taxonID	type	description	language	source
543E8C277C6DFA77FDF2FDD6FA941BEB.taxon	discussion	Review of Afrotropical Nesomyrmex species groups Synoptic list of Afrotropical Nesomyrmex species	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C6DFA77FDF2FDD6FA941BEB.taxon	synonymic_list	Nesomyrmex angulatus species group Nesomyrmex angulatus (Mayr, 1862) [Algeria, Botswana, Comoros, Djibouti, Egypt, Eritrea, Ethiopia, Ghana, Kenya, Libya, Madagascar, Malawi, Mozambique, Niger, Nigeria, Oman, Saudi Arabia, Seychelles, South Africa, Sudan, Uganda, Tanzania, Tunisia, Yemen, Zimbabwe] = Nesomyrmex angulatus ilgii (Forel, 1894) = Nesomyrmex latinodis (Mayr, 1895) = Nesomyrmex angulatus concolor (Santschi, 1914) Nesomyrmex denticulatus (Mayr, 1901) [South Africa] Nesomyrmex evelynae (Forel, 1916) [Burkina Faso, Central African Republic, Ghana, D. R. Congo, Kenya, Uganda] Nesomyrmex grisoni (Forel, 1916) [Central African Republic, D. R. Congo, Ghana] Nesomyrmex inhaca sp. nov. [Mozambique] Nesomyrmex innocens (Forel, 1913) [D. R. Congo, Kenya] Nesomyrmex stramineus (Arnold, 1948) [South Africa, Swaziland] Nesomyrmex cataulacoides species group Nesomyrmex cataulacoides (Snelling, 1992) [Cameroon, Kenya] Nesomyrmex humerosus species group Nesomyrmex humerosus (Emery, 1896) [Kenya, Tanzania, Yemen] Nesomyrmex simoni species group Nesomyrmex antoinetteae Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex braunsi (Forel, 1912) [South Africa] Nesomyrmex cederbergensis Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex entabeni Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex ezantsi Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex inye Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex karooensis Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex koebergensis Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex larsenae Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex mcgregori Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex nanniae Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex njengelanga Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex ruani Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex saasveldensis Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex simoni (Emery, 1895) [South Africa] Nesomyrmex tshiguvhoae Mbanyana & Robertson, 2008 [South Africa] Nesomyrmex vannoorti Mbanyana & Robertson, 2008 [South Africa]	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C6DFA77FDF2FDD6FA941BEB.taxon	discussion	The following newly developed identification key to species groups is loosely based on Bolton (1982) and Mbanyana & Robertson (2008), and also incorporates ideas from Snelling (1992).	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C6EFA76FF50FEACFC111954.taxon	diagnosis	Definition Antennae with 12 segments; anterior clypeal lobe conspicuously convex, rounded and without a small median triangular projection; frontal carinae absent; propodeum always with moderately long to long spines, in profile distinctly longer than their basal width; petiole and postpetiole without large lateral spines, petiole sometimes with small denticles; all dorsal surfaces of body with short, blunt pilosity (with the exception of N. evelynae, which lacks standing hairs on most of first gastral tergite).	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C6EFA76FF50FEACFC111954.taxon	discussion	Comments This group contains seven species that are widely distributed in the Afrotropical region, with one species extending into the Palaearctic and Malagasy regions. All species appear to be arboreal or subarboreal. Species accounts for all group members are provided below in the review of the group. The alpha taxonomy of the group appears straightforward based on the literature (Bolton 1982; Mbanyana & Robertson 2008). However, we find some species delimitations quite problematic (see species accounts below) and it might be necessary to revise the N. angulatus species group in the future after the accumulation of more material from additional Afrotropical localities.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C6EFA76FF50FCC9FB151C01.taxon	diagnosis	Definition Antennae with 11 segments; anterior clypeal lobe conspicuously convex and rounded, without a small median triangular projection; frontal carinae absent; propodeal spines very well developed, long and spiniform; petiole and postpetiole each with a pair of large and conspicuous lateral spines; all dorsal surfaces of body without standing pilosity.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C6EFA76FF50FCC9FB151C01.taxon	discussion	Comments The N. cataulacoides species group holds only one morphologically bizarre and extraordinary species, which is impossible to confuse with any other Nesomyrmex species from the Afrotropical or any other region. The 11 - segmented antennae, lack of standing pilosity on all dorsal surfaces, and the extreme spinosity render N. cataulacoides immediately recognizable. The possession of spines / teeth on the anterior pronotal corners, the anterior and posterior propodeum, and on both waist segments is unique within the genus. The affinities of N. cataulacoides to other Nesomyrmex are difficult to ascertain, mostly due to the extreme morphological specializations. So far it is not possible to associate it closely with any other Nesomyrmex species or species group. Currently, the species is known only from Cameroon and Kenya. Based on the few collections available, this species is strictly arboreal. The observed disjunctive distribution is most likely a sampling artefact due to the scarcity of collecting in Central African canopies, and we expect that N. cataulacoides will be collected in intermediate countries in the future.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C6EFA7AFF50F99CFC0718BA.taxon	diagnosis	Definition Antennae with 12 segments; anterior clypeal lobe short, flat-margined, and never convex, with small median triangular projection; frontal carinae present, but weakly developed; propodeal spines very well developed, long and spiniform; petiole and postpetiole without lateral spines; all dorsal surfaces of body with short, blunt pilosity.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C6EFA7AFF50F99CFC0718BA.taxon	discussion	Comments This group contains only one species, which is morphologically quite unique in the Afrotropical region. Of special importance are the short, flat-margined anterior clypeal lobe with a small median triangular projection and the barrel-shaped petiolar node with its small, triangular node. These characters are in slightly modified ways also seen in several Neotropical and Malagasy species, while they are absent in the other Afrotropical species groups. However, the fact that N. humerosus shares these characters with species from other regions does not necessarily mean that N. humerosus is more closely related to them. It could also be an independent African lineage and similarities with species from other regions might be based on convergence. Despite the fact that N. humerosus does not resemble most species from the N. simoni group, it still shares characters with some species, such as the large eyes and the shape of the dorsal mesosomal outline, and it could be that N. humerosus is just an aberrant N. simoni group member. At present, it is not possible to deduce the biogeographical and phylogenetic affinities of this peculiar species. Currently, N. humerosus is known to occur in Kenya, Tanzania and Yemen. It is a rather rarely collected species and our scarce knowledge is based on just four collection events. Based on a sample collected in Kenya by the first author, it seems to live on vegetation, but it was also sampled from the ground in Tanzania and Yemen. It is possible that the species also occurs in other East African countries, such as Somalia and Mozambique, which are greatly under-sampled.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C62FA7AFF50FD77FB6D1ED7.taxon	diagnosis	Definition Antennae with 12 segments; anterior clypeal lobe conspicuously convex and rounded, without a small median triangular projection; frontal carinae absent; propodeum usually unarmed, rarely with short teeth, in profile no longer than their basal width; petiole and postpetiole without lateral spines; usually all dorsal surfaces of body with (mostly) long, fine or (rarely) short, blunt standing pilosity, sometimes pilosity reduced on a few body parts, but never completely absent from all dorsal surfaces.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C62FA7AFF50FD77FB6D1ED7.taxon	discussion	Comments The 17 species of this group are all endemic to South Africa. In contrast to the members of most other Afrotropical groups, all N. simoni group species nest and live on the ground. Mbanyana & Robertson (2008) revised this group extensively, provided a sound and functional species identification key, and presented detailed descriptions of all species. Consequently, in this study we do not go into further details concerning the N. simoni group and refer to Mbanyana & Robertson (2008).	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C62FA7FFF07FA82FA9418D5.taxon	discussion	Review of the Nesomyrmex angulatus species group The following key is partly based on Bolton (1982) and Mbanyana & Robertson (2008).	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C67FA61FD86FD48FB6719E0.taxon	description	Figs 3 C – D, 4 B, 8 A, 9 A, 14	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C67FA61FD86FD48FB6719E0.taxon	diagnosis	Diagnosis The following character combination distinguishes N. angulatus from the other members of the group: in profile mesosomal dorsum forming a single, uninterrupted flat surface without any trace of metanotal groove; petiolar peduncle short; body colour yellow to very light brown.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C67FA61FD86FD48FB6719E0.taxon	discussion	Diagnostic comments Nesomyrmex angulatus together with N. grisoni are easily separable from the other members of the group on the basis of the dorsal mesosomal outline, which is an uninterrupted, flat surface without any trace of a metanotal groove. The separation of N. angulatus from N. grisoni is less clear though. As Bolton (1982) stated in his revision, the only differentiating character is body colour, which is yellowish in N. angulatus and dark brown to black in N. grisoni. In general, body coloration is a rather weak diagnostic character and extremely variable in many ant species, and it is likely that both species are actually conspecific and the differently coloured forms represent geographic or ecological variants. The latter seems probable if one considers that N. angulatus is predominantly an arid-adapted species, while N. grisoni appears to prefer humid rain forests. Nevertheless, at the moment we hesitate to synonymise the two species and prefer to keep them separate for the following reasons. First, while there is a lot of material of N. angulatus in many museums, there is not much of N. grisoni, making comparative analyses challenging. Secondly, and more importantly, we are not fully convinced that all the material currently listed and identified as N. angulatus represents the same species. On the basis of some recent collections from Kenya and Mozambique we were able to observe a lot of morphological variation within and between localities. As already noted by Bolton (1982), the shape of the petiolar node seems to be especially variable. Consequently, we cannot rule out the possibility of dealing with a complex of more or less cryptic species. At the same time it is possible that N. angulatus is not only a very successful and widespread, but also an extremely variable species. The solution to this problem is not the aim of this study, since it requires the accumulation of an extensive amount of material from all over Africa, the Arabian Peninsula, and the Malagasy region.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C67FA61FD86FD48FB6719E0.taxon	biology_ecology	Biology Nesomyrmex angulatus was collected from a variety of habitat types, such as tropical dry forest, coastal scrub, mangrove forest, savannah, and Acacia woodland. In general it seems as if the species prefers comparatively arid environments. In addition, it is predominantly found on the trunk of trees or the lower vegetation, rarely on the ground, and it nests in pre-existing cavities of dead wood (Bolton 1982; Mbanyana & Robertson 2008).	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C67FA61FD86FD48FB6719E0.taxon	distribution	Distribution This species has by far the widest distribution range within the N. angulatus group, and likely represents the most widespread Nesomyrmex species on a global scale. It is found in the majority of African countries, as well as on the Arabian Peninsula and in most of the Malagasy region.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C79FA63FD96FBBDFD631BCE.taxon	description	Figs 10 B, 11 C – D, 12 A, 15	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C79FA63FD96FBBDFD631BCE.taxon	diagnosis	Diagnosis The following character combination separates N. denticulatus from the remainder of the group: eyes with 10 – 12 ommatidia in longest row; in profile mesosomal dorsum with conspicuously impressed metanotal groove; in dorsal view petiolar node laterally denticulate; subpetiolar process with a conspicuous tooth anteriorly followed by a long cuticular flange which runs back to the postpetiolar junction; dorsum of propodeum with standing hairs; first gastral tergite with standing hairs evenly distributed throughout.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C79FA63FD96FBBDFD631BCE.taxon	discussion	Diagnostic comments The three species, N. denticulatus, N. innocens and N. stramineus, are morphologically very similar and can be well separated from the other species by the laterally denticulate petiolar node. The separation of these three can be challenging though. Nesomyrmex denticulatus is larger in general body size, has larger eyes with more ommatidia, and a subpetiolar process with a conspicuous tooth anteriorly, followed by a long cuticular flange which runs back to the postpetiolar junction, and slightly denser pilosity.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C79FA63FD96FBBDFD631BCE.taxon	biology_ecology	Biology This species usually nests in cavities of branches on trees and bushes previously excavated by woodboring beetles, lepidopteran larvae or termites (Mbanyana & Robertson 2008). It is found in a variety of more arid habitats, such as late succession Fynbos, Succulent Karoo with large bushes, along edges of Southern Afrotemperate Forest, and possibly also Albany Thicket (Mbanyana & Robertson 2008).	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C79FA63FD96FBBDFD631BCE.taxon	distribution	Distribution Nesomyrmex denticulatus is only known from South Africa, where it seems to be relatively common in the Western and Eastern Cape regions.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7BFA63FDF8FDA3FB111C5F.taxon	description	Figs 2 D, 4 A, 10 A, 16	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7BFA63FDF8FDA3FB111C5F.taxon	diagnosis	Diagnosis The following character combination distinguishes N. evelynae from the other species of the group: in profile mesosomal dorsum with conspicuously impressed metanotal groove; petiolar peduncle long; dorsum of propodeum without standing hairs; first gastral tergite lacking standing hairs except for single transverse row on posterior end of tergite.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7BFA63FDF8FDA3FB111C5F.taxon	discussion	Diagnostic comments The recognition of this species within the N. angulatus group is fairly straightforward, since it is the only one that lacks standing hairs on most of the first gastral tergite while all other group species have standing hairs evenly distributed throughout this tergite. It also lacks standing hairs on the propodeum, a character shared only with N. inhaca sp. nov., whereas the other five species have short, standing pilosity on the propodeum.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7BFA63FDF8FDA3FB111C5F.taxon	biology_ecology	Biology It prefers rain forests where it lives in the canopy stratum. Based on canopy fogging samples from Kenya available to the first author, it appears that this species is found commonly on trees, even though in small individual numbers, suggesting smaller colony sizes.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7BFA63FDF8FDA3FB111C5F.taxon	distribution	Distribution Nesomyrmex evelynae is found in Equatorial Africa ranging from Burkina Faso and Ghana in the west through the Central African Republic and the D. R. Congo to Uganda and Kenya in the east. The known distribution is disjunctive since N. evelynae is not known from the countries between Ghana and the Central African Republic and the D. R. Congo. We consider this more of a sampling artefact though, and expect the species to be collected from the countries in between in future sampling projects.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7BFA64FDF6F9D2FAA818D2.taxon	description	Figs 9 B, 17	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7BFA64FDF6F9D2FAA818D2.taxon	diagnosis	Diagnosis The following character combination distinguishes N. grisoni from the other species of the group: in profile mesosomal dorsum forming a single, uninterrupted flat surface without any trace of metanotal groove; petiolar peduncle short; body colour dark brown to black.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7BFA64FDF6F9D2FAA818D2.taxon	discussion	Diagnostic comments As mentioned above, N. grisoni and N. angulatus are straightforwardly distinguishable from the remainder of the group. At the same time they are morphologically very close to each other and only separable on the basis of body colour. For a more in-depth discussion we refer to the species account of N. angulatus.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7BFA64FDF6F9D2FAA818D2.taxon	biology_ecology	Biology Very little information about the natural history of this species is available. It seems to live on vegetation in rain forest habitats.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7BFA64FDF6F9D2FAA818D2.taxon	distribution	Distribution Nesomyrmex grisoni is only known from the Central African Republic, the D. R. Congo and Ghana.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7CFA69FDD7FD4FFCF71E70.taxon	description	urn: lsid: zoobank. org: act: C 33 B 5 A 40 - C 3 C 0 - 4860 - BD 38 - C 99 AC 2 D 774 A 3 Figs 11 A – B, 18	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7CFA69FDD7FD4FFCF71E70.taxon	diagnosis	Diagnosis The following character combination distinguishes N. inhaca sp. nov. from the other members of the N. angulatus group: in profile mesosomal dorsum with conspicuously impressed metanotal groove; in dorsal view petiolar node not laterally denticulate; dorsum of propodeum without standing hairs; first gastral tergite with standing hairs evenly distributed throughout.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7CFA69FDD7FD4FFCF71E70.taxon	etymology	Etymology The new species is named after the type locality, Inhaca Island, to the southeast of Mozambique. The species epithet is a noun in apposition and thus invariant.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7CFA69FDD7FD4FFCF71E70.taxon	materials_examined	Type material Holotype MOZAMBIQUE: pinned worker, Maputo, Inhaca Island, 26 ° 2 ′ 9 ″ S, 32 ° 54 ′ 17 ″ E, 1 m, secondary forest, low vegetation, 21 Jun. 1992, G. D. Alpert leg. (MCZC: MCZ-ENT 00593557). Paratypes MOZAMBIQUE: 6 pinned workers, same collection data as holotype (BMNH: CASENT 0790018; HLMD: HLMD-Hym- 2395; KSMA: CASENT 0790019; MCZC: CASENT 0790017; SAMC: CASENT 0790022; ZFMK: CASENT 0790020).	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7CFA69FDD7FD4FFCF71E70.taxon	description	Worker measurements (n = 7) HL 0.69 – 0.73 (0.71); HW 0.50 – 0.53 (0.51); SL 0.47 – 0.51 (0.50); EL 0.16 – 0.17 (0.17); PH 0.26 – 0.28 (0.27); PW 0.41 – 0.42 (0.41); WL 0.85 – 0.88 (0.87); PSL 0.12 – 0.13 (0.12); PTL 0.14 – 0.16 (0.15); PTH 0.21 – 0.22 (0.22); PTW 0.19 – 0.20 (0.19); PPL 0.18 – 0.20 (0.19); PPH 0.21 – 0.22 (0.21); PPW 0.27 – 0.28 (0.28); CI 71 – 73 (72); SI 95 – 98 (96); OI 30 – 33 (32); DMI 47 – 49 (48); LMI 31 – 32 (32); PSLI 16 – 18 (17); LPeI 64 – 70 (69); DPeI 127 – 141 (131); LPpI 86 – 93 (89); DPpI 140 – 147 (145); PPI 138 – 147 (143). Worker description HEAD. Masticatory margin of mandible with five teeth, decreasing in size from largest, acute apical tooth to smallest basal denticle; clypeus arched-convex to almost triangular, anterior margin with slightly darker, lamellate, flattened ridge all-around; head in full-face view appearing relatively narrow, much longer than broad (CI 71 – 73), sides of head approximately straight, gently broadening behind eye level, narrowest directly behind posterior eye margin, and widest halfway between posterior eye margin and posterior head margin; posterior head margin concave medially; frontal carinae and antennal scrobes absent; antennal scapes moderately long, not reaching posterior head margin (SI 95 – 98). Eyes relatively large (OI 29 – 33), with eight to nine ommatidia in the longest row. MESOSOMA. In lateral view mesosomal outline relatively low (LMI 31 – 32) and flat with conspicuously impressed metanotal groove; promesonotal suture present laterally and completely absent dorsally; pronotum moderately marginate between lateral and dorsal mesosoma, anterodorsal corners not denticulate; propodeum armed with short to moderately long propodeal spines (PSLI 16 – 18), in profile spines distinctly longer than their basal width; propodeal lobes low and rounded. WAIST SEGMENTS AND GASTER. Petiolar peduncle long, anteriorly with a small tooth-like subpetiolar process; in profile petiolar node relatively low and globular, between 1.4 and 1.5 times as high as long (LPeI 64 – 70); anterior face smoothly merging with peduncle and petiolar dorsum without any angles, posterior face slightly better developed; node in dorsal view about 1.3 to 1.4 times as wide as long (DPeI 127 – 141); in dorsal view petiolar node not laterally denticulate; in profile postpetiole globular, about 1.1 to 1.2 times as high as long (LPpI 86 – 93); in dorsal view about 1.4 and 1.5 times as wide as long (DPpI 140 – 147); postpetiole in dorsal view around 1.4 to 1.5 times as wide as petiolar node (PPI 138 – 147). SCULPTURE. Mandibles shagreened to partly smooth and shiny, sometimes with very weak, superficial, irregular, longitudinal rugulae; median clypeal carina present and conspicuous, usually accompanied by one or two lateral, longitudinal, and slightly weaker rugae on each side; cephalic dorsum posteriorly and laterally strongly reticulate-rugose, medially more irregularly longitudinally rugose, ground sculpture conspicuously reticulate-punctulate; mesosoma laterally and dorsally with distinct reticulate-punctulate ground sculpture, lateral mesosoma conspicuously reticulate-rugose, dorsum reticulate-rugose with some irregular, longitudinal elements medially; legs unsculptured, smooth and shining; petiole and postpetiole with irregular reticulate-rugose sculpture superimposed on reticulate-punctulate ground sculpture; sculpture of first gastral tergite variable, some specimens only with reticulate-punctulate ground sculpture, other specimens irregularly rugose / rugulose on top of reticulate-punctulate ground sculpture. PILOSITY AND PUBESCENCE. Head, mesosoma, waist segments and gaster dorsally with sparse, erect, blunt, and moderately long pilosity, hairs shorter on head and mesosoma than on waist segments and gaster; head laterally and ventrally with short appressed to decumbent pubescence; pubescence on mesosoma and waist segments sparse to absent; first gastral tergite with short to moderately long, appressed to decumbent pubescence. COLORATION. Body uniformly yellowish to light brown, in a few specimens legs slightly lighter yellow, almost white.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7CFA69FDD7FD4FFCF71E70.taxon	discussion	Diagnostic comments Within the members of the N. angulatus species group, N. inhaca sp. nov. cannot be mistaken for N. evelynae since the latter is devoid of standing hairs on the first gastral tergite except for a single transverse row on the posterior end of the tergite, and also has very long propodeal spines and a relatively high petiolar node. Nesomyrmex inhaca sp. nov. has short, standing hairs evenly distributed throughout the first gastral tergite, shorter propodeal spines and a much lower petiolar node. The species pair N. angulatus and N. grisoni both do not possess a trace of a metanotal groove, which is obviously present in N. inhaca sp. nov. and thus it is not likely these species will be confused. Nevertheless, N. inhaca sp. nov. is morphologically closer to the trio of species N. denticulatus, N. innocens and N. stramineus. These three are, however, easily separable since they possess a petiolar node with distinct lateral denticles and have standing hairs on the propodeal dorsum, while N. inhaca sp. nov. lacks both the lateral petiolar denticles and the standing hairs on the propodeum. Moreover, N. inhaca sp. nov. has apparently longer antennal scapes based on the much higher SI (94 – 95) compared to the other three species (SI 67 – 77). This is partly due to the fact that N. inhaca sp. nov. has indeed relatively longer antennal scapes, but the comparatively narrow head with low values of HW contributes to these high SI values. Therefore, we suggest being cautious with scape length as the single diagnostic character. Intraspecific variation Since the description is based on just one collection event, the observed intraspecific variation seen in the worker caste is very low. However, the sculpture on the first gastral tergite shows some variability, as described above.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7CFA69FDD7FD4FFCF71E70.taxon	biology_ecology	Biology Nesomyrmex inhaca sp. nov. was sampled from low vegetation in secondary forest at an elevation of 1 m. Apart from this, nothing is known of its natural history.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C7CFA69FDD7FD4FFCF71E70.taxon	distribution	Distribution The new species is so far only known from one collection event on Inhaca Island in the southeast of Mozambique. Despite this apparently restricted distribution to just one island, we are reluctant to declare N. inhaca sp. nov. as endemic to Inhaca Island. With the noticeable exception of the area around Gorongosa, most of Mozambique remains severely under-sampled and our knowledge of local ant communities and species distributions is very poor to non-existent. Consequently, it is possible that N. inhaca sp. nov. will also be found on the mainland.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C71FA6BFD85FB2DFAD3187D.taxon	description	Figs 12 B, 13 A, 19	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C71FA6BFD85FB2DFAD3187D.taxon	diagnosis	Diagnosis The following character combination distinguishes N. innocens from the other members of the group: eyes with 7 – 9 ommatidia in longest row; in profile mesosomal dorsum with conspicuously impressed metanotal groove; propodeal spines short and thick, elongate-triangular and only weakly longer than their basal width; in profile petiolar node nodiform, appearing approximately as long as high; in dorsal view petiolar node laterally denticulate; subpetiolar process without a long cuticular flange running back to the postpetiolar junction; dorsum of propodeum with standing hairs; first gastral tergite with standing hairs evenly distributed throughout.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C71FA6BFD85FB2DFAD3187D.taxon	discussion	Diagnostic comments As noted in the description of N. denticulatus, the latter, N. innocens and N. stramineus are morphologically relatively close. Nesomyrmex innocens and N. stramineus differ from N. denticulatus by generally smaller body size, smaller eyes with less ommatidia, and a subpetiolar process without a long cuticular flange running back to the postpetiolar junction. The separation of N. innocens and N. stramineus is a bit more difficult, as already mentioned by Bolton (1982). Nesomyrmex innocens has shorter and thicker propodeal spines and a lower and thicker petiolar node compared to N. stramineus. It is not clear at the moment whether or not these character states are sufficient to maintain their heterospecificity in the long term. Bolton (1982) had some doubts about this, too, and it is possible that they represent geographical varieties of the same species. However, at present, based on the scarcity of the material, especially of N. innocens, we treat them as two different species.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C71FA6BFD85FB2DFAD3187D.taxon	biology_ecology	Biology Based on the limited data available, N. innocens nests in the stem of trees.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C71FA6BFD85FB2DFAD3187D.taxon	distribution	Distribution and biology This species is only known from very few specimens, collected from the D. R. Congo and Kenya.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C73FA6BFD93FD30FDF11C65.taxon	description	Figs 2 D, 8 B, 13 B, 20	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C73FA6BFD93FD30FDF11C65.taxon	diagnosis	Diagnosis The following character combination distinguishes N. stramineus from the other species of the group: eyes with 7 – 9 ommatidia in longest row; in profile mesosomal dorsum with conspicuously impressed metanotal groove; propodeal spines relatively long and thin, several times longer than their basal width; in profile petiolar node high, rectangular nodiform, appearing around twice as high as long; in dorsal view petiolar node laterally denticulate; subpetiolar process without a long cuticular flange running back to the postpetiolar junction; dorsum of propodeum with standing hairs; first gastral tergite with standing hairs evenly distributed throughout.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C73FA6BFD93FD30FDF11C65.taxon	discussion	Diagnostic comments As pointed out above, N. stramineus is very close to N. denticulatus and N. innocens. For more details on its differentiation from them we refer to the species account of N. innocens.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C73FA6BFD93FD30FDF11C65.taxon	biology_ecology	Biology Nesomyrmex stramineus was sampled in savannah woodlands and Afromontane forests, where it lives in dead wood on trees (Mbanyana & Robertson 2008).	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
543E8C277C73FA6BFD93FD30FDF11C65.taxon	distribution	Distribution This species occurs only in South Africa and Swaziland. It seems to be rather rare since it is only known from a few collection events.	en	Garcia, Francisco Hita, Mbanyana, Nokuthula, Audisio, Tracy Lynn, Alpert, Gary D. (2017): Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique. European Journal of Taxonomy 258: 1-31, DOI: http://dx.doi.org/10.5852/ejt.2017.258
