taxonID	type	description	language	source
C2A9D751DEDC58EFA3B03B61D48DAA07.taxon	description	Figs 1 B, 2, 3, 4, 5, 6 Chinese name 白坭异伪蝎	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
C2A9D751DEDC58EFA3B03B61D48DAA07.taxon	diagnosis	Diagnosis (♀). The new species can be recognized by the following combination of characters: carapace without eyes or eyespots, posterior margin with two setae, chaetotaxy of carapace: 4 - 4 - 2 - 2 - 2, 14; cheliceral palm with four setae only; rallum with nine blades (each with fine pinnate, the basal-most blade shorter than the others); coxa I with six coxal spines (tridentate blades, each blade with a central fan-shaped spine terminally) on a tubercle; pedipalps slender, femur 9.07, chela 5.41 x longer than broad, both chelal fingers with a row of teeth (fixed chelal finger with 19 teeth; movable chelal finger with 17 teeth), slightly retrorse and pointed.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
C2A9D751DEDC58EFA3B03B61D48DAA07.taxon	etymology	Etymology. Named after the village of Baini, near the type locality.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
C2A9D751DEDC58EFA3B03B61D48DAA07.taxon	description	Description. Adult female (male unknown) (Figs 3 - 6). Color (Figs 3, 4): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 4 B, D, 5 A, C): carapace subquadrate, 0.87 x longer than broad, gently narrowed posteriorly; surface smooth, without furrows but with six lyrifissures and the posterior part with squamous sculpturing; no traces of eyes; epistomal process absent, space between median setae slightly recurved; with 14 setae arranged 4: 4: 2: 2: 2, preocular setae absent, most setae heavy, long and gently curved. Chaetotaxy of coxae: P 3, I 4, II 5, III 5, IV 5 - 6; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; coxal spines present on coxa I only, consisting of a tubercle expanded terminally into a characteristic " spray " or " fan " of six elevated processes which extend apically, subequal in length (Figs 4 D, 5 C); bisetose intercoxal tubercle present between coxae III and IV (Fig. 4 D). Chelicera (Figs 4 C, 5 B, E): large, approximately as long as carapace, 2.37 x longer than broad; four setae present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with a medial seta; exterior condylar lyrifissure and exterior lyrifissure exist, palm with two extra setae (close to sub-basal seta). Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 14 acute teeth, distal one largest, plus five small basal teeth, 19 in total; movable finger with 21 retrorse contiguous teeth of equal length; galea absent. Serrula exterior with 18 blades and serrula interior with 12 blades. Rallum in two rows and composed of nine blades with fine pinnate, of which the basal-most blade shorter than the others (Fig. 5 E). Pedipalp (Figs 4 A, 5 D, 6 A, B): long and slender, trochanter 1.68, femur 9.07, patella 3.06, chela 5.41, hand 2.29 x longer than broad; femur 2.62 x longer than patella; movable chelal finger 1.44 x longer than hand and 0.61 x longer than chela. Setae generally long and acuminate; two distal lyrifissures present on patella (Fig. 5 D). Chelal palm robust and slightly constricted towards fingers. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib, isb, eb, esb, and ist clustered at the base of fixed finger, ist slightly distal to esb; it slightly distal to est, situated subdistally; et situated subdistally, very close to chelal teeth; dx situated distal to et, near the tip of fixed finger; sb situated closer to b than to st (Fig. 6 A). Microsetae (chemosensory setae) absent on hand and both palpal fingers. Sensilla absent. Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 19 teeth, slightly retrorse and pointed; movable chelal finger with 17 teeth (slightly smaller than teeth on fixed chelal finger) and a tubercle between the ninth and tenth teeth (Fig. 6 A). Chelal fingers slightly curved in dorsal view (Fig. 6 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I-XII: 2: 4: 4: 6: 6: 7: 7: 6: 7: 5: TT: 0; tergites IX and X each with an unpaired median seta. Sternal chaetotaxy IV-XII: 10: 11: 11: 11: 11: 9: 8: 0: 2. Anterior genital operculum with eight setae plus 14 setae on posterior margin, with a pair of lyrifissures present anterolateral and posteriolateral to genital opening, respectively (Fig. 4 E). Legs (Fig. 6 C, D): generally typical, long, and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.77 x longer than patella and with one lyrifissure at the base of femur; tarsus 2.55 x longer than tibia. Femoropatella of leg IV 4.86 x longer than deep and with one lyrifissure at the base of femur; tibia 6.17 x longer than deep; with basal tactile setae on both tarsal segments: basitarsus 4.22 x longer than deep (TS = 0.24), telotarsus 12.43 x longer than deep and 2.29 x longer than basitarsus (TS = 0.31). Setae of leg I (trochanter to tibia) 2: 10: 9: 12, setae of leg IV (trochanter to basitarsus) 3: 3: 7: 15: 17. Arolium slightly shorter than the claws, not divided; claws simple. Dimensions of female holotype (length / breadth or, in the case of the legs, length / depth in mm): body length 2.72. Pedipalps: trochanter 0.32 / 0.19, femur 1.36 / 0.15, patella 0.52 / 0.17, chela 1.84 / 0.34, hand 0.78 / 0.34, movable finger length 1.12. Chelicera 0.64 / 0.27, movable finger length 0.34. Carapace 0.55 / 0.63. Leg I: trochanter 0.24 / 0.18, femur 0.76 / 0.11, patella 0.43 / 0.10, tibia 0.33 / 0.07, tarsus 0.84 / 0.07. Leg IV: trochanter 0.34 / 0.18, femoropatella 1.02 / 0.21, tibia 0.74 / 0.12, basitarsus 0.38 / 0.09, telotarsus 0.87 / 0.07.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
C2A9D751DEDC58EFA3B03B61D48DAA07.taxon	distribution	Distribution and habitat. This species is only known from the type locality, Liangfeng Cave (Figs 1 B, 2), which is located near a road, 0.6 km southeast of Baini Village (Xishui County). This limestone cave has a medium-sized rectangular entrance (~ 3 m high and 5 m wide) with a large horizontally extending interior space. The interior of the cave is mainly divided into three tunnels, the left tunnel extends ~ 200 m, the middle tunnel extends ~ 500 m, and the right tunnel communicates with the middle tunnel, ~ 100 m in length. Human disturbance in the entrance zone is serious, but the deep zone remains pristine. The specimen was collected under a stone near the wall in the deepest part of the middle tunnel. This space is completely dark, with constant temperature and humidity (temperature ~ 9 ° C, humidity ~ 90 %).	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
A5514B016A2D5F4CB83DB46941CC8861.taxon	description	Figs 1 C, 7, 8, 9, 10, 11, 12 Chinese name 弯指异伪蝎	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
A5514B016A2D5F4CB83DB46941CC8861.taxon	diagnosis	Diagnosis (♂ ♀). The new species can be recognized by the following combination of characters: cheliceral palm with five setae; coxa I with four coxal spines (tridentate blades, each blade with a central fan-shaped spine terminally) on a tubercle; pedipalps slender, femur 9.07 - 10.15 (♂), 8.50 - 8.60 (♀), chela 7.00 - 7.52 (♂), 6.64 - 7.15 (♀) x longer than broad, both chelal fingers with a row of teeth (fixed chelal finger with 31 or 33 teeth; movable chelal finger with 26 or 28 teeth), slightly retrorse and pointed; chela fingers markedly curved in dorsal view.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
A5514B016A2D5F4CB83DB46941CC8861.taxon	etymology	Etymology. The specific name is derived from the Latin word pandus (curved) and refers to the character of the curved chelal fingers.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
A5514B016A2D5F4CB83DB46941CC8861.taxon	description	Description. Adult males (Figs 7 F, 8 A, 9, 10 A, B, 11, 12). Color (Figs 7 F, 8 A, 9, 10 A, B): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 9 B, 10 A, 11 A, C): carapace inverted trapezoid, 0.91 - 0.93 x longer than broad, gently narrowed posteriorly; surface smooth, without furrows but with six lyrifissures and the posterior part with squamous sculpturing; no traces of eyes; epistomal process absent, space between median setae slightly recurved; with 14 setae arranged 4: 4: 2: 2: 2, preocular setae absent, most setae heavy, long, and gently curved. Chaetotaxy of coxae: P 3, I 4, II 4 - 6, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 3 length of medial seta; coxal spines present on coxa I only, consisting of a tubercle expanded terminally into a characteristic " spray " or " fan " of four elevated processes which extend apically, subequal in length (Figs 10 A, 11 C); bisetose intercoxal tubercle present between coxae III and IV (Fig. 10 A). Chelicera (Figs 9 C, 11 B, E): large, approximately as long as carapace, 2.56 - 2.60 x longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with a medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with ten acute teeth, distal one largest; movable finger with 15 or 16 retrorse contiguous teeth of equal length, plus four or five vestigial, rounded, and contiguous basal teeth, 19 - 21 in total; galea absent. Serrula exterior with 19 or 20 blades and serrula interior with 10 or 11 blades. Rallum in two rows and composed of nine blades with fine pinnate, of which the basal-most blade shorter than the others (Fig. 11 E). Pedipalp (Figs 9 A, 11 D, 12 A, B): long and slender, trochanter 1.65 - 1.67, femur 9.07 - 10.15, patella 3.47 - 3.57, chela 7.00 - 7.52, hand 2.60 - 2.96 x longer than broad; femur 2.44 - 2.64 x longer than patella; movable chelal finger 1.59 - 1.72 x longer than hand and 0.62 - 0.64 x longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Fig. 11 D). Chelal palm slightly constricted towards fingers. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib, isb, eb, esb, and ist clustered at the base of fixed finger, esb slightly distal to ist; it slightly distal to est, situated subdistally; et situated subdistally, very close to chelal teeth; dx situated distal to et, near the tip of fixed finger; sb situated closer to b than to st (Fig. 12 A). Microsetae (chemosensory setae) absent on hand and both palpal fingers. Sensilla absent. Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 31 or 33 teeth, slightly retrorse and pointed; movable chelal finger with 24 or 25 teeth (slightly smaller than teeth on fixed chelal finger), plus two or three vestigial, rounded and contiguous basal teeth, 26 - 28 in total; a small tubercle between the fourteenth and fifteenth teeth present (Fig. 12 A). Chelal fingers markedly curved in dorsal view (Fig. 12 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I-XII: 2: 4 - 5: 4 - 5: 6: 6: 6: 6: 6: 3: 2: TT: 0, tergite IX with an unpaired median seta. Sternal chaetotaxy III-XII: 6 - 8: 9 - 10: 9 - 11: 9: 9 - 10: 9 - 10: 7 - 8: 5 - 6: 0: 2. Anterior genital operculum with eight setae, genital opening pit-like, with five or six marginal setae on each side, 18 - 19 in total, with a pair of lyrifissures present anterolateral and posteriolateral to genital opening, respectively (Fig. 10 B). Legs (Fig. 12 C, D): generally typical, long, and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.64 - 1.79 x longer than patella and with one lyrifissure at the base of femur; tarsus 2.42 - 2.60 x longer than tibia. Femoropatella of leg IV 5.00 - 5.17 x longer than deep and with one lyrifissure at the base of femur; tibia 6.55 - 7.30 x longer than deep; with a long tactile seta on both tarsal segments: basitarsus 4.00 - 4.86 x longer than deep (TS = 0.21 - 0.25), telotarsus 13.50 - 14.50 x longer than deep and 2.38 - 2.72 x longer than basitarsus (TS = 0.22 - 0.25). Setae of leg I (trochanter to tibia) 2 - 3: 9 - 11: 9 - 10: 12 - 13, setae of leg IV (trochanter to basitarsus) 2 - 3: 2: 4 - 5: 17 - 19: 10 - 11. Arolium slightly shorter than the claws, not divided; claws simple. Dimensions of adult males (length / breadth or, in the case of the legs, length / depth in mm): body length 1.97 - 2.23. Pedipalps: trochanter 0.28 - 0.30 / 0.17 - 0.18, femur 1.27 - 1.32 / 0.13 - 0.14, patella 0.50 - 0.52 / 0.14 - 0.15, chela 1.73 - 1.75 / 0.23 - 0.25, hand 0.65 - 0.68 / 0.23 - 0.25, movable chelal finger length 1.08 - 1.12. Chelicera 0.64 - 0.65 / 0.25, movable finger length 0.33 - 0.34. Carapace 0.53 - 0.54 / 0.58. Leg I: trochanter 0.20 - 0.21 / 0.15, femur 0.68 - 0.69 / 0.08 - 0.09, patella 0.38 - 0.42 / 0.08, tibia 0.30 - 0.31 / 0.05 - 0.06, tarsus 0.75 - 0.78 / 0.05 - 0.06. Leg IV: trochanter 0.29 - 0.30 / 0.16 - 0.17, femoropatella 0.93 - 0.95 / 0.18 - 0.19, tibia 0.72 - 0.73 / 0.10 - 0.11, basitarsus 0.32 - 0.34 / 0.07 - 0.08, telotarsus 0.81 - 0.87 / 0.06. Adult females (Figs 8 B, 10 C). Mostly same as males, but a little larger; chaetotaxy of coxae: P 3, I 4, II 5, III 5, IV 5; tergal chaetotaxy I-XII: 2: 4: 4: 4 - 6: 6: 6: 6: 6: 4: 2: TT: 0; sternal chaetotaxy IV-XII: 9 - 10: 10 - 12: 9: 10 - 11: 10 - 12: 7 - 8: 6: 0: 2; anterior genital operculum with eight or nine setae, posterior margin with nine or ten marginal setae, 17 - 19 in total; leg IV with a long tactile seta on both tarsal segments: basitarsus 3.78 x longer than deep (TS = 0.24), telotarsus 12.86 - 14.50 x longer than deep and 2.56 - 2.65 x longer than basitarsus (TS = 0.20 - 0.23). Body length 2.10 - 2.41. Pedipalps: trochanter 0.33 / 0.19 - 0.20 (1.65 - 1.74 x), femur 1.36 - 1.38 / 0.16 (8.50 - 8.63 x), patella 0.54 - 0.55 / 0.18 - 0.19 (2.89 - 3.00 x), chela 1.86 / 0.26 - 0.28 (6.64 - 7.15 x), hand 0.72 - 0.75 / 0.26 - 0.28 (2.68 - 2.77 x), movable chelal finger length 1.16. Chelicera 0.73 - 0.76 / 0.26 - 0.28 (2.71 - 2.81 x), movable finger length 0.39. Carapace 0.55 - 0.57 / 0.65 (0.85 - 0.88 x). Leg I: trochanter 0.19 / 0.17 (1.12 x), femur 0.70 - 0.75 / 0.10 - 0.11 (6.82 - 7.00 x), patella 0.43 - 0.44 / 0.09 (4.78 - 4.89 x), tibia 0.33 - 0.35 / 0.06 (5.50 - 5.83 x), tarsus 0.79 - 0.84 / 0.06 (13.17 - 14.00 x). Leg IV: trochanter 0.32 - 0.34 / 0.16 - 0.18 (1.89 - 2.00 x), femoropatella 0.92 - 1.02 / 0.19 - 0.22 (4.64 - 4.84 x), tibia 0.67 - 0.73 / 0.10 - 0.11 (6.64 - 6.70 x), basitarsus 0.34 / 0.09 (3.78 x), telotarsus 0.87 - 0.90 / 0.06 - 0.07 (12.86 - 14.50 x).	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
A5514B016A2D5F4CB83DB46941CC8861.taxon	distribution	Distribution and habitat. This species is only known from the type locality, Daozuo Cave (Figs 1 C, 7), which is located near a road, 1 km southwest of Jinshan Village (Xishui County) and is surrounded by rural and agricultural fields. This limestone cave has a large, rectangular entrance (~ 1 m high and 30 m wide) and a total length of ~ 300 m, only a narrow tunnel leads to the deepest part of the cave, which is a slightly wider, low-temperature, high-humidity, and completely lightless environment (temperature ~ 11 ° C, humidity> 90 %). All specimens were collected under stones in the deepest part of the cave.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
1C08A1E3D2A050C28394CBA818821FC6.taxon	description	Figs 1 D, 13, 14, 15, 16, 17 Chinese name 新桥异伪蝎	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
1C08A1E3D2A050C28394CBA818821FC6.taxon	diagnosis	Diagnosis (♀). The new species can be recognized by the following combination of characters: each cheliceral finger with several small basal teeth between large teeth, most of which appear in pairs, the fingertips blunt, not sharp; rallum with eight blades (each with fine pinnate, the basal-most blade shorter than the others); pedipalps slender, femur 9.71, chela 5.44 x longer than broad, both chelal fingers with a row of teeth (each chelal finger with 23 teeth), slightly retrorse and pointed.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
1C08A1E3D2A050C28394CBA818821FC6.taxon	etymology	Etymology. Named after the village of Xinqiao, near the type locality.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
1C08A1E3D2A050C28394CBA818821FC6.taxon	description	Description. Adult female (male unknown) (Figs 14 - 17). Color (Figs 14, 15): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 15 B, D, 16 A, C): carapace inverted trapezoid, 1.00 x longer than broad, gently narrowed posteriorly; surface smooth, without furrows but with seven lyrifissures and the posterior part with squamous sculpturing; no traces of eyes; epistomal process absent, space between median setae slightly recurved; with 14 setae arranged 4: 4: 2: 2: 2, preocular setae absent, most setae heavy, long and gently curved. Chaetotaxy of coxae: P 3, I 6, II 7 - 9, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; coxal spines present on coxa I only, consisting of a tubercle expanded terminally into a characteristic " spray " or " fan " of six elevated processes which extend apically, subequal in length (Figs 15 D, 16 C); bisetose intercoxal tubercle present between coxae III and IV (Fig. 15 D). Chelicera (Figs 15 C, 16 B, E): large, approximately as long as carapace, 2.38 x longer than broad; five setae present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with a medial seta; exterior condylar lyrifissure and exterior lyrifissure exist, palm with five extra (surrounding an accessory seta). Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with nine acute teeth, distal one largest; movable finger with a slight bump apical tooth and 12 retrorse contiguous teeth of equal length, each finger with several small basal teeth between large teeth, most of which appear in pairs, four on movable finger and six on fixed finger; the fingertips blunt, not sharp; galea represented by a very slight bump on movable finger. Serrula exterior with 17 blades and serrula interior with ten blades. Rallum in two rows and composed of eight blades with fine pinnate, of which the basal-most blade shorter than the others (Fig. 16 E). Pedipalp (Figs 15 A, 16 D, 17 A, B): long and slender, trochanter 1.63, femur 9.71, patella 2.83, chela 5.44, hand 2.25 x longer than broad; femur 2.67 x longer than patella; movable chelal finger 1.44 x longer than hand and 0.60 x longer than chela. Setae generally long and acuminate; two distal lyrifissures present on patella, femur with one (Fig. 16 D). Chelal palm robust and slightly constricted towards fingers. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib, isb, eb, esb, and ist clustered at the base of fixed finger, ist slightly distal to esb; it slightly distal to est, situated subdistally; et situated subdistally, very close to chelal teeth; dx situated distal to et, near the tip of fixed finger; sb situated closer to b than to st (Fig. 17 A). Microsetae (chemosensory setae) absent on hand and both palpal fingers. Sensilla absent. Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 23 teeth, slightly retrorse and pointed; movable chelal finger with 23 teeth (slightly smaller than teeth on fixed chelal finger) and a tubercle between the eleventh and twelfth teeth (Fig. 17 A). Chelal fingers markedly curved in dorsal view (Fig. 17 B). Opisthosoma: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I-XII: 3: 4: 4: 6: 6: 6: 6: 7: 5: 4: TT: 0; tergites VIII and IX each with an unpaired median seta; a lyrifissure on each side of tergites I-IX. Sternal chaetotaxy IV-XII: 9: 12: 11: 12: 12: 9: 8: 0: 2. Anterior genital operculum with six setae plus 12 setae on posterior margin, with a pair of lyrifissures present anterolateral and posteriolateral to genital opening, respectively (Fig. 15 E). Legs (Fig. 17 C, D): generally typical, long, and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.61 x longer than patella and with one lyrifissure at the base of femur; tarsus 2.24 x longer than tibia. Femoropatella of leg IV 4.74 x longer than deep and with one lyrifissure at the base of femur; tibia 6.58 x longer than deep; with basal tactile setae on both tarsal segments: basitarsus 4.44 x longer than deep (TS = 0.28), telotarsus 14.29 x longer than deep and 2.50 x longer than basitarsus (TS = 0.20). Setae of leg I (trochanter to tibia) 2: 12: 11: 19, setae of leg IV (trochanter to basitarsus) 3: 2: 6: 24: 14. Arolium slightly shorter than the claws, not divided; claws simple. Dimensions of female holotype (length / breadth or, in the case of the legs, length / depth in mm): body length 2.01. Pedipalps: trochanter 0.31 / 0.19, femur 1.36 / 0.14, patella 0.51 / 0.18, chela 1.74 / 0.32, hand 0.72 / 0.32, movable finger length 1.04. Chelicera 0.57 / 0.24, movable finger length 0.32. Carapace 0.55 / 0.55. Leg I: trochanter 0.22 / 0.17, femur 0.79 / 0.10, patella 0.49 / 0.09, tibia 0.38 / 0.07, tarsus 0.85 / 0.06. Leg IV: trochanter 0.34 / 0.19, femoropatella 1.09 / 0.23, tibia 0.79 / 0.12, basitarsus 0.40 / 0.09, telotarsus 1.00 / 0.07.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
1C08A1E3D2A050C28394CBA818821FC6.taxon	distribution	Distribution and habitat. This species is only known from the type locality, Sanjie Cave (Figs 1 D, 13), which is located ~ 1.8 km northeast of Xinqiao Village (Fenggang County). This limestone cave has a small oval entrance (~ 1 m high and 2 m wide), ~ 200 meters in length, with a large, elongated exit at the end of the cave (~ 5 m high and 50 m wide). The interior entirety of the cave is large, inclined and extending downwards. The cave ground was covered with stones. The specimen was collected under a stone ~ 100 m from the cave entrance.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
B8502145B46C58B48237EED883C95229.taxon	description	Figs 1 A, 18, 19, 20, 21, 22, 23 Chinese name 五里碑穴伪蝎	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
B8502145B46C58B48237EED883C95229.taxon	diagnosis	Diagnosis (♂ ♀). The new species can be recognized by the following combination of characters: surfaces mostly with fine reticulations; carapace without eyes or eyespots but eye region bulging and convex in dorsal view; anterior margin without protuberances; cheliceral palm with five setae; rallum with 11 blades (each with fine pinnate, the basal-most blade shorter than the others); coxal spines present on coxa I only, comprising a transverse, contiguous series of seven or eight tridentate blades, which arise from a lightly sclerotized or translucent hillock, the central ramus of each blade (except the basal two) sharply acumino-spatulate and extending beyond the lateral rami; pedipalps slender, femur 7.24 (♂), 6.40 (♀), chela 6.21 - 6.22 (♂), 5.68 (♀) x longer than broad, both chelal fingers with a row of teeth (fixed chelal finger with 22 or 24 teeth; movable chelal finger with 16 - 19 teeth), slightly retrorse and pointed; chela fingers straight in dorsal view.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
B8502145B46C58B48237EED883C95229.taxon	etymology	Etymology. Named after the type locality, Wulibei Cave.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
B8502145B46C58B48237EED883C95229.taxon	description	Description. Adult males (Figs 18 D, 19 A, 20, 21 A, B, 22, 23). Color (Figs 18 D, 19 A, 20, 21 A, B): generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale. Cephalothorax (Figs 20 B, 21 A, 22 A, C): carapace subquadrate, 1.02 - 1.03 x longer than broad, gently narrowed posteriorly; surface mostly with fine reticulations, without furrows but with seven or eight lyrifissures; no traces of eyes but eye region bulging and convex in dorsal view; epistome present and with some tiny spinules; with 16 setae arranged s 4 s: 4: 2: 2: 2, most setae heavy, long, and gently curved. Chaetotaxy of coxae: P 3, I 6 - 7, II 4 - 5, III 4, IV 4; manducatory process with two acuminate distal setae, anterior seta less than 1 / 2 length of medial seta; coxal spines present on coxa I only, comprising a transverse, contiguous series of seven or eight tridentate blades, which arise from a lightly sclerotized or translucent hillock, the central ramus of each blade (except the basal two) sharply acumino-spatulate and extending beyond the lateral rami (Figs 21 A, 22 C); bisetose intercoxal tubercle present between coxae III and IV, tear drop-shaped (Fig. 21 A). Chelicera (Figs 20 C, 22 B, E): large, approximately as long as carapace, 2.37 - 2.41 x longer than broad; five setae present on hand, movable finger with a medial seta, all setae acuminate, ventrobasal seta shorter than others; exterior condylar lyrifissure and exterior lyrifissure exist, palm with one extra (between sub-basal seta and an accessory seta). Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 13 - 15 acute teeth, distal one largest; movable finger with 12 retrorse contiguous teeth of equal length, plus three or four round proximal teeth, 15 or 16 in total; galea represented by a very slight bump on movable finger. Serrula exterior with 21 blades and serrula interior with 17 - 20 blades. Rallum in two rows and composed of 11 blades with fine pinnate, of which the basal-most blade shorter than the others (Fig. 22 E). Pedipalp (Figs 20 A, 22 D, 23 A, B): surfaces mostly with fine reticulations; long and slender, trochanter 1.78 - 2.00, femur 7.24, patella 2.44 - 2.47, chela 6.21 - 6.22, hand 2.26 - 2.36 x longer than broad; femur 2.62 - 2.80 x longer than patella; movable chelal finger 1.61 - 1.74 x longer than hand and 0.61 - 0.63 x longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella and femur, respectively (Fig. 22 D). Chelal palm robust and slightly constricted towards fingers. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria, ib, isb, eb, esb, and ist clustered at the base of fixed finger, ist slightly distal to esb, esb close to ist than to eb; it slightly distal to est, situated subdistally and forming a pair; et situated subdistally, very close to chelal teeth; dx situated distal to et, near the tip of fixed finger; sb distinctly closer to b than to st (Fig. 23 A). Microsetae (chemosensory setae) absent on hand and both palpal fingers. Sensilla absent. Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger and well-spaced teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 22 - 24 teeth, slightly retrorse and pointed; movable chelal finger with 16 - 19 teeth (slightly smaller than teeth on fixed chelal finger); a small tubercle between the seventh and eighth teeth present (near trichobothrium t) (Fig. 23 A). Chelal fingers straight in dorsal view (Fig. 23 B). Opisthosoma: generally typical, ovate, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I-XII: 2: 4: 6: 6: 6: 7 - 8: 8: 7: 6: 4: TT: 0, tergites VII-IX each with an unpaired median seta, one lyrifissure present on each side of tergites IV-IX. Sternal chaetotaxy III-XII: 6 - 9: 10 - 14: 13 - 14: 12: 12 - 13: 12 - 13: 10 - 11: 8 - 9: 0: 2, one lyrifissure present on each side of tergite III. Anterior genital operculum with 10 - 12 setae, genital opening pit-like, with seven marginal setae on each side, 24 - 26 in total, with a pair of lyrifissures present anterolateral and posteriolateral to genital opening, respectively (Fig. 21 B). Legs (Fig. 23 C, D): generally typical, long, and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.74 x longer than patella and with one lyrifissure at the base of femur; tarsus 2.11 - 2.26 x longer than tibia. Femoropatella of leg IV 3.70 - 3.88 x longer than deep and with one lyrifissure at the base of femur; tibia 5.92 - 6.00 x longer than deep; with a long tactile seta on both tarsal segments: basitarsus 4.00 x longer than deep (TS = 0.22 - 0.28), telotarsus 11.13 - 12.43 x longer than deep and 2.42 - 2.47 x longer than basitarsus (TS = 0.26 - 0.27). Setae of leg I (trochanter to tibia) 1: 13: 12 - 15: 14 - 17, setae of leg IV (trochanter to basitarsus) 2: 2: 6: 16: 14. Arolium slightly shorter than the claws, not divided; claws simple. Dimensions of adult males (length / breadth or, in the case of the legs, length / depth in mm). Males: body length 2.50. Pedipalps: trochanter 0.32 / 0.16 - 0.18, femur 1.23 / 0.17, patella 0.44 - 0.47 / 0.18 - 0.19, chela 1.68 - 1.74 / 0.27 - 0.28, hand 0.61 - 0.66 / 0.27 - 0.28, movable finger length 1.06. Chelicera 0.64 - 0.65 / 0.27, movable finger length 0.34. Carapace 0.63 / 0.61 - 0.62. Leg I: trochanter 0.24 - 0.25 / 0.18, femur 0.73 - 0.75 / 0.10, patella 0.42 - 0.43 / 0.09, tibia 0.34 - 0.37 / 0.07, tarsus 0.77 - 0.78 / 0.06 - 0.07. Leg IV: trochanter 0.33 - 0.36 / 0.19 - 0.22, femoropatella 0.97 - 1.00 / 0.25 - 0.27, tibia 0.77 - 0.78 / 0.13, basitarsus 0.36 / 0.09, telotarsus 0.87 - 0.89 / 0.07 - 0.08. Adult female (Figs 19 B, 21 C). Mostly same as males; tergal chaetotaxy I-XII: 2: 4: 6: 6: 6: 7: 8: 9: 7: 4: TT: 0; sternal chaetotaxy IV-XII: 10: 13: 11: 12: 12: 11: 8: 0: 2; anterior genital operculum with seven setae, posterior margin with 11 marginal setae, 18 in total; leg IV with a long tactile seta on both tarsal segments: basitarsus 3.60 x longer than deep (TS = 0.25), telotarsus 11.63 x longer than deep and 2.58 x longer than basitarsus (TS = 0.24). Body length 1.93. Pedipalps: trochanter 0.32 / 0.19 (1.68 x), femur 1.28 / 0.20 (6.40 x), patella 0.51 / 0.21 (2.43 x), chela 1.76 / 0.31 (5.68 x), hand 0.66 / 0.31 (2.13 x), movable chelal finger length 1.11. Chelicera 0.70 / 0.30 (2.33 x), movable finger length 0.38. Carapace 0.68 / 0.68 (1.00 x). Leg I: trochanter 0.24 / 0.19 (1.26 x), femur 0.77 / 0.12 (6.42 x), patella 0.45 / 0.11 (4.09 x), tibia 0.36 / 0.08 (4.50 x), tarsus 0.83 / 0.08 (10.38 x). Leg IV: trochanter 0.35 / 0.22 (1.59 x), femoropatella 1.05 / 0.27 (3.89 x), tibia 0.80 / 0.14 (5.71 x), basitarsus 0.36 / 0.10 (3.60 x), telotarsus 0.93 / 0.08 (11.63 x).	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
B8502145B46C58B48237EED883C95229.taxon	distribution	Distribution and habitat. This species is known only from the type locality, Wulibei Cave (Figs 1 A, 18 A-C), which is located ~ 1.2 km east of Yangguan Village (Weining County). This limestone cave has an elongated entrance (~ 2.5 m high and 8 m wide) with some corn stalks scattered nearby. Entrance of the cave has a large muddy cave hall, connected to a small hall through a narrow tunnel, which is a more enclosed, completely dark space, covered with gravel, with temperatures ~ 10 ° C and humidity ~ 90 %. The specimen was collected under a stone in a small cave hall.	en	Gao, Zhizhong, Hou, Yanmeng, Zhang, Feng (2023): Four new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Pseudotyrannochthoniidae) from Guizhou, China. ZooKeys 1139: 33-69, DOI: http://dx.doi.org/10.3897/zookeys.1139.96639, URL: http://dx.doi.org/10.3897/zookeys.1139.96639
