investigated specimens. Strigilis formula variable, 11 – 18 per strigilis article, with generally more spines on 7 th oviger article. Teeth of 10 th oviger article either spatulated (e. g., MNHN-IU- 2016 - 835) or lanceolate (e. g., MNHN-IU- 2016 - 837), sometimes within same barcode cluster.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E24FF8EFD8707970048BCFB.taxon	discussion	Remarks Nymphon ludovici sp. nov. can be discriminated from other aequidigitatum - group species with the combination of the four following characters: (i) the preocular neck is long, (ii) the tarsus is short, (iii) the lateral sense organs on the ocular tubercle tip are low and the ocular tubercle tip is not bifurcate; and (iv) the main and auxiliary claws are subequal in length. The closest species are the Japanese N. plectrum Takahashi et al., 2012, which presents much more setose legs, longer leg setae and lateral sense organs well separated on the ocular tubercle, and N. floridanum Hedgpeth, 1948 from the Caribbean, which has no segmentation at the base of the preocular neck. We were not able to infer whether the segmentation at the preocular neck base corresponds to a mobile articulation. This feature is shared with a number of species from the aequidigitatum complex (Child 1988 b; Takahashi et al. 2012): N. adense Müller, 1989, N. aldabrense Child, 1988, N. giraffa Loman, 1908, N. micronesicum Child, 1982 and N. plectrum Takahashi et al., 2012. Some other species of the aequidigitatum - complex show a constriction at the same location of the ocular neck that does not seem to be a segmentation, e. g., N. natalense Flynn, 1928 (see Flynn 1928). This species is divided into two clusters based on DNA barcoding data (intracluster p-distance = 0 – 0.026, intercluster p-distance = 0.104 – 0.111) (Sabroux et al. 2019 b). Since no morphological character was found to support their separation, the two clusters corresponding to group 1 (holotype MNHN-IU- 2016 - 835 and paratype MNHN-IU- 2016 - 837) and group 2 (paratypes MNHN-IU- 2016 - 562, - 836, - 1118, - 1274 and - 1323) (Appendix), were described as the same species. The species was collected on both the Atlantic and Caribbean coasts.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E24FF8EFD8707970048BCFB.taxon	distribution	Distribution Only known from type locality. Depth range 15 – 57 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E20FFF1FD9E01CD0048BD91.taxon	description	urn: lsid: zoobank. org: act: 2 B 7 EA 480 - 66 C 1 - 4449 - BC 75 - DB 7 D 3459 CBF 7 Fig. 14	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E20FFF1FD9E01CD0048BD91.taxon	materials_examined	Material examined Holotype MARTINIQUE • ♂; Les Anses-d’Arlet; 14 ° 29.7 ʹ N, 61 ° 05.4 ʹ W; depth 19 m; 7 Sep. 2016; st. AB 155; MNHN-IU- 2016 - 889 / MK 411198. Paratypes MARTINIQUE • 2 ♀♀, 1 juv.; same collection data as for holotype; MNHN-IU- 2016 - 1280 • 1 ♀; Bellefontaine; 14 ° 39.7 ʹ N, 61 ° 09.6 ʹ W; depth 12 m; 30 Sep. 2016; st. AB 372; MNHN-IU- 2016 - 1033 • 1 ♂; Rocher du Diamant; 14 ° 26.5 ʹ N, 61 ° 02.4 ʹ W; depth 24 m; 14 Sep. 2016; st. AB 173; MNHN- IU- 2016 - 1159 • 1 ♂ carrying larvae; Pointe de la Baleine; 14 ° 31.1 ʹ N, 61 ° 05.9 ʹ W; depth 17 – 19 m; 30 Sep. 2016; st. AB 369; MNHN-IU- 2016 - 1257 • 3 ♀♀ gr.; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2016 - 1299 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1319 • 1 ♀ preadult?; Presqu’Île de la Caravelle; 14 ° 46.5 ʹ N, 60 ° 51.5 ʹ W; depth 15 m; 22 Sep. 2016; st. AB 350; MNHN-IU- 2016 - 1304 • 1 juv.; Pointe de la Baleine; 14 ° 31.1 ʹ N, 61 ° 05.9 ʹ W; depth 24 m; 30 Sep. 2016; st. AR 413; MNHN-IU- 2016 - 1221.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E20FFF1FD9E01CD0048BD91.taxon	etymology	Etymology From the Latin neologism Martinicus, - a, - um (adj.): Martiniquese.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E20FFF1FD9E01CD0048BD91.taxon	description	Description (holotype, ♂, MNHN-IU- 2016 - 889) BODY. Trunk completely segmented, cuticle smooth. No dorsomedian tubercle. Ocular tubercle near oviger base and positioned anterior to 1 st lateral processes, rounded, as wide at base as high, with 2 large horn-like lateral sense organs on its tip; 4 pigmented eyes. Preocular neck long, as high as base width. Oviger base touching 1 st lateral processes. Lateral processes without ornamentation, well separated by more than once and up to twice their own diameters between 1 st, 2 nd and 3 rd lateral processes and by slightly less than their own diameter between 3 rd and 4 th lateral processes. PROBOScIS. Rounded and large, reaching distal margin of chelifore scape in lateral view, about as long as cephalon and 1 st trunk segment. ABDOMEN. Medium-sized, not reaching beyond lateral processes, directed diagonally. No basal segmentation. CHELIFORE. 2 - articled, reaching far beyond proboscis, carrying scarce setae. Scape 1 - articled, 5.5 times as long as wide. Chela large, palm longer than fingers. Fingers sub-equal in length, bearing simple and bifurcated teeth, 17 on inmovable finger and 15 on movable finger. PALP. 5 - articled. 1 st shortest, as long as wide. 2 nd article longest, about 11 times as long as wide, with a few distal setae. 3 rd article about 0.8 times as long as 2 nd, carrying setae on distal half. 4 th article about half as long as 5 th, carrying setae mostly on ventral side. 5 th article 0.9 times as long as 3 rd, carrying many ventral setae. OvIGER. 10 - articled. 1 st article shorter than wide. 2 nd article about ⅓ as long as wide. 3 rd article about 3 times as long as maximal width. 4 th article more than twice as long as 3 rd. 5 th article longest, about 1.6 times as long as 4 th, setose, widening near distal part. 6 th article about 0.7 times as long as 4 th, setose. 7 th article longest within strigilis, 8 th, 9 th and 10 th articles subequal. Strigilis formula 13: 10: 9: 11. Strigilis spines compound, carrying many lateral teeth, proximalmost shortest and spatulated, median spines longest, oblong to lanceolate), distalmost spatulated, longer than proximalmost. Intermediary position spines roughly intermediary in shape. Terminal claw about as long as 10 th article. LEGS. Slender, setose. Coxa 1 about 1.5 times as long as wide. Coxa 2 about 2.5 times as long as coxae 1 and 3 together. Coxa 3 about 1.5 times as long as wide. Femur about 16 times as long as wide and about twice as long as coxa 2, ventrally carrying one row of hardly conspicuous cement glands and cement gland pores, 45 glands counted. No cement gland tube. Tibia 1 less than 0.1 times as long as femur. Tibia 2 longest, even slenderer, about ¼ as long as tibia 1, setose. Tarsus medium-sized, trapezoid, one spine distally on inner surface. Propodus straight, slender, about 2.5 times as long as tarsus, and 0.15 times as long as tibia 2. Main claw short, about ¼ of propodal length, curved. Auxiliary claws present, curved, slightly longer than main claw. Both main claw and auxiliary claws with small denticles on inner surface. MEASUREMENTS (mm). Trunk 1.59; abdomen 0.20; proboscis 0.75; chelifore scape 0.55; chela palm 0.41; chela fingers 0.31; coxa 1 0.16; coxa 2 0.97; coxa 3 0.21; femur 1.88; tibia 1 2.04; tibia 2 3.00; tarsus 0.19; propodus 0.45; main claw 0.11; auxillary claw 0.15. Sexual dimorphism 5 th ovigeral article distally inflated only in males. Female chela palm not as wide as in males. Individual variability Chela teeth number variable, 12 to 19 on immovable finger, and 14 to 17 on movable finger. Strigilis formula variable, from 9 to 13 spines on each strigilis article, with generally more spines on 1 st strigilis article. In one specimen (MNHN-IU- 2016 - 1304), strigilis formula drops to 8: 7: 7: 7, which may be linked to preadult stage.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E20FFF1FD9E01CD0048BD91.taxon	discussion	Remarks Nymphon martinicum sp. nov. belongs to the aequidigitatum group. It greatly resembles N. floridanum Hedgpeth, 1948, and differs from this species by the ocular tubercle, which is simple and conical in N. floridanum, while it is rounded in Martinique specimens with two conspicuous, horn-like lateral sense organs (Hedgpeth 1948; Child 1992 a; Müller & Krapp 2009). We describe this species as new to science based on those morphological differences, with strong support from CO 1 data: the p-distance between the sequenced specimen and N. floridanum GenBank sequence DQ 390073 (Arango & Wheeler 2007) is equal to 0.147 (see Appendix). This species was collected on both the Atlantic and Caribbean coasts.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E20FFF1FD9E01CD0048BD91.taxon	description	Depth range 12 – 25 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5FFFF4FD7C01230048BA63.taxon	description	urn: lsid: zoobank. org: act: 86 C 9 C 63 A- 9 B 7 A- 4 D 45 - ABAD- 39 D 6 DBA 6 CFD 9 Figs 15 – 16	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5FFFF4FD7C01230048BA63.taxon	materials_examined	Material examined Holotype MARTINIQUE • ♀; Pointe du Diamant; 14 ° 27 ʹ N, 61 ° 04.1 ʹ W; depth 70 m; 9 Sep. 2016; st. AD 214; MNHN-IU- 2016 - 844 / MK 411158. Paratypes MARTINIQUE • 1 ♀; Pointe du Diamant; same collection data as for holotype; MNHN-IU- 2016 - 887 / MK 411196 • 1 ♂, 1 ♀; same collection data as for holotype; MNHN-IU- 2016 - 569 • 1 ♂; E of Le Vauclin; 14 ° 34.2 ʹ N, 60 ° 44.8 ʹ W; depth 90 m; 14 Sep. 2016; st. AD 231; MNHN-IU- 2016 - 1201 • 1 ♂; same collection data as for preceding; MNHN-IU- 2021 - 6623.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5FFFF4FD7C01230048BA63.taxon	etymology	Etymology Noun in apposition. Named after the Ti Mons (from Martinique Creole: ‘ little monster’), a malevolent creature of Martinique folklore which is given birth by placing an egg under his master’s armpit during Lent.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5FFFF4FD7C01230048BA63.taxon	description	Description (holotype, ♀, MNHN-IU- 2016 - 844) BODY. Small species. Trunk completely segmented; cuticle smooth. No dorsomedian ornamentation. Ocular tubercle rounded, about as tall as wide, with 2 small lateral sense organs; four pigmented eyes. Preocular neck broad, medium-sized, about as long as wide. Chelifore insertions well separated. Oviger base touching 1 st lateral processes. Lateral processes well separated by slightly less than their own diameter; 1 st, 2 nd and 3 rd lateral processes about 1.5 times as long as wide, 4 th about as long as wide; unornamented. PROBOScIS. Large, cylindrical, rounded at tip. ABDOMEN. Long, reaching beyond 1 st coxa, oriented diagonally. No basal articulation. CHELIFORE. 2 - articled, slender, reaching beyond proboscis. Scape 1 - articled, with scarce setae, less than 3 times as long as wide. Chela carrying setae; palm slender, about as long as fingers. Fingers slender, subequal in length, with 17 denticled teeth on immovable finger and 15 on movable finger. Some teeth of movable finger curved. PALP. 5 - articled. 1 st article shortest, less than twice as long as wide. 2 nd article longest, 7.5 times as long as wide. 3 rd article 0.8 times as long as 2 nd, with one ventral seta near distal margin. 4 th article 0.4 times as long as 3 rd, with one ventral seta near distal margin. 5 th article about ¼ longer than 4 th, with distal setae. 4 th and 5 th palp articles together about as long as 3 rd. OvIGER. 10 - articled, with scarce setae. 1 st article shortest, shorter than wide. 2 nd and 3 rd articles subequal in length. 4 th article 3 times as long as 2 nd. 5 th article slightly longer than 4 th. 6 th article half as long as 5 th. 7 th article longer than 8 th, 9 th or 10 th, less than 3 times as long as wide. 8 th, 9 th and 10 th articles subequal, about twice as long as wide. Strigilis spines compound, with 2 pairs of long basal teeth, then 3 long teeth at end of long elongation; strigilis formula 5: 4: 4: 4. Terminal claw as long as 10 th article, pectinated with 5 long lateral teeth. LEGS. Slender, with scarce setae. Coxa 1 about 1.5 times as long as wide, carrying setae on distal margin. Coxa 2 longer than coxae 1 and 3 together, widening distally. Coxa 3 about as long as wide, with ventral setae carried distally. Femur about 6.5 times as long as distal width, inflated by ovaries in holotype specimen. Tibia 1 subequal to femur in length, about 12.5 times as long as wide, with one long spine about half its size. Tibia 2 longest, about 1.5 times as long as femur or tibia 1, carrying rows of setae on ventral, dorsal and lateral sides, and 2 ventral spines on distal margin. Tarsus rather long. Propodus less than twice as long as tarsus and ¼ times as long as tibia 2, slightly curved, carrying setae; heel spines absent, 4 sole spines. Main claw less than half as long as propodus, curved. Auxiliary claws curved, about 0.8 times as long as main claw. MEASUREMENTS (mm). Trunk 0.94; abdomen 0.27; proboscis 0.30; chelifore scape 0.31; chela palm 0.31; chela finger 0.29; coxa 1 0.13; coxa 2 0.35; coxa 3 0.13; femur 0.68; tibia 1 0.63; tibia 2 1.01; tarsus 0.15; propodus 0.26; main claw 0.12; auxillary claw 0.09. Sexual dimorphism Male femur narrower, not swollen. Cement glands and cement gland pores inconspicuous, no cement gland tube. Oviger articles longer: in specimen MNHN-IU- 2021 - 6623, 3 rd article longer than wide, 4 th article about 4 times as long as 2 nd article, 5 th article about 1.1 times as long as 4 th. 5 th oviger article curved, with distal apophysis. Individual variability Number of teeth on chelifore fingers 13 to 17 in investigated specimens. Strigilis formula slightly variable although always low, 2 to 5 spines per strigilis article, highest number on 1 st strigilis article. Number of teeth of oviger terminal claw variable, 4 to 5 in investigated specimens.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5FFFF4FD7C01230048BA63.taxon	discussion	Remarks Nymphon timons sp. nov. is the only species of Nymphon in the Madibenthos material that does not belong to the aequidigitatum group. It is a small species for the genus, and is differentiated from other species of Nymphon by the combination of the following characters: tarsus about half as long as propodus; auxiliary claws about 0.8 times as long as main claw; preocular neck medium, about as long as wide; lateral processes separated by slightly less than their own diameter; tibia 2 about 1.5 times as long as tibia 1 or femur; chela teeth denticled; oviger terminal claw pectinated; oviger base touching first lateral processes; fourth and fifth palp articles together about as long as third palp article. To some extent, Nymphon timons sp. nov. shares similarities with N. boogoora Bamber, 2008 from Eastern Australia: both species share the pectinated oviger terminal claw, the long tibiae 2, and the same palp structure (Bamber 2008). However, N. boogoora does not have the chela teeth denticled, and its abdomen does not extand beyond the 4 th lateral processes. Nymphon timons was only collected at two locations: one on the Atlantic coast and the other on the Caribbean coast.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5FFFF4FD7C01230048BA63.taxon	distribution	Distribution Only known from Martinique. Depth range 70 – 90 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5AFFF4FD8D079A0557B961.taxon	type_taxon	Type species Endeis spinosa (Montagu, 1808), by subsequent designation (Child 1998 a).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5AFFF5FD9A045A0079BA4A.taxon	materials_examined	Type material Syntypes: NHMUK ZOO- 1923.10.6.18 - 23 (not examined). Type locality: Madras, India. Material examined MARTINIQUE • 1 ♂; Grande Anse du Diamant; 14 ° 28.6 ʹ N, 60 ° 59.4 ʹ W; depth 1 – 2 m; 9 Sep. 2016; st. AD 215; MNHN-IU- 2016 - 840 / MK 411154 • 1 juv.; same collection data as for preceding; MNHN- IU- 2016 - 559. FRENCH GUIANA • 1 ♂; Île Royale / Île du Diable; Passe des Grenadines; 5 ° 17 ʹ 27.5388 ʺ N, 52 ° 35 ʹ 15.54 ʺ W; depth 6 – 9 m; st. SC 01; MNHN-IU- 2013 - 18550 / KX 535406.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5AFFF5FD9A045A0079BA4A.taxon	discussion	Remarks This is the first record of this species in Martinique. The material only includes one juvenile and one poorly preserved male with no femur left, which strongly impedes identification. However, DNA barcoding data indicated that they belong to the same species as a specimen from French Guiana, MNHN-IU- 2013 - 18550, GenBank reference KX 535406 (Sabroux et al. 2017) with p-distance = 0. This specimen was successfully identified as Endeis flaccida. The species is recognizable by the intestinal tracts having many branched and unbranched diverticula, and the cement gland pores dispatched in an irregular band along the femur (Child 1979). Furthermore, in contrast to the otherwise very similar species E. nodosa Hilton, 1942 (known from Pacific islands and Barbados), E. flaccida does not have a femoral ventral projection (Müller 1992 a). The two specimens were collected at one station on the southern Caribbean coast of Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5AFFF5FD9A045A0079BA4A.taxon	distribution	Distribution Widely distributed and probably pantropical, recorded from Pacific Panama, West Atlantic (Florida, Martinique, Panama, French Guiana), the Indian Ocean (Philippines, Indonesia, India), Taiwan and East Queensland. Depth range 0 – 69 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5BFFF7FDC907770079BBBF.taxon	description	Fig. 17	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5BFFF7FDC907770079BBBF.taxon	materials_examined	Type material Endeis meridionalis (Böhm, 1879). Syntypes: ZMB _ Pyc _ 6 and ZMB _ Pyc _ 7 (not examined). Type localities: Red Sea?, west coast of Sinai Peninsula, Egypt and Singapore (see Dunlop et al. 2007). Material examined MARTINIQUE • 1 ♂; E of Pointe Ferré; 14 ° 27.8 ʹ N, 60 ° 47.8 ʹ W; depth 35 m; 12 Sep. 2016; st. AD 227; MNHN-IU- 2016 - 839 • 1 ♂ ov., 1 ♀ gr., 2 juvs; Pointe du Vauclin; 14 ° 33.6 ʹ N, 60 ° 49.7 ʹ W; depth 2 m; 12 Sep. 2016; st. AD 230; MNHN-IU- 2016 - 560 • 1 ♂; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.2 ʹ W; depth 22 m; 22 Sep. 2016; st. AS 096; MNHN-IU- 2016 - 1043 / MK 411032 • 2 ♂♂; same collection data as for preceding; MNHN-IU- 2016 - 1423 • 1 ♂; Passe du Brigot; 14 ° 33.8 ʹ N, 60 ° 47.6 ʹ W; depth 30 m; 17 Sep. 2016; st. AR 082; MNHN-IU- 2016 - 1125 • 1 ♂ ov.; Le François; 14 ° 37 ʹ N, 60 ° 49.5 ʹ W; depth 1 m; 17 Sep. 2016; st. AB 134; MNHN-IU- 2016 - 1142 / MK 411062 • 1 juv.; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 29 m; 18 Sep. 2016; st. AS 253; MNHN-IU- 2016 - 1265 / MK 411091.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5BFFF7FDC907770079BBBF.taxon	discussion	Remarks This species was regarded as Endeis meridionalis in Sabroux et al. (2019 b) despite the shorter distance between the lateral processes, and the absence of kinked femorae in the Martinique material. The specimens also resemble E. spinosa as drawn by Marcus (1940), but this species was represented in quite different fashions, with a variable shape of the proboscis, distance between the lateral processes, etc. (Hedgpeth 1948; Child 1992 b; Munilla & Soler-Membrives 2014). Since the original description provides little information, and because of the inherent difficulties in identifying species of Endeis, the status of this morphospecies remains doubtful; therefore, we refrain from describing it as new species. Most of the specimens were found on the Atlantic coast, although three were collected at Pointe Diamant on the Carribean coast.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E5BFFF7FDC907770079BBBF.taxon	distribution	Distribution Mostly recorded from the Indian Ocean (Somalia, Réunion, Mauritius (?), Madagascar, India, Malaysia, India, Western Australia) and the Red Sea; also reported in the Pacific (Japan, Moorea, Fiji islands), in the Eastern Atlantic (Cape Verde) and in the Caribbean (Curaçao). Depth range 0 – 90 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E59FFF9FDBB07380048BA6C.taxon	description	Fig. 18	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E59FFF9FDBB07380048BA6C.taxon	materials_examined	Type material Endeis mollis (Carpenter, 1904). Type (s): unknown. Type locality: Cheval Paar, Sri Lanka. Material examined MARTINIQUE • 1 ♀; Baie de Fort-de-France; 14 ° 32.6 ʹ N, 61 ° 04.7 ʹ W; depth 0 – 1 m; 6 Sep. 2016; st. AM 002; MNHN-IU- 2016 - 838 / MK 411153 • 1 ♂ ov.; same collection data as for preceding; MNHN- IU- 2016 - 565 / MK 411116 • 2 ♀♀; same collection data as for preceding; MNHN-IU- 2017 - 216 • 1 ♀; same collection data as for preceding; MNHN-IU- 2017 - 217 / MK 411200.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E59FFF9FDBB07380048BA6C.taxon	discussion	Remarks The general morphology of Martinique specimens corresponds well to the description in Carpenter (1904), as well as that in Müller & Krapp (2009), including the distal spiny tubercle on the lateral processes. However, these specimens are much smaller than other records (the trunk is about half as long as indicated in Müller & Krapp 2009), the ocular tubercle and lateral processes are smaller, and the number of observed cement gland pores is 13 rather than 19 as mentioned by Müller & Krapp (2019). The CO 1 barcoding tree of Sabroux et al. (2019 b) showed a sister-group relationship with a specimen of Endeis mollis from Caribbean Colombia identified by Arango & Wheeler (2007), although the genetic distance is very high (p-distance = 0.157; see Appendix). It is therefore probably another species than E. mollis, but due to the complexity of taxonomy within Endeis, we refrain from describing it as a new species as long as a review for the genus is not available. All specimens were found at the same station, at the entrance of Baie de Fort-de-France.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E59FFF9FDBB07380048BA6C.taxon	distribution	Distribution Endeis mollis is a commonly found pantropical species recorded from the Tropical West Atlantic (Caribbean, Surinam, Brazil), West Africa (Occidental Sahara, off Guinea, Nigeria), Indian Ocean (South Africa, Madagascar, Somalia, Red Sea, Suez, Oman, Pakistan, India, Sri Lanka, Maldives, Christmas Islands, Philippines) and the West Pacific (Australia, Japan mainland, Ryukyu Islands, New Caledonia, Guam, Marianas Islands, Polynesia). There is only one record from the Western Mediterranean, north of Tunis.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E59FFF9FDBB07380048BA6C.taxon	description	Depth range 0 – 105 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E57FFFBFD2C07590048BEC5.taxon	description	Fig. 19	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E57FFFBFD2C07590048BEC5.taxon	materials_examined	Material examined MARTINIQUE • 1 ♀ gr.; Macouba; 14 ° 55.1 ʹ N, 61 ° 09 ʹ W; depth 80 m; 24 Sep. 2016; st. AD 261; MNHN-IU- 2016 - 863 / MK 411175 • 1 leg (♂); Macouba; 14 ° 55 ʹ N, 61 ° 08.9 ʹ W; depth 78 – 80 m; 24 Sep. 2016; st. AD 263; MNHN-IU- 2016 - 1203 / MK 411080.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E57FFFBFD2C07590048BEC5.taxon	discussion	Remarks These specimens resemble E. mollis, but they do not show the spiny tubercle on the lateral processes as represented by Carpenter (1904) and Müller & Krapp (2009). The male femur presents at least 25 pores. Only two specimens were collected: one female and a male leg. The analysis of CO 1 sequences confirmed that the male leg belongs to the same species as the female specimen, making available characters of the cement gland pores. However, in the absence of ovigers, a species description would be incomplete. Therefore we prefer to keep this species as unnamed until new material becomes available. The two specimens were collected at the same location, north of Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E57FFFBFD2C07590048BEC5.taxon	description	Depth range 78 – 80 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E55FFFBFDD500CD00A5B8BB.taxon	materials_examined	Type material Anoplodactylus arcuatus. Holotype: USNM 154801 (not examined). Type locality: Piscadera Bay, Curaçao. Material examined MARTINIQUE • 1 juv.; Vétiver; 14 ° 37.8 ʹ N, 61 ° 07.7 ʹ W; depth 25 m; 8 Sep. 2016; st. AB 161; MNHN- IU- 2016 - 1036 / MK 411030 • 1 ♂, 1 ♂ ov., 1 ♀, 2 ♀♀ gr., 1 juv.; Vétiver; 14 ° 37.9 ʹ N, 61 ° 07.7 ʹ W; depth 10 m; 8 Sep. 2016; st. AS 054; MNHN-IU- 2016 - 1204 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1326 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1327 / MK 411102 • 1 ♀; Pointe de la Baleine; 14 ° 31.1 ʹ N, 61 ° 05.9 ʹ W; depth 17 – 19 m; 30 Sep. 2016; st. AB 369; MNHN- IU- 2016 - 1259 / MK 411087 • 1 ♀; Le Prêcheur; 14 ° 48.5 ʹ N, 61 ° 13.8 ʹ W; depth 56 m; 8 Oct. 2016; st. AS 574; MNHN-IU- 2016 - 1288.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E55FFFBFDD500CD00A5B8BB.taxon	discussion	Remarks Martinique specimens are very similar to those described by Child (1977 b), though the cement gland tubes are situated more proximally on the femorae (at about mid-length). The tubes are not arcuated in the present material; however, this character is known to be strongly variable (Müller & Krapp 2009). All specimens were collected on the Caribbean coast.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E55FFFBFDD500CD00A5B8BB.taxon	distribution	Distribution Anoplodactylus arcuatus is known from the Caribbean (Curaçao, Barbados) and the Gulf of Mexico (Loggerhead Keys, Florida).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E55FFFBFDD500CD00A5B8BB.taxon	description	Depth range Anoplodactylus arcuatus is known to dwell between 2.5 and 18 m. The present material was sampled between 10 and 56 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E55FFFBFDBC00350538BDFA.taxon	type_taxon	Type species Anoplodactylus lentus Wilson, 1878, by subsequent designation (Child 1998 a).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E52FFFDFDD503270369BE97.taxon	description	Fig. 20	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E52FFFDFDD503270369BE97.taxon	materials_examined	Type material Holotype: USNM 154804 (not examined). Type locality: Tongue of the Ocean, off Andros Island, Bahamas. Material examined MARTINIQUE • 3 ♂♂, 1 ♀, 2 juvs; Anse Couleuvre; 14 ° 50.4 ʹ N, 61 ° 13.4 ʹ W; depth 7 m; 1 Oct. 2016; st. AB 463; MNHN-IU- 2016 - 1123 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1390 • 1 ♀; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 29 m; 18 Sep. 2016; st. AS 253; MNHN- IU- 2016 - 1264 / MK 411090 • 1 ♂ ov.; Anse Dufour; 14 ° 31.6 ʹ N, 61 ° 05.4 ʹ W; depth 10 – 20 m; 7 Sep. 2016; st. AD 206; MNHN-IU- 2016 - 1271 • 1 ♂; Baie de Fort-de-France; 14 ° 34.5 ʹ N, 61 ° 05 ʹ W; depth 11 – 15 m; 6 Oct. 2016; st. AR 468; MNHN-IU- 2016 - 1324 • 1 ♂ ov.; Le Prêcheur; 14 ° 49.1 ʹ N, 61 ° 13.8 ʹ W; depth 20 – 25 m; 8 Oct. 2016; st. AS 576; MNHN-IU- 2016 - 1389 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1463.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E52FFFDFDD503270369BE97.taxon	discussion	Remarks Anoplodactylus bahamensis is discriminated from Anoplodactylus micros by the conspicuously larger size of the specimens and the wider opening of the cement gland pores. We observed setae on the ventral side of the proboscis in both species (but not in all specimens), unlike observations in Müller & Krapp (2009). Only one specimen was barcoded, a female. While it is not possible to confirm its identification using the cement glands, the specimen is similar to the male specimens regarding all available characters, including the propodal sole lamina and the dorsomedian tubercles on the lateral processes. This is the first record of this species in Martinique. The species was sampled at five different stations, four of them on the Caribbean coast, one (the barcoded female, MNHN-IU- 2016 - 1264) on the Caribbean coast. Müller & Krapp (2009) mentioned that the species prefers sublittoral areas, where it is peculiarly abundant in sea-grass meadows.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E52FFFDFDD503270369BE97.taxon	distribution	Distribution Caribbean (Belize, Nicaragua, Colombia) and Bahamas.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E52FFFDFDD503270369BE97.taxon	description	Depth range 0.5 – 12 m, extended to 20 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E53FFFEFDE700210079B89A.taxon	materials_examined	Type material Pycnosoma batangense Helfer, 1938. Holotype: ZMB _ Pyc _ 285 (not examined). Type locality: Grand Batanga, Cameroun, Gulf of Guinea. Anoplodactylus intermedius Hilton, 1942. Type (s): unknown. Type locality: Hawaiian Islands. Anoplodactylus stylirostris Hedgpeth, 1948. Holotype: USNM 81094 (not examined). Type locality: off southeast Loggerhead Key, Tortugas, Florida. Anoplodactylus tenuirostris Lebour, 1949. Type (s): unknown. Type locality: Bermuda, the Reach. Material examined MARTINIQUE • 1 ♂; Le Vauclin; 14 ° 32.7 ʹ N, 60 ° 49.8 ʹ W; depth 2 m; 12 Sep. 2016; st. AD 229; MNHN- IU- 2016 - 799 / MK 411122 • 1 ♂ ov.; same collection data as for preceding; MNHN-IU- 2016 - 896 • 1 ♂; E of Le Robert; 14 ° 40.1 ʹ N, 60 ° 51.1 ʹ W; depth 14 m; 18 Sep. 2016; st. AB 191; MNHN-IU- 2016 - 800 / MK 411123 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1144 • 1 ♀; Ste-Luce; 14 ° 27.3 ʹ N, 60 ° 55.5 ʹ W; depth 15 m; 10 Sep. 2016; st. AB 062; MNHN-IU- 2016 - 801 / MK 411124 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 566 • 1 ♀ gr.; Pointe Michel; 14 ° 26.4 ʹ N, 60 ° 49.3 ʹ W; depth 2 m; 12 Sep. 2016; st. AB 120; MNHN-IU- 2016 - 892 • 1 ♀ gr.; Baie de Fort-deFrance; 14 ° 32.6 ʹ N, 61 ° 04.7 ʹ W; depth 0 – 1 m; 6 Sep. 2016; st. AM 002; MNHN-IU- 2016 - 893 • 3 ♀♀; Pointe du Vauclin; 14 ° 33.6 ʹ N, 60 ° 49.7 ʹ W; depth 2 m; 12 Sep. 2016; st. AD 230; MNHN-IU- 2016 - 894 • 1 ♀; Anse d’Arlet; 14 ° 29.4 ʹ N, 61 ° 05 ʹ W; depth 12 m; 6 Sep. 2016; st. AD 203; MNHN-IU- 2016 - 895 • 1 ♂ ov.; Pointe Baham; 14 ° 24.7 ʹ N, 60 ° 50.1 ʹ W; depth 2 m; 11 Sep. 2016; st. AB 117; MNHN- IU- 2016 - 572 • 1 ♂ ov.; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.3 ʹ W; depth 4 – 10 m; 14 Sep. 2016; st. AB 123; MNHN-IU- 2016 - 1068 • 1 juv.; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 00.1 ʹ W; depth 12 m; 26 Sep. 2016; st. AB 360; MNHN-IU- 2016 - 1081 • 3 ♂♂, 1 ♂ ov., 2 ♀♀, 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 1215 • 3 ♂♂, 2 ♀♀, 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 1236 • 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 1403 • 1 ♂ ov.; Presqu’Île de la Caravelle; 14 ° 47.5 ʹ N, 60 ° 57.4 ʹ W; depth 14 m; 4 Oct. 2016; st. AB 562; MNHN- IU- 2016 - 1088 • 1 preadult ♀; Presqu’Île de la Caravelle; 14 ° 45.4 ʹ N, 60 ° 58 ʹ W; depth 0 – 2 m; 22 Sep. 2016; st. AM 027; MNHN-IU- 2016 - 1097 • 1 ♀ gr.; Baie de Sans-Souci; 14 ° 34.7 ʹ N, 60 ° 50.7 ʹ W; depth 0 – 1 m; 9 Oct. 2016; st. AM 325; MNHN-IU- 2016 - 1099 • 1 ♂; NE Pointe Jacob; 14 ° 36.1 ʹ N, 60 ° 49 ʹ W; depth 23 m; 17 Sep. 2016; st. AB 185; MNHN-IU- 2016 - 1101 • 1 ♀; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 02.8 ʹ W; depth 0 – 1 m; 27 Sep. 2016; st. AM 033; MNHN-IU- 2016 - 1110 • 3 ♂♂, 1 ♀; Anse Couleuvre; 14 ° 50.4 ʹ N, 61 ° 13.4 ʹ W; depth 7 m; 1 Oct. 2016; st. AB 463; MNHN-IU- 2016 - 1122 • 1 ♂; Trois Rivières; 14 ° 27.8 ʹ N, 60 ° 57.9 ʹ W; depth 3 – 4 m; 25 Sep. 2016; st. AB 356; MNHN-IU- 2016 - 1164 • 1 ♀; same collection data as for preceding; MNHN-IU- 2016 - 1269 • 2 ♂♂, 2 ♀♀; Trois Rivières; 14 ° 27.7 ʹ N, 60 ° 57.8 ʹ W; depth 4 – 5 m; 25 Sep. 2016; st. AB 354; MNHN-IU- 2016 - 1171 • 1 ♀ gr.; Le Vauclin; 14 ° 32 ʹ N, 60 ° 49 ʹ W; depth 1 m; 16 Sep. 2016; st. AB 130; MNHN-IU- 2016 - 1188 • 1 ♀ gr.; Îlet La Perle; 14 ° 50.5 ʹ N, 61 ° 13.7 ʹ W; depth 23 m; 1 Oct. 2016; st. AS 375; MNHN-IU- 2016 - 1206 • 1 ♀; Presqu’Île de la Caravelle; 14 ° 47.3 ʹ N, 60 ° 57.4 ʹ W; depth 19 m; 21 Sep. 2016; st. AB 301; MNHN-IU- 2016 - 1228 • 4 ♂♂, 4 ♂♂ ov., 3 ♀♀; Le François; 14 ° 37 ʹ N, 60 ° 49.5 ʹ W; depth 1 m; 17 Sep. 2016; st. AB 134; MNHN- IU- 2016 - 1232 • 1 ♀ gr.; Anse Céron; 14 ° 50.1 ʹ N, 61 ° 13.4 ʹ W; depth 0 – 2 m; 3 Oct. 2016; st. AM 039; MNHN-IU- 2016 - 1240 • 1 ♀; Anse des Galets; 14 ° 51.4 ʹ N, 61 ° 12.8 ʹ W; depth 10 m; 7 Oct. 2016; st. AB 567; MNHN-IU- 2016 - 1245 • 1 ♂ ov., 1 ♀ gr.; Baie du Robert; 14 ° 40.2 ʹ N, 60 ° 52.5 ʹ W; depth 11 m; 24 Sep. 2016; st. AS 403; MNHN-IU- 2016 - 1267 • 1 ♂, 1 ♀ gr.; Presqu’Île de la Caravelle; 14 ° 44.5 ʹ N, 60 ° 53.5 ʹ W; depth 5 – 12 m; 17 Sep. 2016; st. AD 241; MNHN-IU- 2016 - 1291 • 1 ♂; same collection data as for preceding; MNHN-IU- 2017 - 3915 • 1 ♀ gr.; Presqu’Île de la Caravelle; 14 ° 45.5 ʹ N, 60 ° 52.8 ʹ W; depth 2 m; 22 Sep. 2016; st. AB 149; MNHN-IU- 2017 - 2558.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E53FFFEFDE700210079B89A.taxon	discussion	Remarks This species, characterized by its slender, tapering proboscis, is widely distributed, particularly common in the Western Atlantic and one of the most abundant of the present study. Already reported from Martinique, from Petite Anse du Diamant (Bourdillon 1955) and Ilet Cabrits, Cap Chevalier, Petite Anse Macabou and Baie de Tartane (Müller 1990 a), this species was collected on both the Atlantic and Caribbean coasts, although none were found along the coast between Saint-Pierre and Fort-de-France where the slope is the steepest.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E53FFFEFDE700210079B89A.taxon	distribution	Distribution Circumtropical species: Tropical West Atlantic (South Florida, Gulf of Mexico, Caribbean, Brazil), Gulf of Guinea, Indo-Pacific (Indonesia, Papua-New Guinea, Philippines, Eastern Australia, New Caledonia, Tonga, Hawaii), East Pacific (Costa Rica and Panama).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E53FFFEFDE700210079B89A.taxon	description	Depth range 0 – 40 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E51FFE0FDE6032700EDBAEA.taxon	description	Fig. 21	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E51FFE0FDE6032700EDBAEA.taxon	materials_examined	Type material Anoplodactylus californicus Hall, 1912. Topotypes (see Child 1987): USNM 231867 (not examined). Type locality: Laguna Beach, California. Anoplodactylus portus Calman, 1927. Syntypes: NHMUK ZOO- 1926.19.9 – 13 (not examined). Type locality: Port Said, Suez Canal. Anoplodactylus carvalhoi Marcus, 1940. Type (s): unknown. Type locality: Bahia de Santos, Brazil. Anoplodactylus projectus Hilton, 1942. Type (s): unknown. Type locality: Pearl Harbour, Oahu, Hawaiian Islands. Material examined MARTINIQUE • 1 ♀; Pointe Michel; 14 ° 26.4 ʹ N, 60 ° 49.3 ʹ W; depth 2 m; 12 Sep. 2016; st. AB 120; MNHN-IU- 2016 - 810 / MK 411132 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1231 • 1 ♀; Pointe Michel; 14 ° 26.4 ʹ N, 60 ° 49.3 ʹ W; depth 0 – 2 m; 12 Sep. 2016; st. AR 121; MNHN-IU- 2016 - 811 / MK 411133 • 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 891 / MK 411199 • 1 ♂; Pointe Baham; 14 ° 24.7 ʹ N, 60 ° 50.1 ʹ W; depth 2 m; 11 Sep. 2016; st. AB 117; MNHN-IU- 2016 - 855 / MK 411168 • 1 juv.; same collection data as for preceding; MNHN-IU- 2016 - 571 • 1 juv; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 02.8 ʹ W; depth 0 – 1 m; 27 Sep. 2016; st. AM 033; MNHN-IU- 2016 - 864 / MK 411176.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E51FFE0FDE6032700EDBAEA.taxon	discussion	Remarks The Madibenthos material matches with the descriptions in Child (1992 b) and Müller & Krapp (2009) of Anoplodactylus californicus for most characters. Instead of lateral processes with rounded tubercles, as represented in Child’s and Müller & Krapp’s specimens, Madibenthos specimens show distal angular thickening. Anoplodactylus califorinicus was already recorded in Martinique (as A. carvalhoi) by Bourdillon (1955) and Müller (1990 a), but Bourdillon’s illustrations show differences from the Madibenthos material. For example, the ocular tubercle is somehow lanceolate (narrower at the base, broader at mid-length, with tapering tip) in the Madibenthos material, while it is represented as conical by Bourdillon. We note that the ocular tubercle shape is highly variable in the illustrations of different authors, more or less tapering, conical or rounded (e. g., Hall 1912; Child 1992 b; Müller & Krapp 2009; Weis & Melzer 2012). Regarding molecular data, p-distances from some previously sequenced material from the Colombian Caribbean identified as A. californicus (GenBank reference DQ 390068; Arango & Wheeler 2007) are relatively low (0.047). Therefore, a revision of this widespread and multisynonymised species is needed. Until then, we only allegedly link Madibenthos material to this species as Anoplodactylus cf. californicus.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E51FFE0FDE6032700EDBAEA.taxon	distribution	Distribution Recorded in the Eastern Pacific (California, Ecuador, Mexico, Chile, Costa Rica, Nicaragua, Panama) and in Hawaii and the Panama Canal. It is also often recorded in the Caribbean Sea (Belize, Caribbean Panama, Bahamas, Martinique, Colombia, Costa Rica, Curaçao, Bonaire, Saint Kitts and Nevis, Virgin Islands, Porto Rico), the Gulf of Mexico and the Western Atlantic in Brazil and Argentina southward to the Magellan Strait. It is also recorded from the Mediterranean, in the Thyrrenean and Levantine Seas, as well as in the Suez Canal.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E51FFE0FDE6032700EDBAEA.taxon	description	Depth range Intertidal – 100 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4EFFE2FDD307DE0084B957.taxon	materials_examined	Type material Phoxichilidium (Anoplodactylus) digitatum Böhm, 1879. Holotype: ZMB _ Pyc _ 12 (not examined). Type locality: Singapore. Anoplodactylus saxatilis Calman, 1923. Type (s): unknown. Type locality: Marble Rocks, Mergui Archipelago, Myanmar. Anoplodactylus investigatoris Calman, 1923. Type (s): unknown. Type locality: Madras, outside harbour, India. Material examined MARTINIQUE • 1 ♂; Baie de Fort-de-France; 14 ° 36.1 ʹ N, 61 ° 04 ʹ W; depth 0 – 1 m; 29 Sep. 2016; st. AM 034; MNHN-IU- 2016 - 875 • 1 ♀; same collection data as for preceding; MNHN-IU- 2016 - 876 / MK 411186 • 1 ♂ ov.; same collection data as for preceding; MNHN-IU- 2016 - 1152 / MK 411065 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1328.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4EFFE2FDD307DE0084B957.taxon	discussion	Remarks This species is widely distributed through the Indian Ocean, with some records in the eastern Mediterranean (Arango & Maxmen 2006). On the other hand, records of this species in Atlantic are rare. It may have been found near the Azores, though the record is unusually deep for the species, and Arnaud (1974) mentioned that the sampled female was “ close to A. digitatus, without being identifical ” without further explanation. The species was already known from Martinique, only recorded from the port of Fort-de-France (Bourdillon 1955; confirmed by the present study).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4EFFE2FDD307DE0084B957.taxon	distribution	Distribution Tropical Indo-Pacific (Indonesia, Madagascar, India, Myanmar, Malaysia, Oman, Singapore, Great Barrier Reef), Eastern Mediterranean and Marmara Sea. Some records from Martinique and Azores (?).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4EFFE2FDD307DE0084B957.taxon	description	Depth range Intertidal to 75 m (most records between 0 and 5 m); one record of Anoplodactylus cf. digitatus in Arnaud (1974) at 600 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4CFFE3FDCF046900F5BC25.taxon	materials_examined	Type material Type (s): not designated? (see Lucena & Christoffersen 2018 a). Type locality: Ilha das Palmas, São Paulo, Brazil. Material examined MARTINIQUE • 1 ♂ ov.; Le Lorrain; 14 ° 50.7 ʹ N, 61 ° 03.8 ʹ W; depth 11 – 14 m; 6 Oct. 2016; st. AS 565; MNHN-IU- 2016 - 1116 • 1 ♂; Le François; 14 ° 37 ʹ N, 60 ° 49.5 ʹ W; depth 1 m; 17 Sep. 2016; st. AB 134; MNHN-IU- 2016 - 1233 / MK 411084.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4CFFE3FDCF046900F5BC25.taxon	discussion	Remarks First record from Martinique. The two specimens were sampled on the Atlantic coast.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4CFFE3FDCF046900F5BC25.taxon	distribution	Distribution Tropical West Atlantic (Gulf of Mexico, Caribbean, Brazil).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4CFFE3FDCF046900F5BC25.taxon	description	Depth range Intertidal to 30 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4DFFE4FE4D0197006BBE97.taxon	materials_examined	Material examined MARTINIQUE • 1 ♀; Baie de Fort-de-France; 14 ° 36.1 ʹ N, 61 ° 04 ʹ W; depth 0 – 1 m; 29 Sep. 2016; st. AM 034; MNHN-IU- 2016 - 854 / MK 411167 • 1 ♀; same collection data as for preceding; MNHN- IU- 2016 - 575 / MK 411117 • 1 ♀; same collection data as for preceding; MNHN-IU- 2016 - 1413 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1414. Type material Holotype: UFPB PYC- 242 (not examined). Type locality: Jericoacoara, Ceará, Brazil.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4DFFE4FE4D0197006BBE97.taxon	discussion	Remarks The specimens were mistakenly listed as Anoplodactylus stictus Marcus, 1940 in Sabroux et al. (2019 b). The distance between the lateral processes and the shape of proboscis extensions (alar rather than cordiform) do not match the descriptions in Marcus (1940) or Lucena & Christoffersen (2018 b). The specimens from Martinique greatly resemble the description of Anoplodactylus ganchiformis Lucena & Christofersen, 2018 from Brazil, including the conical ocular tubercle, the more elongated trunk relative to other species of the Anoplodactylus californicus-digitatus species complex of Arango & Maxmen (2006), the shape of all the processes and the hook at the base of the third article of the oviger. The dorsodistal tubercles of the lateral processes are, however, much lower in Martinique specimens. The specimen MNHN-IU- 2016 - 1413 shows two differences compared to the other specimens assigned to A. ganchiformis: it is slightly more elongated and has completely inconspicuous lateral processes on the dorsodistal tubercles. No CO 1 data could be sequenced for this specimen. All these specimens were collected on ship hulks in the Baie de Fort-de-France.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4DFFE4FE4D0197006BBE97.taxon	distribution	Ditribution Previously recorded in Brazil, and now in Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4DFFE4FE4D0197006BBE97.taxon	description	Depth range 0 – 1 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4AFFE5FDD6002900F5BC2B.taxon	materials_examined	Type material Anoplodactylus glandulifer Stock, 1954. Holotype: NHMD 110667 (material missing; not examined). Type locality: Singapore. Anoplodactylus multiclavus Child, 1977. Holotype: USNM 154807 (not examined). Type locality: Pillsbury Sound, Vessup Bay, St Thomas, Virgin Islands. Anoplodactylus dauphinus Child, 1992. Holotype: USNM 212648 (not examined). Type locality: NW of Clearwater, Florida Middle Ground, 28 ° 32 ʹ N, 84 ° 18 ʹ W. Material examined MARTINIQUE • 1 ♀ gr.; Baie de Fort-de-France; 14 ° 36.1 ʹ N, 61 ° 04 ʹ W; depth 0 – 1 m; 29 Sep. 2016; st. AM 034; MNHN-IU- 2016 - 877 / MK 411187 • 1 ♂ ov.; same collection data as for preceding; MNHN- IU- 2016 - 1148 • 1 ♂ ov.; Baie de Fort-de-France; 14 ° 33 ʹ N, 61 ° 03.7 ʹ W; depth 2 m; 7 Sep. 2016; st. AB 104; MNHN-IU- 2016 - 901 • 1 ♂; SE of Pointe Cerisier; 14 ° 35.6 ʹ N, 60 ° 50.8 ʹ W; depth 2 – 3 m; 15 Sep. 2016; st. AD 235; MNHN-IU- 2016 - 1111 • 1 ♂; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 00.1 ʹ W; depth 12 m; 26 Sep. 2016; st. AB 360; MNHN-IU- 2016 - 1401 • 1 ♀; same collection data as for preceding; MNHN-IU- 2016 - 1402.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4AFFE5FDD6002900F5BC2B.taxon	discussion	Remarks Müller & Krapp (2009) suggested that Anoplodactylus dauphinus and A. multiclavus are two junior synonyms of this species. The only specimen for which barcoding worked is a female (MNHN- IU- 2016 - 877), meaning that it was not possible to validate its identification by the observation of cement glands. However, females attributed to this species are similar to identified males for all other characters (including length of the sole lamina, presence of a propodal heel, shape of the ocular tubercle, shape of chelae carrying teeth, constriction near the tip of the proboscis), which in turn fit with the descriptions of Stock (1954 b) and Müller & Krapp (2009). In addition, both females were collected at localities where males of A. glandulifer were found. We are therefore confident in the attribution of the CO 1 barcode sequence MK 411187 to the species A. glandulifer. This species is mostly known from the Indo-Pacific, although it has regularly been sampled in the Western Atlantic. This is the first record of this species from Martinique; the species was collected on the Atlantic and Caribbean coasts, and in the Baie de Fort-de-France.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4AFFE5FDD6002900F5BC2B.taxon	distribution	Distribution Pantropical: Tropical West Atlantic (Caribbean, Gulf of Mexico), Indian Ocean (Aldabra, Madagascar, Seychelles, Kenya, Oman, Malaysia, Singapore), Australia (Great Barrier Reef and Timor Sea), Southern China Sea (Hong Kong), Pacific (Samoa, Guam, Marshall Islands, Marianas Island).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4AFFE5FDD6002900F5BC2B.taxon	description	Depth range Intertidal to 78 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4BFFE6FDB70195036EBE0C.taxon	materials_examined	Type material Holotype: USNM 171122 (not examined). Type locality: Tobacco Reef, 500 m N of South Water Cay, Carrie Bow Cay, Belize. Material examined MARTINIQUE • 1 ♂; Pointe du Vauclin; 14 ° 33.6 ʹ N, 60 ° 49.7 ʹ W; depth 2 m; 12 Sep. 2016; st. AD 230; MNHN-IU- 2016 - 900 • 1 ♀; NE of Pointe Jacob; 14 ° 36.1 ʹ N, 60 ° 49 ʹ W; depth 23 m; 17 Sep. 2016; st. AB 185; MNHN-IU- 2016 - 1102 / MK 411050 • 1 ♀; Le Lorrain; 14 ° 50.7 ʹ N, 61 ° 03.8 ʹ W; depth 11 – 14 m; 6 Oct. 2016; st. AS 565; MNHN-IU- 2016 - 1117 • 1 ♂; NE of Pointe du Vauclin; 14 ° 34.9 ʹ N, 60 ° 48.4 ʹ W; depth 18 – 20 m; 26 Sep. 2016; st. AS 409; MNHN-IU- 2016 - 1205 / MK 411081 • 1 ♂ ov.; same collection data as for preceding; MNHN-IU - 2017 - 215 • 1 ♂, 1 ♀; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 29 m; 18 Sep. 2016; st. AS 253; MNHN-IU- 2016 - 1262.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4BFFE6FDB70195036EBE0C.taxon	discussion	Remarks This is the first record of this species since its original description from Belize; it was only collected on the Atlantic coast of Martinique. Males are identical to those described by Child (1982 a), except that the ventral spur of coxae 2 on the third and fourth legs are generally slightly less marked than shown by Child, although it remains very prominent. Females have no ventral spurs on coxae 2 of the third and fourth legs, but a conspicuous ventral bulge is present on coxae 2 of all four legs, on the top of which stand the gonopores. We note no other differences from males apart from sexual characters.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4BFFE6FDB70195036EBE0C.taxon	distribution	Distribution Known from Belize and Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E4BFFE6FDB70195036EBE0C.taxon	description	Depth range Intertidal, extended to 23 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E48FFE6FDCA03B90361B942.taxon	materials_examined	Type material Holotype: NHMUK ZOO- 1881.38 (not examined). Type locality: off Bahia, Brazil. Material examined MARTINIQUE • 1 ♀; Canal de Ste Lucie; 14 ° 24.46 ʹ N, 60 ° 53.06 ʹ W; depth 60 m; 11 Sep. 2016; st. AD 226; MNHN-IU- 2016 - 808 / MK 411130 • 1 ♂; Macouba; 14 ° 55.1 ʹ N, 61 ° 09.2 ʹ W; depth 100 m; 24 Sep. 2016; st. AD 262; MNHN-IU- 2016 - 809 / MK 411131.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E48FFE6FDCA03B90361B942.taxon	discussion	Remarks This is the first record of this species from Martinique. One specimen was collected in the north and another in the south.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E48FFE6FDCA03B90361B942.taxon	distribution	Distribution Widespread in the Western Atlantic: North Carolina, Georgia, Florida, Strait of Florida, Caribbean, Yucatan Channel, Gulf of Mexico, Guiana, Surinam, French Guiana, Brazil. Depth range 0 – 90 m, extended to 100 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E48FFE7FDB9044E0426BC70.taxon	materials_examined	Type material Holotype: NHMD 110625 (not examined). Type locality: 800 m off Holetown shore, Barbados, West Indies. Material examined MARTINIQUE • 1 ♀; Bellefontaine; 14 ° 40.4 ʹ N, 61 ° 10.6 ʹ W; depth 70 m; 4 Oct. 2016; st. AD 298; MNHN-IU- 2016 - 1273 / MK 411093.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E48FFE7FDB9044E0426BC70.taxon	discussion	Remarks Anoplodactylus justi has not been found since original description and is known from only one male specimen from Barbados. According to Müller (1992 a), the best criterion for identification of A. justi relies on the shape and position of the cement gland tubes, but these are only present in males. Other characters correspond to Müller’s description.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E48FFE7FDB9044E0426BC70.taxon	distribution	Distribution West Indies. Apart from the present record, known only from the type locality in Barbados.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E48FFE7FDB9044E0426BC70.taxon	description	Depth range Anoplodactylus justi is known to live at a depth of 54 m. The present collection was made at 70 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E49FFEAFDC701440079BC71.taxon	description	urn: lsid: zoobank. org: act: 0 E 6 D 16 F 7 - B 75 F- 4 BF 7 - B 30 C- 934203 E 22322 Figs 22 A – J, 23	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E49FFEAFDC701440079BC71.taxon	materials_examined	Material examined Holotype MARTINIQUE • ♂; Baie du Robert; 14 ° 42 ʹ N, 60 ° 53.8 ʹ W; depth 2 m; 24 Sep. 2016; st. AB 452; MNHN- IU- 2016 - 1071 / MK 411043. Paratypes MARTINIQUE • 1 ♀; Les Anses-d’Arlet; 14 ° 29.6 ʹ N, 61 ° 05.5 ʹ W; depth 8 – 12 m; 7 Sep. 2016; st. AR 103; MNHN-IU- 2016 - 807 / MK 411129 • 1 ♀; Pointe Michel; 14 ° 26.4 ʹ N, 60 ° 49.3 ʹ W; depth 2 m; 12 Sep. 2016; st. AB 120; MNHN-IU- 2016 - 537 • 1 juv.; Case-Pilote; 14 ° 38.9 ʹ N, 61 ° 09.1 ʹ W; depth 0 – 14 m; 27 Sep. 2016; st. AS 365; MNHN-IU- 2016 - 1078 • 1 ♀; Trois Rivières; 14 ° 27.8 ʹ N, 60 ° 57.9 ʹ W; depth 3 – 4 m; 25 Sep. 2016; st. AB 356; MNHN-IU- 2016 - 1202 / MK 411079 • 1 ♂ ov., 1 ♀; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 00.1 ʹ W; depth 12 m; 26 Sep. 2016; st. AB 360; MNHN-IU- 2016 - 1235 • 1 ♂; same collection data as for preceding; MNHN-IU- 2019 - 3401 • 1 juv.; Baie du Robert; 14 ° 42 ʹ N, 60 ° 53.8 ʹ W; depth 1 – 3 m; 24 Sep. 2016; st. AR 453; MNHN-IU- 2016 - 1321 • 1 ♂, 1 ♀, 4 juvs; Ste-Luce; 14 ° 27.3 ʹ N, 60 ° 55.5 ʹ W; depth 15 m; 10 Sep. 2016; st. AB 062; MNHN-IU- 2016 - 1453.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E49FFEAFDC701440079BC71.taxon	etymology	Etymology Apposition. Named after the Madibenthos Expedition.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E49FFEAFDC701440079BC71.taxon	description	Description (holotype, ♂, MNHN-IU- 2016 - 1071) BODY. Small-sized species, slender, cuticle granular. Trunk incompletely segmented with segments 3 and 4 fused. No dorsal ornamentation. Post-ocular neck short. Ocular tubercle about twice as tall as wide, with rounded tip and four pigmented eyes. 1 st and 2 nd lateral processes about 1.5 times as long as wide, 3 rd and 4 th articles about as long as wide, all well separated by about their own diameter, without ornamentation. PROBOScIS. Elongated and cylindrical, widening at mid-length; proboscis tip not reaching further than chelifore scape. ABDOMEN. Long, tapering distally, oriented diagonally. No basal articulation. CHELIFORE. 2 - articled, fingers reaching not far beyond mouth. Scape 1 - articled, slender, about 5 times as long as wide. Chela small, palm and fingers of about same size. Fingers curved, subequal, with few teeth: four teeth on movable finger, six on immovable finger. PALP. Absent. OvIGER. 6 - articled. 1 st article about as long as wide. 2 nd article about 3 times as long as wide, carrying scarce setae. 3 rd article longest, gently curved, about 1.5 times as long as 2 nd and 10 times as long as wide, with scarce setae. 4 th article gently curved, about 3 times as long as wide, with setae on outer surface. 5 th article about ⅔ length of 4 th, less than 3 times as long as wide, with scarce setae on inner and outer surfaces. 6 th article shortest, about ⅔ length of 5 th and twice as long as wide, carrying setae on inner surface. LEGS. Slender. Coxae 1 and 3 subequal, slightly longer than wide, coxa 1 carrying dorsodistal setae. Coxa 2 about 3 times as long as basal width, slightly shorter than coxae 1 and 3 together; small spur present on ventral surface near distal margin carrying gonopore on 3 rd and 4 th legs. Coxa 3 carrying ventral setae. Femur, tibia 1, tibia 2 and propodus longer on 1 st and 2 nd legs than on 3 rd and 4 th legs. Femur widening distally, about 5 times (1 st leg) and 4 times (3 rd leg) as long as distal width, carrying scarce setae; one dorsal spur on distal margin carrying one long seta. Cement gland pore carried on femur as low, straight and elongated aperture. Tibia 1 curved, about 10 times (1 st leg) and 7 times (3 rd leg) as long as basal width and subequal to femur, carrying many setae along and one long dorsal seta at distal margin. Tibia 2 10 times (1 st leg) and 8 times (3 rd leg) as long as wide and subequal to femur and tibia 1, with several setae along and one long dorsal seta on distal fourth. Tarsus short, about as long as wide, carrying setae and one ventral spine on distal margin. Propodus about 0.6 times as long as tibia 2 and about 4 times as long as wide, straight, with setae and one long seta distally; heel carrying 2 heel spines, the distal larger and pectinated; sole carrying 12 short spines and very short lamina distally. Main claw reaching heel spines, slightly curved, slenderer and longer on 1 st and 2 nd legs than on 3 rd and 4 th legs. No auxiliary claw. MEASUREMENTS (mm). Trunk 1.06; abdomen 0.40; proboscis 0.57; chelifore 0.55; coxa 1 0.23; coxa 2 0.45; coxa 3 0.28; femur 1.23 (1 st leg), 1.04 (3 rd leg); tibia 1 1.12 (1 st leg), 0.93 (3 rd leg); tibia 2 1.08 (1 st leg), 0.93 (3 rd leg); tarsus 0.17 (1 st leg), 0.15 (3 rd leg); propodus 0.67 (1 st leg), 0.57 (3 rd leg); main claw 0.56 (1 st leg), 0.42 (3 rd leg). Sexual dimorphism Females larger, without oviger, femur moderately inflated, without cement gland. All four pairs of coxa 2 carrying gonopore on ventral surface, at the tip of a low distal bulge (Fig. 23).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E49FFEAFDC701440079BC71.taxon	discussion	Remarks The propodal pectinated spine is observed in five species: Anoplodactylus madibenthos sp. nov., A. exageratus Stock, 1994 from Indonesia, A. perissoporus Arango & Krapp, 2007 from Australia, A. tenuicorpus Child, 1991 from the Indoacific and A. pectinus Hedgpeth, 1948, which is included in the present report. The three former species, which belong to the same tenuicorpus complex, differ a lot from A. madibenthos by their extreme slenderness and the ovigers which articulate with the first pair of lateral processes. The species Anoplodactylus pectinus (Fig. 22 K – L) is morphologicaly the closest relative to A. madibenthos sp. nov. Anoplodactylus pectinus was sampled in Martinique, and we also consulted the holotype of Hedgpeth (1948). The distance between the lateral processes is larger in A. pectinus (peculiarly between segments 2 and 3), and the proboscis is distally widened in A. pectinus, whereas it is cylindrical in A. madibenthos. The CO 1 analysis also supported a deep divergence between the two species (intraspecific p-distance = 0 – 0.1, interspecific p-distance = 0.174 – 0.180) (Sabroux et al. 2019 b; see Appendix). Specimens were collected on both the Atlantic and Caribbean coasts.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E49FFEAFDC701440079BC71.taxon	distribution	Distribution Only known from Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E49FFEAFDC701440079BC71.taxon	description	Depth range 2 – 15 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E45FFEBFDE003270048B97E.taxon	materials_examined	Type material Anoplodactylus maritimus Hodgson, 1914. Type (s): unknown. Type locality: Sargasso Sea. Anoplodactylus parvus Giltay, 1934. Holotype: USNM 72790 (not examined). Type locality: Bermuda. Material examined MARTINIQUE • 1 ♂; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2016 - 1140.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E45FFEBFDE003270048B97E.taxon	discussion	Remarks This is the first record of this species in Martinique, sampled off Presqu’Île de la Caravelle.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E45FFEBFDE003270048B97E.taxon	distribution	Distribution Amphi-Atlantic, including Virginia, Bermuda, the Caribbean, Gulf of Mexico, Brazil, Sargasso Sea, Middle Atlantic and Macaronesia (Madeira, Canary Islands, Azores).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E45FFEBFDE003270048B97E.taxon	description	Depth range 0 – 120 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E45FFECFDE0044A0048BC0F.taxon	materials_examined	Type material Holotype: NL ZMA. PYC. P. 1091 (not examined). Type locality: off the mouth of the Amazon River, Brazil. Material examined MARTINIQUE • 1 ♂; Rocher du Diamant; 14 ° 26.5 ʹ N, 61 ° 02.4 ʹ W; depth 26 – 32 m; 14 Sep. 2016; st. AS 071; MNHN-IU- 2016 - 1051 / MK 411037 • 1 ♂; Anse Couleuvre; 14 ° 50.4 ʹ N, 61 ° 13.4 ʹ W; depth 7 m; 1 Oct. 2016; st. AB 463; MNHN-IU- 2016 - 1124 / MK 411055 • 1 juv.; Trois Rivières; 14 ° 27.5 ʹ N, 60 ° 58.2 ʹ W; depth 17 – 19 m; 9 Oct. 2016; st. AS 579; MNHN-IU- 2016 - 1128 / MK 411057 • 1 juv.; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 29 m; 18 Sep. 2016; st. AS 253; MNHN-IU- 2016 - 1263.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E45FFECFDE0044A0048BC0F.taxon	discussion	Remarks This is the first record of this species in Martinique. The four specimens were collected at very different locations around the Island, on both the Atlantic and Caribbean coasts.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E45FFECFDE0044A0048BC0F.taxon	distribution	Distribution Brazil, French Guiana, Guyana, Caribbean (Barbados, Martinique, Panama, Aruba), Panama Canal.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E45FFECFDE0044A0048BC0F.taxon	description	Depth range 0 – 100 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E42FFEFFDD501A00079BB6C.taxon	description	Figs 20, 24	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E42FFEFFDD501A00079BB6C.taxon	materials_examined	Material examined Neotype MARTINIQUE • ♂; Les Anses-d’Arlet; 14 ° 29.7 ʹ N, 61 ° 05.4 ʹ W; depth 19 m; 7 Sep. 2016; st. AB 155; MNHN-IU- 2016 - 1325 / MK 411100. Other material MARTINIQUE • 1 ♀, 3 juvs; same collection data as for neotype; MNHN-IU- 2016 - 1230 • 1 ♀; same collection data as for neotype; MNHN-IU- 2016 - 1387 / MK 411110 • 1 ♀; same collection data as for neotype; MNHN-IU- 2016 - 1388 • 1 ♂; Anse Noire; 14 ° 32 ʹ N, 61 ° 05.3 ʹ W; depth 13 m; 6 Sep. 2016; st. AB 150; MNHN-IU- 2016 - 802 / MK 411125 • 1 ♂; Grande Anse d’Arlets; 14 ° 30.5 ʹ N, 61 ° 06.1 ʹ W; depth 20 – 23 m; 6 Sep. 2016; st. AB 152; MNHN-IU- 2016 - 803 • 1 preadult ♀; Pointe du Vauclin; 14 ° 33.6 ʹ N, 60 ° 49.7 ʹ W; depth 2 m; 12 Sep. 2016; st. AD 230; MNHN-IU- 2016 - 804 / MK 411126 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 568 • 1 ♀; Grande Anse d’Arlets; 14 ° 29.9 ʹ N, 61 ° 05.4 ʹ W; depth 28 m; 7 Sep. 2016; st. AB 157; MNHN-IU- 2016 - 536 • 1 ♂, 1 ♂ ov.; Baie de St-Pierre; 14 ° 45.1 ʹ N, 61 ° 11 ʹ W; depth 17 m; 4 Oct. 2016; st. AB 388; MNHN-IU- 2016 - 1059 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1386 • 1 ♂; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 00.1 ʹ W; depth 12 m; 26 Sep. 2016; st. AB 360; MNHN-IU- 2016 - 1080 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1218 • 1 ♂; Le Prêcheur; 14 ° 49.1 ʹ N, 61 ° 13.8 ʹ W; depth 20 – 25 m; 8 Oct. 2016; st. AS 576; MNHN-IU- 2016 - 1087 • 1 ♂, 1 ♀; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2016 - 1139 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1250 • 1 preadult ♂; same collection data as for preceding; MNHN-IU- 2016 - 1391 • 1 ♀; Grande Anse du Diamant; 14 ° 27.9 ʹ N, 61 ° 01.4 ʹ W; depth 17 m; 26 Sep. 2016; st. AB 358; MNHN- IU- 2016 - 1211 • 1 ♀, 1 juv.; Anse des Galets; 14 ° 51.4 ʹ N, 61 ° 12.8 ʹ W; depth 10 m; 7 Oct. 2016; st. AB 567; MNHN-IU- 2016 - 1246 • 2 ♀♀, 4 ♀♀ gr.; Pointe de la Baleine; 14 ° 31.1 ʹ N, 61 ° 05.9 ʹ W; depth 17 – 19 m; 30 Sep. 2016; st. AB 369; MNHN-IU- 2016 - 1258 • 1 ♀; Anse Couleuvre; 14 ° 50.4 ʹ N, 61 ° 13.4 ʹ W; depth 7 m; 1 Oct. 2016; st. AB 463; MNHN-IU- 2019 - 3390.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E42FFEFFDD501A00079BB6C.taxon	description	Description (neotype, ♂, MNHN-IU- 2016 - 1325) BODY. Small-sized species; trunk segmentation incomplete with 3 rd and 4 th segments fused; cuticle smooth. No dorsomedian ornamentation. Ocular tubercle less than twice as tall as wide, distal tip triangular, with two pointy lateral sense organs above four large pigmented eyeseyes. Two setae laterally positioned on ocular tubercle between anterior and posterior eyes. Post-ocular neck medium-sized. Lateral processes about as long as wide, well separated by less than their own diameter, 3 rd and 4 th the closest. Lateral processes ornamented with a rounded tubercle, bearing one seta. PROBOScIS. Cylindrical, with setae on ventral surface, constricted medially and distally. ABDOMEN. Medium-sized, diagonally oriented, reaching beyond lateral processes of 4 th trunk segment. CHELIFORE. 2 - articled, reaching beyond mouth. Scape 1 - articled, about 4 times as long as wide, carrying setae distally. Chela carrying setae, fingers slightly longer than palm. Fingers subequal, crossing at tip, with few teeth. PALP. Absent. OvIGER. 6 - articled, with scarce setae. 1 st article about 1.5 times as long as wide. 2 nd article about ¼ longer than 1 st article, slenderer. 3 rd article longest, 1.5 times as long as 1 st, with constriction at base. 4 th and 5 th articles about half as long as 3 rd, 5 th most setose. 6 th article smallest, bud-like, with distal seta. LEGS. Stout. Coxa 1 slightly longer than wide, carrying setae at distal margin. Coxa 2 about ⅔ longer than coxa 1 or 3, with small ventrodistal spur on 3 rd and 4 th legs. Femur longest, more than 3 times as long as median width, with one long dorsal seta on distal margin, and one short cement gland tube, about mid-sized and slightly constricted at base. Tibia 1 about ¾ of femur length, with one long seta near distal margin. Tibia 2 about ⅔ of femur size, with one long seta on distal half rising from low tubercle. Tarsus short, trapezoid, with one ventrodistal spine. Propodus about 0.9 times as long as tibia 2, sole straight, with strong heel carrying 2 large heel spines followed by 2 smaller spines; sole carrying 2 small spines on proximal 3 rd directed distally. Lamina on distal part of sole, about ⅔ of sole length. Main claw almost reaching heel, straight with terminal bending. Auxiliary claws present, very small. MEASUREMENTS (mm). Trunk 0.44; abdomen 0.18; proboscis 0.25; chelifore 0.26; coxa 1 0.10; coxa 2 0.17; coxa 3 0.13; femur 0.41; tibia 1 0.32; tibia 2 0.3; tarsus 0.05; propodus 0.22; main claw 0.13. Sexual dimorphism Females with dilated femorae, no oviger. Individual variability Number of propodus heel spines variable, with 2 or 3 large spines and 2 or 3 smaller spines. Sole spines also variable in number, 1 or 2.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E42FFEFFDD501A00079BB6C.taxon	discussion	Remarks Because the holotype is considered to be lost (Müller & Krapp 2009 and MNHN collections data), we propose the selection of a new specimen from type locality (Martinique, southern Caribbean coast) to designate as a neotype. According to Müller & Krapp (2009), the two similar species Anoplodactylus micros and A. bahamensis can be distinguished based on the setae on the ventral side of the proboscis in A. bahamensis. However, both species exhibit setae in Madibenthos material. The species differ in two aspects: (i) the most obvious is the size of the specimens, the trunk of A. micros in the present material and in the original description being about half as long as A. bahamensis as measured by Child (1977 b); (ii) the cement gland opening is larger in A. bahamensis than in A. micros, although in some specimens this distinction may be tricky (Fig. 20). Regarding molecular data, the two species are clearly distinct (interspecific p-distance = 0.077 – 0.1). Interestingly, within the material identified as A. micros the presence of two clusters (Sabroux et al. 2019) with p-distances similar to that between A. micros and A. bahamensis (intercluster p-distance = 0.077 – 0.079) suggests that there may be at least two cryptic species. We thus informally describe two groups: group 1 includes the neotype, MNHN-IU- 2016 - 1325, and specimens MNHN-IU- 2016 - 802 and MNHN-IU- 2016 - 1387, and group 2 is only represented by MNHN-IU- 2016 - 804 (see Appendix for complete list). The species Anoplodactylus micros was originally collected in Petite Anse du Diamant on the southern Caribbean coast (Bourdillon 1955). Here, the species was mostly found on the Caribbean coast, except six specimens sampled at Pointe du Vauclin and Presqu’Île de la Caravelle.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E42FFEFFDD501A00079BB6C.taxon	distribution	Distribution So far only recorded from the Colombian Caribbean and Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E42FFEFFDD501A00079BB6C.taxon	description	Depth range 2 – 30 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E41FFD0FDD406400093BBE3.taxon	materials_examined	Type material Holotype: NL ZMA. PYC. P. 1076 (not examined). Type locality: Santa Marta Bay, Curaçaõ. Material examined MARTINIQUE • 1 ♀; Presqu’Île de la Caravelle; 14 ° 44.1 ʹ N, 60 ° 50.8 ʹ W; depth 16 m; 18 Sep. 2016; st. AB 189; MNHN-IU- 2016 - 797 / MK 411120 • 1 ♂ ov.; E of Le Vauclin; 14 ° 34.2 ʹ N, 60 ° 44.8 ʹ W; depth 90 m; 14 Sep. 2016; st. AD 231; MNHN-IU- 2016 - 890 • 1 ♂ ov.; Baie de Fort-de-France; 14 ° 34.3 ʹ N, 61 ° 02.7 ʹ W; depth 7 m; 7 Oct. 2016; st. AB 394; MNHN-IU- 2016 - 1046 / MK 411034 • 1 ♂ ov., 1 juv.; Le Robert; 14 ° 43.2 ʹ N, 60 ° 50.6 ʹ W; depth 23 m; 25 Sep. 2016; st. AB 405; MNHN-IU- 2016 - 1048 • 1 ♀; Ste-Marie; 14 ° 46.3 ʹ N, 60 ° 57.5 ʹ W; depth 20 m; 20 Sep. 2016; st. AB 260; MNHN-IU- 2016 - 1065 • 1 ♂; Baie du Robert; 14 ° 42 ʹ N, 60 ° 53.8 ʹ W; depth 2 m; 24 Sep. 2016; st. AB 452; MNHN-IU- 2016 - 1072 • 1 ♀; CasePilote; 14 ° 38.9 ʹ N, 61 ° 09.1 ʹ W; depth 14 m; 27 Sep. 2016; st. AS 365; MNHN-IU- 2016 - 1079 • 1 ♀; Le Prêcheur; 14 ° 49.1 ʹ N, 61 ° 13.8 ʹ W; depth 20 – 25 m; 8 Oct. 2016; st. AS 576; MNHN-IU- 2016 - 1086 • 1 ♂, 1 ♀; Case-Pilote; 14 ° 38.3 ʹ N, 61 ° 08.4 ʹ W; depth 12 m; 8 Sep. 2016; st. AB 159; MNHN-IU- 2016 - 1092 • 1 ♀; same collection data as for preceding; MNHN-IU- 2017 - 223. • 1 juv.; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.3 ʹ W; depth 19 – 21 m; 9 Sep. 2016; st. AS 057; MNHN-IU- 2016 - 1094 • 1 juv.; Banc du Diamant; 14 ° 26.4 ʹ N, 61 ° 01.7 ʹ W; depth 18 m; 15 Sep. 2016; st. AB 177; MNHN-IU- 2016 - 1106 • 1 ♂; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.4 ʹ W; depth 14 m; 14 Sep. 2016; st. AB 175; MNHN-IU- 2016 - 1130 • 2 ♂♂; Presqu’Île de la Caravelle; 14 ° 48.4 ʹ N, 60 ° 52.8 ʹ W; depth 23 – 25 m; 20 Sep. 2016; st. AB 197; MNHN-IU- 2016 - 1138 • 1 ♂; Schoelcher; 14 ° 37.7 ʹ N, 61 ° 07.2 ʹ W; depth 4 m; 27 Sep. 2016; st. AB 460; MNHN-IU- 2016 - 1155 / MK 411066 • 1 ♀; Rocher du Diamant; 14 ° 26.5 ʹ N, 61 ° 02.4 ʹ W; depth 24 m; 14 Sep. 2016; st. AB 173; MNHN-IU- 2016 - 1157 • 1 ♂; Pointe Michel; 14 ° 26.4 ʹ N, 60 ° 49.3 ʹ W; depth 2 m; 12 Sep. 2016; st. AB 120; MNHN-IU- 2016 - 1179 • 1 juv.; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 00.1 ʹ W; depth 12 m; 26 Sep. 2016; st. AB 360; MNHN-IU- 2016 - 1185 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1216 • 1 ♂; Grande Anse d’Arlets; 14 ° 30.5 ʹ N, 61 ° 06.1 ʹ W; depth 20 – 23 m; 6 Sep. 2016; st. AB 152; MNHN-IU- 2016 - 1191 • 1 ♂; Presqu’Île de la Caravelle; 14 ° 47.3 ʹ N, 60 ° 57.4 ʹ W; depth 19 m; 21 Sep. 2016; st. AB 301; MNHN-IU- 2016 - 1227 • 1 ♂; Grande Anse du Diamant; 14 ° 27.9 ʹ N, 61 ° 01.2 ʹ W; depth 15 m; 15 Sep. 2016; st. AB 179; MNHN-IU- 2016 - 1242 • 1 ♂ ov.; Presqu’Île de la Caravelle; 14 ° 46.5 ʹ N, 60 ° 51.5 ʹ W; depth 15 m; 22 Sep. 2016; st. AB 350; MNHN- IU- 2016 - 1301.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E41FFD0FDD406400093BBE3.taxon	discussion	Remarks This species was first recorded in Martinique by Müller (1990 a) from Petite Anse Macabou and was abundantly sampled in the present study.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E41FFD0FDD406400093BBE3.taxon	distribution	Distribution Tropical West Atlantic: Gulf of Mexico, Caribbean, South Florida, Bahamas, Brazil. The distribution may be extended with misidentifications as Anoplodactylus robustus (Dohrn, 1881) in the Tropical Western Atlantic (Stock 1979). Depth range 0 – 41 m, extended to 90 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7EFFD1FDD906D50079BBDD.taxon	description	Fig. 22 K – L	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7EFFD1FDD906D50079BBDD.taxon	materials_examined	Type material Holotype: USNM 81095 (examined). Type locality: Loggerhead Key, Tortugas, Florida (Gulf of Mexico). Material examined MARTINIQUE • 1 ♂ ov.; Case-Pilote; 14 ° 38.3 ʹ N, 61 ° 08.4 ʹ W; depth 12 m; 8 Sep. 2016; st. AB 159; MNHN-IU- 2016 - 805 / MK 411127 • 1 ♂; Anse Noire; 14 ° 32 ʹ N, 61 ° 05.3 ʹ W; depth 13 m; 6 Sep. 2016; st. AB 150; MNHN-IU- 2016 - 806 / MK 411128 • 1 ♂; Passe du Marin; 14 ° 26.8 ʹ N, 60 ° 54.3 ʹ W; depth 15 m; 9 Sep. 2016; st. AB 165; MNHN-IU- 2016 - 897 • 1 ♂, 1 ♀; Grande Anse d’Arlets; 14 ° 29.9 ʹ N, 61 ° 05.4 ʹ W; depth 28 m; 7 Sep. 2016; st. AB 157; MNHN-IU- 2016 - 898 • 1 ♀, 2 juvs; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.3 ʹ W; depth 19 – 21 m; 9 Sep. 2016; st. AS 057; MNHN-IU- 2016 - 899 • 1 ♂, 1 ♀; SteLuce; 14 ° 27.3 ʹ N, 60 ° 55.5 ʹ W; depth 15 m; 10 Sep. 2016; st. AB 062; MNHN-IU- 2016 - 1040 • 1 ♀; Les Anses-d’Arlet; 14 ° 28.5 ʹ N, 61 ° 05.1 ʹ W; depth 29 m; 26 Sep. 2016; st. AD 271; MNHN-IU- 2016 - 1093 • 1 juv.; Trois Rivières; 14 ° 27.5 ʹ N, 60 ° 58.2 ʹ W; depth 17 – 19 m; 9 Oct. 2016; st. AS 579; MNHN- IU- 2016 - 1126 • 1 juv.; Fond Boucher; 14 ° 39.3 ʹ N, 61 ° 09.4 ʹ W; depth 14 m; 27 Sep. 2016; st. AS 363; MNHN-IU- 2016 - 1161 • 2 ♂♂, 2 juvs; Trois Rivières; 14 ° 27.5 ʹ N, 60 ° 58.2 ʹ W; depth 17 m; 9 Oct. 2016; st. AB 578; MNHN-IU- 2016 - 1167 • 1 ♂ ov.; Pointe de la Baleine; 14 ° 31.1 ʹ N, 61 ° 05.9 ʹ W; depth 17 – 19 m; 30 Sep. 2016; st. AB 369; MNHN-IU- 2016 - 1254 / MK 411086.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7EFFD1FDD906D50079BBDD.taxon	discussion	Remarks This species is reported for the first time from Martinique, where it was only sampled on the Caribbean coast. The species is morphologically close to A. madibenthos sp. nov., with the same characteristic pectinated propodal spine; however, this species is much more elongated, with a wider separation between the lateral processes.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7EFFD1FDD906D50079BBDD.taxon	distribution	Distribution Pantropical: Tropical West Atlantic (Caribbean, Gulf of Mexico, Florida), Indian Ocean (Madagascar, Indonesia), Easter Australia, Pacific Ocean (Philippines, Ryukyu Archipelago, Fiji Islands, Caroline Islands). It is possible that some of these records were mistaken with Anoplodactylus madibenthos sp. nov.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7EFFD1FDD906D50079BBDD.taxon	description	Depth range 0 – 34 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7FFFD3FDDE06EF0462BE87.taxon	materials_examined	Type material Phoxichilidium petiolatum Krøyer, 1844. Syntypes: NHMD 110693 and 110694 (material missing, not examined). Type locality: Øresundsbroen Strait, Denmark. Phoxichilidium mutilatum Frey & Leuckart, 1847. Type (s): unknown. Type locality: Helgoland, Germany. Anoplodactylus petiolatus ssp. hedgpethi Băcescu, 1959. Type (s): not designated. Type locality: West Atlantic. Anoplodactylus guyanensis Child, 1977. Holotype: USNM 154805 (not examined). Type locality: NE of Georgetown (07 ° 40 ʹ N, 57 ° 34 ʹ W), Guyana. Material examined MARTINIQUE • 1 ♂; Rocher du Diamant; 14 ° 26.7 ʹ N, 61 ° 02.3 ʹ W; depth 4 – 10 m; 14 Sep. 2016; st. AB 123; MNHN-IU- 2016 - 1069 • 1 ♀; Trois Rivières; 14 ° 27.5 ʹ N, 60 ° 58.2 ʹ W; depth 17 – 19 m; 9 Oct. 2016; st. AS 579; MNHN-IU- 2016 - 1127 / MK 411056 • 1 ♀; Bellefontaine; 14 ° 40.2 ʹ N, 61 ° 10 ʹ W; depth 40 m; 2 Oct. 2016; st. AD 286; MNHN-IU- 2016 - 1195 / MK 411078 • 1 ♂; Anse Couleuvre; 14 ° 50.4 ʹ N, 61 ° 13.4 ʹ W; depth 7 m; 1 Oct. 2016; st. AB 463; MNHN-IU- 2016 - 1382.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7FFFD3FDDE06EF0462BE87.taxon	discussion	Remarks This is the first record of this species from Martinique, only from the Caribbean coast. Anoplodactylus petiolatus subsp. hedgpethi was originally proposed by Băcescu (without designating type material) in order to mark the difference between the West Atlantic material described by Hedgpeth (1948) and Mediterranean / Black Sea material. Differences noted by Băcescu include thinner legs, different propodal armature of the propodal sole and the longer coxae 2 of the fourth legs. In this regard, our material seems to be similar to Hedgpeth’s material.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7FFFD3FDDE06EF0462BE87.taxon	distribution	Distribution Widespread in Atlantic: West Atlantic (off New York state, North Carolina, Georgia, Bahamas, Gulf of Mexico, Caribbean, Guyana, Surinam, French Guiana, Brazil, Uruguay, Argentina), East Atlantic (northern Norway, Northern Sea, English Channel, British Isles, Irish Sea, Celtic Sea, Galicia, Portugal, Gibraltar region, West African Atlantic, Cape Verde, Azores). It is also recorded from the entire Mediterranean, Marmara Sea and Black Sea. There are additional records from India and Chile.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7FFFD3FDDE06EF0462BE87.taxon	description	Depth range 4 – 1520 m; one record at 4825 m potentially linked to sampling contamination (Child 1982 b).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7DFFD3FE12003803D5BACB.taxon	materials_examined	Type material Holotype: MCZ CRU- 12259 (not examined). Type locality: near channel to Key West, Florida. Material examined MARTINIQUE • 1 ♂; Le François; 14 ° 37 ʹ N, 60 ° 49.5 ʹ W; depth 1 m; 17 Sep. 2016; st. AB 134; MNHN- IU- 2016 - 1143 / MK 411063 • 1 ♀; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 00.1 ʹ W; depth 12 m; 26 Sep. 2016; st. AB 360; MNHN-IU- 2016 - 1220.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7DFFD3FE12003803D5BACB.taxon	discussion	Remarks This is the first record of this species from Martinique. The two collected specimens were sampled at two localities on both the Caribbean and Atlantic coasts.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7DFFD3FE12003803D5BACB.taxon	distribution	Distribution So far only recorded from Key West (Florida), Caribbean Colombia, Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7DFFD3FE12003803D5BACB.taxon	description	Depth range 0.5 m. We extend the depth-range to 12 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7DFFD4FDD007FD0079BB96.taxon	materials_examined	Type material Syntype (?) (see Dunlop et al. 2007): ZMB _ Pyc _ 39 (not examined). Type locality: Naples Bay, Italy. Material examined MARTINIQUE • 1 ♀; Case-Pilote; 14 ° 38.3 ʹ N, 61 ° 08.4 ʹ W; depth 12 m; 8 Sep. 2016; st. AB 159; MNHN-IU- 2016 - 796 / MK 411119 • 1 ♂; same collection data as for preceding; MNHN-IU- 2016 - 1415 • 1 ♀; Case-Pilote; 14 ° 38.6 ʹ N, 61 ° 08.5 ʹ W; depth 8 m; 8 Sep. 2016; st. AB 108; MNHN-IU- 2016 - 798 / MK 411121 • 1 ♂; Vétiver; 14 ° 37.9 ʹ N, 61 ° 07.7 ʹ W; depth 10 m; 8 Sep. 2016; st. AS 054; MNHN- IU- 2016 - 1096 / MK 411048 • 1 ♀; Grande Anse du Diamant; 14 ° 28 ʹ N, 61 ° 00.1 ʹ W; depth 12 m; 26 Sep. 2016; st. AB 360; MNHN-IU- 2017 - 224.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7DFFD4FDD007FD0079BB96.taxon	discussion	Remarks This species is relatively rare compared to its closest relative, Anoplodactylus monotrema Stock, 1979; similar rarity was mentioned by Arnaud (1987) for the fauna of the Mediterranean Sea. Previous records of A. robustus from the Western Atlantic were regarded as doubtful or incorrect by Stock (1979), who suggested that most of the older records of this species were mistaken with A. monotrema. Child (1979), however, considered that the Brazilian specimens identified as Halosoma robustum by Marcus (1940) should be treated as A. robustus. All our specimens come from three localities on the Caribbean coast. They match very well with Stock’s description in showing angulous proboscis corners around the mouth, five-articled ovigers and multiple cement gland pores.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7DFFD4FDD007FD0079BB96.taxon	distribution	Distribution Most records are from the Western Mediterranean (Gibraltar Strait, Alboran Sea, Balearic Sea, Gulf of Lion, Tyrrhenian Sea). Additional records from Brazil, the Caribbean and Galápagos, some of which were regarded as possibly mistaken with Anoplodactylus monotrema by Stock (1979) (see remarks above).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7DFFD4FDD007FD0079BB96.taxon	description	Depth range 4 – 44 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7AFFD6FD60072A006BBE0C.taxon	description	Fig. 25	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7AFFD6FD60072A006BBE0C.taxon	materials_examined	Material examined MARTINIQUE • 1 juv.; Baie de Sans-Souci; 14 ° 34.7 ʹ N, 60 ° 50.7 ʹ W; depth 0 – 1 m; 9 Oct. 2016; st. AM 325; MNHN-IU- 2016 - 1100 / MK 411049 • 1 ♀; Trois Rivières; 14 ° 27.8 ʹ N, 60 ° 57.9 ʹ W; depth 3 – 4 m; 25 Sep. 2016; st. AB 356; MNHN-IU- 2016 - 1165 / MK 411068.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7AFFD6FD60072A006BBE0C.taxon	discussion	Remarks These specimens were collected at two localities on Atlantic and Caribbean coasts. This species differs from most of other Anoplodactylus by the combination of the following characters: lateral processes separated by more than their own diameter, less than twice; low rounded tubercles on the lateral processes; chela fingers simple; leg articles elongated; auxillary claw present, short; length of propodal lamina about half that of propodal sole. The species furthermore differs from A. dissitus Child, 2004 in showing no tubercle on coxae 1; from A. trispinosus Stock, 1951 in having slenderer leg articles (Stock 1975 a; Child 2004; Müller & Krapp 2009). This species could be assigned to A. petiolatus (Krøyer, 1864), A. arcuatus Child, 1977 or another undescribed species. Without male specimens it is, however, impossible to provide a definitive conclusion.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E7AFFD6FD60072A006BBE0C.taxon	description	Depth range 0 – 3 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E78FFD6FD6003B90064BBBA.taxon	description	Fig. 26	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E78FFD6FD6003B90064BBBA.taxon	materials_examined	Material examined MARTINIQUE • 1 ♂; Basse Pointe; 14 ° 53.1 ʹ N, 61 ° 06.9 ʹ W; depth 35 m; 10 Oct. 2016; st. AD 626; MNHN-IU- 2016 - 1163 / MK 411067.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E78FFD6FD6003B90064BBBA.taxon	discussion	Remarks This specimen differs from other species of Anoplodactylus by the combination of the following characters: lateral processes separated by about their own diameter or more, less than twice their own diameter, and with one low dorsomedian tubercle; chela fingers simple; propodal lamina about half of propodal length; cement gland tube as a medium dorsomedian spur slightly anterior to half length of femur; propodal auxillary claw present, minute. The single available specimen was collected in northern Martinique. It is damaged, with the ocular tubercle missing. It may be related to Anoplodactylus stictus Marcus, 1940, which it resembles in the characters above and in showing a constriction at the base of the 3 rd article oviger (Lucena & Christoffersen 2018 b), but several other characters show differences with the original description (Marcus 1940), including the position of the cement gland tubes and the more elongated lateral processes, post-ocular neck and oviger articles.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E78FFD6FD6003B90064BBBA.taxon	description	Depth range 35 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E78FFD9FD6007060048BD5B.taxon	description	Fig. 27	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E78FFD9FD6007060048BD5B.taxon	materials_examined	Material examined MARTINIQUE • 1 ♀; Canal de Ste Lucie; 14 ° 23.1 ʹ N, 60 ° 50.2 ʹ W; depth 65 m; 11 Sep. 2016; st. AD 224; MNHN-IU- 2016 - 539 / MK 411113 • 1 ♀; Baie de St-Pierre; 14 ° 45.1 ʹ N, 61 ° 11 ʹ W; depth 17 m; 4 Oct. 2016; st. AB 388; MNHN-IU- 2016 - 1060 / MK 411039.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E78FFD9FD6007060048BD5B.taxon	discussion	Remarks These two specimens were collected on the Caribbean coast and southern Martinique; only females are available, preventing us from providing a complete description. The specimens differ from other species of Anoplodactylus by the combination of the following characters: body stout; ocular tubercle low, rounded; trunk without dorsomedian ornamentation; lateral processes well separated by less than their own diameter, and without ornamentation; proboscis barrel-shaped, truncated; legs stout; no propodal lamina; auxillary claw minute. The specimens strongly resemble A. pycnosoma (Helfer, 1938), a species only recorded in the Indo-Pacific (Bamber 2000), from which it differs by the absence of angulate distal projections on the proboscis; it also differs from A. dentimanus Stock, 1979 from the West Indies (Stock 1979) by the greater space between the lateral processes, the more elongate chela palms and the truncated proboscis end. Finally, the specimens are similar to Anoplodactylus nanus Krapp, Koçak & Katagan, 2008 from the Eastern Mediterranean (Krapp et al. 2008), except that their low dorsodistal femoral tubercles carry one seta.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E78FFD9FD6007060048BD5B.taxon	description	Depth range 17 – 65 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E77FFD9FDB600BC0271BC7B.taxon	type_taxon	Type species Pentapycnon charcoti Bouvier, 1910 by monotypy.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E77FFD9FDAC014503D8B892.taxon	materials_examined	Type material Syntypes: MNHN PY 566 - 569 (examined). Type locality: Cayenne, French Guiana. Material examined MARTINIQUE • 1 ♀; Pointe Baham; 14 ° 24.7 ʹ N, 60 ° 50.1 ʹ W; depth 2 m; 11 Sep. 2016; st. AB 117; MNHN-IU- 2016 - 4187 / MK 411112.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E77FFD9FDAC014503D8B892.taxon	discussion	Remarks Pentapycnon geayi is one of the few polymerous species in the world, and one of two known outside of the Southern Ocean, with the other being Pentacolossendeis reticulata Hedgpeth, 1943 (Arnaud & Bamber 1987). The two other known species of Pentapycnon have only been reported from the Southern Ocean (Child 1995). The species is here recorded for the first time in Martinique. The single specimen was collected on the southern Atlantic coast of Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E77FFD9FDAC014503D8B892.taxon	distribution	Distribution Recorded from Tropical West Atlantic (Puerto Rico, French Guiana, Surinam, Brazil) and Spanish Mediterranean coast (Gibraltar and Alboran Sea).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E77FFD9FDAC014503D8B892.taxon	description	Depth range 5 – 70 m. We extend this depth range to 2 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E74FFDAFDBF0326027CBECD.taxon	type_taxon	Type species Pycnogonum littorale (Strøm, 1762), by monotypy.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E74FFDCFD9A03FE0195B89A.taxon	description	urn: lsid: zoobank. org: act: FEDED 36 A-EFA 3 - 413 B- 8 E 4 E-BE 4010 D 1 A 5 AE Fig. 28	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E74FFDCFD9A03FE0195B89A.taxon	materials_examined	Material examined Holotype MARTINIQUE • ♂; Grand’Rivière; 14 ° 52 ʹ N, 61 ° 11.7 ʹ W; depth 6 m; 3 Oct. 2016; st. AB 556; MNHN- IU- 2016 - 10338 / MK 411029.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E74FFDCFD9A03FE0195B89A.taxon	etymology	Etymology Latinized name, 2 nd decl. (masc.), genitive singular. Named after the Martinique poet Aimé Césaire.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E74FFDCFD9A03FE0195B89A.taxon	description	Description (holotype, ♂, MNHN-IU- 2016 - 10338) BODY. Trunk completely segmented, stout, reticulated. Dorsomedian tubercles on 1 st to 3 rd segments, rounded, very granulous, carrying small setae. Ocular tubercle slightly higher than dorsomedian tubercles, rounded, carrying four pigmented eyes. Lateral processes shorter than wide, separated by a narrow space all along their length, with bead at their distal margin and one spine on posterior side. PROBOScIS. About as long as distance from 1 st to 3 rd trunk segments, gently bent downward, tapering, truncated, very granular getting smoother at its tip, with four rows of spines (2 lateral, 2 dorsolateral) on proximal half, each separated at regular distance. ABDOMEN. Trapezoidal and angulous dorsally, larger at tip than base. CHELIFORE. Absent. PALP. Absent. OvIGER. 8 - articled, cuticle reticulated. 1 st article shortest, about as long as wide, fused with 2 nd. 2 nd article larger than 1 st, as long as wide. 3 rd article shorter than 2 nd, about as long as wide, with distal blunt spine on inner surface. 4 th and 5 th articles subequal and longest, 5 th carrying a blunt distal spine on inner surface. 6 th, 7 th and 8 th articles subequal, about 0.8 times as long as 4 th or 5 th. 6 th and 7 th articles with blunt distal spine on inner surface, 8 th article with 3 blunt spines on distal half. Terminal claw as long as 8 th article. LEGS. Stout, cuticle reticulated. Coxa 1 very wide dorsally, wider than long, with large bead at distal margin carrying some tiny spines, forming low dorsodistal tubercle on 4 th leg. Coxa 2 wider than long, bearing several tiny spines, with white coxal pellicula on dorsal surface. Coxa 3 smaller than 1 or 2, bearing some tiny normal spines. Femur and tibia 1 subequal; femur about 1.4 times as long as wide, dorsally granulated on distal half; on ventral and dorsal surface, several blunt or slightly bifurcated spines mounted on tiny tubercles, plus tiny normal spines; dorsal side of distal margin forming short rounded spurs. Tibia 1 about 1.6 times as long as wide, with dorsal surface granulous, bearing a few blunt spines ventrally, mounted on small tubercles, and tiny normal spines dorsally. Tibia 2 short, less than half as long as femur or tibia 1, about as long as wide, with many blunt or slightly bifurcated spines distally on ventral surface, longer blunt spine distally on dorsal surface. Tarsus shortest, with spines on ventral side. Propodus about 0.9 times as long as femur or tibia 1 and twice as long as tibia 2, 3 times as long as wide, straight except for most proximal part, which is bent with strong angle marked dorsally; spines of sole simple or slightly bifurcated at tip. Main claw about half as long as propodus. No auxiliary claw. MEASUREMENTS (mm). Trunk 1.52; abdomen 0.49; proboscis 1.18; coxa 1 0.28; coxa 2 0.29; coxa 3 0.26; femur 0.49; tibia 1 0.53; tibia 2 0.24; tarsus 0.14; propodus 0.47; main claw 0.209. Sexual dimorphism No female currently available. Individual variability No other material available.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E74FFDCFD9A03FE0195B89A.taxon	discussion	Remarks Pycnogonum cesairei sp. nov. differs from most other species of the genus by the following combination of characters: the cuticula is strongly reticulated; the proboscis is conical, bent forward; there is no tubercle on coxa 1 (except on the fourth legs); the tibia 2 is very short, about as long as wide, and shorter than the propodus. Three other species of Pycnogonum share these characters: P. tesselatum Stock, 1968 from Pakistan, which has, however, a proboscis broader at the tip and a rounded abdomen (Stock 1968 a; George et al. 2020); P. clarki Staples, 2002 from Australia, which differs from P. cesairei by its more rounded dorsal tubercles, a more deeply incised dichotomy of propodus spines, the presence of a bulge on the posterodorsal end of the abdomen, the presence of tubercles on the lateral processes of the fourth trunk segment and no bifurcated spine on the legs (Staples 2002); and P. madagascariensis Bouvier, 1911 from Madagascar. For this latter species, we reexamined the syntypes of Bouvier (1911 a): the proboscis of P. madagascariensis is barrel-shaped (Arnaud 1971), the tibia 2 of P. madagascariensis is shorter than wide, while tibia 2 is about as long as wide in P. cesairei, and the abdomen of P. madagascariensis presents a distinctive posterodorsal bulge (visible in Arnaud 1971) with two distal rounded tips lateral to the anus, both characters not being present in P. cesairei. It also seems that none of these species have an angulous propodus, in contrast to P. cesairei. However, the closest species to Pycnogonum cesairei sp. nov. may well be P. calculum Bamber, 1995 from the Falkland Islands, which also presents a rather angulous propodus, has a proboscis of the same size, has the same body proportions and carries blunt spines. The coloration indicated by Bamber (light brown culticle with yellowish reticulation; Bamber 1995 b) also corresponds to that observed on the specimen from Martinique. However, P. calculum is described with a small rounded dorsomedian tubercle on the fourth trunk segment, which is absent in P. cesairei, as well as on all lateral processes, while P. cesairei only carries one on coxa 1 of the fourth leg. In Caribbean waters, the new species can be mistaken at first glance with Pycnogonum reticulatum Hedgpeth, 1948, for which we could examine the holotype of Hedgpeth (1948), but the two species are readily distinguished by the wider proboscis and longer tibiae 2 of P. reticulatum. The single sampled specimen was collected on the northern coast of Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E74FFDCFD9A03FE0195B89A.taxon	distribution	Distribution Only known from Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E74FFDCFD9A03FE0195B89A.taxon	description	Depth range 6 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E73FFDDFDC203260442BA35.taxon	description	Fig. 29	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E73FFDDFDC203260442BA35.taxon	materials_examined	Type material Pycnogonum ornans Stock, 1992. Holotype: NL ZMA. PYC. 3441 and ZMA. PYC. P. 1432 (not examined). Type locality: off Barra Sêca, Espírito Santo state, Brazil. Material examined MARTINIQUE • 1 ♀; Ste-Marie; 14 ° 46.3 ʹ N, 60 ° 57.5 ʹ W; depth 20 m; 20 Sep. 2016; st. AB 260; MNHN- IU- 2016 - 1064 / MK 411042.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E73FFDDFDC203260442BA35.taxon	discussion	Remarks This specimen is similar to the description given by Stock (1992 a) for Pycnogonum ornans, a monotypic species described from Brazil. In particular, propodal sole spines are found to be bifid as in Stock’s description; it also agrees in the distance between the lateral processes, ornamentation of the trunk, the lateral processes, the coxae 1 and the proboscis. However, the Martinique specimen differs from Stock’s description by the overall stouter appearance of the animal, as well as the less pronounced ornamentation of the trunk, lateral processes and coxae, and the less elongate propodus.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E73FFDDFDC203260442BA35.taxon	distribution	Distribution Pycnogonum ornans is only known from Brazil. This would be the first record of this species in the Caribbean.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E73FFDDFDC203260442BA35.taxon	description	Depth range Pycnogonum ornans is known to dwell at 38 – 47 m. The present specimen was sampled at 20 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E73FFC0FDCC079E0064BA3D.taxon	description	Fig. 30	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E73FFC0FDCC079E0064BA3D.taxon	materials_examined	Type material Pycnogonum pusillum Dohrn, 1881. Syntypes (?) (see Dunlop et al. 2007): ZMB _ Pyc _ 52 (not examined). Type locality: Naples Bay, Italy. Material examined MARTINIQUE • 1 ♀; Passe du Brigot; 14 ° 33.8 ʹ N, 60 ° 47.6 ʹ W; depth 17 m; 16 Sep. 2016; st. AB 183; MNHN-IU- 2016 - 815 • 1 ♀; Le Vauclin; 14 ° 30.5 ʹ N, 60 ° 48.5 ʹ W; depth 2 m; 10 Oct. 2016; st. AB 419; MNHN-IU- 2016 - 860.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E73FFC0FDCC079E0064BA3D.taxon	discussion	Remarks The Martinique specimens resemble the original illustration in Dohrn (1881) by their body shape, the conical proboscis being distally truncated and the relative sizes of the legs articles and main propodal claws; they similarly present the same tubercles on the lateral processes and first coxae as represented by Stock (1952), also illustrated by Lehmann et al. (2014) and Munilla & Soler-Membrives (2014). Unlike in Stock (1952), however, the specimens have a tubercle posterior to the ocular tubercle that is barely visible in dorsal view, and the auxiliary claws are longer (about half as long as the main claw). Since there is no available molecular data for comparison, we regard these two females as belonging to Dohrn’s species for now. The specimens are also morphologically very close to the Australian species Pycnogonum carinatum Staples, 2002, from which they chiefly differ by the presence of auxiliary claws in the Martinique material, as well as by the more granulous dorsal surface of the legs and the cleft spines on the propodal sole in the type material P. carinatum (Staples 2002).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E73FFC0FDCC079E0064BA3D.taxon	distribution	Distribution Pycnogonum pusillum is known from the Western Mediterranean (Balearic Sea, Adriatic Sea, Thyrrenian Sea, Ionian Sicily), and the Eastern Atlantic (Morocco, Angola, South Africa, Cap Verde).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E73FFC0FDCC079E0064BA3D.taxon	description	Depth range Pycnogonum pusillum was recorded from 0.5 to 1200 m. The present material was sampled at 2 and 17 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E6EFFC0FDB507CD0201B932.taxon	type_taxon	Type species Rhynchothorax mediterraneus Costa, 1861, by monotypy.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E6EFFC1FDC70486055BBB5F.taxon	materials_examined	Type material Holotype: USNM 170996 (not examined). Type locality: Carrie Bow Cay, Belize. Material examined MARTINIQUE • 1 juv.; Le Robert; 14 ° 43.2 ʹ N, 60 ° 50.6 ʹ W; depth 23 m; 25 Sep. 2016; st. AB 405; MNHN-IU- 2016 - 852 / MK 411165.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E6EFFC1FDC70486055BBB5F.taxon	discussion	Remarks The holotype of Rhynchothorax crenatus is a juvenile (Child 1982). Later, Child (1996 a) found adults of R. mediterraneus Costa, 1861 and considered that they were the adult form of R. crenatus. Since there is no clear argument to justify that these two species are conspecific, we prefer to regard R. crenatus as valid for the moment. It is worth noting, however, that the Martinique specimen is also a juvenile, so we cannot refute Child’s hypothesis. DNA barcode sequences will be useful to further investigate this question, when a sufficient sampling of DNA barcoding sequences from R. mediterraneus becomes available. Rhynchothorax mediterraneus is a species mostly known from the Mediterranean, with some records in the Indo-Pacific (Stock 1974; Miyazaki & Stock 1995) and Brazil (e. g., Stock 1992 a), but it has only been recorded in the Caribbean by Child (1996 a). Rhynchothorax crenatus is recorded for the first time outside of Belize. The single specimen was collected on the Atlantic coast of Martinique.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E6EFFC1FDC70486055BBB5F.taxon	distribution	Distribution Only recorded from Belize and Martinique. Depth range Recorded in literature at 27 m. The Martinique specimen was found at 23 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E6FFFC3FDA806100064B895.taxon	description	urn: lsid: zoobank. org: act: DF 2 C 9474 - DD 03 - 4190 - A 39 F- 37 F 9 F 8 E 43435 Fig. 31	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E6FFFC3FDA806100064B895.taxon	materials_examined	Material examined Holotype MARTINIQUE • 1 subadult?; Canal de Ste Lucie; 14 ° 22.7 ʹ N, 60 ° 51.6 ʹ W; depth 65 m; 11 Sep. 2016; st. AD 222; MNHN-IU- 2016 - 817 / MK 411138.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E6FFFC3FDA806100064B895.taxon	etymology	Etymology From the Latin ‘ sidereus ’, - a, - um (adj.), ‘ starry’, in memory of long working nights sorting samples and of the nice starry sky above. The name also refers to the many tubercular projections of the specimen, which gives it a meteor-like shape.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E6FFFC3FDA806100064B895.taxon	description	Description (holotype, MNHN-IU- 2016 - 817) BODY. Trunk and legs cuticle granulated, incompletely segmented with 3 rd and 4 th segments fused. 1 st to 3 rd segments with mediodorsal conical granulose tubercle near posterior margin. Ocular tubercle grossly club-shaped, slender at base, leaning forward, with 4 large pigmented eyes, ornamented with 3 cones above eyes, one slender anterolateral pair corresponding to lateral sense organs plus single cone on dorsomedian surface. Cephalon ornamented with 2 tall, slender, conical tubercles situated posterior to ocular tubercle base. Lateral projections of cephalon (on which palps articulate) conspicuously diverging from cephalic segment, anteriorly directed. Lateral processes of 1 st, 2 nd and 3 rd segments as long as wide, all well separated by less than half their diameter at base; lateral processes on 4 th segment shorter than wide, touching 3 rd pair at base. All lateral processes unornamented. PROBOScIS. Surface smooth, reaching 3 rd palp article, without conspicuous dorsomedian tubercle, with 2 pairs of dorsolateral bulges at about mid-length and ⅔ of proboscis length, latter most conspicous. Distal part down-curved. Lips bilateral, vertical, with inner and outer antimeres. ABDOMEN. Horizontal, cylindrical, blunt, about 3 times as long as wide, extending lightly beyond coxae 2. No basal segmentation. CHELIFORE. Absent. PALP. 5 - articled. 1 st article shorter than wide, with inconspicuous articulation. 2 nd article longest, about 7 times as long as wide, with dorsal tubercle at distal end, tip with spine. 3 rd article about 0.6 times as long as 2 nd and 4 times as long as 4 th, carrying scarce setae, with large spine on inner side distally. 4 th article about as long as wide, carrying many setae. 5 th article shortest, bud-like, carrying many setae. OvIGER. 10 - articled. 1 st article about as long as wide, glabrous. 2 nd article slightly longer than 1 st, more than 1.5 times as long as wide, glabrous. 3 rd article slightly longer than 2 nd, about 4 / 3 times as long as wide, glabrous. 4 th article longest, about 2.5 times as long as 3 rd and 5 times as long as wide, with distal seta. 5 th article half as long as 4 th and more than 1.5 times as long as wide, curved, carrying distal seta. 6 th article about 0.9 times as long as 4 th and about 3 times as long as wide, carrying proximal seta. 7 th article about 4 / 3 times as long as wide. 8 th and 9 th articles subequal, about as long as wide. 10 th article larger, rounded, about as long as wide, typical of the genus in having lamina on inner strigilis surface in addition to strigilis spine and terminal claw. Striglis typical, formula 3: 3: 3: 2, with bifurcated spine on each article. Terminal claw inner surface apparently flattened, about as long as strigilis lamina. LEGS. Slender. Coxa 1 bearing one dorsomedian and 2 lateral granulated projections on 1 st, 2 nd and 3 rd legs, only one anterior projection on 4 th leg. Coxa 2 of about same size, slenderer, unornamented. Coxa 3 small, unornamented. Femur approximately 4 times as long as wide. Tibiae 1 and 2 slenderer than femur, bearing long dorsomedian spine at distal end. Tibia 1 of about same size as femur, tibia 2 slightly shorter than femur or tibia 1. Tarsus medium-sized. Propodus about ⅙ shorter than tibia 2, setose, sole gently curved, bearing 4 spines. Main claw about half as long as propodus. No auxiliary claw. Gonopores inconspicuous. MEASUREMENTS (mm). Trunk 0.47; abdomen 0.16; proboscis 0.31; palp 0.33; coxa 1 0.07; coxa 2 0.07; coxa 3 0.06; femur 0.24; tibia 1 0.22; tibia 2 0.19; tarsus 0.04; propodus 0.18; main claw 0.08.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E6FFFC3FDA806100064B895.taxon	discussion	Remarks Rhynchothorax sidereus sp. nov. is distinguished from most other species of the genus by the following combination of characters: presence of pigmented eyes; presence of one dorsodistal spur on second palp article; two conical tubercles posterior to ocular tubercle base; three tubercles on coxa 1 of first, second and third legs; no ornamentation on lateral processes. These characters are shared with R. orientalis Child, 1988 from Philippines. Rhynchothorax sidereus sp. nov. differs from Child’s (1988 a) specimen in showing no bifid extremity of the dorsomedian tubercles or any anterior projection of the ocular tubercle. The species R. sidereus is also morphologically similar to R. swir Staples, 2019 from the southwest Indian Ridge, but the former has distinctive tall tubercles on the ocular tubercle and is without conical tubercles on the lateral processes (Staples 2019).	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E6FFFC3FDA806100064B895.taxon	distribution	Distribution Only known from type locality.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
03A95E063E6FFFC3FDA806100064B895.taxon	description	Depth range 65 m.	en	Sabroux, Romain, Hassanin, Alexandre, Corbari, Laure (2022): Sea spiders (Arthropoda: Pycnogonida) collected during the Madibenthos Expedition from Martinique shallow waters. European Journal of Taxonomy 851 (1): 1-141, DOI: 10.5852/ejt.2022.851.1999, URL: http://dx.doi.org/10.5852/ejt.2022.851.1999
