taxonID	type	description	language	source
03914E40E956FF98FF3DBCEFFB69FA83.taxon	type_taxon	Type-species. Scolopocryptops melanostoma Newport, 1845, by subsequent designation by Lucas (1849).	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E956FF98FF3DBCEFFB69FA83.taxon	diagnosis	Diagnosis. The most recent one is provided by Schileyko et al. 2020: 10. Range. China, Taiwan, Japan, Korea, Vietnam, Philippines, Sunda Archipelago, New Guinea, Fiji, West Africa, North, Central and South America, Caribbean islands.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E956FF98FF3DBCEFFB69FA83.taxon	discussion	Remarks. The new general list of Scolopocryptops ’ species is presented in Appendix 1; the comparative morphological data for the main part of East Asian species are summarized in Appendix 5.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E956FF9CFF3DBA64FF16FE26.taxon	description	Figs 2 – 4	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E956FF9CFF3DBA64FF16FE26.taxon	materials_examined	Material. SON LA Province, Ta Xua NR: 1 spm (SVR. TX. 2019.033), regenerating forest, 21.31472 ° N 104.52052 ° E, 1555 m a. s. l., 05.10.2019, col. Le X. Son; 2 spms (SVR. TX. 001, SVR. TX. 007), bamboo forest, 21.35278 ° N 104.6740 ° E, 777 m a. s. l., 09.02.2017, col. Le X. Son, Ha K. Loan; 1 spm (SVR. TX. 005), mixed forest, 21.36528 ° N 104.6619 ° E, 831 m a. s. l., 09.022017, col. Vu T. Ha; 1 spm (SVR. TX. 015), 21.33978 ° N 104.6898 ° E, 09.022017, col. Nguyen D. Hung. CAO BANG Province, Phia Oac — Phia Den NP: 1 spm (SVR. PO. 020), natural forest, 22.60941 ° N 105.86414 ° E, 1709 m a. s. l., 12.07.2017, col. Le X. Son; 1 spm (SVR. PO. 213), natural forest, 22.600946 ° N 105.87015 ° E, 1605 m a. s. l., 04.06.2019, col. Le X. Son; 1 spm (SVR. PO. 236), mixed forest, 22.60321 ° N 105.87344 ° E, 1508 m a. s. l., 08.06.2019, col. Le X. Son; 1 spm (SVR. PO. 203.1), mixed forest, 22.60901 ° N 105.86814 ° E, 1596 m a. s. l., 04.06.2019, col. A. Abramov. NGHE AN Province, Pu Hoat NR: 1 spm (SVR. PH. 024), natural forest, 19.77022 ° N 104.78436 ° E, 1042 m a. s. l., 17.05.2019, col. Le X. Son; 1 spm (SVR. PH. 050), natural forest, 19.75619 ° N 104.78569 ° E, 1103 m a. s. l., 19.05.2019, col. Le X. Son; 2 spms (SVR. PH. 073, SVR. PH. 074), mixed forest, 19.77069 ° N 104.78408 ° E, 1009 m a. s. l., 25.05.2019, col. Le X. Son & Ngo T. Dung. QUANG NAM Province: 1 spm (SVR. STh. 00085) Song Thanh NR, regenerated forest, 15.54008 ° N 107.38300 ° E, 1165 m a. s. l., 01.05.2019, col. Le X. Son. KON TUM Province: 1 spm (SVR. TNh. 116) Thach Nham forest, mixed forest, 14.71878 ° N 108.31711 ° E, 1129 m a. s. l., 04.06.2016, col. Le X. Son; 1 spm (SVR. TNh. 025) Thach Nham forest, natural forest, 14.75550 ° N 108.29761 E, 1363 m a. s. l., 07.06.2016, col. Le X. Son; 3 spms (SVR. TNh. 028 - 029 - 030) Thach Nham forest, natural forest, 14.75397 ° N 108.29878 E, 1350 m a. s. l., 07.06.2016, col. Le X. Son; 1 spm (SVR. TNh. 043) Thach Nham forest, mixed forest, 14.74731 ° N 108.30117 ° E, 1256 m a. s. l., 08.06.2016, col. Le X. Son; 1 spm (SVR. TNh. 046) Thach Nham forest, mixed forest, 14.74914 ° N 108.30017 ° E, 1241 m a. s. l., 08.06.2016, col. Le X. Son; 1 spm (SVR. TNh. 056) Thach Nham forest, mixed forest, 14.74328 ° N 108.30481 ° E, 1213 m a. s. l., 09.06.2016, col. Le X. Son; 1 spm (SVR. TNh. T 3) Thach Nham forest, natural forest, 14.72544 ° N 108.31506 ° E, 1035 m a. s. l., 08.05.2015, col. Nguyen Huu Thuc; 1 spm (SVR. NL. 030), Ngoc Linh NR, pine forest, 15.12594 ° N 107.81256 ° E, 1160 m a. s. l., 22.09.2019, col. Le X. Son. HA TINH Province: 1 ad (Rc 6686 in ZMMU), Vu Quang, under log. 1200 m a. s. l., 08.08.1997, col. М. V. Kalyakin; 1 ad (Rc 6738 in ZMMU), Vu Quang village, М 13, 08.1997, col. М. V. Kalyakin. Comparative material. 4 ad (Rc 7106 in ZMMU), Central China, Shaanxi Province, Panda area, Fo ping NR, 1600 m a. s. l., 06 – 11.04.1999, col. V. Siniaev & A. Plutenko.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E956FF9CFF3DBA64FF16FE26.taxon	diagnosis	Diagnosis. 2 basal antennal articles virtually lacking setae; cephalic plate marginate laterally; tergites 8 – 18 with complete paramedian sutures, ultimate tergite with complete lateral margination; LBS 7 lacks spiracles; ultimate legs lacking setae (more rarely their distal articles setose), ultimate prefemur with two typical (ventral and dorsomedial) spinous processes.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E956FF9CFF3DBA64FF16FE26.taxon	description	Description of adult SVR. PO. 020 [data from ZMMU specimens in square brackets where it differs]. Body length 36.8 mm, the width of LBS 10 ca 3.33 mm. Antennae (Figs 2 AB) composed of 17 articles (left one of 15 ones, it seems to be damaged / regenerated) of them 2 basal ones practically not setose [at least] dorsally, following articles densely covered by minute setae. Cephalic plate (Fig. 2 A) nearly as long as wide, not setose, [very sparsely] and definitely punctate, marginate laterally (posterior margination absent). Forcipular segment (Fig. 2 B): coxosternite, trochanteroprefemora and basal part of tarsungulae coarsely and sparsely punctate. Coxosternite with a [very] short median suture which reaches its middle [no more than 1 / 3 of coxosternite length, Fig. 2 E], several transverse sutures cross the median one in anterior third of coxosternite. Anterior margin of coxosternite strongly sclerotised and definitely divided by a median diastema into two standard low lobes (so it does not look strictly straight); process of trochanteroprefemur short, with a distinct basal suture; tarsungulum long, pointed. Tergites 1 – 20 sparsely punctate, tergites 21 – 23 not punctate. Tergite 1 with anterior transverse suture, its anterior margin covered by cephalic plate (Fig. 2 A). Tergite 2 short, half as long as tergite 3. Tergites 8 – 18 [7 – 19] with complete paramedian sutures, the following tergites lacking sutures (Figs 2 C, 3 A). Lateral margination (Figs 2 C, 3 A) incomplete [nearly complete] on tergites 4 – 22, only tergite 23 marginate completely. Tergite 23 nearly as long as wide, its posterior margin much convex in the middle (Fig. 4 A). Tergites 2, 6 – 13, 15 – 19, 21 with some [lacking any] dark pigment (at least at posterior and lateral margins; Figs 2 C, 3 A). Sternites lacking paramedian sutures [with very shallow rounded depression in the center], 1 – 16 ones sparsely punctate (Fig. 2 D), 2 – 20 with transverse sutures. Sternite 23 somewhat wider than long, trapeziform, its posterior margin slightly concave (Fig. 4 B). Coxopleuron (Figs 3 A – C) densely covered by irregularly located coxal pores; coxopleural process [relatively short] long and pointed, much extended over the posterior margin of tergite 23. Legs 1 – 21 with monopartite tarsus; legs (Figs 2 CD, 3 B) 1 – 22 with one tarsal spur, 1 – 19 with two tibial, 20 – 21 with one tibial and 22 lacking any spur; legs 1 – 22 with two pretarsal accessory spines. All legs not setose (in some specimens, tibia and tarsus of leg 22 sparsely setose) [leg 22 with both tibia and tarsus densely setose]. Ultimate legs (Fig. 3 D) not setose (in some specimens, tibia and both tarsal articles densely setose, Fig. 3 E) [tibia and tarsal articles densely setose in both ZMMU specimens]; prefemur with two spinous processes of standard both structure and disposition — the larger ventral and smaller dorso-medial ones. Variability. The setosity of tibia and tarsus of legs 22 and 23 varies considerably (from the total absence of setae to the presence of very dense ones). In all IEBR specimens from Northern Vietnam, the ultimate legs are practically not setose (Fig. 3 D) while in the IEBR Highland’s material as well as in ZMMU’s one (two adults Rc 6686, 6738) from Central Vietnam (Ha Tinh Province) the ultimate both tibia and tarsus are densely setose (Fig. 3 E). Variability of this character may also reflect a sexual dimorphism.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E956FF9CFF3DBA64FF16FE26.taxon	discussion	Remarks. According to Attems (1930) and Chao (2002), tergites 5 – 20 have two complete paramedian sutures, but Song et al. (2004) reported these sutures at tergites 4 – 21 and oblique sutures in the anterior corners of tergites 3 – 4. The recently studied specimens have complete paramedian sutures at tergites 8 – 18 (Fig. 2 C). Adult Rc 6686 (ZMMU) has LBS 7 with a spiracle at right pleuron only (Fig. 2 F) — this spiracle is much smaller than the well developed neighboring ones (on LBS 5 and 8, Fig. 2 F) and seems to be rudimentary. The same situation was described for S. nigrimaculatus Song, Song & Zhu, 2004 in its original description. This fact reduces the systematic value of as character as the number of spiracles, reducing the difference between S. rubiginosus and S. broelemanni esulcata (see Remarks to the latter). Range. This species occurs in China, Taiwan, Korea, Japan and North America (Minnesota, Wisconsin to Texas) (Shelley 2002; Chao & Chang 2003; Song et al. 2004). In Vietnam it was recorded from Vu Quang NP (Ha Tinh Province) only (Schileyko, 2007), and the present study demonstrates that it widely occurs through this country (Fig. 4).	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E95DFF96FF3DBF8EFEB3FD62.taxon	description	Figs 5 – 7	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E95DFF96FF3DBF8EFEB3FD62.taxon	materials_examined	Material. CAO BANG Province, Phia Oac — Phia Den NP: 1 spm (SVR. PO. 014), natural forest, 22.59842 ° N 105.89417 ° E, 1371 m a. sl., 10.07.2017, col. Le X. Son; 1 spm (SVR. PO. 018), natural forest, 22.61319 ° N 105.86698 ° E, 1803 m a. s. l., 12.07.2017, col. Le X. Son; 1 spm (SVR. PO. 038), natural forest, 22.61414 ° N 105.86822 ° E, 1750 m a. s. l., 13.07.2017, col. Le X. Son; 1 spm (SVR. PO. 051), mixed forest, 22.60610 ° N 105.87730 ° E, 1610 m a. s. l., 15.07.2017, col. Le X. Son; 1 spm (SVR. PO. 061), natural forest, 22.60610 ° N 105.87730 ° E, 1541 m a. s. l., 16.07.2017, col. Le X. Son; 1 spm (SVR. PO. 064), natural forest, 22.60644 ° N 105.87550 ° E, 1667 m a. s. l., 18.07.2017, col. Le X. Son; 2 spms (SVR. PO. 073, SVR. PO. 077), bamboo forest, 22.61497 ° N 105.86206 ° E, 1855 m a. s. l., 18.07.2017, col. Le X. Son; 1 spm (SVR. PO. 00139), natural forest, 22.60856 ° N 105.86458 ° E, 1694 m a. s. l., 04.06.2018, col. Le X. Son; 1 spm (SVR. PO. 153), mixed forest, 22.60625 ° N 105.86775 ° E, 1633 m a. s. l., 05.06.2018, col. Le X. Son; 1 spm (SVR. PO. 202), mixed forest, 22.60901 ° N 105.87074 ° E, 1596 m a. s. l., 04.06.2019, col. Le X. Son; 1 spm (SVR. PO. 205), mixed forest, 22.60892 ° N 105.87064 ° E, 1595 m a. s. l., 04.06.2019, col. Le X. Son; 2 spms (SVR. PO. 211, SVR. PO. 212), natural forest, 22.60935 ° N 105.87027 ° E, 1604 m a. s. l., 04.06.2019, col. Le X. Son; 1 spm (SVR. PO. 234), mixed forest, 22.60321 ° N 105.87344 ° E, 1508 m a. s. l., 08.06.2019, col. Le X. Son; 1 spm (SVR. PO. 247), pine forest, 22.58684 ° N 105.85835 ° E, 1008 m a. s. l., 10.06.2019, col. Le X. Son; 1 spm (SVR. PO. 125), bamboo forest, 22.61342 ° N 105.8648 ° E, 1848 m a. s. l., 03.06.2019, col. Le X. Son; 8 spms (SVR. PO. 002 – 009), mixed forest, 22.60836 ° N 105.86978 ° E, 1611 m a. s. l., 08.2020, col. Nguyen D. Anh. SON LA Province: 1 spm (SVR. TX. 017) Ta Xua NR, regenerating forest, 21.32525 ° N 104.52722 ° E, 1417 m a. s. l., 05.10.2019, col. Le X. Son. QU ẢNG NAM Province, Song Thanh NR: 1 spm (SVR. STh. 101), natural forest, 15.57269 ° N 107.35642 ° E, 1038 m a. s. l., 03.05.2019, col. Le X. Son; 1 spm (SVR. STh. 102), natural forest, 15.57181 ° N 107.40994 ° E, 1007 m a. s. l., 03.05.2019, col. Le X. Son; 1 spm (SVR. STh. 122), regenerating forest, 15.53931 ° N 107.38408 ° E, 1144 m a. s. l., 04.05.2019, col. Le X. Son; 1 spm (SVR. STh. 132), regenerating forest, 15.54053 ° N 107.38272 ° E, 1154 m a. s. l., 09.05.2019, col. Le X. Son. VINH PHUC Province, Tam Dao: 2 spms (Rc 6341 in ZMMU), 1000 – 1200 m, subtropical forest, 18.08. - 04.09.1989, col. D. Popkov; 4 juv (Rc 6343 in ZMMU), 800 – 1200 m, subtropical montane forest, 12 – 22.04.1986., col. S. I. Golovatch & L. N. Medvedev. GIA LAI Province: 1 ad (Rc 6432 in ZMMU), An Khe, environs of Buon Luoi Biological Station, tropical forest, litter, 12.1989, col. Y. M. Zaitsev. LAO CAI Province: 2 spms (Rc 6614, 6615 in ZMMU), Sa Pa District, 6 - 8 km W of Sa Pa, 1800 m a. s. l., rotten logs with soil inside, 07 – 11.12.1996, col. M. V. Kalyakin; 1 ad + 1 juv (Rc 7165 in ZMMU), Hoang Lien NP, ca 2000 m a. s. l., subtropical forest, 16 – 30.07.2007, col. S. I. Golovatch. LAM DONG Province, Bi Doup — Nui Ba NP, environs of Long Lanh: 1 ad + 1 sad (Rc 7225 in ZMMU), Exped. Rus-Viet. Trop. Cent., 1400 – 1900 m a. s. l., 01 – 22.04.2008, col. D. Fedorenko; 6 spms (Rc 7497 in ZMMU), 12 ° 10 ’ 44 ’’ N 108 ° 40 ’ 44 ’’ E, 1400 – 1600 m a. s. l., 28.04 – 10.05.2009, col. D. Fedorenko. Comparative material. Japan, Honshu, Nagano-ken, Sanada-cho, Sugadaira, 1 spm (Rc 6503 in ZMMU), 22 – 24.06.1994, col. O. Gorbunov & J. Arita.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E95DFF96FF3DBF8EFEB3FD62.taxon	diagnosis	Diagnosis. 2 basal antennal articles virtually lacking setae dorsally; cephalic plate marginate laterally; tergal paramedian sutures absent, ultimate tergite with complete lateral margination; LBS 7 lacks spiracles; ultimate legs lacking setae (more rarely their distal articles setose), ultimate prefemur with two spinous processes of typical both structure and disposition.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E95DFF96FF3DBF8EFEB3FD62.taxon	description	Description of adult SVR. PO. 202 [data on ZMMU material in square brackets where it differs]. Body length 35.6 [up to 60] mm, width of LBS 10 ca 3.56 mm. Antennae (Figs 5 AB) composed of 17 articles of them 2 basal ones very sparsely setose dorsally [2 dorsally and 4.5 - 5.5 laterally, Fig. 5 E], following articles densely covered by minute setae. Cephalic plate (Fig. 5 A) nearly as long as wide, sparsely [deeply and densely] punctuate and marginate laterally, posterior margination absent. Forcipular segment (Fig. 5 B): coxosternite, trochanteroprefemora and the base of tarsungulae coarsely and sparsely punctate. Coxosternite with a median suture which reaches its middle, a few transverse sutures cross median one in anterior third of coxosternite [all these sutures are poorly developed in not full-grown specimens]. Anterior margin of coxosternite strongly sclerotised, practically stright and is definitely divided by a median diastema into two standard very low lobes; process of trochanteroprefemur small, with a distinct basal suture; tarsungulum long, pointed. Tergites 1 – 20 sparsely [densely and strongly] punctate, tergites 21 - 23 not punctate; tergite 1 with anterior margin covered by the cephalic plate, tergite 2 short, as long as 1 / 3 of tergite 3 (Fig. 5 A). All tergites lacking complete paramedian sutures (Figs 5 DF). Incomplete lateral margination at tergites 5 – 22 [nearly complete at tergites (10) 16 – 22, Fig. 5 F], only tergite 23 marginate completely. Tergite 23 nearly as long as wide [and with a narrow longitudinal depression in posterior half], its posterior margin much convex in the middle. Sternites 2 – 20 (Figs 5 BC) sparsely [densely and strongly] punctuate, with some transverse sutures near posterior margin; sternite 23 (Fig. 6 B) trapeziform and much narrower than penultimate one, its posterior margin slightly concave. Coxopleuron (Figs 6 AB) densely covered by irregularly located pores [coxal pore field long and wide, reaching both sternite 22 and coxopleuron’s posterior margin]; sharply pointed coxopleural process relatively short, slightly extended over the posterior margin of tergite 23. Legs 1 – 21 with monopartite tarsus; legs (Figs 5 BC, 6 AB) 1 – 22 [1 – 21] with one tarsal spur, 1 – 20 [1 – 19 (21)] with two tibial spurs, 21 – 22 with one tibial spur; legs 1 – 22 with two pretarsal accessory spines. All legs not setose. Ultimate legs (Figs 6 A – C) not setose (sparsely setose in a few specimens) [in some specimens tibia and tarsus densely setose, Fig. 6 E]; basal part of prefemur with two typical spinous processes — the larger ventral and the smaller dorso-medial ones. Variability. ZMMU specimens (collected in Vinh Phuc, Gia Lai, Lao Cai and Lam Dong Provinces; Schileyko 1995: 74) have the maximal body length up to 60 mm (one of two adults Rc 6341 from Tam Dao), being much larger than IEBR material (recently collected in Cao Bang, Son La and Quang Nam Provinces); in other respects, all studied specimens are very similar.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E95DFF96FF3DBF8EFEB3FD62.taxon	discussion	Remarks. Attems (1930: 260) wrote that S. sexspinosus (Say, 1821) (actually S. spinicaudus in part) has all antennal articles setose which data have been confirmed by Chao (2002) based on Taiwanese material. However, in all studied specimens from both IEBR and ZMMU two basal antennal articles are definitely not setose (Figs 5 AB). Schileyko (2007: 74) wrote in Remarks to this species: “ Based on all taxonomic characters, the above specimens fit well Shinohara’s [1990] concept of Scolopocryptops nipponicus, a species synonymised with S. spinicaudus Wood, 1862 by Shelley [2002] ”. In some ZMMU specimens (one of six adults of Rc 7497, adult Rc 7225, adult Rc 6342, adult + juvenille Rc 7165) tarsus and tibia (and sometimes femur) of the ultimate legs are covered by quite long and well-developed setae (Fig. 6 E) which are slightly less dense at femur comparing to more distal articles. As these setae are present only in a few specimens of the same sample (for example in Rc 7497 of four adults which kept the ultimate legs, only one specimen has these legs setose) one can suppose them to be, very probably, a result of the sexual dimorphism. The recent literature (Qiao et al. 2021) states that a “ lack ” of tergal paramedian sutures (in fact a lack of the complete ones) is the main diagnostic character to distinguish S. spinicaudus from S. rubiginosus. In fact, all IEBR material as well as the majority of ZMMU specimens of S. spinicaudus do not have these sutures (except for extremely short rudiments at very anterior and posterior margins of practically all tergites). However at least two adults of this species (both of ZMMU) — the largest (ca 50 mm) one of Rc 7497 and large (ca 40 mm) one Rc 6342 — demonstrate tergites 4 (8) – (18) 21 with well-developed but incomplete (i. e. somewhat interrupted in the middle) paramedian sutures (Fig. 6 F). Thus S. spinicaudus and S. rubiginosus differ from each other less than it was thought before. Also, the recent authors (for example Song et al. 2004) wrote that tergites of S. spinicaudus have “ incomplete ” lateral margination, but re-investigation of the ZMMU material, made by the second author, demonstrates this margination to be practically complete (at least in LBS of posterior body half, Fig. 5 F). This fact supports as well the closest similarity of this species and S. rubiginosus, thus the further analyses may possibly show an identity of two these species. Another noteworthy point is that the largest adult of the mentioned above Rc 7497 (ZMMU) has a pair of the well (!) developed spiracles on LBS 7 (Fig. 6 D); thus this specimen seems to be quite similar to S. broelemanni esulcata. Also this fact much reduces taxonomical value of such character as the number of spiracle pairs (at least in the genus Scolopocryptops). Range. Canada, Southern USA, Mexico, Korea, Japan, China (Attems 1930, Shelley 2002, Schileyko 2007). In Vietnam (Fig. 7) this species occurs from Northern to Central regions (Lao Cai, Vinh Phuc and Gia Lai Provinces; Schileyko 2007).	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E958FF8BFF3DBC00FE5DFECA.taxon	description	Figs 8 – 10	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E958FF8BFF3DBC00FE5DFECA.taxon	materials_examined	Material. CAO BANG Province, Phia Oac — Phia Den NP: 1 spm (SVR. PO. 002), natural forest, 22.60047 ° N 105.88313 ° E, 1325 m a. s. l., 09.07.2017, col. Le X. Son; 1 spm (SVR. PO. 005), natural forest, 22.59875 ° N 105.89417 ° E, 1452 m a. s. l., 10.07.2017, col. Le X. Son. VINH PHUC Province, Tam Dao NP: 1 spm (SVR. TD. 002), bamboo forest, 21.45378 ° N 105.6491 ° E, 1029 m a. s. l., 08.6.2018, col. Nguyen D. Hung; 2 spms (SVR. TD. 008, SVR. TD. 009), bamboo forest, 21.45758 ° N 105.6523 ° E, 977 m a. s. l., 08.8.2018, col. Nguyen D. Hung. NGHE AN Province, Pu Hoat NR: 2 spms (SVR. PH. 011, SVR. PH. 012), natural forest, 19.75372 ° N 104.78836 ° E, 985 m a. s. l., 16.05.2019, col. Le X. Son; 1 spm (SVR. PH. 048), natural forest, 19.75542 ° N 104.78569 ° E, 1079 m a. s. l., 19.05.2019, col. Le X. Son.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E958FF8BFF3DBC00FE5DFECA.taxon	diagnosis	Diagnosis. 2 basal antennal articles sparsely setose; cephalic plate marginate laterally; tergal paramedian sutures absent, ultimate tergite with complete lateral margination; LBS 7 lacks spiracles; ultimate legs with the distal articles densely setose and femur setose, ultimate prefemur with two typical (larger ventral and smaller dorsomedial) spinous processes.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E958FF8BFF3DBC00FE5DFECA.taxon	description	Description of adult SVR. PH. 048. Body length 34.4 mm, the width of LBS 10 ca 3.27 mm. Antennae composed of 17 articles, of them 2 basal ones sparsely setose, following articles densely covered by minute setae (Figs 8 AB). Cephalic plate (Fig. 8 A) nearly round, not setose, definitely punctate and marginate laterally, lacking posterior margination. Forcipular segment (Fig. 8 B): coxosternite, trochanteroprefemora and the basal part of tarsungulae coarsely and sparsely punctate. Coxosternite with a median suture which reaches its middle; some transverse sutures cross the median one at anterior third of coxosternite. Anterior margin of coxosternite strongly sclerotised and is definitely divided by a median diastema into two very low lobes; process of trochanteroprefemur small with a distinct basal suture; tarsungulum long, pointed. Tergites (Figs 8 A, C, 9 A) lacking paramedian sutures, not punctate, anterior margin of tergite 1 slightly covered by cephalic plate (Fig. 8 A), tergite 2 very short, as long as 1 / 3 – 1 / 2 of tergite 3. Lateral margination incomplete on tergites 5 – 22, tergite 23 marginate completely. Tergite 23 (Fig. 9 A) nearly as long as wide, its posterior margin much convex in the middle. Sternites (Figs 8 B, D, 9 C) lacking paramedian sutures, 2 – 17 ones sparsely punctate; sternites 2 – 19 with several transverse sutures. Sternite 23 (Fig. 9 C) wider than long, tongue-shaped, its posterior margin slightly concave. Coxopleuron (Figs 9 BC) densely covered by irregularly located pores; coxopleural process pointed and comparatively short, slightly extending over the posterior margin of tergite 23. Legs (Figs 8 B – D, 9 C) 1 – 21 with one tarsal spur, 1 – 19 with two tibial spurs, 20 – 21 with one tibial spur; leg 1 – 22 with two pretarsal accessory spines. Ultimate legs (Fig. 9 D): prefemur and femur sparsely setose, tibia and tarsus densely setose; prefemur with two typical spinous processes — the larger ventral and the smaller dorso-medial. Variability. Studied eight specimens show no visible variations. Range. This species was originally described from Korea by Takakuwa (1938), and recorded from Japan (Miyoshi 1982) and Taiwan (Chao 2002; Chao & Chang 2003); this is the first record of S. capillipedatus from Vietnam (Fig. 10).	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E958FF8BFF3DBC00FE5DFECA.taxon	discussion	Remarks. The studied IEBR specimens fit well with Taiwanese material described by Chao (2002); however, this species is extremely close to S. spinicaudus in all morphological aspects except for the presence of “ dense ” (vs “ sparse ” in S. spinicaudus) setae on the ultimate both tibia and tarsus. It should be noted, that some (not all) specimens of S. spinicaudus (kept in both ZMMU and IEBR) have ultimate tarsus, tibia and (rarely) femur densely setose (Fig. 6 E), sharing the main diagnostic character of S. capillipedatus. This fact deletes any real (i. e. morphological) differences between these two species except for the presence (vs. the virtual absence) of the sparse setae at two basal antennal articles in S. capillipedatus and the slight difference in shape of the ultimate sternite (tongue-shaped vs. trapeziform in S. spinicaudus). Thus, the recent identificational Scolopocryptops keys (Chao 2002, Chao & Chang 2003, 2008, Qiao et al 2021, Xiao et al 2021) do not allow to distinguish reliably “ S. capillipedatus ” from S. spinicaudus. Summing up, we consider S. capillipedatus as questionable (“ molecular ”) species with validity not supported reliably by morphology (see couplet 5 in the key below). At the moment we kept S. capillipedatus as a valid name because of shortage of the investigated material, but the further analyses will highly likely show its identity to S. spinicaudus.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E944FF8AFF3DBF8EFE29F8AA.taxon	description	Figs 11 – 13 Locus typicus. Hoang Lien NP, Lao Cai Province, Northwestern Vietnam.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E944FF8AFF3DBF8EFE29F8AA.taxon	materials_examined	Material. LAO CAI Province, Hoang Lien NP, natural forest: Holotype (SVR. HL. 002), 22.35063 ° N 103.77477 ° E, 1927 m a. s. l., 16.09.2020, col. Do T. Thinh; 2 Paratypes (SVR. HL. 007, SVR. HL. 008), 22.34563 ° N 103.77552 ° E, 2004 m a. s. l., 16.09.2020, col. Le X. Son; 1 spm (HL. 133 in VAST), 22.34603 ° N 103.7777 ° E, 2000 m a. s. l., 29.11.2018, col. Nguyen Duc Hung; Bat Xat NR, mixed bamboo forest, 22.6158 ° N 103.6359 ° E, 1686 m a. s. l., 1 spm (BX. 026), 18.10.2018, col. Sung A. De.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E944FF8AFF3DBF8EFE29F8AA.taxon	diagnosis	Diagnosis. 2 basal antennal articles virtually lacking setae; cephalic plate marginate laterally; tergites lacking paramedian sutures, tergites 6 – 20 with a well-developed “ drop-like ” longitudinal median depression, ultimate tergite with complete lateral margination; LBS 7 lacks spiracles; ultimate legs not setose, ultimate prefemur with two typical (ventral and dorso-medial) spinous processes.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E944FF8AFF3DBF8EFE29F8AA.taxon	description	Description of the adult holotype SVR. HL. 002. Body length 35.3 mm, the width of LBS 10 ca 2.7 mm. Antennae (Figs 11 AB) composed of 17 antennomeres of them 2 basal ones not setose, following articles densely covered by minute setae. Cephalic plate (Fig. 11 A) as long as wide, sparsely punctuate and marginate laterally, posterior margination absent. Forcipular coxosternite (Fig. 11 B): coxosternite and basal part of trochanteroprefemora coarsely and sparsely punctate. Coxosternite with a median suture which reaches its middle, some transverse sutures cross the median one in anterior third of coxosternite. Strongly sclerotised anterior margin of coxosternite definitely divided by a median diastema into two standard very low lobes; process of trochanteroprefemur small with a distinct basal suture; tarsungulum long, pointed. Tergites lacking paramedian sutures, sparsely punctuate; anterior margin of tergite 1 covered by cephalic plate. Lateral margination incomplete on tergites 4 – 22 (Figs 11 CD, 13 B), only tergite 23 marginate completely (Figs 12 BC). Tergites 6 – 20 with a long (quasi-complete) “ drop-like ” longitudinal median depression which expanded to tergal posterior end (Figs 11 CD, 13 B) and is bordered by paramedian keels. Tergite 23 (Fig. 12 B) nearly as long as wide, its posterior margin much convex in the middle. Sternites lacking paramedian sutures, trapeziform, with many transverse / oblique sutures (Figs 11 B, 12 A – D). Sternite 23 (Fig. 12 D) trapeziform, its posterior margin slightly concave. Coxopleuron (Figs 12 CD) densely covered by irregularly located pores; coxopleural process pointed and long, extended over the posterior margin of tergite 23. Legs (Figs 11 D, 12 A – C) 1 – 21 with 1 tarsal spur; 1 – 19 with two tibial spurs, 20 – 21 with 1 tibial spur; leg 1 – 22 with two pretarsal accessory spines. Ultimate legs (Fig. 12 E) not setose; prefemur with two spinous processes — very large ventral one and much smaller dorso-medial one, both disposed nearly at prefemur’s middle. Variability. Incomplete lateral margination is from tergite 2 in specimen SVR. HL. 007 and longitudinal median depression is present on tergite 2 - 22 in specimens SVR. HL. 008 and BX. 026.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E944FF8AFF3DBF8EFE29F8AA.taxon	etymology	Etymology. Named after Hoang Lien NP — the type locality of this species.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E944FF8AFF3DBF8EFE29F8AA.taxon	discussion	Remarks. S. hoanglieni has a much thinner body than its congeners. The new species definitely differs from S. rubiginosus by the absence (vs. presence) of tergal paramedian sutures and from S. capillipedatus by not setose ultimate legs (vs. densely setose ultimate both tibia and tarsus). The new species is relatively similar to S. spinicaudus in the absence of tergal paramedian sutures and not setose ultimate legs, but is easily distinguished from the latter by tergites with a very characteristic longitudinal depression (Fig. 11 C) which character seems to be unique, at least within Scolopocryptopinae.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E940FF8EFF3DBF8EFCA2FB7D.taxon	materials_examined	Comparative material. E Indonesia, West Papua Province, South Bird’s Neck: 1 ad (Rc 7503 in ZMMU), Kaimana 47 km E, Triton bay, environs Kamaka village, lake Kamakawalar, 03 ° 45 ’ 33 ” S, 134 ° 12 ’ 05 ” E, 90 m, primeval lowland rainforest on limestone, 09.09.2010, leg. M. Kalninsh; 1 ad (Rc 7504 in ZMMU), Kaimana 7 – 9 km NW, 25 – 200 m, primeval lowland rainforest on limestone, 05.09.2010, leg. D. Telnov.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E940FF8EFF3DBF8EFCA2FB7D.taxon	diagnosis	Diagnosis. 4 – 6 basal antennal articles virtually lacking setae; cephalic plate not marginate laterally; tergites 3 (4) – 21 (22) with paramedian sutures, ultimate tergite lacks lateral margination; LBS 7 lacks spiracles; femur, tibia and tarsus of the ultimate legs densely setose.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E940FF8EFF3DBF8EFCA2FB7D.taxon	description	Description. For morphological details see Schileyko (2014) and Schileyko & Stoev (2016) (the most recent data on this species) who described specimens from Venezuela and East Indonesia (West Papua Province) respectively. Range (by Schileyko & Stoev 2016). Mexico, Central America (Guatemala, Honduras, Costa Rica, Panama), Greater Antilles (Puerto Rico, Haiti), Lesser Antilles (Martinique, Saint Vincent and Grenadines, Trinidad), South America (Venezuela, Colombia, Ecuador, Peru, Brazil), Australasia (Fiji Islands), Indochina (Nicobar Island, Vietnam), Taiwan, Philippines, Eastern Indonesia, Papua New Guinea.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E940FF8EFF3DBF8EFCA2FB7D.taxon	discussion	Remarks. Only Attems (1953) reported this widespread species from Vietnam (Langbiang Mountains in Lam Dong Province). Attems (1953) considered the complete absence of leg’s accessory spines to be a diagnostic character for S. melanostoma, but Schileyko & Stoev (2016: 250) described the corresponding rudiments (recognizable at x 85 magnification) in material from West Papua. In fact, the main diagnostic character of this species is the absence of any lateral margination of the tergites, including the ultimate.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E940FF81FF3DBA3DFDC1FD62.taxon	description	Figs 14 – 16	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E940FF81FF3DBA3DFDC1FD62.taxon	materials_examined	Material. LAM DONG Province: collected in locus typicus, (sub) adult syntypes 1 and 2 (MY 2063 in NHMW) and 1 adult non-type spm (MY 8714 in NHMW).	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E940FF81FF3DBA3DFDC1FD62.taxon	diagnosis	Diagnosis. The only Vietnamese Scolopocryptops with spiracles on LBS 7 (diagnostic character of the former genus Dinocryptops). The original description (direct translation by the second author). “ Body length 24 mm. Cephalic plate marginate laterally. Tergite 1 with well-developed anterior transverse suture, anterior margin of tergite 1 covered by cephalic plate. Ultimate tergite lacks median suture, with a single rounded depression in posterior third; this tergite has definite lateral margination and characteristic small pointed process at each posterior corner. Legs 1 – 21 with one tarsal spur, 1 – 20 with two tibial spurs, leg 21 with a single tibial one, leg 22 lacking spurs. In all other characters / moments corresponds to the detailed description of S. broelemanni by Brolemann ”. Composite re-description of two syntypes MY 2063 (NHMW). Antennae) composed of 17 articles (in not damaged antenna) of them 2 basal ones dorsally (Figs 14 A, 15 A) and 1 basal ventrally (Figs 14 B, 15 B) with the single long setae, following articles densely covered by minute setae. Cephalic plate virtually as long as wide, very sparsely setose, with very definite lateral and less developed posterior margination (Figs 14 AF, 15 A). Forcipular segment (Figs 14 B, 15 B): anterior half of coxosternite with a few hardly visible transverse sutures (median one is not recognizable). Anterior margin of coxosternite visibly sclerotised and definitely divided by a median diastema into two standard low lobes, so it does not look straight; process of trochanteroprefemur short; tarsungulum long, pointed. Tergite 1 with anterior transverse suture, its anterior margin covered by cephalic plate (Figs 14 A, 15 A). Tergite 2 short, half (or somewhat less) as long as tergite 3. Tergal paramedian sutures not recognizable (Figs 14 C, 15 C). Lateral margination incomplete on tergites 4 – 10 (11) (Fig. 14 D) and nearly complete on following tergites, only tergite 23 marginate completely (Fig. 15 D). Tergite 23 nearly as long as wide, its posterior margin much convex in the middle. Sternites lacking sutures, with very shallow rounded depression in the center (Fig. 15 E). Sternite 23 tongueshaped, its rounded posterior margin slightly concave in the middle (Figs 14 E, 15 F). LBS 7 with a pair of standardly developed spiracles (Figs 14 D, 15 E). Coxopleuron (Figs 14 E, 15 F) covered by irregularly located coxal pores of various sizes. Coxopleural processes conical, very short (much shorter than sternite 23) and diverging slightly, they visibly extended over the posterior margin of tergite 23 (Fig. 15 D). Posterior margin of pleuron forming an acute angle, its tip definitely pointed (Fig. 14 E). Legs 1 – 21 with monopartite tarsus (Fig. 15 E); legs 1 – 21 with one tarsal and two tibial spurs and 22 lacking any spur (Fig. 14 E) legs 1 – 22 with two pretarsal accessory spines. All legs practically not setose. Ultimate legs not setose; prefemur (Figs 14 CE) with two spinous processes of standard both structure and disposition — the larger ventral and smaller dorso-medial ones. Variability. In syntype 1 both sternite 23 and the coxopleural process are relatively longer than in syntype 2 (cf. Figs 14 E and 15 F). The ultimate legs are attached only in syntype 1. Range. Known from the type locality only.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
03914E40E940FF81FF3DBA3DFDC1FD62.taxon	discussion	Remarks. S. broelemanni esulcata is known only from three specimens kept in NHMW, collected by C. Dawydoff in the Lang Biang Mountains (Fig. 16). S. broelemanni esulcata is the only Vietnamese member of the former genus Dinocryptops which name has been synonymised with Scolopocryptops by Edgecombe et al. (2012: 777 – 778) based on both morphology and molecular analysis (this synonymy has been omitted by Bonato et al. 2016). The study of digital images of both syntypes showed that they have well developed (not rudimentary) spiracles on LBS 7 (Figs 14 D, 15 E) — the only diagnostic character of former Dinocryptops. New data on the occasional presence of spiracles on LBS 7 (Fig. 2 F, 6 D) in both S. rubiginosus and S. spinicaudus reduce the difference between them and S. broelemanni esulcata. Thus, a character such as the number of spiracles must be used with much care (when used alone) to reliably identify Scolopocryptops species. Discussion on validity of S. broelemanni esulcata. Available data on this subspecies are scarce; the original description is very short and incomplete, lacking both illustrations and information about some diagnostic characters (number of glabrous basal antennal articles, presence / conditions of tergal sutures, pilosity of distal articles of the ultimate legs etc.). Judging from the digital images, the NHMW specimens fit the descriptions of S. broelemanni Kraepelin, 1903 by Kraepelin (1903: 77) and Attems (1930: 256) well. In the original description of S. broelemanni esulcata Attems (1938: 338) indicated, that this subspecies is, on the whole, very similar to S. broelemanni, differing from the latter mainly in the absence (vs. presence) of the median suture on the ultimate tergite. This character is present in various groups of scolopendromorphs (Scolopendridae, Plutoniumidae, Scolopocryptopinae), being species-specific for the overwhelming majority of their species. Digital images of three NHMW specimens do not recognize this suture (Fig. 15 D), so they are fully consistent with S. broelemanni esulcata. There is also a significant geographical distance between the range of S. broelemanni esulcata (Southern Vietnam), and the nominotypical subspecies described from “ Chou-San ” (Zhoushan, formerly romanized as Chusan, Zhejiang Province in East China). Summing up, we consider S. broelemanni esulcata Attems, 1938 to be a valid subspecies, but its taxonomic status may be changed by examining representative relevant material. In 2005, NHMW specimens were examined by Amazons Chagas Junior, who identified them as “ Dinocryptops brolemanni (sic!) (Kraepelin, 1903) ” (Fig. 14 G). In his PhD thesis Chagas (2008) proposed the synonymy of S. broelemanni esulcata to S. broelemanni broelemanni and considered (p. 100) both of them as representative (s) of some new (but unnamed) genus — “ Gen. Nov. 1 ” (any nomenclatural acts proposed there have no formal status — disclaimer to dissertation). He referred to both of these subspecies as “ Gen. Nov. 1 broelemanni (Kraepelin, 1903) Comb. nova ” (p. 101), but also as “ Dinocryptops broelemanni ” (pp. 25, 27 etc). Considering both subspecies to be the same form, Chagas (2008: 101) gave a composite description of the NHMW specimens as well as one specimen of Scolopocryptops broelemanni broelemanni from “ Guangdong Sheng, Nanking ” (actually Nanking or Nanjing is disposed in Jiangsu Province, not in Guangdong Province) kept in National Museum of Natural History (Smithsonian Institution, Washington D. C.) (No 31325, “ Cook & Loomis, 23 - X- 1919 ”). This description is sufficiently detailed, but there are no data on the presence or absence of a median suture on tergite 23, a diagnostic character of S. broelemanni esulcata (it is clearly seen in the corresponding figure 27 f on p. 183 that tergite 23 is devoid of this suture). In discussing the validity of S. broelemanni esulcata Chagas (2008: 138) wrote that both subspecies share common characters such as a spiracle on LBS 7 and lateral margination of the cephalic plate, but again omitted the diagnostic character mentioned above. Thus the synonymy proposed by Chagas (2008) remains in question.	en	Le, Son X., Schileyko, Arkady A., Nguyen, Anh D. (2023): A review of Vietnamese Scolopocryptops Newport, 1844 (Chilopoda: Scolopendromorpha), with a description of S. hoanglieni n. sp. and the updated generic list of species. Zootaxa 5228 (4): 441-447, DOI: https://doi.org/10.11646/zootaxa.5228.4.3
