taxonID	type	description	language	source
E43287C1FFB11E04FF20FA9FFA39F92A.taxon	type_taxon	Type species: Pseudoligota varians Cameron, 1920, fixed by subsequent designation by Blackwelder, 1952: 327.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB11E04FF20FA9FFA39F92A.taxon	diagnosis	Diagnosis. The genus Pseudoligota can be distinguished from other Japanese gyrophaenin genera by the combination of the following character states: body minute (adults BL: 0.8 – 1.2 mm) and slightly limuloid; hypomeron not visible in lateral view; mesoventrite without medial longitudinal carina; meso- and metaventral processes between mesocoxae fused; inner face of lacinia with single row of setae; apex of galea with four rows of setae; labium with ligula bifid, divided at distal 2 / 3 – 3 / 4 of its length; prelabium with single medial seta (modified after Ashe 1984).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB11E01FF20F8F8FF56FE3E.taxon	description	(Figs 1 A, 2 A – C, 4 A – B, 5 A)	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB11E01FF20F8F8FF56FE3E.taxon	materials_examined	Material examined. Type material. Holotype: male, “ Onsen. Japan / lg. Sauter. [handwritten] // antennata / Brh. Typ [handwritten] // Oligota antennata / Brh. Typus [handwritten] // Chicago NHMus / M. Bernhauer / Collection. ” (abdominal segments VIII ‒ X and aedeagus were dissected and mounted in Euparal by MM) (FMNH). See details on Fig. 4. Additional specimens studied. JAPAN: Hokkaido: [Hokkaido]: 4 exs., Kannonzawa-rindô, Toyama, Minami-ku, Sapporo-shi, 8 VIII 2021, T. Nozaki leg. (KUM); Shikoku: [Ehime-ken]: 1 ex., Matsuyama Castle, Kuromon Trail, 17 V 2018, P. Jałoszyński leg. (pcPJ); Kyushu: [Ôita-ken]: 17 exs., Shônaichô-Asono, Yufu-shi, 20 V 2021, T. Hashizume leg. (KUM); [Nagasaki-ken]: 3 exs., Mitsushimamachi-Sumo, Tsushima-shi, 11 IX 2021, T. Hashizume leg. (KUM); [Kagoshima-ken]: 1 ex., Koseda, Yaku-shima Is., 22 VII 2021, T. Hashizume leg. (KUM); 6 exs., Kurio, Yaku-shima Is., 21 VII 2021, T. Hashizume leg. (KUM).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB11E01FF20F8F8FF56FE3E.taxon	diagnosis	Diagnosis. This species can be distinguished from its congeners by a combination of the following characters: body black, antennae and legs yellow, elytra and male tergite VII unmodified, male tergite VIII with broad blunt median tooth at posterior margin, and the shape of the median lobe of aedeagus. Pseudoligota picea Pace, 2003 and P. picetoides Pace, 2003 have somewhat similar median lobes of the aedeagus, but males of these species have carinae or granules at sutural elytral margins and an elongate tubercle in the center of tergite VII.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB11E01FF20F8F8FF56FE3E.taxon	description	Redescription. Measurements. (n = 5): BL 0.90 – 1.27; FBL: 0.52 – 0.66; HL: 0.17 – 0.20; HW: 0.26 – 0.29; PL: 0.19 – 0.25; PW: 0.38 – 0.46; EL: 0.19 – 0.26; EW: 0.43 – 0.56. (Holotype: BL ≈ 1.1; PL: 0.24; PW: 0.43; HTL: 0.24). Body (Fig. 1 A) black; legs and antennae excluding terminal segment yellow; antennal terminal segment darker than proximal segments. Head. Transverse, about 1.50 times as wide as long. Sparsely punctured with indistinct shallow punctures. Antenna 11 - segmented, segments I – III longer than wide, segments IV – V about as wide as long, VI – X transverse, segment XI 1.5 times as long as wide. Thorax. Pronotum transverse, about 1.83 times as wide as long, widest at posterior angles, base rounded; punctures somewhat denser, more distinct than those of head; microsculpture indistinct. Elytra transverse, slightly wider than pronotum; punctures larger than those of pronotum; microsculpture slightly stronger than that on pronotum, composed of transverse reticulation; carinae or granules on sutural margins absent. Hind wings developed. Abdomen narrowed posteriad, with punctures finer than those on elytra, with diamond-shaped reticulation. Tergite VII unmodified. Male. Tergite VIII (Fig. 2 A) with broad blunt median tooth at posterior margin. Median lobe of aedeagus (Fig. 2 B) of complicated shape; parameral process bilobed, parameral lobe of parameral process long, apex pointed, bending paramerally in the apical 2 / 5; abparameral lobe short, apex rounded, exceeding the flexure of parameral lobe; long thin lobe on left side in abparameral view; flagellum very long, emerging to outside from behind the parameral process, basal part of flagellum relatively small and slightly oval. Female. Similar to male in general appearance. Tergite VIII without broad blunt tooth at posterior margin. Spermatheca (Fig. 2 C) simple; distal bulb rounded with tongue-shaped lobe, wall of the distal end protrudes inward; duct curved.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB11E01FF20F8F8FF56FE3E.taxon	distribution	Distribution. Japan: (Kyushu; Hokkaido, Tsushima Is., Yaku-shima Is. — new record); India, Peninsular Malaysia.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB11E01FF20F8F8FF56FE3E.taxon	discussion	Remarks. Pseudoligota antennata was originally described as Oligota antennata by Bernhauer (1907). Then, Holobus was given a full generic rank by Coiffait & Saiz (1967) and recently this species was catalogued as Holobus antennatus (Smetana 2004; Schülke & Smetana 2015). However, after studying the holotype of Oligota antennata, it was determined that this species should belong to the genus Pseudoligota. Pseudoligota affinis was described from northernmost India by Cameron (1939). The original description of P. affinis (see Cameron (1939 )) and illustrations of the aedeagus of P. affinis in Ashe (1984) are consistent with the characteristics of P. antennata, and the two species are indistinguishable. Therefore, P. affinis should be regarded as a junior synonym of P. antennata. Bionomics. Most specimens were obtained from fungi on dead wood in deciduous and evergreen forests (Fig. 5 A – B).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB41E03FF20FDCCFA95FE3E.taxon	description	(Figs 1 B, 3 A – C, 5 C)	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB41E03FF20FDCCFA95FE3E.taxon	materials_examined	Type material. Holotype: male (KUM), “ JAPAN: Okinawa-ken, / Ishigaki-shi, Sakieda, / Maese-dake, / 26 III 2022, / T. Nozaki leg. // HOLOTYPE / Pseudoligota / nozakii / des. T. Hashizume, / S. Yamamoto & / M. Maruyama, 2022 ”. Paratypes. JAPAN: [Okinawa-ken]: 4 males, 2 females, 4 unsexed, same data as holotype. (KUM).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB41E03FF20FDCCFA95FE3E.taxon	diagnosis	Diagnosis. This species is similar to P. antennata, it can be distinguished by its smaller body, the male tergite VIII without a broad blunt tooth at the posterior margin, and the abparameral lobe of the parameral process of median lobe of aedeagus not reaching the flexure of the parameral lobe of parameral process. This species can also be distinguished from other Pseudoligota spp. by a combination of the black body coloration, the unmodified elytra, male tergite VII, and male tergite VIII, the relatively complicated general shape of the median lobe of the aedeagus, and the absence of inward protrusion on the wall of distal end of the spermatheca.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB41E03FF20FDCCFA95FE3E.taxon	description	Description. Measurements. (n = 5): BL 0.84 – 0.96; FBL: 0.51 – 0.58; HL: 0.16 – 0.18; HW: 0.25 – 0.26; PL: 0.20 – 0.22; PW: 0.38; EL: 0.19 – 0.21; EW: 0.41 – 0.46. Body (Fig. 1 B) black; legs and antennae excluding terminal segment yellow; antennal terminal segment darker than proximal segments. Head. Transverse, about 1.53 times as wide as long. Sparsely punctured with indistinct shallow punctures. Antenna 11 - segmented, segments I – III longer than wide, segments IV – X transverse, segment XI about 1.5 times as long as wide. Thorax. Pronotum transverse, about 1.85 times as wide as long, widest at posterior angles, base rounded; punctures somewhat denser, more distinct than those of head; microsculpture indistinct. Elytra transverse, slightly wider than pronotum; punctures larger than those of pronotum; microsculpture slightly stronger than that on pronotum, composed of transverse reticulation; carinae or granules on sutural margins absent. Hind wings developed. Abdomen narrowed posteriad, with punctures finer than those on elytra, with diamond-shaped sculpture. Tergite VII unmodified. Male. Tergite VIII (Fig. 3 A) without broad blunt tooth at posterior margin. Median lobe of aedeagus (Fig. 3 B) of complicated shape; parameral process bilobed, parameral lobe of parameral process long, apex pointed, bending paramerally in the apical 2 / 5; abparameral lobe short, apex rounded, not reaching the flexure of parameral lobe; long thin lobe on left side in abparameral view; flagellum very long, emerging to outside from behind the parameral process, basal part of flagellum large and slightly oval. Female. Similar to male in general appearance. Spermatheca (Fig. 3 C) simple; distal bulb rounded with tongueshaped lobe, wall of the distal end without inward protrusion; duct curved.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB41E03FF20FDCCFA95FE3E.taxon	distribution	Distribution. Japan: Ryukyus: Ishigaki-jima Is.	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
E43287C1FFB41E03FF20FDCCFA95FE3E.taxon	etymology	Etymology. This specific name of the new species is dedicated to Tsubasa Nozaki who is the collector of the type series. Bionomics. All specimens were obtained from fungi on dead wood in evergreen forest (Fig. 5 C – D).	en	Hashizume, Takuto, Yamamoto, Shûhei, Maruyama, Munetoshi (2023): Taxonomic notes on the genus Pseudoligota Cameron (Coleoptera: Staphylinidae: Aleocharinae) from Japan. Zootaxa 5227 (1): 100-108, DOI: https://doi.org/10.11646/zootaxa.5227.1.4
