identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0A7C56947F60536A9697407BD5220B17.text	0A7C56947F60536A9697407BD5220B17.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stegosauridae L.Trevisan 1971	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Stegosauridae indet. Buffetaut et al. 2001</p>
            <p>Material.</p>
            <p>SM2011-1-001 (renumbered from KPS2- 1 in Buffetaut et al. (2001)), a single posterior dorsal vertebra (Fig. 3A).</p>
            <p>Locality and age.</p>
            <p>Ban Khok Sanam locality, Kham Muang District, Kalasin Province; the lower Phu Kradung Formation, Late Jurassic.</p>
            <p>Previous study.</p>
            <p> This the first evidence of a thyreophoran dinosaur in Southeast Asia. The specimen is identified as belonging to the family  Stegosauridae , which is more advanced than primitive taxa, such as huayangosaurid  Huayangosaurus from the Middle Jurassic of China (Buffetaut et al. 2001). The specimen has a much higher neural arch and more upright transverse processes, indicating that it should be referred to the  Stegosauridae (Buffetaut et al. 2001). </p>
            <p>Comments.</p>
            <p> The anterior part of the centrum and the neural arch of SM2011-1-001 has been destroyed, but likely to possess centra of the dorsal vertebrae longer than wide as most stegosaurians, except for  Miragaia longicollum (Maidment et al. 2008; Mateus et al. 2009). </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/0A7C56947F60536A9697407BD5220B17	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
B92F9B51D7F95D00A7A5EF0572561C6D.text	B92F9B51D7F95D00A7A5EF0572561C6D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mandschurosaurus laosensis " Hoffet 1944	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> "  Mandschurosaurus laosensis " Hoffet, 1944</p>
            <p>Material.</p>
            <p>unnumbered specimen consists of vertebrae, ilium, and femora.</p>
            <p>Locality and age.</p>
            <p> Muong Phalane, Savannakhet Province of Laos;  Grès Supérieurs Formation (≈ the Khok Kruat Formation), Aptian-Albian. </p>
            <p>Previous study.</p>
            <p> Mandschurosaurus was the first dinosaur named from China, its material collected from the Late Cretaceous Yuliangze Formation (Maastrichtian) in Heilongjiang (Amur) River area between China and Russia (Godefroit et al. 2011). It was initially referred to the genus "  Trachodon "  Trachodon amurense (Riabinin 1925), but was later re-assigned in 1930 to a new genus as  Mandschurosaurus amurensis (Godefroit et al. 2011).  M. amurensis is a large hadrosaurid (duck-billed dinosaurs), based on a poorly preserved and incomplete skeleton, and is often considered as a nomen dubium (Horner et al. 2004). Hoffet described the Laotian ornithopod material as "  Mandschurosaurus laosensis " (Hoffet 1944), which was considered Late Cretaceous (Senonian) in age (Buffetaut 1991). However, some palaeontologists consider "  M. laosensis " a nomen dubium (Buffetaut 1991; Horner et al. 2004). "  M. laosensis " is potentially a non-hadrosaurid iguanodontian, such as  Siamodon ,  Ratchasimasaurus , and  Sirindhorna from rocks of the same age in Thailand. Hoffet also suggested that another ilium which is more robust than "  M. laosensis ", indicated the presence of a second taxon (Buffetaut 1991; Allain et al. 1999). </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/B92F9B51D7F95D00A7A5EF0572561C6D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
6D9069BBBBD95916AE770F2411FB4AC5.text	6D9069BBBBD95916AE770F2411FB4AC5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Napaisaurus guangxiensis Ji & Zhang 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Napaisaurus guangxiensis Ji &amp; Zhang, 2022</p>
            <p>Material.</p>
            <p>The holotype, FS-20-007 to 008, a right ischium and ilium.</p>
            <p>Locality and age.</p>
            <p>Napai Basin, Fusui County; Xinlong Formation, Early Cretaceous (Aptian).</p>
            <p>Previous study.</p>
            <p>This is the first named basal iguanodontian taxon from southern China, based on characteristics of the ilium and ischium which differ from other known iguanodontian taxa (Ji and Zhang 2022).</p>
            <p>Comments.</p>
            <p> The authors did not perform a phylogenetic analysis of the taxon. It is necessary to compare with the unpublished ischium and ilium of  S. khoratensis . </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/6D9069BBBBD95916AE770F2411FB4AC5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
A2E6CC7C3B1E5F3B94E07E6F16A5FC13.text	A2E6CC7C3B1E5F3B94E07E6F16A5FC13.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psittacosaurus sattayaraki Buffetaut & Suteethorn 1992	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Psittacosaurus sattayaraki Buffetaut &amp; Suteethorn, 1992</p>
            <p>Material.</p>
            <p>holotype SM2016-1-163 (renumbered from TF 2449a by Buffetaut and Suteethorn (1992)), right dentary (Fig. 3F); SM2016-1-164 (renumbered from TF 2449b by Buffetaut and Suteethorn (1992)) maxilla fragment.</p>
            <p>Locality and age.</p>
            <p>Ban Dong Bang Noi, Lat Yai Subdistrict, Mueang District, Chaiyaphum Province; late Early Cretaceous Khok Kruat Formation (Aptian).</p>
            <p>Previous study.</p>
            <p> Apart from the ornithopods mentioned above, another valid taxon from the Khok Kruat Formation is a small basal ceratopsian.  P. sattayaraki was described from a well-preserved dentary (SM2016-1-163) and a maxilla fragment (SM2016-1-164), and it is the southernmost known occurrence of this genus (Buffetaut and Suteethorn 1992). However, the incompleteness of the material makes the validity of the taxa questionable (Sereno 2000; You and Dodson 2004). Buffetaut, Suteethorn, and Khansubha (2007) defended  P. satayaraki as a species of  Psittacosaurus , based on the tooth and dentary morphology clearly exhibiting fan-shaped tooth crowns bearing bulbous primary ridge, and a markedly convex alveolar border in lateral or medial view and an incipient ventral flange of the dentary. The taxon has been accepted as a valid species of  Psittacosaurus in other reviews (Averianov et al. 2006; Lucas 2006). </p>
            <p>Comment.</p>
            <p> Although  Psittacosaurus was abundant in the Early Cretaceous of Eastern Asia (especially China, Mongolia, and Siberia), it is worth noting that material of  Psittacosaurus seems to be scarce in Southeast Asia. In Thailand, only fragmentary materials were discovered in Chaiyaphum and Khon Kaen Provinces, and have not been found in other Khok Kruat localities (Manitkoon et al. 2022). </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/A2E6CC7C3B1E5F3B94E07E6F16A5FC13	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
DE4F771F0B0B5E6B844A1855F089B07E.text	DE4F771F0B0B5E6B844A1855F089B07E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psittacosaurus sp. Buffetaut et al. 2007	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Psittacosaurus sp. Buffetaut et al., 2007</p>
            <p>Material.</p>
            <p>SM2016-1-080, a right femur (Fig. 3E); unnumbered material including isolated tooth, a dentary fragment, a dorsal vertebra, and a fragmentary sacrum.</p>
            <p>Locality and age.</p>
            <p>SM2016-1-080 was collected from the banks of the Nam Phong River, Ban Bueng Klang Village, Nam Phong District, Khon Kaen Province; other materials were collected from Phu Hin Rong, Mancha Khiri District, Khon Kaen Province; all specimens belong to late Early Cretaceous Khok Kruat Formation (Aptian-Albian).</p>
            <p>Previous study.</p>
            <p> Additional postcranial specimens referred to as  Psittacosaurus have been found in Khon Kaen Province (Buffetaut et al. 2007). The alveolar margin of the dentary fragment is markedly convex resembling the holotype of  P. sattayaraki , but it is a larger individual (Buffetaut et al. 2007). </p>
            <p>Comment.</p>
            <p>Only the femur is still kept in the Sirindhorn Museum.</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/DE4F771F0B0B5E6B844A1855F089B07E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
885B90BAABEB5011A86E1B4A0473050D.text	885B90BAABEB5011A86E1B4A0473050D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ratchasimasaurus suranareae ? Psittacosaurid indet. Mo et al. 2016	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Ratchasimasaurus suranareae?Psittacosaurid indet. Mo et al., 2016</p>
            <p>Material.</p>
            <p>Unnumbered specimens including distal end of a right femur.</p>
            <p>Locality and age.</p>
            <p>Napai Basin, Guangxi Zhuang Autonomous Region; Xinlong Formation, Early Cretaceous (Aptian).</p>
            <p>Previous study.</p>
            <p>Mo et al. (2016) reported an incomplete femur, which resembles a psittacosaurid. Although the condition of the specimen is poor, this group is widely distributed in the early Cretaceous of China, Mongolia, Siberia, Thailand, and Laos. It is possible that this basal ceratopsian could be present in the Xinlong Formation (Buffetaut and Suteethorn 1992; Buffetaut et al. 2007; Mo et al. 2016).</p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/885B90BAABEB5011A86E1B4A0473050D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
373228C245C758EAA85062FC76C5C5D4.text	373228C245C758EAA85062FC76C5C5D4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ratchasimasaurus suranareae Basal neornithischian indet. (‘ Dan Luang neornithischian’)	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> 
Ratchasimasaurus suranareae Basal neornithischian indet. ('Dan Luang 
neornithischian'
)
</p>
            <p>Material.</p>
            <p>SM2016-1-081, a left femur (Buffetaut et al. 2003, 2006, 2014) (Fig. 5).</p>
            <p>Locality and age.</p>
            <p>Dan Luang locality, Kamcha-I District, Mukdahan Province; upper Phu Kradung Formation,?Early Cretaceous.</p>
            <p>Previous study.</p>
            <p> This is the first basal neornithischian specimen to have been discovered in Thailand, excavated in 1996, but it has not yet been described (Buffetaut and Suteethorn 1998a; Buffetaut et al. 2001, 2002, 2003, 2006). Buffetaut and Suteethorn considered that it is generally similar to  Yandusaurus (=  Hexinlusaurus multidens ) from China (Buffetaut and Suteethorn 1998a). </p>
            <p>Description.</p>
            <p> The left femur is robust and almost complete, except the distal end is eroded. It is 12.08 cm in length, and has a transverse mid-shaft diameter of 1.57 cm. The shaft of the femur is bowed in the lateral view resembling that of those early ornithopods and basal neornithischians, such as  Hexinlusaurus multidens (He and Cai 1984),  Agilisaurus louderbacki (Peng 1992),  Hypsilophodon foxii (Galton 2009) and the Phu Noi neornithischian (Manitkoon et al. 2019). The femoral head is mostly intact, but the finished, articular surface is unpreserved. The anterior end of the greater trochanter is slightly convex, while the posterior end is strongly convex. The greater trochanter lies upon the same plane as the femoral head. The lesser trochanter is distinguished from the greater trochanter by a deep groove. The fourth trochanter, located on the medial margin, is incomplete, its position being in the proximal half of the femur as in PRC 150. An oval fossa occurs between the base of the fourth trochanter and the femoral shaft for muscle insertion. The distal portion of the femoral shaft shows mediolaterally expansion towards the distal condyles and, although it was damaged, posteriorly the distal condyles are separated by a caudal intercondylar groove. </p>
            <p>Comments.</p>
            <p> The Dan Luang locality has yielded mamenchisaurid teeth (Suteethorn et al. 2013), a possible mamenchisaurid rib, theropod teeth, crocodyliform osteoderms and large teeth resembling the pholidosaurid  Chalawan thailandicus (Buffetaut and Ingavat 1980; Martin et al. 2014), shell fragments of the basal trionychoid turtle  Basilochelys macrobios (Tong et al. 2009), petrified wood, and amber. The site belongs to the upper part of the Phu Kradung Formation (?basal Cretaceous), based on the appearance of large pholidosaurid crocodylomorphs and large trionychoid turtles. This contrasts with the semi-aquatic fauna, such as the small xinjiangchelyid turtles  Phunoichelys thirakhupti , and  Kalasinemys prasarttongosothi (Tong et al. 2015, 2019b), and teleosaurid  Indosinosuchus potamosiamensis (Martin et al. 2019), from the lower Phu Kradung localities, such as Phu Noi. We suggest that SM2016-1-081 belongs in a basal position in Neornithischia, as with the older Phu Noi taxon, but more specimens are needed to increase our understanding of the basal neornithischians from the upper Phu Kradung Formation. </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/373228C245C758EAA85062FC76C5C5D4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
04FB58BC51415E6292C05A8155902DA4.text	04FB58BC51415E6292C05A8155902DA4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ratchasimasaurus suranareae Basal neornithischian indet. (‘ Khok Sanam neornithischian’)	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> 
Ratchasimasaurus suranareae Basal neornithischian indet. ('Khok Sanam 
neornithischian'
)
</p>
            <p>Material.</p>
            <p>WNM-Ks-001, an isolated tooth (Fig. 4C-E).</p>
            <p>Locality and age.</p>
            <p>Khok Sanam locality, Kham Muang District, Kalasin Province; the lower Phu Kradung Formation, Late Jurassic.</p>
            <p>Comments.</p>
            <p> A dentary tooth shows the fan-shaped crown, and has the characters of the posterior teeth present in the dentary teeth of PRC 149 (Buffetaut et al. 2014) and of  Nanosaurus agilis (Carpenter and Galton 2018). Although the enamelled surface is thin, a well-developed ornamentation of ridges is still noticeable on the lingual side as the enamel is asymmetrically distributed. The margin of the laterally compressed crowns bears distinct denticles. The labial side of the tooth bears no enamel and shows worn facets. The ornamented lingual side does not show a prominent median ridge, contrasting with the European Early Cretaceous  Hypsilophodon foxii (Galton 2009) and other derived neornithischians. There is moderately developed labiolingual expansion (  ‘cingulum’ ) at the base of the crown, as seen in basal neornithischians (Barrett et al. 2005). The cylindrical root has been preserved and curves to the labial side. </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/04FB58BC51415E6292C05A8155902DA4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
D9E9B138B0E55E4FB5720E4B7D4BCC16.text	D9E9B138B0E55E4FB5720E4B7D4BCC16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ratchasimasaurus suranareae Basal neornithischian indet. (‘ Phu Noi neornithischian’)	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> 
Ratchasimasaurus suranareae Basal neornithischian indet. ('Phu Noi 
neornithischian'
)
</p>
            <p>References.</p>
            <p>Buffetaut et al. 2014; Manitkoon et al. 2019.</p>
            <p>Material.</p>
            <p>PRC 149 (renumbered from PN 13-09 in Buffetaut et al. (2014)), a lower jaw (Figs 3B, 4A, B); PRC 150, an articulated postcranial skeleton (Manitkoon et al. 2019) (Fig. 3C).</p>
            <p>Locality and age.</p>
            <p>Phu Noi locality, Kham Muang District, Kalasin Province; the lower Phu Kradung Formation, Late Jurassic.</p>
            <p>Previous study.</p>
            <p> Buffetaut and his team reported a lower jaw (PRC 149) from the Phu Noi locality. The fan-shaped teeth with a strongly ridged crown and an asymmetric enamel distribution suggests that the specimen belongs to a small ornithopod dinosaur (Buffetaut et al. 2014). A well-preserved articulated small ornithischian skeleton without skull (PRC 150) was reported from the same locality (Manitkoon et al. 2019). The pre-acetabular process of the ilium is narrow, long with a perfectly rounded termination and slightly ventrally curved. The post-acetabular process is short and high. A combination of characters resembles those Jurassic basal neornithischians from China, such as  Agilisaurus louderbacki and  Hexinlusaurus multidens (Manitkoon et al. 2019). </p>
            <p>Comments.</p>
            <p> Many taxa of basal neornithischians, once considered as early members of ornithopods, have been reclassified as the basal neornithischians (Boyd 2015; Madzia et al. 2018, 2021;  Dieudonné et al. 2020). The similarities to the Chinese taxa, for example, the axial skeleton, is composed of 15 dorsal and five sacrals; the Brevis shelf of the ilium is visible in lateral view along the entire length; a distinct supra-acetabular flange on the pubic peduncle of the ilium; prepubis tip extends beyond the distal end of the pre-acetabular process of ilium, suggesting PRC 149 is a basal neornithischian outside the clade  Ornithopoda . Numerous limb bones of various sizes from basal neornithischians were also found at the Phu Noi site, indicating that these dinosaurs were abundant in this area. It is the oldest neornithischian known, so far, from southeast Asia.  Agilisaurus louderbacki and  Hexinlusaurus multidens exhibit symmetrically distributed enamel (Peng 1992; Barrett et al. 2005), but the dentary teeth of PRC 149 and other isolated teeth from Phu Noi are different in having asymmetrically distributed enamel on the teeth as in  Yandusaurus hongheensis from the upper Shaximiao Formation,  Nanosaurus agilis (=  Drinker ,  Othnielia ,  Othnielosaurus ) from the Late Jurassic Morrison Formation, and a variety of more derived neornithischians (Barrett et al. 2005; Butler et al. 2008; Galton 2009; Carpenter and Galton 2018). Preliminary comparisons seem to indicate that it is a new taxon (Buffetaut et al. 2014; Manitkoon et al. 2019). </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/D9E9B138B0E55E4FB5720E4B7D4BCC16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
5069D8BC509D5D0BB7DC4995EE8629A5.text	5069D8BC509D5D0BB7DC4995EE8629A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ratchasimasaurus suranareae Iguanodontian indet. (‘ Khok Pha Suam iguanodontian’)	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> 
Ratchasimasaurus suranareae Iguanodontian indet. ('Khok Pha Suam 
iguanodontian'
)
</p>
            <p>Material.</p>
            <p>Isolated teeth and postcranial material including: cervical vertebra (PRC 155); dorsal vertebra (SM2021-1-113) (Fig. 3G); proximal caudal vertebra (SM2021-1-114); distal caudal vertebrae (PRC 156 and SM2021-1-115); chevron (PRC 157); metacarpal (SM2021-1-116); left femur (SM2021-1-117) (Fig. 3H); right femur (SM2021-1-118); tibia (SM2021-1-119); fibula (PRC 158); and phalanx (SM2021-1-120).</p>
            <p>Locality and age.</p>
            <p>Khok Pha Suam, Na Kham Subdistrict, Si Muang Mai District, Ubon Ratchathani Province; late Early Cretaceous Khok Kruat Formation (Aptian-Albian).</p>
            <p>Previous study.</p>
            <p>Teeth of iguanodontians are common at Khok Pha Suam, but fragmentary (Manitkoon et al. 2022). Some isolated postcranial bones belonging to iguanodontians were discovered, including vertebrae and limb bones from different individuals, based on the great discrepancy in size of the left and right femur (Manitkoon et al. 2022; Samathi and Suteethorn 2022). Samathi and Suteethorn assumed that most of the Khok Pha Suam iguanodontian material belonged to a single taxon, and found its phylogenetic position to be a non-hadrosauriform styracosternan (Samathi and Suteethorn 2022).</p>
            <p>Comment.</p>
            <p> The teeth of Thai iguanodontians exhibit a robust primary ridge displaced distally relative to the crown apicobasal axis, which is a derived feature of iguanodontians amongst ornithopods (Norman 2004, 2014). They also possess mammillate marginal denticles, which is a synapomorphy of taxa closer to hadrosaurids than basal ankylopollexians, such as  Camptosaurus (Fanti et al. 2016). </p>
            <p> The maxillary teeth of Thai forms, including  S. khoratensis (Fig. 8A),  S. nimngami (Fig. 8C), and the Khok Pha Suam taxon (Fig. 8E), are diagnostic of the level of ankylopollexian iguanodontian by displaying the prominent primary ridge, accessory ridges, and the vertical channels marking the positions occupied by successional tooth crowns (Norman 2014). The different maxillary teeth characteristics are as follows:  S. khoratensis : lanceolate-shaped crown, primary ridge separates the labial surface unevenly, distal portion of the labial surface bears weak subsidiary ridges and is slightly broader than the mesial portion (Shibata et al. 2015);  S. nimngami : diamond-shaped crown, primary ridge in a median position, no/one short weak accessory ridge is present in what is presumably the mesial half of the crown, in the apical part. (Buffetaut and Suteethorn 2011); Khok Pha Suam iguanodontian: possibly diamond-shaped crown, primary ridge divides the crown into two asymmetrical halves, at least four weak accessory ridges in what is presumably the mesial portion along the apicobasal axis. The more derived hadrosauroids usually lose the accessory ridges on the crowns of maxillary teeth, and show a shifting of the primary ridge on the maxillary tooth crown to the mid-line (You et al. 2003). </p>
            <p> The dentary teeth of the Thai forms, including  S. khoratensis (Fig. 8B),  S. nimngami (Fig. 8D), and Khok Pha Suam taxon (Fig. 8F) (not preserved in  R. suranareae ) possess a prominent primary ridge. The crowns allowed the teeth to interlock, resulting in the more elaborate structure of the dental battery. The different characteristics are as follow:  S. khoratensis : wide with leaf-shaped crown, the secondary ridge is positioned mesial to and is less prominent than the primary ridge, with no other accessory ridges (Shibata et al. 2015);  S. nimngami : leaf-shaped crown, the secondary ridge is positioned mesial to and is less prominent than the primary ridge, at least one faint accessory ridge is present on the mesial side, crown appears to be curved apicobasally; Khok Pha Suam iguanodontian: leaf-shaped crown, the less prominent secondary ridge is positioned mesial to and is less prominent than the primary ridge and at least two weak accessory ridges are present on the mesial and the distal portion. The dentary teeth of  S. nimngami and the Khok Pha Suam iguanodontian show accessory ridges, which are absent in  S. khoratensis . This character appeared in basal hadrosauroids (  Prieto-Márquez et al. 2016). In this respect,  S. khoratensis is probably more advanced than  S. nimngami and the Khok Pha Suam iguanodontian. </p>
            <p> So far, three taxa of styracosternan iguanodontians, including  S. ninngami ,  R. suranareae , and  S. khoratensis , have been described from the Khok Kruat Formation in Nakhon Ratchasima Province plus one Laotian taxon "  M. laosensis " from the  Grès Supérieurs Formation of Laos. If Khok Pha Suam iguanodontian is one of the previously-named taxa from Nakhon Ratchasima, this would provide a geographic distribution of about 400 km to the far east (Fig. 1). However, the Khok Pha Suam locality is closer to Savannakhet than Nakhon Ratchasima. The comparison between them has to be very careful, and overlapping elements are required. It would be significant if the Khok Pha Suam iguanodont is a new taxon, as it would mean that there was a diversity of up to five to six species in the region. It is necessary to compare the postcranial material between the Khok Pha Suam taxon and  S. khoratensis . </p>
            <p> If a high diversity in iguanodontians is present in Southeast Asia, then careful consideration and more materials will be required. This may be similar to the case of  Edmontosaurus , the duck-billed edmontosaurine that was widely distributed in the Late Cretaceous (Campanian-Maastrichtian) ranging from Colorado to Alaska of North America, where several genera were consolidated into two species under a single genus, based on ontogenetic variation, morphometrics and several other factors (Campione and Evans 2011; Takasaki et al. 2020). </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/5069D8BC509D5D0BB7DC4995EE8629A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
FF8D1845268454A196C103D64A4D21FC.text	FF8D1845268454A196C103D64A4D21FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ratchasimasaurus suranareae Iguanodontian indet. (‘ Savannakhet iguanodontian B’)	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> 
Ratchasimasaurus suranareae Iguanodontian indet. ('Savannakhet iguanodontian 
B'
)
</p>
            <p>Material.</p>
            <p>unnumbered specimen consists of series of dorsal vertebrae, rib, pubis, and ischium.</p>
            <p>Locality and age.</p>
            <p> Ban Lamthouay, Tang Vay District, Savannakhet Province; The  Grès Supérieurs Formation (≈ the Khok Kruat Formation), Aptian-Albian. </p>
            <p>Comments.</p>
            <p> These unpublished materials are kept in the Dinosaur Museum of Savannakhet. It is necessary to compare these with the unpublished postcranial material of  S. khoratensis . </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/FF8D1845268454A196C103D64A4D21FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
D9591B58FABD566484004F1BF470246A.text	D9591B58FABD566484004F1BF470246A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ratchasimasaurus suranareae Iguanodontian indet. Mo et al. 2016	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Ratchasimasaurus suranareae Iguanodontian indet. Mo et al., 2016</p>
            <p>Material.</p>
            <p>Unnumbered specimens including cervical vertebra, dorsal vertebra, distal end of left humerus, distal end of left femur, and isolated teeth.</p>
            <p>Locality and age.</p>
            <p>Napai Basin, Guangxi Zhuang Autonomous Region; Xinlong Formation, Early Cretaceous (Aptian).</p>
            <p>Previous study.</p>
            <p>These poorly preserved specimens cannot be identified more precisely. However, some lower teeth bear a strong median primary ridge and at least one subsidiary ridge suggesting a relatively advanced iguanodontian (Mo et al. 2016).</p>
            <p>Comments.</p>
            <p> There are many taxa of iguanodontians in Thailand and Laos, but it cannot be concluded whether these materials belong to  Napaisaurus guangxiensis or not. </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/D9591B58FABD566484004F1BF470246A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
1CBAB95C6C7656B0A046A4315AB772F8.text	1CBAB95C6C7656B0A046A4315AB772F8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ratchasimasaurus suranareae Psittacosaurid indet. (‘ Savannakhet psittacosaurid’)	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> 
Ratchasimasaurus suranareae Psittacosaurid indet. ('Savannakhet 
psittacosaurid'
)
</p>
            <p>Material.</p>
            <p>unnumbered specimen of left mandible.</p>
            <p>Locality and age.</p>
            <p> Ban Lamthouay, Tang Vay District, Savannakhet Province;  Grès Supérieurs Formation (≈ the Khok Kruat Formation), Aptian-Albian. </p>
            <p>Comments.</p>
            <p> The unpublished specimen of psittacosaurid indet. was reported and the cast of this specimen is displayed in the Dinosaur Museum of Savannakhet (Allain et al. 1999; Buffetaut et al. 2007; Cavin et al. 2019). Detailed comparison with  Psittacosaurus sattayaraki of Thailand is needed after a full description of this material has been completed. </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/1CBAB95C6C7656B0A046A4315AB772F8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
AEB51D09A7B65C4B976251EA8F9E8BED.text	AEB51D09A7B65C4B976251EA8F9E8BED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ratchasimasaurus suranareae Shibata et al. 2011	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Ratchasimasaurus suranareae Shibata et al., 2011</p>
            <p>Material.</p>
            <p>holotype NRRU-A2064, a left dentary (Fig. 3J).</p>
            <p>Locality and age.</p>
            <p>Ban Pong Malaengwan, Khok Kruat Subdistrict, Nakhon Ratchasima Province; late Early Cretaceous Khok Kruat Formation (Aptian).</p>
            <p>Previous study.</p>
            <p> R. suranareae is a hadrosauroid (Madzia et al. 2020; Shibata et al. 2015) and the material comprises a complete toothless left dentary with 18 alveoli (Shibata et al. 2011). One autapomorphy of  R. suranareae is its elongated and dorsoventrally shallow dentary ramus, with a ratio of length (from the rostral to the caudal margin)/height (at the middle of the dentary) of 6.9 (Shibata et al. 2011). It shows both primitive and derived characters for iguanodontians, such as a caudally inclined coronoid process and alveolar trough with a primitive crown impression, and a derived buccal shelf between the tooth row and the coronoid process (Shibata et al. 2011). </p>
            <p>Comment.</p>
            <p> The length of  R. suranareae dentary is 19.81 cm, which is relatively small when compared to other skull material of iguanodontians from Thailand. It is not possible to determine if it is an immature or mature individual (Shibata et al. 2011). This compares with the nearly complete right dentary of  Sirindhorna khoratensis (NRRU3001-167), which is about 38 cm in length with 20 alveoli (Shibata et al. 2015). In iguanodontians, tooth number increases during growth and single teeth also becoming relatively wider, as in  Dysalotosaurus lettowvorbecki and  Zalmoxes robustus , and there is a slight ontogenetic increase of dentary tooth positions from 10 to 13 and 8 to 10, respectively (Weishampel et al. 2003;  Hübner and Rauhut 2010). In hadrosauroids, there are many ontogenetic changes occurring in the skull and mandible, and the dentary experiences an elongation of the mandibular ramus during growth (Bell 2011; Campione and Evans 2011; Prieto-Marquez and Guenther 2018). However, the ratio of length/height of the dentary of  R. suranareae is 6.9, and approximately 5 in  S. khoratensis (NRRU3001-167), contradicting the ontogenetic tend of hadrosauroids mentioned above, if  R. suranareae is a younger stage of  S. khoratensis . </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/AEB51D09A7B65C4B976251EA8F9E8BED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
91D3D1F3B05D5AF78A7DCA97CD96601F.text	91D3D1F3B05D5AF78A7DCA97CD96601F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Siamodon nimngami Buffetaut & Suteethorn 2011	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Siamodon nimngami Buffetaut &amp; Suteethorn, 2011</p>
            <p>Material.</p>
            <p>holotype PRC-4, a left maxilla (Fig. 3I) and the referred materials, an isolated maxillary tooth (PRC-5) and a braincase (PRC-6).</p>
            <p>Locality and age.</p>
            <p>Ban Saphan Hin, Khok Kruat Subdistrict, Nakhon Ratchasima Province; late Early Cretaceous Khok Kruat Formation (Aptian) (Buffetaut and Suteethorn 2011).</p>
            <p>Previous study.</p>
            <p> S. nimngami shows a combination of iguanodontian features: maxilla shaped like an isosceles triangle, with the dorsal process located at about mid-length of the bone; a strong longitudinal bulge on the medial surface of the maxilla; maxillary teeth bear a strong median primary ridge, one short weak subsidiary ridge or no subsidiary ridge; and mamillated denticles on the crown margins similar to  Gongpoquansaurus mazongshanensis and  Probactrosaurus mongoliensis from China (Buffetaut and Suteethorn 2011). There may have been as many as 25 tooth positions in the maxilla (Buffetaut and Suteethorn 2011). It differs from hadrosaurids in the jugal, which forms a tab-like process, whereas in hadrosaurids, the expanded anterior end of the jugal contacts and overlaps a large sutural area on the maxilla (Buffetaut and Suteethorn 2011).  S. nimngami and  R. suranareae were mentioned as members of hadrosauroids (Shibata et al. 2015), but the phylogenetic analysis recovered  S. nimngami at the base of Hadrosauriformes (Madzia et al. 2020). </p>
            <p>Additional material and description.</p>
            <p> WNM-Sp-001 (Fig. 8I, J), an isolated dentary tooth from the same locality is referred to  S. nimngami . It is well preserved and apparently from a right dentary. It generally resembles a previously reported  S. khoratensis dentary tooth (Shibata et al. 2015). The crown of the tooth is leaf-shaped with enamel covering only the lingual surface. The ratio of apicobasal length / mesiodistal width: 1.87. One prominent primary ridge situated slightly distal to the mid-line makes the crown asymmetric. The secondary ridge is positioned mesial and it is less prominent. In contrast to  S. khoratensis dentary teeth, where the crowns appear to be apicobasally erect, whereas in  S. nimngami , the dentary teeth appear to be curved apicobasally. Shibata et al. (2015) noted that there were no other accessory ridges on the crown of  S. khoratensis , but WNM-KS-001 shows at least one very faint accessory ridge on the mesial side. Small denticles are present on the mesial and distal margins of the upper half of the crown. </p>
            <p>Comment.</p>
            <p> Some palaeontologists consider  S. nimngami a nomen dubium as its material does not show any autapomorphic characters, and it might be referable to some of the other taxa from the same area (Norman 2014). However, Shibata et al. (2015) showed the possibility of three iguanodontians in the Khok Kruat Formation. The holotype specimen of  S. nimngami and  S. khoratensis maxilla (NRRU-A2048) are similar in size. They likely to belong to same growth stage, but have distinct physical characteristics suggesting that they represent different animals, while comparisons to  R. suranareae are not feasible as overlapping material has not been discovered. </p>
            <p> There has been some disagreement about the type locality of  S. nimngami given by Shibata and his team, and Buffetaut and Suteethorn; however, we would like to confirm that, after corroboration from the holotype collector, Mr. Witaya Nimngam, we now know that the type locality of  S. nimngami is at Ban Saphan Hin. This is far from the type locality of  S. khoratensis and definitely not from Ban Nong Rangka as previously suggested (Fig. 7). </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/91D3D1F3B05D5AF78A7DCA97CD96601F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
8DD77FC142E2565E9261BF3E51DABEB6.text	8DD77FC142E2565E9261BF3E51DABEB6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sirindhorna khoratensis Shibata et al. 2015	<html xmlns:mods="http://www.loc.gov/mods/v3">
    <body>
        <div>
            <p> Sirindhorna khoratensis Shibata et al., 2015</p>
            <p>Material.</p>
            <p>holotype NRRU3001-166, an articulated braincase including referred skull elements: a braincase articulating with a left postorbital (NRRU-A2035), dorsal half of a braincase (NRRU3001-65), caudal portion of a braincase (NRRU3001-179), a right premaxilla (NRRU-A3623), a left maxilla (NRRU-A2048) (Fig. 3K), a right maxilla (NRRU-A2047), a right jugal (NRRU3001-7), a right quadrate (NRRU3001-175), a predentary (NRRU3001-169), a left dentary (NRRU3001-14), a right dentary (NRRU3001-167) (Fig. 3L), a right surangular (NRRU3001-137), isolated maxillary teeth (NRRU-A1956, A3630, A3649, NRRU3001-157, 163), an isolated dentary tooth (NRRU3001-28).</p>
            <p>Locality and age.</p>
            <p> Ban Saphan Hin (a different site from the  S. nimngami was found), Suranaree Subdistrict, Nakhon Ratchasima Province; late Early Cretaceous Khok Kruat Formation (Aptian). </p>
            <p>Previous study.</p>
            <p> It is known from the presence of several braincases and dentaries that at least four individuals are known. The holotype material, a braincase, shows an autapomorphy: a sagittal crest extending along the entire dorsal surface of the parietal and reaching the frontoparietal suture (Shibata et al. 2015). Referred materials display a unique combination of characters, such as antorbital fossa of the maxilla not visible, a slightly rostrally deepening dentary ramus, and dentary teeth with primary and secondary ridges, but no accessory ridges (Shibata et al. 2015). It was recovered in the basal position of non-hadrosaurid hadrosauroids (Shibata et al. 2015). However, later analysis recovered it near the base of Hadrosauromorpha, more advanced than  R. suranareae (Madzia et al. 2020). The  S. khoratensis maxilla (NRRU-A2048) has a low-angled triangular shape and the caudally positioned lacrimal process, and its 24 alveoli are rostrocaudally arranged and slightly curved caudolaterally. It is different from the isosceles triangular shape with a dorsal process positioned at the middle of the maxilla in  S. nimngami (Shibata et al. 2015).  R. suranareae has a low and elongated dentary ramus and a robust coronoid process that differs from the robust and straight dentary ramus with a subvertical coronoid process seen in  S. khoratensis (Shibata et al. 2015). </p>
            <p>Comment.</p>
            <p> S. khoratensis is considered to be the best-preserved iguanodontian ornithopod in Southeast Asia (Shibata et al. 2015). In addition to the published cranial material, the Ban Saphan Hin locality has also yielded a postcranial skeleton that is assumed to belong to  S. khoratensis because the five discovered braincases show no features to imply the presence of different taxa (Shibata et al. 2018). CT-scanning revealed the brain morphology of  S. khoratensis , which has general endocast features resembling those of non-hadrosaurid hadrosauroids (Shibata et al. 2018). </p>
        </div>
    </body>
</html>
	https://treatment.plazi.org/id/8DD77FC142E2565E9261BF3E51DABEB6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Manitkoon, Sita;Deesri, Uthumporn;Warapeang, Prapasiri;Nonsrirach, Thanit;Chanthasit, Phornphen	Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit, Chanthasit, Phornphen (2023): Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications. Fossil Record 26 (1): 1-25, DOI: http://dx.doi.org/10.3897/fr.26.e93456, URL: http://dx.doi.org/10.3897/fr.26.e93456
