taxonID	type	description	language	source
583A87D7A33DFFD3FF29FE87CDEED697.taxon	discussion	Males (unknown in Kurzenkotilla ariana, K. indica, K. niveosignata, K. scrobiculata, K. visrara, Standfussidia taprobane, Storozhenkotilla binghami, and K. cicatricifera)	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33DFFD3FF29FE87CDEED697.taxon	discussion	Females (unknown in Kurzenkotilla rufodorsata, K. semiviolacea, and Storozhenkotilla nathani)	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33CFFD4FF29FCA2CA60D5E9.taxon	materials_examined	Type species: Artiotilla ariana Lelej in Lelej & Kabakov, 1980 by original designation.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33CFFD4FF29FCA2CA60D5E9.taxon	diagnosis	Diagnosis. MALE. Head width less than pronotal width, rounded posterad; occipital carina dorsally well developed, sharp and slightly projecting over pronotum. Eye weakly notched inside. Ocelli small. Mandible bidentate without subbasal tooth beneath and without subbasal inner tooth. F 2 longer than F 1. Prementum with longitudinal mesal ridge. Notauli and parapsids visible. Tegula longer than pronotal length laterad, projecting over axilla. Pterostigma large, closed. Fore tarsus without comb of strong spines. Mid and hind tibia with two rows of weak spines. Mesoscutellum non-modified. Propodeal posterior slope with strong longitudinal medial carina. Metasomal segment 1 widely transverse. T 1 dorsally and T 2 – 3 (4) each with band of pale setae posterad. T 2 with long lateral felt line, S 2 without any lateral felt line. S 8 (hypopygium) not modified, T 7 generally with smooth triangular area posteromesally. Gonostylus wide, basiventrally setose. Volsella usually shorter than penial valve and consisting of wide cuspis and short finger-like digitus; penial valves symmetrical, apically setose. FEMALE. Head large, somewhat rounded posterad, width equal to pronotal width; behind eyes elongated; eyes large, distance between postero-dorsal margin of eye and that of vertex less than eye longitudinal diameter, projecting over head. Hypostomal carina slightly widened in medial portion. Prementum not tuberculate. F 1 1.5 – 2.0 × F 2. Mandible rather wide, bidentate, without subbasal inner tooth, preapical tooth obtuse (tridentate in K. annamensis and K. harmandi). Mesosoma with concave mesopleuron, scutellar scale lacking. Mesopleuron with developed vertical (supracoxal) carina. Mid and hind tibia with two rows of weak spines. Metasomal segment 1 wide transverse; T 2 with lateral felt line. S 2 latero-apically swollen, swell in type species K. ariana ending in tooth. T 2 basally with larger or smaller pale spots disposed transversely, T 3 with wide pale band or lateral spots. T 6 without distinct pygidial plate, apical part microgranulated and covered by dense setae. S 6 with two strong apical tubercles. Species included. Kurzenkotilla annamensis Lelej, 2005, ♁ ♀; K. ariana (Lelej in Lelej & Kabakov, 1980), ♀; K. harmandi (André, 1898), ♁ ♀, comb. nov.; K. cicatricifera (André, 1894), comb. nov., ♀; K. niveosignata (André, 1894), ♀; K. rufodorsata (Cameron, 1897), ♁, comb. nov.; K. scrobiculata (Hammer, 1962), ♀; K. semiviolacea (André, 1896), ♁, comb. nov.; and K. visrara (Cameron, 1898), ♀.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33CFFD4FF29FCA2CA60D5E9.taxon	distribution	Distribution. Palaearctic (Afghanistan) and Oriental Regions.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33CFFD4FF29FCA2CA60D5E9.taxon	discussion	Remarks. Kurzenkotilla Lelej is closely related to Storozhenkotilla Lelej but, in addition to setal markings of T 2, differs by lacking a pygidial plate in the female (weakly carinated pygidial plate present in Storozhenkotilla). By having two pale discal setal spots on T 2, the female of Kurzenkotilla resembles that of Macromyrme Lelej, 1984, but differs by lacking a subbasal inner tooth on the non-acuminate mandible (with developed subbasal inner tooth on an acuminate mandible in Macromyrme), and by having larger eyes (eyes very small in Macromyrme). Males of Kurzenkotilla are similar to those of Storozhenkotilla and, as yet, there are no clear characters that can be used to separate all Kurzenkotilla from all Storozhenkotilla, which is why species from both genera are included in the same key above. Males of the apparently closely related Macromyrme are currently unknown, but they will likely be similar to these genera as well.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33BFFD4FF29FE32CB88D323.taxon	materials_examined	Material examined. THAILAND: Chiang Mai: Samoeng, Ban Ang Khai, 12 – 15. V. 2003, M. Maruyama leg., flight interception trap, 2 ♁ [TUA]. VIETNAM: Hoa Binh, Tonkin, 1919, 1 ♀ [RBINS]; Ninh Thuan, Nui Chua N. P., northeast part, malaise traps, 90 – 150 m, 23 – 30. V. 2007, 1 ♁ [RMNH].	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33BFFD4FF29FE32CB88D323.taxon	distribution	Distribution. * Thailand, South Vietnam.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33BFFD4FF29FE32CB88D323.taxon	description	Description. FEMALE. See Lelej (2005). MALE. Head width less than pronotal width, not rounded posterad. Preoccipital carina dorsally well developed, sharp and slightly projecting behind head. Eyes well notched inside. Ocelli small, ratio POD: OOD = 0.75. Mandible bidentate without basal tooth beneath and without subbasal tooth on inner border. F 2 1.25 × F 1. Prementum with raised mesal longitudinal carina. Notauli well visible in posterior half, parapsids weak. Tegula longer than pronotal length laterad, projecting over axilla. Propodeal posterior slope with strong longitudinal medial carina which ends dorsally in tubercle. Metasomal segment 1 widely transverse. T 2 with long lateral felt line, S 2 without any lateral felt line. Sternum 8 (hypopygium) not modified. T 7 obscurely angulate laterally, apical border smooth with wide medial smooth triangle marked by blunt carinae laterally. Gonostylus wide, basiventrally setose. Volsella slightly longer than penial valve and consists of wide cuspis and short fingerlike digitus; penial valves symmetrical, posterior margin with vertical row of setae, postero-ventrally bidentate, pre-apical tooth sharp, larger than apical tooth. Head orange-brown with mandible, clypeus, and postgenal area blackened. Antennae black, except scape with obscure yellow-brown apical area, scape and pedicel with brown and whitish setae. Head densely punctured, setae interspersed pale yellow and dark brown. Mesosoma orange-brown, except mesopleuron ventral half, metapleuron, and propodeum black; setae predominantly silvery-white laterally, dorsal setae largely orange-brown. Tegula moderately punctate, laterally and posteriorly smooth. Pronotum, mesoscutum, mesoscutellum, and mesopleuron coarsely areolate-punctate; propleuron with large shallow punctures; metapleuron smooth; propodeum areolate. Legs with dense whitish setae. Metasoma blackish-brown with distinct purple lustre. T 1 – 2 densely punctured, sparser on disk of T 2. T 1 – 3 and S 2 – 5 with apical fringes of whitish setae; T 4 – 6 with setae black, with few scattered erect whitish setae on discs laterally. Wings fusco-violaceous.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33BFFD4FF29FE32CB88D323.taxon	discussion	Remarks. The male of this species was recognized only after the sex association of Kurzenkotilla harmandi, comb. nov. (see below) was discovered. The newly recognized females of K. harmandi initially appeared to be conspecific with K. annamensis, but closer examination revealed that they represented different species, particularly due to the differences in the metasomal setal pattern and the constricted mesonotum of K. harmandi. The male here recognized as K. annamensis occurs in southern Vietnam, where the type and other examined female of K. annamensis have been examined, but K. harmandi have not been seen. Furthermore, this male has different metasomal setal markings (as discussed in the key above) than K. harmandi. Additionally, it has the mesoscutum clearly wider than long, while the male of K. harmandi has the mesoscutum only scarcely wider than long; likewise, females of K. annamensis have a wider mesonotum than that of K. harmandi.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33AFFD5FF29FF7FC9B7D58B.taxon	description	(Figs 5 – 6) Artiotilla ariana Lelej in Lelej & Kabakov, 1980: 191, ♀, holotype, ♀, Afghanistan, western border of Farah province, 70 km south of Nazar-Khan, Shand, 1. XI. 1972, O. Kabakov leg. [ZIN], examined.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33AFFD5FF29FF7FC9B7D58B.taxon	materials_examined	Material examined. Holotype only.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33AFFD5FF29FF7FC9B7D58B.taxon	distribution	Distribution. Afghanistan.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33AFFD5FF29FF7FC9B7D58B.taxon	discussion	Remarks. This rare species has not yet been found in the Oriental Region and is the westernmost known species in the genus Kurzenkotilla.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A339FFD6FF29FF7FCC61D392.taxon	materials_examined	Material examined. LAOS: Sayaboury, Sayaboury, 6. XII. 1965, 1 ♀ [BPBM]; Vientiane, Tat Mun waterfall, el. 183 m, 18 ° 01 ′ 03 ″ N 102 ° 27 ′ 38 ″ E, 7. VI. 2013, G. R. Ballmer leg., 1 ♀ [UCRC, UCRCENT # 428259]. THAILAND: Chaiyaphum: Tat Tone N. P., Staff house at Takfah waterfall, 15 ° 56.461 ′ N 102 ° 5.955 ′ E, 242 m, malaise trap, T. Jaruphan & O. Budsawong leg., 5 – 12. IX. 2006, 1 ♁, [QSBG]; Tha Hin Ngom Village, 16.001 ° N 102.011 ° E, 560 m, malaise trap, K. A. Williams et al. leg., 18 – 28. III. 2019, 1 ♁, [CSCA, DNA voucher 19 E 658]; Chiang Mai, Omkoi District, dry dipterocarp forest, 17 ° 50 ′ 49.9 ″ N 98 ° 22 ′ 33.0 ″ E, 950 – 1010 m, R. Mizuno leg.: 19. XII. 2016, 1 ♀, [THNHM]; 20. VI. 2017, 1 ♀ [THNHM]; 27. VI. 2017, 1 ♀ [THNHM]; 7. VII. 2017, 1 ♀, [THNHM, DNA voucher 19 E 961]; 19. VII. 2019, 1 ♀ [THNHM]; Khonkaen, Nam Phong N. P., office, 16 ° 37.377 ′ N 102 ° 34.454 ′ E, 344 m, K. Jaidee leg., malaise trap, 19 – 26. VII. 2006, 4 ♁, [CSCA, EMUS, QSBG]; Loei: Phu Kradeung N. P., mixed deciduous forest N of NA Noy Forest Unit, 16 ° 48.17 ′ N 101 ° 47.66 ′ E, 276 m, malaise trap, S. Gong-Iasae leg., 14 – 20. XI. 2006, 2 ♁, [QSBG]; Nakhon Ratchasima, Kog Nong Bun Nak-Rat, 28. VII. 2009, V. Bezborodov leg., 1 ♀, [IBSS].	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A339FFD6FF29FF7FCC61D392.taxon	description	Description. FEMALE. Body length 11 – 13 mm. Coloration. Head, mesosoma, and S 1 orange-brown; legs, antenna, and metasoma blackish. Body setae generally sparse and silvery, except frons, vertex, and mesosomal dorsum with sub-erect golden and blackish setae; T 1, T 2 disc, and T 4 – 5 setae dense black dorsally; T 1 – 6 laterally, S 1 – 6, T 2 disc lateral spots, T 2 fringe, T 3 entirely, and T 6 with dense pale golden setae; distance between T 2 lateral spots 0.6 × spot diameter. Head. Width behind eye subequal to pronotal width. Frons, vertex, and gena punctures coarsely confluent. Mandible apex apparently tridentate, middle tooth blunt and confluent with lower tooth. Clypeus basomesal longitudinal carina and mesally notched transverse pre-apical carina. Antennal scrobe with thick dorsal carina. Antennal tubercle dorsally smooth, anteriorly micropunctate. Genal and postgenal carinae distinct, forming blunt tooth where they connect with hypostomal carina. F 1 2.9 × pedicel length, F 2 2.2 × pedicel length. Mesosoma. Length 1.15 × width (measured at posterolateral propodeal corners). Dorsum of mesosoma with coarse confluent punctures; wavy carina separating dorsal and lateral faces of mesonotum. Side of mesosoma with micropunctures and short setae. Mesopleural lamella a short thick carina. Humeral carina distinct. No trace of scutellar scale. Lateral and posterior propodeal faces separated by row of distinct teeth. Metasoma. T 1 anterior face with separated punctures, posteriorly with dense setae obscuring sculpture. T 2 disc and S 2 with dense coarse subconfluent punctures; S 2 obscurely concave anterolaterally, anteromesally with longitudinal ridge. T 3 – 5 and S 3 – 5 with dense setae obscuring sculpture. S 1 with thick longitudinal lamella, notched sub-posteriorly. T 2 felt line deeply linear, 0.7 × T 2 total length. T 6 convex without pygidial plate, smooth. S 6 posterior margin distinctly bituberculate. MALE. See André (1898).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A339FFD6FF29FF7FCC61D392.taxon	distribution	Distribution. Laos, Thailand.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A339FFD6FF29FF7FCC61D392.taxon	discussion	Remarks. This species was previously known from males only and was treated as a member of the genus Mutilla, until the female was discovered in this paper. The sex association is initially based on identical ITS 1 sequences of a male (GenBank # OP 854799) from Chaiyaphum, Thailand, and a female (GenBank # OP 854800) from Chiang Mai, Thailand. In previous studies on Mutillidae sex associations using ITS (Pilgrim & Pitts 2006; Pilgrim et al., 2008; Williams et al. 2012), intraspecific sequence comparisons generally revealed genetic distances below 2 %, while interspecific sequence comparisons generally revealed genetic distances above 2 %. The identical ITS 1 sequences, coupled with similar body size, coloration, and overlapping distribution are adequate evidence to recognize the previously unrecognized female of Mutilla harmandi. This female was not previously named and is clearly a member of the genus Kurzenkotilla, permitting us to recognize a new combination for K. harmandi, and to recognize the male sex for the genus. This species was named in honor of Dr. François-Jules Harmand (1845 – 1921). Dr. J. Harmand explored in Indo- China region between 1875 and 1878 (Geiser & Nagel 2013, Huang et al. 2015) and collected insects at a mountain of La-khôn (Harmand, 1880). Some of his collections have a label with the same data “ LAKHON, HARMAND 1878 ” and were deposited in MNHN and SDEI. According to Harmand’s (1880) map, there was a province named La-khôn. Meanwhile, a place named La-khôn was in Lakhôn province and located on a bank of the Mekong [= “ Mèkhông ” in Harmand (1880)] (Huang et al. 2015: Fig. 18). Firstly, “ Montagnes de Lakhon ” lies in the vicinity of Tha Khaek, Khammuane Province, Laos (Geiser & Nagel, 2013) and La-khôn was in the vicinity of Mekong (Harmand, 1880) which is the boundary of modern Laos and Thailand (Huang et al. 2015).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A337FFD8FF29FF7FCA55D3EB.taxon	description	(Figs 21 – 28)	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A337FFD8FF29FF7FCA55D3EB.taxon	diagnosis	Diagnosis. FEMALE. Head and mesosoma ferruginous, rugoso-punctate, mandible apically and basally black, middle portion ferruginous, clypeus with tuft of suberect golden setae. T 1 apically with fringe of golden setae at postero-lateral edges, T 2 basally with two conjugended spots of golden setae, not reaching lateral edge of T 2, T 2 laterally with long felt line of golden setae, T 2 apically with fringe of golden setae, T 3 with band of golden setae, T 6 with tuft of erect long golden setae around pygidial plate, metasoma laterally with long erect golden setae, S 1 to S 6 with fringe of golden setae. MALE. Unknown.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A337FFD8FF29FF7FCA55D3EB.taxon	materials_examined	Material examined. Holotype. INDIA, Karnataka, Kodagu district, Bettathur, 12.417675 ° N, 75.666364 ° E, 958 m, 23. XII. 2019, P. Girish Kumar & Party, ♀ ZSI / WGRC / IR / INV. 21260 [ZSIK]. INDIA, Kerala, Kannur district, Kottiyoor, 11.900077 ° N, 75.887628 ° E, 882 m, 12. IV. 2022, Siva K. leg., ♀ ZSI / WGRC / IR / INV. 21261 [ZSIK].	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A337FFD8FF29FF7FCA55D3EB.taxon	description	Description. FEMALE. Body length 18.1 – 18.9 mm. Body stout, large-sized, moderately convex, slightly broadest at metasoma. Head oval, somewhat flattened, rugoso-punctate, posterior margin curved, genal carina well developed; eyes small, oval; longitudinal eye diameter in fronto-lateral view, 1.86 × distance between eyes. Clypeus straight, with medial deep emargination; mandible pointed towards the apex, bidentate; lower half of gena bordered beneath by a carina; hypostomal carina well developed. Mesosoma rectangular, with slightly concave laterally; rugoso-punctate as in head, lateral margin crenulated, propodeum posteriorly abrupt; propleuron rugoso-punctate as in head; metapleuron and allied lateral propodeal face smooth; mid and hind tibia with two rows of spines, one row with four and other one with six spines. T 1 wide, slightly narrower than T 2; T 2 with long lateral felt line, 0.75 × T 2; pygidium rugose with some small punctures. Coloration and setation. Antennae, head, clypeus, legs and mesosoma ferruginous with lateral margin darker than mesosoma dorsal; mandible apically and basally black, middle portion ferruginous; tibial spine dark brown; vertex with sparse suberect dark brown setae, towards lateral and frons with golden setae; clypeus with tuft of suberect golden setae; scape with golden setae and flagellomeres with short golden setae; mesosoma dorsally with dark brown setae throughout, apically towards propodeum with long erect golden setae; legs with long golden setae throughout, tarsomeres with dense dark brown setae; metasoma black, T 1 apically with golden fringe at posterolateral edges, T 2 basally with two conjoined spots of golden setae, not reaching lateral edge of T 2, T 2 laterally with long felt line of golden setae, T 2 apically with fringe of golden setae, T 3 with band of golden setae, T 6 with tuft of erect long golden setae around pygidial plate, metasoma laterally with long erect golden setae, S 1 – 6 with fringe of golden setae. Measurements. Ratio of pedicel length, F 1 length, F 2 length, and F 3 length, 0.22: 0.49: 0.24: 0.29; longitudinal eye diameter in fronto-lateral view: 1.18 mm, mesosoma maximal width in dorsal view: 3.51 mm, mesosoma length in dorsal view: 4.19 mm. MALE (unknown).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A337FFD8FF29FF7FCA55D3EB.taxon	distribution	Distribution. India (Karnataka, Kerala).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A337FFD8FF29FF7FCA55D3EB.taxon	discussion	Remarks. The female from Kerala differs by having slightly lighter golden setae, the genal carina not developed, the mesosoma dorsally with irregular dark patches, and the mesopleuron darker. Other than K. visrara, whose apparently erroneous type locality is Sri Lanka (see below), this is the southwesternmost representative of the genus Kurzenkotilla. Based on its southern distribution and broadly transverse band on T 2, we initially thought it may actually belong in the genus Storozhenkotilla. Furthermore, this seems to be the only Kurzenkotilla species with a sculptured apparent pygidial plate. The species seems to fit more closely with Kurzenkotilla, however, based on the moderately large head and eye, the weakly defined genal carina, and the black T 1 fringe. Furthermore, the transverse setal band on T 2 more closely resembles two coalescent spots, due to the black triangular patch of setae basomesally. This band is also more centrally located on the T 2 disc than that seen in Storozhenkotilla, in K. cicatricifera, the T 2 band extends posteriorly to the midpoint of T 2, while the band is restricted to the basal quarter of T 2 in the genus Storozhenkotilla.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FF7FCAB4D610.taxon	materials_examined	Material examined. PAKISTAN: Baluchistan, near Deragazichan village, Indus River valley, 26. III. 2003, S. Ovchinnikov leg., 1 ♀ [IBSS]; Punjab, Lahore, 7. VII. 1946, 1 ♀ [UMMZ]. INDIA: Rajasthan, Bharatpur, 22. X. 1995, R. Carradori leg., 1 ♀ [MSNF]	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FF7FCAB4D610.taxon	distribution	Distribution. * Pakistan, India (Bihar, Delhi, Haryana, Madhya Pradesh, Maharashtra, Odisha, Rajasthan *).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FF7FCAB4D610.taxon	discussion	Remarks. The male of this species might eventually be recognized to be K. semiviolacea, comb. nov., based on their co-occurrence in both India and Pakistan.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FD29CD5CD162.taxon	description	(Figs 29 – 31)	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FD29CD5CD162.taxon	materials_examined	Material examined. INDIA: Rajasthan, Jodphur, 26 ° 13.42 ′ N 73 ° 01.96 ′ E, 190 m, malaise in garden, 11 – 16. VIII. 2008, M. E. Irwin & D. R. Priyadarsanan leg., 1 ♁ [EMUS]; New Delhi, IARI, 28 ° 37 ′ 51 ″ N 77 ° 9 ′ 50 ″ E, 230 m, 3. VII. 2005, J. Heraty leg., 1 ♁ [UCRC, UCRCENT # 434275]; Uttar Pradesh, Aligarth Muslim University, 27 ° 55 ′ 00 ″ N 78 ° 4 ′ 41 ″ E, 190 m, 28. VI. 2005, J. Heraty leg., 1 ♁ [UCRC, UCRCENT # 441172]. NEPAL: Jumla district, Sinja Khola, 1900 – 2000 m, W. Schwaller leg., 10. VI. 1998, 1 ♁ [IBSS]; Bagmati, 16. VII. 1974, S. Yamaguchi & T. Aoki leg., 1 ♁ [NARO].	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FD29CD5CD162.taxon	distribution	Distribution. * Nepal, India (Delhi, Rajasthan, Uttar Pradesh).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FD29CD5CD162.taxon	discussion	Remarks. In both coloration and structure, this species is similar to K. semiviolacea, comb. nov. Prior to this paper, the species was included in the genus Mutilla (Lelej 2005), but its similarity to the males of K. annamensis and K. harmandi, and its occurrence in northern India and Nepal suggest that it belongs in Kurzenkotilla.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FABBCAF5D335.taxon	materials_examined	Material examined. NEPAL: Jumla District: lower Sinja Khola, 1700 – 1900 m, 9. VI. 1998, G. Miksch leg., 2 ♀ [IBSS]; Mahakali / Darchula Ghamna, Chamliya Khola, 29 º 41 ′ 25 ″ N 80 º 41 ′ 04 ″ E, 30. V. 2005, 830 m, at light, A. Weigel leg., 1 ♀ [IBSS]; Unknown district: Nepal, I. Yoneta leg., 1 ♀ [SEHU] .. INDIA: Uttarakhand, Gobind Ghat Village, 30 º 37.5 ′ N 79 º 33.5 ′ E, 1750 – 1900 m, 17 – 23. V. 1999, Yu. Marusik leg., 2 ♀ [IBSS]; Dehra Dun, 28. IX. 1944, 1 ♀ [UMMZ].	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FABBCAF5D335.taxon	distribution	Distribution. * Nepal, India (Uttarakhand, Himachal Pradesh).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FABBCAF5D335.taxon	discussion	Remarks. The male of this species might eventually be recognized to be K. rufodorsata, comb. nov., based on their co-occurrence in both India and Nepal.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FF7FCCACD627.taxon	description	(Figs 32 – 35)	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FF7FCCACD627.taxon	materials_examined	Material examined. INDIA: Agra, 2. III. 1958, 1 ♁ [NSMT]; PAKISTAN: Margalla Hills National Park, 20. VI. 2005, Malaise trap, H. Khurram & M. Fida, 1 ♁ [IBSS].	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FF7FCCACD627.taxon	distribution	Distribution. Pakistan, India (Himalaya).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FF7FCCACD627.taxon	discussion	Remarks. In both color and structure, this species is similar to K. rufodorsata, comb. nov. Based on similarity to S. aurofasciata, Lelej et al. (2007) transferred the species to Storozhenkotilla. The species, however, lives outside the range for other species in Storozhenkotilla and more closely resembles K. harmandi and K. annamensis in coloration, and clypeal and pygidial morphology, so we here transfer the species to Kurzenkotilla.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FD66CD2BD03D.taxon	materials_examined	Material examined. INDIA: Rajasthan: Bharatpur, VIII. 1985, A. Riedel leg., 1 ♀ [IBSS]; Jaipur, 21. IV. 1945, 1 ♀ [UMMZ]; Chandigarh: Sukhna Lake, 15. X. 2007, Sk. Yamane leg., 1 ♀ [IBSS].	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FD66CD2BD03D.taxon	distribution	Distribution. * India (Rajasthan, Chandigarh), Sri Lanka.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FD66CD2BD03D.taxon	discussion	Remarks. Although AL examined large volumes of mutillid material from Sri Lanka, the female of Mutilla visrara was not discovered there. Possibly, the type locality of this species is wrong. The late Børge Petersen, who studied most of the types of Cameron’s Oriental species, wrote AL (Petersen, in litt., 1980), that several of Cameron’s mutillid species described from Sri Lanka are likely based on material from outside of Sri Lanka.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FB4CCB88D2FF.taxon	materials_examined	Type species: Mutilla europaea Linnaeus, 1758 by subsequent designation (Latreille 1810).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FB4CCB88D2FF.taxon	diagnosis	Diagnosis. FEMALE. In the Oriental and eastern Palaearctic regions, this genus is unique in having the mesosomal sides parallel, T 1 broad, and the T 2 disc with uniformly black setae. Other diagnostic features include: mandible unarmed ventrally, head usually wider than mesosoma, mesosomal width equal to or narrower than pronotum in dorsal view, and T 6 lacking defined pygidial plate. MALE. In the Oriental region, males can be separated from other Mutillinae genera by having the mandible unarmed ventrally, the clypeus with a medial projection, the tegula elongate, the mesoscutellum unarmed, T 1 broad, T 7 tuberculate or carinate, and the paramere with a ventral brush of long setae.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FB4CCB88D2FF.taxon	discussion	Remarks. Seventy species are currently placed in the genus Mutilla, mostly from the Afrotropical and Palaearctic regions (Bischoff 1920, Lelej 2002, Pagliano et al. 2020). The eastern Palaearctic Mutilla mikado Cameron, 1900 penetrates into Oriental areas of China. Other Oriental species that were previously placed in Mutilla (Lelej 2005) are transferred into the genera Kurzenkotilla or Storozhenkotilla in this paper.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDDFF29F90DCC15D679.taxon	description	(Figs 1 – 4)	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDDFF29F90DCC15D679.taxon	materials_examined	Material examined. RUSSIA: * Kuril Islands, SW Kunashir, Ivanovskij Cape, 28 – 29.09.2013, Yu. & L. Sundukov, 2 ♀ [IBSS]. CHINA: * Sichuan, 35 km W Zhangle, 3620 – 3720 m, 12 – 13. VII. 2001, S. Murzin, 1 ♀ [IBSS]; * Shaanxi, S of Huaxian, SE Damingzhen, Huashan Mts., 2010 – 2155 m, 24. V. 2009, I. Belousov & I. Kabak, 1 ♀ [IBSS].	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDDFF29F90DCC15D679.taxon	distribution	Distribution. Russia: Eastern Siberia (south), Far East (Amurskaya oblast, Khabarovskii krai, Primorskii krai, Sakhalin, * Kuril Islands (Kunashir )); China (Gansu, Heilongjiang, Jilin, Neimenggu, Shanxi, * Shaanxi, * Sichuan, Zhejiang, Yunnan), Japan (Hokkaido, Honshu, Kyushu, Shikoku), North Korea, South Korea.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDDFF29F90DCC15D679.taxon	discussion	Remarks. Unlike the other species treated here, Mutilla mikado is predominantly a Palaearctic species and only occurs in a few Oriental areas of China. In some older papers (e. g., Chen 1957), this species was treated as a subspecies of Mutilla europaea Linnaeus, 1758. More recently, Mutilla europaea was recorded as a parasite of Bombus breviceps Smith, 1852, in Yunnan, China, but the mutillid in question was later determined to be Mutilla mikado, which is the only Mutilla species known to occur in Oriental regions of China.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFDFFF29FF7FCD43D7BD.taxon	materials_examined	Type species: Standfussidia taprobane Lelej, 2005, ♀, by original designation.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFDFFF29FF7FCD43D7BD.taxon	diagnosis	Diagnosis. MALE. Unknown. FEMALE. Head width equal to mesosomal maximum width. Eyes small, rounded convex, but not hemispherical, closer to mandibular base than posterior head border. F 1 1.25 × longer its maximum width, relation of pedicel and F 1 – 3 0.5: 0.6: 0.6: 0.6. Scape curved. Mandible tridentate. Frons lacking carina between tubercles and without carina between tubercle and inner eye border. Maxillary palps shorter than antenna length. Mesosoma length equals its maximum width, mesoscutellum narrowed posterad, propodeum distinctly widened posterad with abrupt posterior slope. Mesosoma dorsally without any traces of sutures, mesopleuron distinctly convex with vertical row of long setae, scutellar scale lacking, posterior dorsal border not dentate nor serrate. Legs not shortened, fore tarsus with pectinate comb, mid and hind tibia with two rows of strong spines. Metasoma distinctly flattened dorso-ventrally, segment 2 very long, segment 1 very wide not constricted posterad. T 2 with rather short lateral felt line; S 2 without any lateral felt line. T 6 with narrow pygidial plate, carinated laterally, with granulated and striate sculpture. Body setae usual, not elongated. Species included. Standfussidia taprobane Lelej, 2005.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFDFFF29FF7FCD43D7BD.taxon	distribution	Distribution. Oriental region: Sri Lanka.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFDFFF29FF7FCD43D7BD.taxon	discussion	Remarks. This genus differs from all other Oriental Mutillini by the coloration and pygidial structure. The well-defined triangular pygidial plate, and a single basomesal setal spot on T 2 and a fore tarsal comb are more similar to members of the tribe Smicromyrmini. The wide T 1 shape, however, is diagnostic for the Mutillini. The eventual discovery of males for this genus could necessitate a change in its tribal placement in the future.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFDFFF29FBC7CD5FD18E.taxon	materials_examined	holotype, ♀, Sri Lanka: Monaragala District: Angunakolapelessa, 27 – 28. III. 1981, K. V. Krombein, T. Wijesinhe, L. Weeratunge leg. [USNM], examined; Pagliano et al. 2020: 236, Fig. 362, ♀.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFDFFF29FBC7CD5FD18E.taxon	materials_examined	Material examined. Paratypes: SRI LANKA, 2 ♀ with the same label as holotype; Anuradhapura District: Padaviya, irrigation bungalow, 18 – 19. V. 1976, K. Krombein et al. leg., 1 ♀; Monaragala District: Mau Ara, 16 km E of Uda Walawe, 100 m, 24 – 26. IX. 1977, K. Krombein et al. leg., 1 ♀; Angunakolapelessa, 30. IX – 1. X. 1977, K. Krombein et al. leg., 1 ♀; Hambantota District: Palatupana tank, 10 – 16 m, 29. III – 2. IV. 1981, K. Krombein et al. leg., 1 ♀ [USNM, IBSS].	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFDFFF29FBC7CD5FD18E.taxon	distribution	Distribution: Sri Lanka.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFDFFF29FBC7CD5FD18E.taxon	discussion	Remarks. The females of Standfussidia taprobane have been collected in the dry zone of Sri Lanka.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFC0FF29F9A8CB0AD775.taxon	materials_examined	Type species: Mutilla aurofasciata André, 1907 by original designation.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFC0FF29F9A8CB0AD775.taxon	diagnosis	Diagnosis. MALE. Head width less than pronotal width, not rounded posterad; preoccipital carina dorsally well developed, sharp and slightly projecting over head. Eyes weakly notched inside. Ocelli small. Mandible bidentate without subbasal tooth beneath and without subbasal inner tooth. F 2 1.3 × F 1. Prementum not tuberculate. Notauli and parapsids visible. Tegula longer than pronotal length laterad, projecting over axilla. Pterostigma large closed. Mesoscutellum non-modified. Propodeal posterior slope with strong longitudinal medial carina. Metasomal segment 1 wide transverse. T 2 with long lateral felt line, S 2 without any lateral felt line. S 8 (hypopygium) not modified, T 7 without carina. Gonostylus wide, basiventrally setose. Volsella shorter than penial valve and consists of wide cuspis and short process-like digitus; penial valves symmetrical, apically setose. T 1 dorsally and T 2 – 3 (4) posterad with band of pale setae. FEMALE. Head large, widened posterad, width slightly less than pronotal width; behind eye elongated; eye small. Genal carina well developed, hypostomal carina widened medially. Prementum not tuberculate. F 1 1.5 × F 2. Mandible bidentate, without subbasal inner tooth. Mesosoma with concave mesopleuron, scutellar scale lacking. Mesopleuron with well developed vertical (supracoxal) carina. Promesonotal suture carinated in type species. Fore tarsus without comb of strong spines. Mid and hind tibia with two rows of spines. Metasomal segment 1 wide transverse; T 2 with lateral felt line. T 2 basally with pale band which medially emarginated posterad and apical narrow pale band; T 3 with wide pale band. T 6 with distinct pygidial plate weakly carinated lateroapically, shiny or shagreened with a few rugae. S 6 with two strong apical tubercles. Species included. Storozhenkotilla aurofasciata (André, 1907), ♁ ♀; S. binghami (Lelej, 2005), ♀, comb. nov.; and S. nathani Lelej, sp. nov., ♁.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFC0FF29F9A8CB0AD775.taxon	distribution	Distribution. Oriental region (South India, Sri Lanka).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFC0FF29F9A8CB0AD775.taxon	discussion	Remarks. Storozhenkotilla is most similar to Mutilla Linnaeus, 1758. The male differs by having the gonostylus weakly setose (densely setose in Mutilla and many Kurzenkotilla), and by having a lateral clypeal projection (medial projection in Mutilla). In having setal markings on the basal part of T 2, females of Storozhenkotilla Lelej resemble those of Kurzenkotilla Lelej but easily differ by having setal markings forming a transverse pale band basally on T 2 (with two pale setal spots disposed transversely in Kurzenkotilla).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32FFFC0FF29FC8ECB72D2C7.taxon	description	(Figs 44 – 54) Mutilla aurofasciata André, 1907: 255, ♀, holotype, ♀, Pankulam / Ceylon [Sri Lanka] / W. Horn / Holotype / Mutilla aurofasciata ♀ nov. sp. [SDEI], examined.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32FFFC0FF29FC8ECB72D2C7.taxon	materials_examined	Material examined. SRI LANKA: Mannar District: 0.8 km NE Kokmotte Bungalow, Wilpattu National Park, 21 – 25. V. 1976, K. V. Krombein, P. B. Karunaratne, S. Karunaratne, D. W. Balasooriya leg. / Mutilla aurofasciata André, B. Petersen det. 1980, 1 ♀ [IBSS]. INDIA: Karnataka, Bangalore, 916 m, 29. X. 1978, K. D. Ghorpade leg., A 719 / Mutilla aurofasciata André, B. Petersen det. 1980, 1 ♀ [IBSS]; Tamil Nadu: Kodaikanal, 6500 ft [1981 m], Pulney Hills [Palni Hills], IV – V. 1953, P. S. Nathan, K. V. Krombein collection, 1975 – 1976, 1 ♁ [USNM]; Madras, Coimbatore, 1400 ft., VIII. 1963, P. Susai Nathan, 1 ♀ [FSCA]; Thirunelveli district, Kodumundiyar dam site, 26. IX. 2018, P. Girish Kumar, 1 ♁, ZSI / WGRC / IR / INV. 21298 [ZSIK]; Pondicherry: Auroville, 10 km N Pondicherry [Puducherry], 24. X. 2004, F. Burger, 1 ♁; Auroville, 10 km N Pondicherry, Canians, 25. XI. 2005, F. Burger, 1 ♀; Auroville, 10 km N Pondicherry, 7. I. 2007, F. Burger, 1 ♀; same place, 12. II. 2007, F. Burger, 1 ♀; same place, 20. II. 2007, F. Burger, 1 ♀; same place, 5. III. 2007, F. Burger, 1 ♀; same place, 13. III. 2007, F. Burger, 1 ♀; same place, 23. III. 2007, F. Burger, 1 ♀ [IBSS].	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32FFFC0FF29FC8ECB72D2C7.taxon	distribution	Distribution. Sri Lanka, South India (Karnataka, * Pondicherry, Tamil Nadu).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32FFFC0FF29FC8ECB72D2C7.taxon	discussion	Remarks. This is the type species for the genus Storozhenkotilla and the females are easily separated from all other Oriental Mutillini species by the transverse suture between the pronotum and mesonotum. Males are less obviously distinct from other Mutillini, but they do seem to be the only Oriental Mutillini with T 7 uniformly punctate and the metasoma without any metallic luster.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32FFFC1FF29F911C993D6E5.taxon	materials_examined	Material examined. SRI LANKA: Mannar District: 0.8 km NE Kokmotte Bungalow, Wilpattu National Park, 21 – 25. V. 1976, K. V. Krombein, P. B. Karunaratne, S. Karunaratne, D. W. Balasooriya, 2 ♀ [USNM]. INDIA: Tamil Nadu, Tiruchirappali District, Pudukkottai: XII. 1991, T. R. S. Nathan, 1 ♀ [SEMC]; XI. 1992, T. R. S. Nathan, 1 ♀ [SEMC]; XI. 2002, T. R. S. Nathan, 1 ♀ [FSCA]. INDIA: Karnataka, 15 km N Bangalore [Bengaluru], KT, 23 – 24. VII. 1996, K. Werner & L. Lorenz, 5 ♀ [MSNF].	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32FFFC1FF29F911C993D6E5.taxon	distribution	Distribution. * Sri Lanka, India (Karnataka, Tamil Nadu, West Bengal).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32FFFC1FF29F911C993D6E5.taxon	discussion	Remarks. The type of this species, from West Bengal, is the only specimen of Storozhenkotilla known from northern India. As suggested in the remarks for K. visrara above, there are doubts about the veracity of type localities for many of Cameron’s species. It seems possible that the type could be mislabeled and the genus Storozhenkotilla truly is restricted to a southern Indian and Sri Lankan distribution. Previous studies (Lelej 2005) considered K. cicatricifera comb. nov. as a member of this genus that could be separated from S. binghami by minor color and sculpture differences (see couplet 5 in key to females above). The form that was previously considered S. cicatricifera (Figs 57, 58) is here treated as a color variant of S. binghami, but further studies could eventually reveal it as a discrete species.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32EFFC3FF29F91BC960D5B1.taxon	description	(Figs 59 – 64)	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32EFFC3FF29F91BC960D5B1.taxon	diagnosis	Diagnosis. Body antennae and legs black with bluish metallic luster on head, pronotum and metallic blue metasoma. Propodeal posterior slope with strong longitudinal medial carina which ends dorsally by tooth. T 7 laterally angulate, apical border smooth with wide medial smooth triangle without carina. T 1 – 6 and S 2 – 5 with apical fringe of whitish setae.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32EFFC3FF29F91BC960D5B1.taxon	materials_examined	Material examined. Holotype, ♁, INDIA: Karnataka, Mysore [Mysuru], Shimoga dist [rict], Agumbe Ghat, 600 m, V. 2004, TRSN coll. [IBSS].	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32EFFC3FF29F91BC960D5B1.taxon	description	Description. FEMALE (unknown). MALE. Head width less than pronotal width, not rounded posterad. Preoccipital carina dorsally well developed, sharp and slightly projecting over head. Eyes well notched inside. Ocelli small, ratio POD: OOD = 0.7. Mandible bidentate without basal tooth beneath and without subbasal tooth on inner border. F 2 1.3 × F 1. Prementum not tuberculate. Notauli well visible in posterior half, parapsids weak. Tegula longer than pronotal length laterad, projecting over axilla. Propodeal posterior slope with strong longitudinal medial carina which ends dorsally by tooth. Metasomal segment 1 wide transverse. T 2 with long lateral felt line, S 2 without any lateral felt line. Sternum 8 (hypopygium) not modified. T 7 laterally angulate, apical border smooth with wide medial smooth triangle without carina. Gonostylus wide, basiventrally setose. Volsella slightly shorter than penial valve and consists of wide cuspis and short finger-like digitus; penial valves symmetrical, posterior margin with vertical row of setae, postero-ventrally bidentate, pre-apical tooth bluntly rounded. Head black with bluish metallic luster. Antennae entirely black, scape and pedicel with dense whitish setae. Head densely punctured, sparsely around ocelli, with dense whitish setae all over. Mesosoma black, pronotum with bluish metallic luster, with sparse whitish setae denser on mesopleuron, mesoscutum with sparse black setae. Tegula sparsely punctured, laterally and posteriorly smooth. Pronotum, mesoscutum and mesoscutellum coarsely punctured; propleuron, mesopleuron and lateral propodeal face coarsely punctured, metapleuron smooth; propodeum posterad with sharp slope. Legs with dense whitish setae. Metasoma metallic blue. T 1 – 2 densely punctured, sparser on disk of T 2. T 1 – 6 and S 2 – 5 with apical fringe of whitish setae. The wings fusco-violaceous, basal radial abscissa sharply oblique, apically curved; first transverse radio-medial vein oblique, second almost straight, third sharply angled in the middle; first recurrent vein is received in basal third, second at apical third of cell.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32EFFC3FF29F91BC960D5B1.taxon	distribution	Distribution. India (Karnataka).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32EFFC3FF29F91BC960D5B1.taxon	etymology	Etymology. The specific name is dedicated to P. Susai Nathan (1891 – 1976), Indian naturalist, entomologist, natural history specimens collector, who collected the Mutillidae in South India (Tamil Nadu).	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32EFFC3FF29F91BC960D5B1.taxon	discussion	Remarks. This species may be the opposite sex of S. binghami (Lelej, 2005), which was also collected in Karnataka.	en	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
