identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
583A87D7A33DFFD3FF29FE87CDEED697.text	583A87D7A33DFFD3FF29FE87CDEED697.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mutillini Latreille 1802	<div><p>Key to the Oriental genera and species of tribe Mutillini</p> <p>Males (unknown in Kurzenkotilla ariana, K. indica, K. niveosignata, K. scrobiculata, K. visrara, Standfussidia taprobane, Storozhenkotilla binghami, and K. cicatricifera)</p> <p>1. Mandible strongly dilated apically, tridentate; clypeus with sparse short setae, not obscuring sculpture (China, Japan, Korea, Russia)................................................................... Mutilla mikado Cameron, 1900</p> <p>- Mandible not strongly dilated apically, clearly bidentate; clypeus with long dense setae, obscuring sculpture, at least mesally................................................................................................... 2</p> <p>2 (1). Frons, vertex, and pronotum pale reddish orange; clypeus with especially long and dense setae that cover mandibles; ventral setae of paramere especially dense and long (Laos, Thailand, Vietnam)...................... 3 (Kurzenkotilla, in part)</p> <p>- Head and mesosoma more extensively blackened, with at least pronotum blackened laterally; clypeal setae not so long, usually not covering mandibles; ventral paramere setae shorter and sparser (India, Nepal, Pakistan, Sri Lanka)................ 4</p> <p>3 (2). Pale metasomal setae predominantly silver; T2 disc lacking basolateral circular patch of appressed setae; fringes of T4–5 with setae entirely black (Thailand, Vietnam)............................................. K. annamensis Lelej, 2005</p> <p>- Pale metasomal setae usually predominantly golden; T2 disc with basolateral circular patch of appressed golden setae; fringes of T4–5 with scattered pale golden setae (Laos, Thailand).................... K. harmandi (André, 1898), comb. nov.</p> <p>4 (2). Anterior border of clypeus strongly emarginated and bordered by strong tubercle; clypeus with long dense golden setae; T1–3(4) posterad with narrow band of golden setae; T7 not angulate lateroapically; T7 densely punctate throughout (India, Sri Lanka)......................................................... Storozhenkotilla aurofasciata (André, 1907)</p> <p>- Anterior border of clypeus shallowly concave and bordered by weak tubercle; clypeus with sparse yellowish or whitish setae; T1–3(4) posterad with narrow band or fringe of whitish or yellowish setae; T7 angulate lateroapically; T7 densely punctate, medially partly smooth................................................................................ 5</p> <p>5 (4). Mesosoma totally black; metasoma strongly violaceous (India)................ Storozhenkotilla nathani Lelej, sp. nov.</p> <p>- Mesosoma black with ferruginous-red mesoscutum, mesoscutellum, and pronotum dorsally; metasoma weakly violaceous.................................................................................. 6 (Kurzenkotilla, in part)</p> <p>6 (5). Fore wing with distal portion dark brown-violaceous, basally yellow-brown; T7 medially with wide smooth basal area and carinated apical triangle (India, Nepal)............................... K. rufodorsata (Cameron, 1897), comb. nov.</p> <p>- Fore wing uniformly pale brown; T7 medially with narrow basal smooth line and weakly carinated apical triangle (India, Pakistan)......................................................... K. semiviolacea (André, 1896), comb. nov.</p> <p>Females (unknown in Kurzenkotilla rufodorsata, K. semiviolacea, and Storozhenkotilla nathani)</p> <p>1. T2 disc with setae uniformly black basally, without appressed whitish or golden setal patches; T6 with no trace of pygidial plate, uniformly punctate and covered with short whitish setae (China, Japan, Korea, Russia)................................................................................................... Mutilla mikado Cameron, 1900</p> <p>- T2 disc with appressed patch or patches of whitish or golden setae in the basal half; T6 with apparent pygidial plate, mesally mostly bare, sculpture rugose, shagreened, or smooth....................................................... 2</p> <p>2 (1). T2 basally with one mesal spot of white setae; T6 with flat elongate pygidial plate defined throughout its length by lateral carinae (Sri Lanka)...................................................... Standfussidia taprobane Lelej, 2005</p> <p>- T2 basally with transverse pale band or with two ovate pale spots disposed transversely; T6 generally convex, at most with weakly defined pygidial plate having short carinae lateroapically.............................................. 3</p> <p>3 (2). Promesonotal suture carinated and distinct; dorsal and lateral pronotal surfaces separated by thick raised oblique carina; T2 disc with complete transverse band of golden setae, confluent with golden T1 fringe (India, Sri Lanka).................................................................................. Storozhenkotilla aurofasciata (André, 1907)</p> <p>- Promesonotal suture not carinated, indistinct; dorsal and lateral pronotal surfaces not separated by thick raised carina; T2 disc with mesally interrupted transverse band, with separated discal spots disposed transversely, or with mesally constricted transverse band that is separated from black T1 fringe mesally................................................ 4</p> <p>4 (3). T2 disc with mesally interrupted transverse band of golden setae basally, band confluent with golden T1 fringe; pygidial plate weakly rugose, defined by distinct short lateral carinae................................. 5 (Storozhenkotilla, in part)</p> <p>- T2 disc with white or golden setal patches usually separated from T1 fringe; if T2 patches or band confluent with T1 fringe, then T1 fringe composed of black setae mesally; pygidial plate smooth or obscurely shagreened, with lateral carina indistinct or obliterated; head as wide or wider than mesosoma........................................... 6 (Kurzenkotilla)</p> <p>5 (4). Head dorsally reddish-orange, mesosoma laterally blackish with dorsomesal redish-orange area, densely areolate-punctate dorsally (India).................................................................. S. binghami (Lelej, 2005)</p> <p>- Head mostly black with obscure reddish-brown dorsal area; mesosoma entirely black, coarsely rugoso-punctate dorsally (India, Sri Lanka). S. binghami (Lelej, 2005), comb. nov. (Mutilla valida Cameron, 1897, nom. praeocc., non Smith, 1855)</p> <p>6 (4). Head orange-brown; T2 disc with golden patches large or confluent, distance between spots less than their diameter or spots confluent; T2 fringe and T3 with wide band of golden setae................................................... 7</p> <p>- Head black; T2 basally with two small spots of yellowish setae, distance between spots 1.5 × their diameter; T2 fringe mostly composed of black setae, T3 with two separated patches of whitish or yellowish setae............................. 10</p> <p>7 (6). T2 disc with mesally constricted transverse band of golden setae (India)........ K. cicatricifera (André, 1894), comb. nov.</p> <p>- T2 disc with two separated spots of golden setae........................................................... 8</p> <p>8 (7). Mandible bidentate; F1 1.75 × F2; legs concolorous with pale orange-brown mesosoma (Afghanistan).................................................................................. K. ariana (Lelej in Lelej &amp; Kabakov, 1980)</p> <p>- Mandible apparently tridentate, middle tooth blunt and somewhat confluent with lower tooth; F1 1.5 × F2; legs generally darker than mesosoma (Southeastern Asia)................................................................ 9</p> <p>9 (8). Basal spots larger, distance between them ~0.3 × their diameter; mesosoma scarcely longer than wide with propodeum and mesonotum of similar widths (Thailand, Vietnam)..................................... K. annamensis Lelej, 2005</p> <p>- Basal spots smaller, distance between them ~0.6 × their diameter; mesosoma clearly longer than wide with propodeum laterally expanded (Laos, Thailand)....................................................... K. harmandi (André, 1898)</p> <p>10 (6). T1 dorsally with black setae (India, Nepal)...................................... K. scrobiculata (Hammer, 1962)</p> <p>- T1 dorsally with more or less dense orange setae.......................................................... 11</p> <p>11 (10). T1 dorsally with dense ferruginous setae; S2 with three longitudinal carinae, lateral ones ending apically by tubercle; T4 generally with two spots of whitish setae, often sparser than those on T3 (India, Pakistan)... K. niveosignata (André, 1899)</p> <p>- T1 dorsally with sparser ferruginous setae; S2 without three longitudinal carinae; T4 at most with scattered whitish setae (India, Sri Lanka).............................................................. K. visrara (Cameron, 1898)</p></div> 	https://treatment.plazi.org/id/583A87D7A33DFFD3FF29FE87CDEED697	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33CFFD4FF29FCA2CA60D5E9.text	583A87D7A33CFFD4FF29FCA2CA60D5E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kurzenkotilla Lelej 2005	<div><p>Genus Kurzenkotilla Lelej, 2005</p> <p>Kurzenkotilla Lelej, 2005: 170, ♀; Nithya &amp; Dey 2013: 289, Brothers &amp; Lelej 2017: 96; Terine et al. 2020a: 3; Terine et al. 2020b: 446; Pagliano et al. 2020: 225</p> <p>Type species: Artiotilla ariana Lelej in Lelej &amp; Kabakov, 1980 by original designation.</p> <p>Diagnosis. MALE. Head width less than pronotal width, rounded posterad; occipital carina dorsally well developed, sharp and slightly projecting over pronotum. Eye weakly notched inside. Ocelli small. Mandible bidentate without subbasal tooth beneath and without subbasal inner tooth. F2 longer than F1. Prementum with longitudinal mesal ridge. Notauli and parapsids visible. Tegula longer than pronotal length laterad, projecting over axilla. Pterostigma large, closed. Fore tarsus without comb of strong spines. Mid and hind tibia with two rows of weak spines. Mesoscutellum non-modified. Propodeal posterior slope with strong longitudinal medial carina. Metasomal segment 1 widely transverse. T1 dorsally and T2–3(4) each with band of pale setae posterad. T2 with long lateral felt line, S2 without any lateral felt line. S8 (hypopygium) not modified, T7 generally with smooth triangular area posteromesally. Gonostylus wide, basiventrally setose. Volsella usually shorter than penial valve and consisting of wide cuspis and short finger-like digitus; penial valves symmetrical, apically setose.</p> <p>FEMALE. Head large, somewhat rounded posterad, width equal to pronotal width; behind eyes elongated; eyes large, distance between postero-dorsal margin of eye and that of vertex less than eye longitudinal diameter, projecting over head. Hypostomal carina slightly widened in medial portion. Prementum not tuberculate. F1 1.5–2.0 × F2. Mandible rather wide, bidentate, without subbasal inner tooth, preapical tooth obtuse (tridentate in K.annamensis and K. harmandi). Mesosoma with concave mesopleuron, scutellar scale lacking. Mesopleuron with developed vertical (supracoxal) carina. Mid and hind tibia with two rows of weak spines. Metasomal segment 1 wide transverse; T2 with lateral felt line. S2 latero-apically swollen, swell in type species K. ariana ending in tooth. T2 basally with larger or smaller pale spots disposed transversely, T3 with wide pale band or lateral spots. T6 without distinct pygidial plate, apical part microgranulated and covered by dense setae. S6 with two strong apical tubercles.</p> <p>Species included. Kurzenkotilla annamensis Lelej, 2005, ♁ ♀; K. ariana (Lelej in Lelej &amp; Kabakov, 1980), ♀; K. harmandi (André, 1898), ♁ ♀, comb. nov.; K. cicatricifera (André, 1894), comb. nov., ♀; K. niveosignata (André, 1894), ♀; K. rufodorsata (Cameron, 1897), ♁, comb. nov.; K. scrobiculata (Hammer, 1962), ♀; K. semiviolacea (André, 1896), ♁, comb. nov.; and K. visrara (Cameron, 1898), ♀.</p> <p>Distribution. Palaearctic (Afghanistan) and Oriental Regions.</p> <p>Remarks. Kurzenkotilla Lelej is closely related to Storozhenkotilla Lelej but, in addition to setal markings of T2, differs by lacking a pygidial plate in the female (weakly carinated pygidial plate present in Storozhenkotilla). By having two pale discal setal spots on T2, the female of Kurzenkotilla resembles that of Macromyrme Lelej, 1984, but differs by lacking a subbasal inner tooth on the non-acuminate mandible (with developed subbasal inner tooth on an acuminate mandible in Macromyrme), and by having larger eyes (eyes very small in Macromyrme). Males of Kurzenkotilla are similar to those of Storozhenkotilla and, as yet, there are no clear characters that can be used to separate all Kurzenkotilla from all Storozhenkotilla, which is why species from both genera are included in the same key above. Males of the apparently closely related Macromyrme are currently unknown, but they will likely be similar to these genera as well.</p> </div>	https://treatment.plazi.org/id/583A87D7A33CFFD4FF29FCA2CA60D5E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33BFFD4FF29FE32CB88D323.text	583A87D7A33BFFD4FF29FE32CB88D323.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kurzenkotilla annamensis Lelej 2005	<div><p>Kurzenkotilla annamensis Lelej, 2005</p> <p>(Figs 7–13)</p> <p>Kurzenkotilla annamensis Lelej, 2005: 171, ♀, holotype, ♀, Vietnam, Prov. Nghe Anh [currently Prov. Khu Bon Cu], Con River, Naca, 200 m, 8.IX.1963, N. Kabakov leg. [IBSS], examined.</p> <p>Material examined. THAILAND: Chiang Mai: Samoeng, Ban Ang Khai, 12–15.V.2003, M. Maruyama leg., flight interception trap, 2♁ [TUA]. VIETNAM: Hoa Binh, Tonkin, 1919, 1♀ [RBINS]; Ninh Thuan, Nui Chua N.P., northeast part, malaise traps, 90–150 m, 23–30.V.2007, 1♁ [RMNH].</p> <p>Distribution. * Thailand, South Vietnam.</p> <p>Description. FEMALE. See Lelej (2005). MALE. Head width less than pronotal width, not rounded posterad. Preoccipital carina dorsally well developed, sharp and slightly projecting behind head. Eyes well notched inside. Ocelli small, ratio POD:OOD=0.75. Mandible bidentate without basal tooth beneath and without subbasal tooth on inner border. F2 1.25 ×F1. Prementum with raised mesal longitudinal carina. Notauli well visible in posterior half, parapsids weak. Tegula longer than pronotal length laterad, projecting over axilla. Propodeal posterior slope with strong longitudinal medial carina which ends dorsally in tubercle. Metasomal segment 1 widely transverse. T2 with long lateral felt line, S2 without any lateral felt line. Sternum 8 (hypopygium) not modified. T7 obscurely angulate laterally, apical border smooth with wide medial smooth triangle marked by blunt carinae laterally. Gonostylus wide, basiventrally setose. Volsella slightly longer than penial valve and consists of wide cuspis and short fingerlike digitus; penial valves symmetrical, posterior margin with vertical row of setae, postero-ventrally bidentate, pre-apical tooth sharp, larger than apical tooth.</p> <p>Head orange-brown with mandible, clypeus, and postgenal area blackened. Antennae black, except scape with obscure yellow-brown apical area, scape and pedicel with brown and whitish setae. Head densely punctured, setae interspersed pale yellow and dark brown. Mesosoma orange-brown, except mesopleuron ventral half, metapleuron, and propodeum black; setae predominantly silvery-white laterally, dorsal setae largely orange-brown. Tegula moderately punctate, laterally and posteriorly smooth. Pronotum, mesoscutum, mesoscutellum, and mesopleuron coarsely areolate-punctate; propleuron with large shallow punctures; metapleuron smooth; propodeum areolate. Legs with dense whitish setae. Metasoma blackish-brown with distinct purple lustre. T1–2 densely punctured, sparser on disk of T2. T1–3 and S2–5 with apical fringes of whitish setae; T4–6 with setae black, with few scattered erect whitish setae on discs laterally. Wings fusco-violaceous.</p> <p>Remarks. The male of this species was recognized only after the sex association of Kurzenkotilla harmandi, comb. nov. (see below) was discovered. The newly recognized females of K. harmandi initially appeared to be conspecific with K. annamensis, but closer examination revealed that they represented different species, particularly due to the differences in the metasomal setal pattern and the constricted mesonotum of K. harmandi. The male here recognized as K. annamensis occurs in southern Vietnam, where the type and other examined female of K. annamensis have been examined, but K. harmandi have not been seen. Furthermore, this male has different metasomal setal markings (as discussed in the key above) than K. harmandi. Additionally, it has the mesoscutum clearly wider than long, while the male of K. harmandi has the mesoscutum only scarcely wider than long; likewise, females of K. annamensis have a wider mesonotum than that of K. harmandi.</p> </div>	https://treatment.plazi.org/id/583A87D7A33BFFD4FF29FE32CB88D323	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A33AFFD5FF29FF7FC9B7D58B.text	583A87D7A33AFFD5FF29FF7FC9B7D58B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kurzenkotilla ariana (Lelej 1980)	<div><p>Kurzenkotilla ariana (Lelej in Lelej &amp; Kabakov, 1980)</p> <p>(Figs 5–6)</p> <p>Artiotilla ariana Lelej in Lelej &amp; Kabakov, 1980: 191, ♀, holotype, ♀, Afghanistan, western border of Farah province, 70 km south of Nazar-Khan, Shand, 1.XI.1972, O. Kabakov leg. [ZIN], examined.</p> <p>Material examined. Holotype only.</p> <p>Distribution. Afghanistan.</p> <p>Remarks. This rare species has not yet been found in the Oriental Region and is the westernmost known species in the genus Kurzenkotilla.</p> </div>	https://treatment.plazi.org/id/583A87D7A33AFFD5FF29FF7FC9B7D58B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A339FFD6FF29FF7FCC61D392.text	583A87D7A339FFD6FF29FF7FCC61D392.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kurzenkotilla harmandi (Andre 1898) Lelej & Williams & Terine & Okayasu & Parikh & Kumar 2023	<div><p>Kurzenkotilla harmandi (André, 1898), comb. nov.</p> <p>(Figs 14–20)</p> <p>Mutilla harmandi André, 1898: 29, ♁, holotype, ♁, Lakhon [Laos / Thailand], [1878], J. Harmand leg. [MNHN]; 1902: 39, ♁.</p> <p>Material examined. LAOS: Sayaboury, Sayaboury, 6.XII.1965, 1♀ [BPBM]; Vientiane, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.460556&amp;materialsCitation.latitude=18.0175" title="Search Plazi for locations around (long 102.460556/lat 18.0175)">Tat Mun</a> waterfall, el. 183 m, 18°01′03″N 102°27′38″E, 7.VI.2013, G.R. Ballmer leg., 1♀ [UCRC, UCRCENT#428259]. THAILAND: Chaiyaphum: Tat Tone N.P., Staff house at Takfah waterfall, 15°56.461′N 102°5.955′E, 242 m, malaise trap, T. Jaruphan &amp; O. Budsawong leg., 5–12.IX.2006, 1♁, [QSBG]; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.011&amp;materialsCitation.latitude=16.001" title="Search Plazi for locations around (long 102.011/lat 16.001)">Tha Hin Ngom Village</a>, 16.001°N 102.011°E, 560 m, malaise trap, K.A. Williams et al. leg., 18–28.III.2019, 1♁, [CSCA, DNA voucher 19E658]; Chiang Mai, Omkoi District, dry dipterocarp forest, 17°50′49.9″N 98°22′33.0″E, 950–1010 m, R. Mizuno leg.: 19.XII.2016, 1♀, [THNHM]; 20.VI.2017, 1♀ [THNHM]; 27.VI.2017, 1♀ [THNHM]; 7.VII.2017, 1♀, [THNHM, DNA voucher 19E961]; 19.VII.2019, 1♀ [THNHM]; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=102.574234&amp;materialsCitation.latitude=16.62295" title="Search Plazi for locations around (long 102.574234/lat 16.62295)">Khonkaen</a>, Nam Phong N.P., office, 16°37.377′N 102°34.454′E, 344 m, K. Jaidee leg., malaise trap, 19–26.VII.2006, 4♁, [CSCA, EMUS, QSBG]; Loei: Phu Kradeung N.P., mixed deciduous forest N of NA <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.794334&amp;materialsCitation.latitude=16.802834" title="Search Plazi for locations around (long 101.794334/lat 16.802834)">Noy Forest Unit</a>, 16°48.17′N 101°47.66′E, 276 m, malaise trap, S. Gong-Iasae leg., 14–20.XI.2006, 2♁, [QSBG]; Nakhon Ratchasima, Kog Nong Bun Nak-Rat, 28.VII.2009, V. Bezborodov leg., 1♀, [IBSS].</p> <p>Description. FEMALE. Body length 11–13 mm. Coloration. Head, mesosoma, and S1 orange-brown; legs, antenna, and metasoma blackish. Body setae generally sparse and silvery, except frons, vertex, and mesosomal dorsum with sub-erect golden and blackish setae; T1, T2 disc, and T4–5 setae dense black dorsally; T1–6 laterally, S1–6, T2 disc lateral spots, T2 fringe, T3 entirely, and T6 with dense pale golden setae; distance between T2 lateral spots 0.6 × spot diameter. Head. Width behind eye subequal to pronotal width. Frons, vertex, and gena punctures coarsely confluent. Mandible apex apparently tridentate, middle tooth blunt and confluent with lower tooth. Clypeus basomesal longitudinal carina and mesally notched transverse pre-apical carina. Antennal scrobe with thick dorsal carina. Antennal tubercle dorsally smooth, anteriorly micropunctate. Genal and postgenal carinae distinct, forming blunt tooth where they connect with hypostomal carina. F1 2.9 × pedicel length, F2 2.2 × pedicel length. Mesosoma. Length 1.15 × width (measured at posterolateral propodeal corners). Dorsum of mesosoma with coarse confluent punctures; wavy carina separating dorsal and lateral faces of mesonotum. Side of mesosoma with micropunctures and short setae. Mesopleural lamella a short thick carina. Humeral carina distinct. No trace of scutellar scale. Lateral and posterior propodeal faces separated by row of distinct teeth. Metasoma. T1 anterior face with separated punctures, posteriorly with dense setae obscuring sculpture. T2 disc and S2 with dense coarse subconfluent punctures; S2 obscurely concave anterolaterally, anteromesally with longitudinal ridge. T3–5 and S3–5 with dense setae obscuring sculpture. S1 with thick longitudinal lamella, notched sub-posteriorly. T2 felt line deeply linear, 0.7 × T2 total length. T6 convex without pygidial plate, smooth. S6 posterior margin distinctly bituberculate. MALE. See André (1898).</p> <p>Distribution. Laos, Thailand.</p> <p>Remarks. This species was previously known from males only and was treated as a member of the genus Mutilla, until the female was discovered in this paper. The sex association is initially based on identical ITS1 sequences of a male (GenBank # OP854799) from Chaiyaphum, Thailand, and a female (GenBank # OP854800) from Chiang Mai, Thailand. In previous studies on Mutillidae sex associations using ITS (Pilgrim &amp; Pitts 2006; Pilgrim et al., 2008; Williams et al. 2012), intraspecific sequence comparisons generally revealed genetic distances below 2%, while interspecific sequence comparisons generally revealed genetic distances above 2%. The identical ITS1 sequences, coupled with similar body size, coloration, and overlapping distribution are adequate evidence to recognize the previously unrecognized female of Mutilla harmandi. This female was not previously named and is clearly a member of the genus Kurzenkotilla, permitting us to recognize a new combination for K. harmandi, and to recognize the male sex for the genus.</p> <p>This species was named in honor of Dr. François-Jules Harmand (1845–1921). Dr. J. Harmand explored in Indo- China region between 1875 and 1878 (Geiser &amp; Nagel 2013, Huang et al. 2015) and collected insects at a mountain of La-khôn (Harmand, 1880). Some of his collections have a label with the same data “LAKHON, HARMAND 1878” and were deposited in MNHN and SDEI. According to Harmand’s (1880) map, there was a province named La-khôn. Meanwhile, a place named La-khôn was in Lakhôn province and located on a bank of the Mekong [= “Mèkhông” in Harmand (1880)] (Huang et al. 2015: Fig. 18). Firstly, “Montagnes de Lakhon” lies in the vicinity of Tha Khaek, Khammuane Province, Laos (Geiser &amp; Nagel, 2013) and La-khôn was in the vicinity of Mekong (Harmand, 1880) which is the boundary of modern Laos and Thailand (Huang et al. 2015).</p></div> 	https://treatment.plazi.org/id/583A87D7A339FFD6FF29FF7FCC61D392	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A337FFD8FF29FF7FCA55D3EB.text	583A87D7A337FFD8FF29FF7FCA55D3EB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kurzenkotilla cicatricifera (Andre 1894) Lelej & Williams & Terine & Okayasu & Parikh & Kumar 2023	<div><p>Kurzenkotilla cicatricifera (André, 1894), comb. nov.</p> <p>(Figs 21–28)</p> <p>Mutilla cicatricifera André, 1894: 466, 477, ♀, holotype, type locality: “Kanara” [India: Karnataka], Wroughton leg. [MNHN]; André 1902: 38, ♀, India; photographs examined</p> <p>Diagnosis. FEMALE. Head and mesosoma ferruginous, rugoso-punctate, mandible apically and basally black, middle portion ferruginous, clypeus with tuft of suberect golden setae. T1 apically with fringe of golden setae at postero-lateral edges, T2 basally with two conjugended spots of golden setae, not reaching lateral edge of T2, T2 laterally with long felt line of golden setae, T2 apically with fringe of golden setae, T3 with band of golden setae, T6 with tuft of erect long golden setae around pygidial plate, metasoma laterally with long erect golden setae, S1 to S6 with fringe of golden setae. MALE. Unknown.</p> <p>Material examined. Holotype. INDIA, Karnataka, Kodagu district, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=75.88763&amp;materialsCitation.latitude=11.900077" title="Search Plazi for locations around (long 75.88763/lat 11.900077)">Bettathur</a>, 12.417675°N, 75.666364°E, 958 m, 23.XII.2019, P. Girish Kumar &amp; Party, ♀ ZSI/WGRC/IR/INV.21260 [ZSIK]. INDIA, Kerala, Kannur district, Kottiyoor, 11.900077°N, 75.887628°E, 882 m, 12.IV.2022, Siva K. leg., ♀ ZSI/WGRC/IR/INV.21261 [ZSIK].</p> <p>Description. FEMALE. Body length 18.1–18.9 mm. Body stout, large-sized, moderately convex, slightly broadest at metasoma. Head oval, somewhat flattened, rugoso-punctate, posterior margin curved, genal carina well developed; eyes small, oval; longitudinal eye diameter in fronto-lateral view, 1.86 × distance between eyes. Clypeus straight, with medial deep emargination; mandible pointed towards the apex, bidentate; lower half of gena bordered beneath by a carina; hypostomal carina well developed. Mesosoma rectangular, with slightly concave laterally; rugoso-punctate as in head, lateral margin crenulated, propodeum posteriorly abrupt; propleuron rugoso-punctate as in head; metapleuron and allied lateral propodeal face smooth; mid and hind tibia with two rows of spines, one row with four and other one with six spines. T1 wide, slightly narrower than T2; T2 with long lateral felt line, 0.75 × T2; pygidium rugose with some small punctures.</p> <p>Coloration and setation. Antennae, head, clypeus, legs and mesosoma ferruginous with lateral margin darker than mesosoma dorsal; mandible apically and basally black, middle portion ferruginous; tibial spine dark brown; vertex with sparse suberect dark brown setae, towards lateral and frons with golden setae; clypeus with tuft of suberect golden setae; scape with golden setae and flagellomeres with short golden setae; mesosoma dorsally with dark brown setae throughout, apically towards propodeum with long erect golden setae; legs with long golden setae throughout, tarsomeres with dense dark brown setae; metasoma black, T1 apically with golden fringe at posterolateral edges, T2 basally with two conjoined spots of golden setae, not reaching lateral edge of T2, T2 laterally with long felt line of golden setae, T2 apically with fringe of golden setae, T3 with band of golden setae, T6 with tuft of erect long golden setae around pygidial plate, metasoma laterally with long erect golden setae, S1–6 with fringe of golden setae.</p> <p>Measurements. Ratio of pedicel length, F1 length, F2 length, and F3 length, 0.22:0.49:0.24:0.29; longitudinal eye diameter in fronto-lateral view: 1.18 mm, mesosoma maximal width in dorsal view: 3.51 mm, mesosoma length in dorsal view: 4.19 mm.</p> <p>MALE (unknown).</p> <p>Distribution. India (Karnataka, Kerala).</p> <p>Remarks. The female from Kerala differs by having slightly lighter golden setae, the genal carina not developed, the mesosoma dorsally with irregular dark patches, and the mesopleuron darker. Other than K. visrara, whose apparently erroneous type locality is Sri Lanka (see below), this is the southwesternmost representative of the genus Kurzenkotilla. Based on its southern distribution and broadly transverse band on T2, we initially thought it may actually belong in the genus Storozhenkotilla. Furthermore, this seems to be the only Kurzenkotilla species with a sculptured apparent pygidial plate. The species seems to fit more closely with Kurzenkotilla, however, based on the moderately large head and eye, the weakly defined genal carina, and the black T1 fringe. Furthermore, the transverse setal band on T2 more closely resembles two coalescent spots, due to the black triangular patch of setae basomesally. This band is also more centrally located on the T2 disc than that seen in Storozhenkotilla, in K. cicatricifera, the T2 band extends posteriorly to the midpoint of T2, while the band is restricted to the basal quarter of T 2 in the genus Storozhenkotilla.</p> </div>	https://treatment.plazi.org/id/583A87D7A337FFD8FF29FF7FCA55D3EB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FF7FCAB4D610.text	583A87D7A334FFDBFF29FF7FCAB4D610.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kurzenkotilla niveosignata (Andre 1894)	<div><p>Kurzenkotilla niveosignata (André, 1894)</p> <p>(Figs 36–37)</p> <p>Mutilla niveosignata André, 1894: 465, 480, ♀, syntypes: Poona (Wroughton leg.) and Orissa (J. Taylor leg.) [India: Maharashtra, Odisha], [MNHN]; Dalla Torre 1897: 66, ♀; Bingham 1897: 4, 13, ♀; André 1902: 40, ♀.</p> <p>Kurzenkotilla niveosignata: Lelej 2005: 35, ♀; Nithya &amp; Dey 2013: 289, ♀; Terine et al. 2020a: 3; Terine et al. 2020b: 446; Pagliano et al. 2020: 225.</p> <p>Material examined. PAKISTAN: Baluchistan, near Deragazichan village, Indus River valley, 26.III.2003, S. Ovchinnikov leg., 1 ♀ [IBSS]; Punjab, Lahore, 7.VII.1946, 1♀ [UMMZ]. INDIA: Rajasthan, Bharatpur, 22.X.1995, R. Carradori leg., 1♀ [MSNF]</p> <p>Distribution. * Pakistan, India (Bihar, Delhi, Haryana, Madhya Pradesh, Maharashtra, Odisha, Rajasthan *).</p> <p>Remarks. The male of this species might eventually be recognized to be K. semiviolacea, comb. nov., based on their co-occurrence in both India and Pakistan.</p> </div>	https://treatment.plazi.org/id/583A87D7A334FFDBFF29FF7FCAB4D610	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FD29CD5CD162.text	583A87D7A334FFDBFF29FD29CD5CD162.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kurzenkotilla rufodorsata (Cameron 1897) Lelej & Williams & Terine & Okayasu & Parikh & Kumar 2023	<div><p>Kurzenkotilla rufodorsata (Cameron, 1897), comb. nov.</p> <p>(Figs 29–31)</p> <p>Mutilla rufodorsata Cameron, 1897: 65, ♁, syntypes, “Agra” [India: Uttar Pradesh], Rothney leg. [OUMNH]; Bingham 1897: 10, 51, ♁ (India: North-West Provinces); André 1902: 71, ♁, India; Lelej 2005: 36; Terine et al. 2020a: 3; Pagliano et al. 2020: 230.</p> <p>Material examined. INDIA: Rajasthan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=78.07806&amp;materialsCitation.latitude=27.916666" title="Search Plazi for locations around (long 78.07806/lat 27.916666)">Jodphur</a>, 26°13.42′N 73°01.96′E, 190m, malaise in garden, 11–16.VIII.2008, M.E. Irwin &amp; D.R. Priyadarsanan leg., 1♁ [EMUS]; New Delhi, IARI, 28°37′51″N 77°9′50″E, 230 m, 3.VII.2005, J. Heraty leg., 1♁ [UCRC, UCRCENT#434275]; Uttar Pradesh, Aligarth Muslim University, 27°55′00″N 78°4′41″E, 190 m, 28.VI.2005, J. Heraty leg., 1♁ [UCRC, UCRCENT#441172]. NEPAL: Jumla district, Sinja Khola, 1900– 2000 m, W. Schwaller leg., 10.VI.1998, 1♁ [IBSS]; Bagmati, 16.VII.1974, S. Yamaguchi &amp; T. Aoki leg., 1♁ [NARO].</p> <p>Distribution. * Nepal, India (Delhi, Rajasthan, Uttar Pradesh).</p> <p>Remarks. In both coloration and structure, this species is similar to K. semiviolacea, comb. nov. Prior to this paper, the species was included in the genus Mutilla (Lelej 2005), but its similarity to the males of K. annamensis and K. harmandi, and its occurrence in northern India and Nepal suggest that it belongs in Kurzenkotilla.</p> </div>	https://treatment.plazi.org/id/583A87D7A334FFDBFF29FD29CD5CD162	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A334FFDBFF29FABBCAF5D335.text	583A87D7A334FFDBFF29FABBCAF5D335.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kurzenkotilla scrobiculata (Hammer 1962)	<div><p>Kurzenkotilla scrobiculata (Hammer, 1962)</p> <p>(Figs 40–41)</p> <p>Mutilla scrobiculata Hammer, 1962: 4, ♀, holotype, ♀, Bijrani, Nainital Dist [rict], W[est] Himalayas [India, Uttarakhand], [NZC].</p> <p>Kurzenkotilla scrobiculata: Lelej 2005: 35, ♀; Nithya &amp; Dey 2013: 289, ♀; Terine et al. 2020a: 3; Pagliano et al. 2020: 225.</p> <p>Material examined. NEPAL: Jumla District: lower Sinja Khola, 1700–1900 m, 9.VI.1998, G. Miksch leg., 2♀ [IBSS]; Mahakali / Darchula Ghamna, Chamliya Khola, 29º41′25″N 80º41′04″E, 30.V. 2005, 830 m, at light, A. Weigel leg., 1♀ [IBSS]; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.68445&amp;materialsCitation.latitude=29.690277" title="Search Plazi for locations around (long 80.68445/lat 29.690277)">Unknown district</a>: Nepal, I. Yoneta leg., 1♀ [SEHU].. INDIA: Uttarakhand, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=79.558334&amp;materialsCitation.latitude=30.625" title="Search Plazi for locations around (long 79.558334/lat 30.625)">Gobind Ghat Village</a>, 30º37.5′N 79º33.5′E, 1750–1900 m, 17–23.V.1999, Yu. Marusik leg., 2♀ [IBSS]; Dehra Dun, 28.IX.1944, 1 ♀ [UMMZ].</p> <p>Distribution. * Nepal, India (Uttarakhand, Himachal Pradesh).</p> <p>Remarks. The male of this species might eventually be recognized to be K. rufodorsata, comb. nov., based on their co-occurrence in both India and Nepal.</p> </div>	https://treatment.plazi.org/id/583A87D7A334FFDBFF29FABBCAF5D335	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FF7FCCACD627.text	583A87D7A333FFDCFF29FF7FCCACD627.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kurzenkotilla semiviolacea (Andre 1896) Lelej & Williams & Terine & Okayasu & Parikh & Kumar 2023	<div><p>Kurzenkotilla semiviolacea (André, 1896), comb. nov.</p> <p>(Figs 32–35)</p> <p>Mutilla semiviolacea André 1896: 17, ♁, type locality “Himalaya” [India], [HNHM]; André 1902: 41; Ramakrishna 1916: 541.</p> <p>Storozhenkotilla semiviolacea: Lelej et al. 2007: 56, ♁; Terine et al. 2020a: 3; Pagliano et al. 2020: 236.</p> <p>Material examined. INDIA: Agra, 2.III.1958, 1♁ [NSMT]; PAKISTAN: Margalla Hills National Park, 20.VI.2005, Malaise trap, H. Khurram &amp; M. Fida, 1♁ [IBSS].</p> <p>Distribution. Pakistan, India (Himalaya).</p> <p>Remarks. In both color and structure, this species is similar to K. rufodorsata, comb. nov. Based on similarity to S. aurofasciata, Lelej et al. (2007) transferred the species to Storozhenkotilla. The species, however, lives outside the range for other species in Storozhenkotilla and more closely resembles K. harmandi and K. annamensis in coloration, and clypeal and pygidial morphology, so we here transfer the species to Kurzenkotilla.</p> </div>	https://treatment.plazi.org/id/583A87D7A333FFDCFF29FF7FCCACD627	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FD66CD2BD03D.text	583A87D7A333FFDCFF29FD66CD2BD03D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kurzenkotilla visrara (Cameron 1898)	<div><p>Kurzenkotilla visrara (Cameron, 1898)</p> <p>(Figs 38–39)</p> <p>Mutilla visrara Cameron, 1898: 1, ♀, syntypes, ♀, Trincomali, Ceylon, R.A. Yerbury leg. [Yerburgh] coll. [Sri Lanka], [OUMNH]; André 1902: 41, ♀.</p> <p>Kurzenkotilla visrara: Lelej 2005: 35, ♀; Pagliano et al. 2020: 225.</p> <p>Material examined. INDIA: Rajasthan: Bharatpur, VIII.1985, A. Riedel leg., 1♀ [IBSS]; Jaipur, 21.IV.1945, 1♀ [UMMZ]; Chandigarh: Sukhna Lake, 15.X.2007, Sk. Yamane leg., 1♀ [IBSS].</p> <p>Distribution. * India (Rajasthan, Chandigarh), Sri Lanka.</p> <p>Remarks. Although AL examined large volumes of mutillid material from Sri Lanka, the female of Mutilla visrara was not discovered there. Possibly, the type locality of this species is wrong. The late Børge Petersen, who studied most of the types of Cameron’s Oriental species, wrote AL (Petersen, in litt., 1980), that several of Cameron’s mutillid species described from Sri Lanka are likely based on material from outside of Sri Lanka.</p> </div>	https://treatment.plazi.org/id/583A87D7A333FFDCFF29FD66CD2BD03D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDCFF29FB4CCB88D2FF.text	583A87D7A333FFDCFF29FB4CCB88D2FF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mutilla Linnaeus 1758	<div><p>Genus Mutilla Linnaeus, 1758</p> <p>Type species: Mutilla europaea Linnaeus,1758 by subsequent designation (Latreille 1810).</p> <p>Diagnosis. FEMALE. In the Oriental and eastern Palaearctic regions, this genus is unique in having the mesosomal sides parallel, T1 broad, and the T2 disc with uniformly black setae. Other diagnostic features include: mandible unarmed ventrally, head usually wider than mesosoma, mesosomal width equal to or narrower than pronotum in dorsal view, and T6 lacking defined pygidial plate. MALE. In the Oriental region, males can be separated from other Mutillinae genera by having the mandible unarmed ventrally, the clypeus with a medial projection, the tegula elongate, the mesoscutellum unarmed, T1 broad, T7 tuberculate or carinate, and the paramere with a ventral brush of long setae.</p> <p>Remarks. Seventy species are currently placed in the genus Mutilla, mostly from the Afrotropical and Palaearctic regions (Bischoff 1920, Lelej 2002, Pagliano et al. 2020). The eastern Palaearctic Mutilla mikado Cameron, 1900 penetrates into Oriental areas of China. Other Oriental species that were previously placed in Mutilla (Lelej 2005) are transferred into the genera Kurzenkotilla or Storozhenkotilla in this paper.</p> </div>	https://treatment.plazi.org/id/583A87D7A333FFDCFF29FB4CCB88D2FF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A333FFDDFF29F90DCC15D679.text	583A87D7A333FFDDFF29F90DCC15D679.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mutilla mikado Cameron 1900	<div><p>Mutilla mikado Cameron, 1900</p> <p>(Figs 1–4)</p> <p>Mutilla mikado Cameron, 1900; 75, ♀, type locality [“ Japan ”]</p> <p>Mutilla rugiceps maesta Chen, 1957: 151, ♀, holotype, ♀, Kirin [Heilongjiang], Kaolintze [Gaolinzi], 16.vi.1940 [Heude Museum, Shanghai (now Shanghai Entomological Museum, Chinese Academy of Sciences)]. Junior subjective synonym of M. mikado according to Lelej 1985: 161.</p> <p>Mutilla europaea: Su et al. 2019: 104, parasite of Bombus breviceps (China: Yunnan). Misidentification.</p> <p>Material examined. RUSSIA: * Kuril Islands, SW Kunashir, Ivanovskij Cape, 28– 29.09.2013, Yu. &amp; L. Sundukov, 2♀ [IBSS]. CHINA: * Sichuan, 35 km W Zhangle, 3620–3720 m, 12–13.VII.2001, S. Murzin, 1♀ [IBSS]; * Shaanxi, S of Huaxian, SE Damingzhen, Huashan Mts., 2010–2155 m, 24.V.2009, I. Belousov &amp; I. Kabak, 1♀ [IBSS].</p> <p>Distribution. Russia: Eastern Siberia (south), Far East (Amurskaya oblast, Khabarovskii krai, Primorskii krai, Sakhalin, *Kuril Islands (Kunashir)); China (Gansu, Heilongjiang, Jilin, Neimenggu, Shanxi, * Shaanxi, * Sichuan, Zhejiang, Yunnan), Japan (Hokkaido, Honshu, Kyushu, Shikoku), North Korea, South Korea.</p> <p>Remarks. Unlike the other species treated here, Mutilla mikado is predominantly a Palaearctic species and only occurs in a few Oriental areas of China. In some older papers (e. g., Chen 1957), this species was treated as a subspecies of Mutilla europaea Linnaeus, 1758. More recently, Mutilla europaea was recorded as a parasite of Bombus breviceps Smith, 1852, in Yunnan, China, but the mutillid in question was later determined to be Mutilla mikado, which is the only Mutilla species known to occur in Oriental regions of China.</p> </div>	https://treatment.plazi.org/id/583A87D7A333FFDDFF29F90DCC15D679	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFDFFF29FF7FCD43D7BD.text	583A87D7A330FFDFFF29FF7FCD43D7BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Standfussidia Lelej 2005	<div><p>Genus Standfussidia Lelej, 2005</p> <p>Standfussidia Lelej, 2005: 107, 197 (description), ♀; Brothers &amp; Lelej 2017: 96; Pagliano et al. 2020: 236.</p> <p>Type species: Standfussidia taprobane Lelej, 2005, ♀, by original designation.</p> <p>Diagnosis. MALE. Unknown.</p> <p>FEMALE. Head width equal to mesosomal maximum width. Eyes small, rounded convex, but not hemispherical, closer to mandibular base than posterior head border. F1 1.25 × longer its maximum width, relation of pedicel and F1–3 0.5:0.6:0.6:0.6. Scape curved. Mandible tridentate. Frons lacking carina between tubercles and without carina between tubercle and inner eye border. Maxillary palps shorter than antenna length. Mesosoma length equals its maximum width, mesoscutellum narrowed posterad, propodeum distinctly widened posterad with abrupt posterior slope. Mesosoma dorsally without any traces of sutures, mesopleuron distinctly convex with vertical row of long setae, scutellar scale lacking, posterior dorsal border not dentate nor serrate. Legs not shortened, fore tarsus with pectinate comb, mid and hind tibia with two rows of strong spines. Metasoma distinctly flattened dorso-ventrally, segment 2 very long, segment 1 very wide not constricted posterad. T2 with rather short lateral felt line; S2 without any lateral felt line. T6 with narrow pygidial plate, carinated laterally, with granulated and striate sculpture. Body setae usual, not elongated.</p> <p>Species included. Standfussidia taprobane Lelej, 2005.</p> <p>Distribution. Oriental region: Sri Lanka.</p> <p>Remarks. This genus differs from all other Oriental Mutillini by the coloration and pygidial structure. The well-defined triangular pygidial plate, and a single basomesal setal spot on T2 and a fore tarsal comb are more similar to members of the tribe Smicromyrmini. The wide T1 shape, however, is diagnostic for the Mutillini. The eventual discovery of males for this genus could necessitate a change in its tribal placement in the future.</p> </div>	https://treatment.plazi.org/id/583A87D7A330FFDFFF29FF7FCD43D7BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFDFFF29FBC7CD5FD18E.text	583A87D7A330FFDFFF29FBC7CD5FD18E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Standfussidia taprobane Lelej 2005	<div><p>Standfussidia taprobane Lelej, 2005</p> <p>(Figs 42–43)</p> <p>Standfussidia taprobane Lelej, 2005: 198, ♀,</p> <p>holotype, ♀, Sri Lanka: Monaragala District: Angunakolapelessa, 27–28.III.1981, K.V. Krombein, T. Wijesinhe, L. Weeratunge leg. [USNM], examined; Pagliano et al. 2020: 236, Fig. 362, ♀.</p> <p>Material examined. Paratypes: SRI LANKA, 2♀ with the same label as holotype; Anuradhapura District: Padaviya, irrigation bungalow, 18–19.V.1976, K. Krombein et al. leg., 1♀; Monaragala District: Mau Ara, 16 km E of Uda Walawe, 100 m, 24–26.IX.1977, K. Krombein et al. leg., 1♀; Angunakolapelessa, 30.IX–1.X.1977, K. Krombein et al. leg., 1♀; Hambantota District: Palatupana tank, 10–16 m, 29.III–2.IV.1981, K. Krombein et al. leg., 1♀ [USNM, IBSS].</p> <p>Distribution: Sri Lanka.</p> <p>Remarks. The females of Standfussidia taprobane have been collected in the dry zone of Sri Lanka.</p> </div>	https://treatment.plazi.org/id/583A87D7A330FFDFFF29FBC7CD5FD18E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A330FFC0FF29F9A8CB0AD775.text	583A87D7A330FFC0FF29F9A8CB0AD775.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Storozhenkotilla Lelej 2005	<div><p>Genus Storozhenkotilla Lelej, 2005</p> <p>Storozhenkotilla Lelej, 2005: 37, 176 (description), ♀, ♁; Lelej et al. 2007: 56; Brothers &amp; Lelej 2017: 96; Pagliano et al. 2020: 236.</p> <p>Type species: Mutilla aurofasciata André, 1907 by original designation.</p> <p>Diagnosis. MALE. Head width less than pronotal width, not rounded posterad; preoccipital carina dorsally well developed, sharp and slightly projecting over head. Eyes weakly notched inside. Ocelli small. Mandible bidentate without subbasal tooth beneath and without subbasal inner tooth. F2 1.3 × F1. Prementum not tuberculate. Notauli and parapsids visible. Tegula longer than pronotal length laterad, projecting over axilla. Pterostigma large closed. Mesoscutellum non-modified. Propodeal posterior slope with strong longitudinal medial carina. Metasomal segment 1 wide transverse. T2 with long lateral felt line, S2 without any lateral felt line. S8 (hypopygium) not modified, T7 without carina. Gonostylus wide, basiventrally setose. Volsella shorter than penial valve and consists of wide cuspis and short process-like digitus; penial valves symmetrical, apically setose. T1 dorsally and T2–3(4) posterad with band of pale setae.</p> <p>FEMALE. Head large, widened posterad, width slightly less than pronotal width; behind eye elongated; eye small. Genal carina well developed, hypostomal carina widened medially. Prementum not tuberculate. F1 1.5 × F2. Mandible bidentate, without subbasal inner tooth. Mesosoma with concave mesopleuron, scutellar scale lacking. Mesopleuron with well developed vertical (supracoxal) carina. Promesonotal suture carinated in type species. Fore tarsus without comb of strong spines. Mid and hind tibia with two rows of spines. Metasomal segment 1 wide transverse; T2 with lateral felt line. T2 basally with pale band which medially emarginated posterad and apical narrow pale band; T3 with wide pale band. T6 with distinct pygidial plate weakly carinated lateroapically, shiny or shagreened with a few rugae. S6 with two strong apical tubercles.</p> <p>Species included. Storozhenkotilla aurofasciata (André, 1907), ♁ ♀; S. binghami (Lelej, 2005), ♀, comb. nov.; and S. nathani Lelej, sp. nov., ♁.</p> <p>Distribution. Oriental region (South India, Sri Lanka).</p> <p>Remarks. Storozhenkotilla is most similar to Mutilla Linnaeus, 1758. The male differs by having the gonostylus weakly setose (densely setose in Mutilla and many Kurzenkotilla), and by having a lateral clypeal projection (medial projection in Mutilla). In having setal markings on the basal part of T2, females of Storozhenkotilla Lelej resemble those of Kurzenkotilla Lelej but easily differ by having setal markings forming a transverse pale band basally on T2 (with two pale setal spots disposed transversely in Kurzenkotilla).</p> </div>	https://treatment.plazi.org/id/583A87D7A330FFC0FF29F9A8CB0AD775	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32FFFC0FF29FC8ECB72D2C7.text	583A87D7A32FFFC0FF29FC8ECB72D2C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Storozhenkotilla aurofasciata (Andre 1907)	<div><p>Storozhenkotilla aurofasciata (André, 1907)</p> <p>(Figs 44–54)</p> <p>Mutilla aurofasciata André, 1907: 255, ♀, holotype, ♀, Pankulam / Ceylon [Sri Lanka] / W. Horn / Holotype / Mutilla aurofasciata ♀ nov. sp. [SDEI], examined.</p> <p>Storozhenkotilla aurofasciata: Lelej 2005: 37, Figs 99–102, ♁, 176 (description of ♁), India: Karnataka, Tamil Nadu; Terine et al. 2020a: 3; Pagliano et al. 2020: 236.</p> <p>Material examined. SRI LANKA: Mannar District: 0.8 km NE Kokmotte Bungalow, Wilpattu National Park, 21–25.V.1976, K.V. Krombein, P.B. Karunaratne, S. Karunaratne, D.W. Balasooriya leg. / Mutilla aurofasciata André, B. Petersen det. 1980, 1♀ [IBSS]. INDIA: Karnataka, Bangalore, 916 m, 29.X.1978, K.D. Ghorpade leg., A719 / Mutilla aurofasciata André, B. Petersen det. 1980, 1♀ [IBSS]; Tamil Nadu: Kodaikanal, 6500 ft [1981 m], Pulney Hills [Palni Hills], IV–V.1953, P.S. Nathan, K.V. Krombein collection, 1975–1976, 1♁ [USNM]; Madras, Coimbatore, 1400 ft., VIII.1963, P. Susai Nathan, 1♀ [FSCA]; Thirunelveli district, Kodumundiyar dam site, 26.IX.2018, P. Girish Kumar, 1♁, ZSI/WGRC/IR/INV.21298 [ZSIK]; Pondicherry: Auroville, 10 km N Pondicherry [Puducherry], 24.X.2004, F. Burger, 1♁; Auroville, 10 km N Pondicherry, Canians, 25.XI.2005, F. Burger, 1 ♀; Auroville, 10 km N Pondicherry, 7.I.2007, F. Burger, 1♀; same place, 12.II.2007, F. Burger, 1♀; same place, 20.II.2007, F. Burger, 1♀; same place, 5.III.2007, F. Burger, 1♀; same place, 13.III.2007, F. Burger, 1♀; same place, 23.III.2007, F. Burger, 1♀ [IBSS].</p> <p>Distribution. Sri Lanka, South India (Karnataka, * Pondicherry, Tamil Nadu).</p> <p>Remarks. This is the type species for the genus Storozhenkotilla and the females are easily separated from all other Oriental Mutillini species by the transverse suture between the pronotum and mesonotum. Males are less obviously distinct from other Mutillini, but they do seem to be the only Oriental Mutillini with T7 uniformly punctate and the metasoma without any metallic luster.</p> </div>	https://treatment.plazi.org/id/583A87D7A32FFFC0FF29FC8ECB72D2C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32FFFC1FF29F911C993D6E5.text	583A87D7A32FFFC1FF29F911C993D6E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Storozhenkotilla binghami (Lelej 2005) Lelej & Williams & Terine & Okayasu & Parikh & Kumar 2023	<div><p>Storozhenkotilla binghami (Lelej, 2005), comb. nov.</p> <p>(Figs 55–58)</p> <p>Mutilla binghami Lelej, 2005: 35. Replacement name for Mutilla valida Cameron, 1897; Terine et al. 2020a: 3; Pagliano et al. 2020: 227.</p> <p>Mutilla valida Cameron, 1897: 71, ♀ (nom. praeocc., non Smith, 1855), type locality: ”Barrackpore” [India: West Bengal], Rothney leg. [OUMNH]; Bingham 1897: 4, 14, ♀; André, 1902: 71, ♀, India.</p> <p>Material examined. SRI LANKA: Mannar District: 0.8 km NE Kokmotte Bungalow, Wilpattu National Park, 21– 25.V.1976, K.V. Krombein, P.B. Karunaratne, S. Karunaratne, D.W. Balasooriya, 2♀ [USNM]. INDIA: Tamil Nadu, Tiruchirappali District, Pudukkottai: XII.1991, T.R.S. Nathan, 1♀ [SEMC]; XI.1992, T.R.S. Nathan, 1♀ [SEMC]; XI.2002, T.R.S. Nathan, 1♀ [FSCA]. INDIA: Karnataka, 15 km N Bangalore [Bengaluru], KT, 23–24.VII.1996, K. Werner &amp; L. Lorenz, 5♀ [MSNF].</p> <p>Distribution. * Sri Lanka, India (Karnataka, Tamil Nadu, West Bengal).</p> <p>Remarks. The type of this species, from West Bengal, is the only specimen of Storozhenkotilla known from northern India. As suggested in the remarks for K. visrara above, there are doubts about the veracity of type localities for many of Cameron’s species. It seems possible that the type could be mislabeled and the genus Storozhenkotilla truly is restricted to a southern Indian and Sri Lankan distribution. Previous studies (Lelej 2005) considered K. cicatricifera comb. nov. as a member of this genus that could be separated from S. binghami by minor color and sculpture differences (see couplet 5 in key to females above). The form that was previously considered S. cicatricifera (Figs 57, 58) is here treated as a color variant of S. binghami, but further studies could eventually reveal it as a discrete species.</p> </div>	https://treatment.plazi.org/id/583A87D7A32FFFC1FF29F911C993D6E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
583A87D7A32EFFC3FF29F91BC960D5B1.text	583A87D7A32EFFC3FF29F91BC960D5B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Storozhenkotilla nathani Lelej & Williams & Terine & Okayasu & Parikh & Kumar 2023	<div><p>Storozhenkotilla nathani Lelej, sp. nov.</p> <p>(Figs 59–64)</p> <p>Diagnosis. Body antennae and legs black with bluish metallic luster on head, pronotum and metallic blue metasoma. Propodeal posterior slope with strong longitudinal medial carina which ends dorsally by tooth. T7 laterally angulate, apical border smooth with wide medial smooth triangle without carina. T1–6 and S2–5 with apical fringe of whitish setae.</p> <p>Material examined. Holotype, ♁, INDIA: Karnataka, Mysore [Mysuru], Shimoga dist[rict], Agumbe Ghat, 600 m, V.2004, TRSN coll. [IBSS].</p> <p>Description. FEMALE (unknown).</p> <p>MALE. Head width less than pronotal width, not rounded posterad. Preoccipital carina dorsally well developed, sharp and slightly projecting over head. Eyes well notched inside. Ocelli small, ratio POD:OOD=0.7. Mandible bidentate without basal tooth beneath and without subbasal tooth on inner border. F2 1.3 × F1. Prementum not tuberculate. Notauli well visible in posterior half, parapsids weak. Tegula longer than pronotal length laterad, projecting over axilla. Propodeal posterior slope with strong longitudinal medial carina which ends dorsally by tooth. Metasomal segment 1 wide transverse. T2 with long lateral felt line, S2 without any lateral felt line. Sternum 8 (hypopygium) not modified. T7 laterally angulate, apical border smooth with wide medial smooth triangle without carina. Gonostylus wide, basiventrally setose. Volsella slightly shorter than penial valve and consists of wide cuspis and short finger-like digitus; penial valves symmetrical, posterior margin with vertical row of setae, postero-ventrally bidentate, pre-apical tooth bluntly rounded.</p> <p>Head black with bluish metallic luster. Antennae entirely black, scape and pedicel with dense whitish setae. Head densely punctured, sparsely around ocelli, with dense whitish setae all over. Mesosoma black, pronotum with bluish metallic luster, with sparse whitish setae denser on mesopleuron, mesoscutum with sparse black setae. Tegula sparsely punctured, laterally and posteriorly smooth. Pronotum, mesoscutum and mesoscutellum coarsely punctured; propleuron, mesopleuron and lateral propodeal face coarsely punctured, metapleuron smooth; propodeum posterad with sharp slope. Legs with dense whitish setae. Metasoma metallic blue. T1–2 densely punctured, sparser on disk of T2. T1–6 and S2–5 with apical fringe of whitish setae. The wings fusco-violaceous, basal radial abscissa sharply oblique, apically curved; first transverse radio-medial vein oblique, second almost straight, third sharply angled in the middle; first recurrent vein is received in basal third, second at apical third of cell.</p> <p>Distribution. India (Karnataka).</p> <p>Etymology. The specific name is dedicated to P. Susai Nathan (1891–1976), Indian naturalist, entomologist, natural history specimens collector, who collected the Mutillidae in South India (Tamil Nadu).</p> <p>Remarks. This species may be the opposite sex of S. binghami (Lelej, 2005), which was also collected in Karnataka.</p> </div>	https://treatment.plazi.org/id/583A87D7A32EFFC3FF29F91BC960D5B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Lelej, Arkady S.;Williams, Kevin A.;Terine, Joshua B.;Okayasu, Juriya;Parikh, Grishma R.;Kumar, Girish P.	Lelej, Arkady S., Williams, Kevin A., Terine, Joshua B., Okayasu, Juriya, Parikh, Grishma R., Kumar, Girish P. (2023): Review of the tribe Mutillini (Hymenoptera: Mutillidae) from the Oriental Region. Zootaxa 5228 (4): 455-476, DOI: https://doi.org/10.11646/zootaxa.5228.4.5
