taxonID	type	description	language	source
645687B47377EF0119BFEFC1ABECFC70.taxon	diagnosis	Diagnosis: — Acrobolbus mashpianus is similar to Acrobolbus setaceus in its toothed leaf margins but differs by the strongly asymmetrically bilobed leaves that are more deeply bifid (1 / 2 – 4 / 5 vs. 1 / 4 – 1 / 3) and the longer teeth of the leaf margin (5 - 8 cells vs. 1 - 3 cells long).	en	Burghardt, Michael (2022): Acrobolbus mashpianus (Acrobolbaceae, Marchantiophyta) sp. nov. from the Northern Andes of Ecuador-Notes on the Bryophytes of Ecuador VII. Phytotaxa 577 (1): 133-138, DOI: 10.11646/phytotaxa.577.1.7, URL: http://dx.doi.org/10.11646/phytotaxa.577.1.7
645687B47377EF0119BFEFC1ABECFC70.taxon	materials_examined	Type: — ECUADOR. Pichincha: Metropolitan District of Quito, Parrish Pacto, Mashpi Ecological Reserve, Jungle Swing trail, elev. 913 m, 0 ° 09 ’ 58.13 ’’ N 78 ° 52 ’ 50.36 ’’ W, 22 February 2019, M. Burghardt MB 11033 (holotype: QCNE!).	en	Burghardt, Michael (2022): Acrobolbus mashpianus (Acrobolbaceae, Marchantiophyta) sp. nov. from the Northern Andes of Ecuador-Notes on the Bryophytes of Ecuador VII. Phytotaxa 577 (1): 133-138, DOI: 10.11646/phytotaxa.577.1.7, URL: http://dx.doi.org/10.11646/phytotaxa.577.1.7
645687B47377EF0119BFEFC1ABECFC70.taxon	description	Plants prostrate, scattered among other bryophytes; soft-textured, whitish green, turning brownish in the herbarium. Shoots small, up to 1.5 cm long and 1.3 – 1.5 mm wide with leaves; microphyllous stoloniferous base lacking. Stems light brown, slightly dorsoventrally flattened; near base ca. 85 – 140 × 60 – 100 µm thick, in cross-section about 4 – 6 cells high; differentiation in cortex and medulla lacking, cells thin-walled, ca. 10 – 23 × 11 – 17 µm, without trigones or only tiny ones; stem surface papillose to striate-papillose. Branching sparse, lateral-intercalary of the Plagiochila - type; rhizoids scattered along the stem. Shoots with contiguous to subimbricate foliation. Leaves succubous, obliquely to widely spreading, plane; ventral leaf margin not decurrent; dorsal margin not to shortly decurrent by few cells; leaves cuneate to occasionally subrectangular, widest above the middle, asymmetrically bilobed to 4 / 5, occasionally only to 1 / 2, with the dorsal lobe distinctly smaller than the ventral lobe; ca. 450 – 720 × 270 – 400 µm, 1.5 – 2.4 times as long as wide; lobes elongate triangular to lanceolate, acuminate, margins plane and crenulated, especially towards the (2 –) 4 – 7 (– 8) - celled uniseriate lobe tips; lobes usually widely diverging, sometimes subparallel to somewhat connivent. Leaf margins beset with one to four elongate triangular to more commonly ciliate teeth, teeth 5 – 8 cells long, consisting of a short 2 - celled base and a 4 – 7 cell long uniseriate tip. Leaf areolation ± regular; cells in the center of the ventral leaf lobe isodiametric to distinctly elongated with scattered cells shorter than wide, (30 –) 35 – 42 (– 52) × (29 –) 30 – 38 (– 47) µm, (0.9 –) 1.1 – 1.4 (– 1.8) times as long as wide; cells of the lobe tips (uniseriate part) slightly elongated, (29 –) 30 – 35 (– 40) × (20 –) 22 – 24 (– 26) µm, (1.1 –) 1.2 – 1.5 (– 1.7) times as long as wide; cell walls thin, trigones small, triangular with convex sides, intermediate thickenings absent, leaf surfaces striate-papillose with some papillae rounded. Oil bodies of the Jungermannia - type, brown, 4 – 6 per cell, ellipsoidal to sometimes globose, surface granular. Underleaves vestigial, one-to-two-celled. Asexual reproduction possibly by fragmenting leaf lobes and teeth. Male and female plants were not seen.	en	Burghardt, Michael (2022): Acrobolbus mashpianus (Acrobolbaceae, Marchantiophyta) sp. nov. from the Northern Andes of Ecuador-Notes on the Bryophytes of Ecuador VII. Phytotaxa 577 (1): 133-138, DOI: 10.11646/phytotaxa.577.1.7, URL: http://dx.doi.org/10.11646/phytotaxa.577.1.7
645687B47377EF0119BFEFC1ABECFC70.taxon	biology_ecology	Habitat and distribution: — Acrobolbus mashpianus is only known from the type collection from the Mashpi Ecological Reserve in the Province of Pichincha. For a detailed description of the locality see Burghardt (2020). The holotype was collected in a humid evergreen premontane forest, where it grew on a tree trunk in appressed mats of a rich liverwort community. Associated species include Ceratolejeunea cornuta (Lindenberg 1829: 23) Stephani (1895: 65), Leptoscyphus trapezoïdeus (Montagne 1843: 251) Söderström (2013: 27), Plagiochila rudischusteri Robinson (1988: 199), Prionolejeunea aemula (Gottsche 1845: 338) Evans (1904: 219), P. muricatoserrulata (Spruce 1884: 155) Stephani (1913: 223), and P. trachyodes (Spruce 1895: 338) Stephani (1913: 215).	en	Burghardt, Michael (2022): Acrobolbus mashpianus (Acrobolbaceae, Marchantiophyta) sp. nov. from the Northern Andes of Ecuador-Notes on the Bryophytes of Ecuador VII. Phytotaxa 577 (1): 133-138, DOI: 10.11646/phytotaxa.577.1.7, URL: http://dx.doi.org/10.11646/phytotaxa.577.1.7
645687B47377EF0119BFEFC1ABECFC70.taxon	etymology	Etymology: — The name of the new species refers to the type locality.	en	Burghardt, Michael (2022): Acrobolbus mashpianus (Acrobolbaceae, Marchantiophyta) sp. nov. from the Northern Andes of Ecuador-Notes on the Bryophytes of Ecuador VII. Phytotaxa 577 (1): 133-138, DOI: 10.11646/phytotaxa.577.1.7, URL: http://dx.doi.org/10.11646/phytotaxa.577.1.7
