taxonID	type	description	language	source
4F26309B80C35222A171AA0CE35F26AB.taxon	materials_examined	Material examined. Holotype: 1 mature (CTU-EW. 189. h 01), Kien Giang Province, Phu Quoc island, industrial tree plantations (10 ° 06 ' 11 " N, 104 ° 00 ' 51 " E), 20 m, 6 November 2016, coll. Lam HD & Trinh TKB. Paratypes: 9 matures (CTU-EW. 189. p 02), same data as for holotype. Non-types: 8 matures (CTU-EW. 189.03), same data as for holotype.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
4F26309B80C35222A171AA0CE35F26AB.taxon	diagnosis	Diagnosis. Small to medium-sized, length 51 - 54 mm, diameter 2.2 - 2.5 mm, segments 85 - 89. Prostomium epilobous. Four pairs of spermathecal pores in lateroventral intersegments 5 / 6 / 7 / 8 / 9. Male pores in xviii, without copulatory pouches. Genital markings arranged in two transverse line in front of and behind the setal ring xviii. Intestinal caeca simple. Holandric. Testis sacs in xi and xii, separated. Septa 8 / 9 / 10 absent.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
4F26309B80C35222A171AA0CE35F26AB.taxon	etymology	Etymology. " catenatus ", an adjective in apposition, to emphasize the chained seminal chamber of diverticula.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
4F26309B80C35222A171AA0CE35F26AB.taxon	description	Description. Body cylindrical, small to medium-sized, length 51 - 54 mm, diameter 2.2 - 2.5 mm, segments 85 - 89, weight 1.2 - 1.4 g. Body uniformly whitish grey, clitellum darkish brown. Prostomium 1 / 2 epilobous. First dorsal pore in 5 / 6. Pre-clitellum setae stouter and sparser than post-clitellum setae; setal numbers: 47 - 54 in viii, 46 - 48 in xxx, no setae between two male porophores; setal distance: aa> ab, zz> zy. Clitellum within xiv-xvi, slightly flattened ventrally, without setae and dorsal pores. Female pore single, in mid-ventral xiv. Four pairs of spermathecal pores in lateroventral intersegments 5 / 6 / 7 / 8 / 9. No genital markings in the spermathecal region. Male porophores small in xviii, without copulatory pouches; ventral distance between two male pores about 0.25 - 0.28 x body circumference. About 8 - 12 small genital markings ventrally arranged in two transverse lines in front of and behind the setal ring xviii. Septa 5 / 6 / 7 / 8 thick, 8 / 9 / 10 absent, 10 / 11 / 12 thick. Oesophageal gizzard within viii-x. Intestinal origin at xv. Intestinal caeca simple, within xxvii-xxvi. Last hearts in xiii. Pharyngeal micronephridia in 5 / 6 / 7. Typhlosole lamelliform. Lymph glands absent. Spermathecae paired in vi-ix. Ampulla clavate; ducts not clearly distinct from ampulla. Diverticula slightly waved, attached to the base of ampulla; seminal chamber small, chained-shaped with 2 or 3 parts. No accessory glands. Holandric. Testis sacs in x and xi, connected. Seminal vesicles well developed in xi and xii. Ovaries in 12 / 13. Ovisacs invisible. Prostate glands deeply lobuled within xvi-xx; ducts U-shaped. No accessory glands.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
BA7F06852F4C5D538BD32FF8DBC149CF.taxon	materials_examined	Material examined. Holotype: 1 mature (CTU-EW. 188. h 01), Kien Giang Province, Phu Quoc island, natural forests (10 ° 20 ' 51.3 " N, 103 ° 58 ' 58.0 " E), 44.2 m, 7 November 2016, coll. Lam HD & Trinh TKB. Paratypes: 9 matures (CTU-EW. 188. p 02) same data as for holotype. Non-types: 9 matures (CTU-EW. 188.03) same data as for holotype; 4 matures (CTU-EW. 188.04), Kien Giang Province, Phu Quoc island, natural forests (10 ° 20 ' 50 " N, 103 ° 55 ' 07 " E), 57 m, 7 November 2016, coll. Lam HD & Trinh TKB.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
BA7F06852F4C5D538BD32FF8DBC149CF.taxon	diagnosis	Diagnosis. Medium-sized, length 74 - 145 mm, average diameter 2.5 - 4.7 mm, segments 116 - 145. Prostomium epilobous. Two pairs of spermathecal pores in ventral intersegments 7 / 8 / 9. Male pores in xviii, without copulatory pouches. Two pairs of genital markings present in ventral 17 / 18 and 18 / 19. Intestinal caeca simple. Holandric. Testis sacs in xi, separated. Septa 8 / 9 / 10 / 11 absent.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
BA7F06852F4C5D538BD32FF8DBC149CF.taxon	etymology	Etymology. " phuquocensis ", named for the type locality.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
BA7F06852F4C5D538BD32FF8DBC149CF.taxon	description	Description. Body cylindrical, medium size, length 74 - 145 mm, average diameter 2.5 - 4.7 mm, segments 116 - 145, weight 0.6 - 3.7 g. Body uniformly greyish brown except darkish brown clitellum. Prostomium 2 / 3 epilobous. First dorsal pore in 12 / 13. Pre-clitellum setae stouter and sparser than post-clitellum ones; setal numbers: 42 - 61 in viii, 50 - 77 in xxx, 5 - 9 between male porophores in xviii; setal distance: aa> ab, zz> zy. Clitellum within xiv-xvi, without setae and dorsal pores. Female pore single, in mid-ventral xiv. Two pairs of spermathecal pores in ventral intersegments 7 / 8 / 9. Genital markings absent in the spermathecal region. Male pores located in the setal ring xviii, without copulatory pouches; ventral distance between two male pores about 0.28 x body circumference. Genital markings roundly pad-shaped and variable in the male region, normally with two pairs in 17 / 18 and 18 / 19, or a pair in xix (near 19 / 20). Septa 5 / 6 / 7 / 8 thick, 8 / 9 / 10 / 11 absent, 11 / 12 / 13 thick. Oesophageal gizzard within viii-xi. Intestinal origin at xv. Intestinal caeca simple, within xxvii-xxiv. Last hearts in xiii. Pharyngeal micronephridia in 5 / 6 / 7. Typhlosole lamelliform. Lymph lobuled, from 15 / 16. Two pairs of spermathecae in viii and ix. Ampulla large, heart-shaped with transverse wrinkles; ducts short and stout. Diverticula waved, attached to the base of ampulla; seminal chamber ellipsoid. Accessory glands bean-shaped, surrounding spermathecal ducts. Holandric. Testis sacs in xi, separated, the anterior pair poorly developed. Seminal vesicles well developed in xi and xii. Ovaries in 12 / 13. Ovisacs invisible. Prostate glands deeply lobuled in xvi-xx; ducts strongly coiled. Accessory glands present.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
2955009C0F355B8A83AFD634C00747A8.taxon	materials_examined	Material examined. Holotype: 1 C (CTU-EW. 190. h 01), Kien Giang Province, Phu Quoc island, natural forests (10 ° 22 ' 53 " N, 104 ° 00 ' 22 " E), 38 m, 7 November 2016, coll. Trinh TKB. Paratypes: 9 C (CTU-EW. 190. p 02) same data as for holotype. Non-types: 11 C (CTU-EW. 190.03) same data as for holotype.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
2955009C0F355B8A83AFD634C00747A8.taxon	diagnosis	Diagnosis. Medium-sized, length 139 - 170 mm, diameter 4.4 - 5.8 mm, segments 149 - 151. Prostomium epilobous. First dorsal pore in 12 / 13. Two pairs of spermathecal pores in ventral intersegments 7 / 8 / 9. Male pores located behind the setal ring xviii, on the posterior edge of genital markings. Two pairs of genital markings present in xvii and xviii. Intestinal caeca simple. Holandric. Testis sacs in xi, separated. Septa 8 / 9 / 10 / 11 absent.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
2955009C0F355B8A83AFD634C00747A8.taxon	etymology	Etymology. " poropapillatus ", an adjective in apposition, to emphasize the position of male pores on the genital markings.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
2955009C0F355B8A83AFD634C00747A8.taxon	description	Description. Body cylindrical, medium-sized, length 139 - 170 mm, diameter 4.4 - 5.8 mm, segments 149 - 151, weight 1.4 - 2.2 g. Body uniformly greyish brown except darkish brown clitellum. Prostomium 1 / 2 epilobous. First dorsal pore in 12 / 13. Pre-clitellum setae stouter and sparser than post-clitellum ones; setal numbers: 56 - 59 in viii, 84 - 85 in xxx, 17 - 20 between two male porophores in xviii; setal distance: aa = ab, zz = zy. Clitellum xiv-xvi, without setae and dorsal pores. Female pore single, in mid-ventral xiv. Two pairs of spermathecal pores in intersegments 7 / 8 / 9. Male pores behind the setal ring xviii, on the posterior edge of genital markings, without copulatory pouches; ventral distance between two male pores about 0.33 x circumference. Two pairs of genital markings present in xvii and xviii, slightly concave inside body wall. Septa 5 / 6 / 7 / 8 thick, 8 / 9 / 10 / 11 absent, 11 / 12 / 13 thin. Oesophageal gizzard within viii-xi. Intestinal origin at xv. Intestinal caeca simple, within xxii-xxvii. Last hearts in xiii. Pharyngeal micronephridia in 5 / 6 / 7. Typhlosole lamelliform. Lymph glands lobuled, from 15 / 16. Two pairs of spermathecae in viii and ix. Ampulla clavate, sometimes constricted; ducts short. Diverticula shorter than ampulla, folded several times, attached to the base of ampulla; seminal chamber oval. Each spermatheca with a bean-shaped accessory gland. Holandric. Testis sacs in xi, separated. Seminal vesicles well developed in xi and xii. Ovaries in 12 / 13. Ovisacs invisible. Prostatic glands deeply lobuled in xvi-xxi; ducts long. Two pairs of accessory glands present.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
F2DDD825F8C45E018C968B08512C1066.taxon	materials_examined	Material examined. 1 mature (CTU-EW. 002.15), residential gardens (10 ° 10 ' 43 " N, 103 ° 58 ' 24 " E), 7 m, 5 November 2016, coll. Lam HD & Trinh TKB.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
F2DDD825F8C45E018C968B08512C1066.taxon	diagnosis	Diagnosis. Medium-sized. Prostomium epilobous. Clitellum within xiv-xvii. Three pairs of spermathecal pores in ventral intersegments 6 / 7 / 8 / 9. No genital markings. Male pores in xviii with penis seta. Spermathecae with two diverticula. Holandric. Intestinal caeca absent. Septa 4 / 5 / 6 absent. Oesophageal gizzard within v-vi.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
196E96B3AFBD5AC3BA5C50D3496168DA.taxon	materials_examined	Material examined. 3 matures (CTU-EW. 011.13), residential gardens (10 ° 10 ' 52 " N, 103 ° 58 ' 08 " E), 10 m, 5 November 2016, coll. Lam HD & Trinh TKB.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
196E96B3AFBD5AC3BA5C50D3496168DA.taxon	diagnosis	Diagnosis. Medium-sized. Prostomium epilobous. First dorsal pore in 12 / 13. Four pairs of spermathecal pores in lateroventral intersegments 5 / 6 / 7 / 8 / 9. Male pores located inside shallow copulatory pouches in xviii. Two pairs of genital markings present in xvii and xix. Intestinal caeca simple. Holandric. Testis sacs connected. Septum 8 / 9 thick, 9 / 10 absent.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
DDBCC598140550F7BF150B90323146C0.taxon	materials_examined	Material examined. 22 matures (CTU-EW. 004.46) natural forests, (10 ° 10 ' 48 " N, 103 ° 58 ' 15 " E), 20 m, 5 November 2016, coll. Lam HD & Trinh TKB; 30 matures (CTU-EW. 004.47) residential gardens (10 ° 11 ' 06 " N, 103 ° 58 ' 15 " E), 13.4 m, 6 November 2016, coll. Lam HD & Trinh TKB; 13 matures (CTU-EW. 004.48) industrial tree plantations, (10 ° 06 ' 11 " N, 104 ° 00 ' 51 " E), 20 m, 5 November 2016, coll. Lam HD & Trinh TKB.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
DDBCC598140550F7BF150B90323146C0.taxon	diagnosis	Diagnosis. Medium-sized. Prostomium 2 / 3 epilobous. First dorsal pore in 12 / 13. Three pairs of spermathecal pores in lateroventral intersegments 6 / 7 / 8 / 9. Male pores located inside copulatory pouches in xviii. Male region strongly concave to form an ellipsoid or rounded area. Genital markings absent in the spermathecal region, but two pairs present in 17 / 18 and 18 / 19, in line with male pores. Intestinal caeca simple. Holandric. Testis sacs separated. Septa 8 / 9 / 10 absent.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
0761082F34585355BB8E7AB4B688FB29.taxon	materials_examined	Material examined. 21 matures (CTU-EW. 136.02), residential gardens, (10 ° 12 ' 20 " N, 103 ° 57 ' 53 " E), 26 m, 6 November 2016, coll. Lam HD & Trinh TKB.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
0761082F34585355BB8E7AB4B688FB29.taxon	diagnosis	Diagnosis. Medium-sized. Prostomium epilobous. First dorsal pore in 12 / 13. Two pairs of spermathecal pores in lateroventral intersegments 7 / 8 / 9. Male pores located inside copulatory pouches in xviii. Genital markings absent in both spermathecal and male regions. Intestinal caeca simple. Holandric. Testis sacs connected. Septa 8 / 9 / 10 absent.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
BEAC577DFE1A5D65B52E9AC8172BA55C.taxon	materials_examined	Material examined. 7 matures (CTU-EW. 014.02), residential gardens (10 ° 09 ' 29 " N, 104 ° 00 ' 01 " E), 16 m, 5 November 2016, coll. Lam HD & Trinh TKB; 5 matures (CTU-EW. 014.03), natural forests (10 ° 21 ' 08 " N, 103 ° 55 ' 42 " E), 82 m, 7 November 2016, coll. Lam HD & Trinh TKB. Diagnosis. Medium-sized. Prostomium epilobous. First dorsal pore in 12 / 13. Three pairs of spermathecal pores in lateroventral intersegments 6 / 7 / 8 / 9. Male pores located inside copulatory pouches in xviii. Two pairs of genital markings in xvii and xix, in line with male pores. Intestinal caeca lobuled. Holandric. Testis sacs in xi, separated. Septa 8 / 9 / 10 absent.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
BEAC577DFE1A5D65B52E9AC8172BA55C.taxon	description	Re-description. Body cylindrical, medium-sized, length 168 - 220 mm, diameter 6.50 - 7.58 mm, segments 99 - 133, weight 5.5 - 13.2 g. Body greyish brown, ventrum paler than dorsum, clitellum darkish brown. Prostomium 3 / 4 epilobous. First dorsal pore in 12 / 13. Pre-clitellum setae stouter and sparser than post-clitellum setae; setal number: 65 - 77 in viii, 94 - 97 in xxx, 4 - 10 between two male porophores in xviii; setal distance: aa> ab, zz> zy. Clitellum within xiv-xvi, without setae and dorsal pores. Female pore single, in mid-ventral xiv. Three pairs of spermathecal pores in lateroventral intersegments 6 / 7 / 8 / 9. Four pairs of pad-shaped genital markings in vi-ix, in line with spermathecal pores. Male pores located inside copulatory pouches in xviii. Ventral distance between two openings of copulatory pouches about 0.3 x body circumference. Two pairs of dish-shaped genital markings present in 17 / 18 and 18 / 19, in line with the openings of copulatory pouches. Septa 5 / 6 / 7 / 8 thick, 8 / 9 / 10 / 11 absent, 11 / 12 / 13 thick. Oesophageal gizzard within viii-xi. Intestinal origin at xv. Intestinal caeca within xxvii-xxix, ventrally lobuled. Last hearts in xiii. Pharyngeal micronephridia grouped in 5 / 6 / 7. Typhlosole lamelliform. Lymph glands lobuled, from 16 / 17. Spermathecae paired in vi-ix. Ampulla large, oval-shaped; ducts as long as 1 / 3 ampulla. Diverticula strongly coiled, attached to the middle of ampulla duct; seminal chamber oval. No accessory glands. Holandric. Testis sacs in xi, separated. Seminal vesicles well developed in xi and xii. Ovaries in 12 / 13; ovisacs inviable. Prostate glands deeply lobuled in xvi-xxi; ducts spherical, enlarged basally. Two pairs of large accessory glands present.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
FB172F8EC3AD5792AFA1C66770A30CD7.taxon	materials_examined	Material examined. 1 mature (CTU-EW. 191. h 01), 9 matures (CTU-EW. 191. p 02), 24 matures (CTU-EW. 191.03), Kien Giang Province, Phu Quoc island, industrial tree plantations (10 ° 09 ' 31 " N, 104 ° 00 ' 38 " E), 16 m, 5 November 2016, coll. Lam HD & Trinh TKB.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
FB172F8EC3AD5792AFA1C66770A30CD7.taxon	diagnosis	Diagnosis. Small to medium-sized. Prostomium epilobous. First dorsal pore in 12 / 13. Three pairs of spermathecal pores in dorsal intersegments 6 / 7 / 8 / 9. Male pores located inside copulatory pouches in xviii. No genital markings. Penial setae present. Intestinal caeca simple. Holandric. Testis sacs connected in xi. Septa 8 / 9 / 10 absent.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
FB172F8EC3AD5792AFA1C66770A30CD7.taxon	description	Re-description. Body cylindrical, small to medium-sized, length 81 - 118 mm, diameter 3.3 - 3.7 mm, segments 106 - 142, weight 0.5 - 0.8 g. Body uniformly whitish grey, clitellum darkish brown. Prostomium 1 / 2 epilobous. First dorsal pore in 12 / 13. Pre-clitellum setae stouter and sparser than post-clitellum setae; setal numbers: 44 - 47 in viii, 54 - 58 in xxx, 7 - 8 between two openings of copulatory pouches; setal distance aa> ab, zz ≥ zy. Clitellum within xiv-xvi, slightly flattened ventrally, without setae and dorsal pores. Female pore single, in mid-ventral xiv. Three pairs of spermathecal pores in dorsal intersegments 6 / 7 / 8 / 9, located near dorsal line. Genital markings absent in the spermathecal region. Male pores located inside copulatory pouches in xviii; ventral distance between two openings of copulatory pouches about 0.3 x body circumference. No genital markings in the male region. Septa 5 / 6 / 7 / 8 thick, 8 / 9 / 10 / 11 absent, 11 / 12 / 13 thin. Oesophageal gizzard within viii-xi. Intestinal origin at xv. Intestinal caeca simple, within xxvii-xxv. Last hearts in xiii. Typhlosole lamelliform. Lymph glands lobuled, from 17 / 18. Three pairs of spermathecae in vii-ix. Ampulla clavate; ducts long. Diverticula thin, waved, shorter than ampulla, attached to the base of ampulla; seminal chamber ellipsoid. No accessory glands. Holandric. Testis sacs in xi, connected. Seminal vesicles xi and xii well developed. Ovaries in 12 / 13. Ovisacs invisible. Prostate glands racemose, within xvii-xxii; ducts spherical, enlarged basally, ending at bell-shaped copulatory pouches. Each copulatory pouch with a penial seta. Two pairs of accessory glands attached to copulatory pouches.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
F1D6B7558C455C93A1AC74BEA2E32E8E.taxon	materials_examined	Material examined. 10 matures (CTU-EW. 006.29), industrial tree plantations (10 ° 06 ' 11 " N, 104 ° 00 ' 51 " E), 20 m, 5 November 2016, coll. Lam HD & Trinh TKB; 15 matures (CTU-EW. 018.024), residential gardens (10 ° 16 ' 04 " N, 103 ° 56 ' 28 " E), 6 November 2016, coll. Lam HD & Trinh TKB; 7 matures (CTU-EW. 018.025), natural forests (10 ° 22 ' 53 " N, 104 ° 00 ' 22 " E), 38 m, 07 November 2016, coll. Lam HD & Trinh TKB.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
F1D6B7558C455C93A1AC74BEA2E32E8E.taxon	diagnosis	Diagnosis. Medium-sized. Prostomium epilobous. First dorsal pore in 11 / 12. Three pairs of spermathecal pores in lateroventral intersegments 6 / 7 / 8 / 9. Male pores located inside copulatory pouches in xviii. Two to four penial setae. Intestinal caeca simple. No genital markings. Holandric. Testis sacs connected. Septa 8 / 9 / 10 absent.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
8F45FD3F065E57E39EE382A15E0DCCF1.taxon	materials_examined	Material examined. 15 matures (CTU-EW. 009.13) natural forests, (10 ° 10 ' 48 " N, 103 ° 58 ' 15 " E), 20 m, 5 November 2016, coll. Lam HD & Trinh TKB.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
8F45FD3F065E57E39EE382A15E0DCCF1.taxon	diagnosis	Diagnosis. Medium-sized. Prostomium epilobous. First dorsal pore in 12 / 13. Three pairs of spermathecal pores in lateroventral intersegments 6 / 7 / 8 / 9. Male pores located inside copulatory pouches in xviii. Male region not concave. Genital markings absent in the spermathecal region, but two pairs disc-shaped in 17 / 18 and 18 / 19, in line with the openings of copulatory pouches. Intestinal caeca simple. Holandric. Testis sacs separated. Septa 8 / 9 / 10 absent.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
FA5A816800885299B995B11C60BF5FDA.taxon	materials_examined	Material examined. 10 matures (CTU-EW. 005.030), industrial tree plantations (10 ° 09 ' 31 " N, 104 ° 00 ' 38 " E), 25 m, 5 November 2016, coll. Lam HD & Trinh TKB; 21 matures (CTU-EW. 005.031), residential gardens (10 ° 12 ' 09 " N, 103 ° 58 ' 06 " E), 36 m, 5 November 2016, coll. Lam HD & Trinh TKB.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
FA5A816800885299B995B11C60BF5FDA.taxon	diagnosis	Diagnosis. Medium-sized. Prostomium epilobous. First dorsal pore in 11 / 12. Two pairs of spermathecal pores in lateral intersegments 6 / 7 / 8. Male pores located inside copulatory pouches in xviii. Genital markings absent in the male region, but 1 - 3 located near each spermathecal pore. Intestinal caeca simple. Holandric. Testis sacs separated. Accessory glands present, sac-shaped. Septa 8 / 9 / 10 absent.	en	Nguyen, Tung T., Lam, Dang H., Trinh, Binh T. K., Nguyen, Anh D. (2020): The megascolecid earthworms (Annelida, Oligochaeta, Megascolecidae) in the Phu Quoc island, Vietnam, with descriptions of three new species. ZooKeys 932: 1-25, DOI: http://dx.doi.org/10.3897/zookeys.932.50314, URL: http://dx.doi.org/10.3897/zookeys.932.50314
