taxonID	type	description	language	source
1F070A1EC6723C5AA1DCFF2E9271FE65.taxon	description	Figs. 2 A, 4 A, 5 A – C, 7 A, 9	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6723C5AA1DCFF2E9271FE65.taxon	materials_examined	Type material. HOLOTYPE, male, in NZAC, labeled: \ W. Lewis Pass, 2000 ’. 23.3. 65 N. A. Walker \ DSIR \ No \ NZ PB \. PARATYPES (47 specimens, in NZAC, dissected 10 exx.), 2 males and 5 females labeled same as holotype; 2 males and 2 females labeled: \ Lewis Pass 762 m Canterbury \ 19 Mar 65 G. W. Ramsay J. I. Townsend \ moss 65 / 283 \ DSIR \ No \ NZ PB \; 1 male labeled: \ Lewis Pass Canterbury 23 Mar 65 Litter 65 / 108 \ Litter 65 / 108 \ DSIR \ No \ NZ PB \; 2 males and 1 female labeled: \ Leafmould \ Lewis Pass 18 – 24.11.61 G. Kuschel \ DSIR \ No \ NZ PB \; 1 male and 3 females labeled: \ Maruia Saddle 1900 ’ 2 - 6 - 65 moss J. I. Townsend \ DSIR \ No \ NZ PB \; 1 female labeled: \ E. S. Brookes leaf mould Sept. 1948 \ Maruia Springs Lewis Pass Westland \ A. E. Brookes Collection \ Teneral \ NZ PB \; 1 male and 2 females labeled: \ Howard Val BR 7 Oct 68 S. Edridge \ moss 68 / 143 \ DSIR \ No \ NZ PB \; 2 females labeled: \ Col. T. Hall 27 - 3 - 1915 \ Howard Nelson South Is. \ T. Broun Collection \ A. E. Brookes Collection \ DSIR \ No \ NZ PB \; 3 females labeled: \ L. Rotoiti 2000 ’ 27.7.65 Grey Duck Bay moss under Mt. beech \ DSIR \ No \ NZ PB \; 2 females labeled: \ L. Rotoiti Nelson leaf litter 21 - 3 - 65 N. A. Walker \ DSIR \ No \ NZ PB \; 1 female labeled: \ L. Rotoiti NN 23 Dec 64 J. I. Townsend \ Litter 64 / 49 \ DSIR \ No \ NZ PB \; 1 male and 1 female labeled: \ Mt. Hope Nelson 3.3.1915 \ Tachys N. Sp. \ T. Broun Collection \ A. E. Brookes Collection \ DSIR \ No \ NZ PB \; 1 male labeled: \ Hope 10.12.16 \ not. 2677 \ T. Broun Collection \ A. E. Brookes Collection \ DSIR \ No \ NZ PB \; 1 female labeled: \ var. 2677 \ Hope 10.12.16 \ T. Broun Collection \ A. E. Brookes Collection \ DSIR \ No \ NZ PB \; 1 male labeled: \ Glenhope 10 - 12 - 14 \ not. 2677 \ T. Broun Collection \ A. E. Brookes Collection \ DSIR \ No \ NZ PB \; 1 male and 1 female labeled: \ Glenhope 10 - 12 - 14 \ var. 2677 \ T. Broun Collection \ A. E. Brookes Collection \ DSIR \ No \ NZ PB \; 1 female labeled: \ Coll. T. Hall 27.3.1915 \ Glenhope Nelson 10.12.1914 \ 2677 var. \ T. Broun Collection \ A. E. Brookes Collection \ NZ PB \; 1 male labeled: \ Lamb Val. 1400 ’ Glenhope 19.7.65 moss under open silver and black beech J. I. Townsend \ DSIR \ No \ NZ PB \; 5 females labeled: \ Kaka — Glenhope Rd. 28.7.64 J. I. Townsend \ NZ PB \; 1 male labeled: \ nr. ft. Hope Saddle 1800 ’ 19.7.65 mixed moss J. I. Townsend \ DSIR \ No \ NZ PB \; 1 female labeled: \ New Zealand BR Nelson Lakes NP Speargrass Tck Mt Robert \ On mossy log 30 - Nov- 05 \.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6723C5AA1DCFF2E9271FE65.taxon	etymology	Etymology. The specific epithet is a Latinized eponym in the genitive case, and is based on the surname of Captain Thomas Broun, a New Zealand entomologist of the 19 th century in recognition of his significant contributions to the study of New Zealand coleopterans, including publications of descriptions of New Zealand anilline beetles.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6723C5AA1DCFF2E9271FE65.taxon	materials_examined	Type locality. New Zealand, South Island, Canterbury, Southern Alps, Lewis Pass. Recognition. Adults of this species (Fig. 2 A) are practically indistinguishable from the adults of many Nesamblyops species based on external characters (cf. with Figs. 2 B – C) and are distinguished from them by the structure of the male genitalia.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6723C5AA1DCFF2E9271FE65.taxon	description	Description. Large for genus (SBL range 1.76 – 2.02 mm, mean 1.89 ± 0.078 mm, n = 17). Habitus. Body form (Fig. 2 A) markedly convex, elongate ovoid, general proportions wide (WE / SBL 0.41 ± 0.011), head narrow relative to pronotum (WH / WPm 0.68 ± 0.016), proportions of pronotum in comparison to elytra average for genus (WPm / WE 0.75 ± 0.017). Color. Body color brunneorufous, appendages testaceous. Prothorax. Pronotum (Fig. 4 A) moderately long in comparison to elytra (LP / LE 0.42 ± 0.013) and moderately transverse (WPm / LP 1.23 ± 0.026), with lateral margins arcuately constricted posteriorly (WPm / WPp 1.30 ± 0.025). Anterior angles indistinct, posterior angles obtuse (108 – 116 °), rounded. Width between posterior angles greater than between anterior angles (WPa / WPp 0.87 ± 0.024). Basal margin almost rectilinear, slightly convex at middle. Elytra. Ovoid, narrowly depressed along suture, comparatively long (LE / SBL 0.59 ± 0.008) and moderately wide (WE / LE 0.69 ± 0.020). Humeri completely rounded. Lateral margins slightly divergent at basal half, subparallel at middle and evenly rounded to apex in apical third. Male genitalia. Median lobe of aedeagus (Fig. 5 C) moderately arcuate and moderately twisted. Shaft subparallel basally, tapering in apical half. Apex moderately curved dorsally with narrowly rounded tip. Apical orifice long, occupies almost half of the shaft length. Ventral margin of median lobe straight. Walls of shaft without poriferous canals. Dorsal copulatory sclerites in form of characteristic “ bird’s head ” plate, surrounded by distinct scaled membraneous field. Spines of internal sac absent. Left paramere (Fig. 5 A) comparatively wide, with shortly attenuate apex, bearing three long setae. Right paramere (Fig. 5 B) of moderate length, bearing three long setae, which are slightly longer the length of paramere. Ring sclerite as in Fig. 7 A. Female internal genitalia. Not examined. Geographical distribution. This species is distributed from the Lewis Pass of North Canterbury along the Spenser Mountains northward up to the Hope Range in the Nelson area (Crosby et al. 1998) (Fig. 9, green circles). So, the range of this species includes both sides of the Buller River valley. Habitat. Specimens were collected from leaf litter, leaf mould, moss in Nothofagus forests (labels mention silver N. menziesii, black N. solandri var. solandri, and mountain N. solandri var. cliffortioides beeches). Relationships. The structure of the male genitalia of N. brouni suggests its relatedness to the other species having dorsal copulatory sclerite in a shape of “ bird’s head ”, such as N. tararua, N. distinctus, and N. townsendi. Based on the details of the sclerite configuration and the state of the scaled membraneous field it is postulated that N. brouni is a sister taxon to N. distinctus.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6753C5BA1DCFDFF91EFFCCD.taxon	description	Figs. 2 B, 4 B, 5 D – F, 7 B, 10	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6753C5BA1DCFDFF91EFFCCD.taxon	materials_examined	Type material. HOLOTYPE, male, in NZAC, labeled: \ Wooded Pk. East Summit Dun Mountain Nelson \ 14 Sept 71 G. W. Ramsay \ Litter 71 / 116 \ DSIR \ No \ NZ PB \. PARATYPES (146 specimens, in NZAC, dissected 9 exx.), 81 males and females labeled same as holotype; 1 male and 3 females labeled: \ Wooded Pk. Dun Track Sdle Nelson 14 Sept. 71 \ G. Ramsay \ litter 71 / 109 \ DSIR \ No \ NZ PB \; 1 female labeled: \ Dun Mt. 2000 ’ 28.11.43 E. S. Gourlay \ E. S. Gourlay Acc. 1970 Ent. Div. \ NZ PB \; 2 males and 2 females labeled: \ Dun Mt. 2500 ’ 13.11.61 Nelson G. Kuschel \ 61 / 1 \ DSIR \ No \ NZ PB \; 1 female labeled: \ Leafmould \ Dun Mt. 2500 ’ 13.11.61 G. Kuschel \ NZ PB \; 1 male and 2 females labeled: \ Dun Track Nelson 14 Sept 71 \ G. Ramsay \ Litter 71 / 112 \ NZ PB \; 1 male labeled: \ Just beyond 3 rd. House Dun Mt. Nelson J. I. Townsend \ DSIR \ No \ NZ PB \; 1 female labeled: \ Pelorus Bridge 100 ’ 15.10. 65 L. P. Marchaut \ moss in bush 65 / 512 \ NZ PB \; 1 female labeled: \ Pelorus Val. 29 - 4 - 64 litter J. I. Townsend \ DSIR \ No \ NZ PB \; 4 males and 2 females labeled: \ Upper Maitai, 7 - 3 - 49 E. S. Gourlay \ E. S. Gourlay Acc. 1970 Ent. Div. \ NZ PB \; 1 female labeled: \ Upper Maitai, (c.) 19 - 5 - 41 E. S. Gourlay \ E. S. Gourlay Acc. 1970 Ent. Div. \ Nesamblyops oreobius Broun E. B. Britton det. 1960 \ NZ PB \; 1 female labeled: \ Upper Maitai, 13 - 2 - 57 E. S. Gourlay \ NZ PB \; 16 males and females labeled: \ Upper Maitai 3.5.50 E. S. Gourlay \ Carabidae \; 3 males and 3 females labeled: \ Maungatapu Sdle. 2400 ’ \ Nel. 15.11.67 J. I. Townsend \ moss sample 67 / 83 \ DSIR \ No \ NZ PB \; 1 female labeled: \ Maungatapu 3000 ’ 17.4.38 E. S. Gourlay \ E. S. Gourlay Acc. 1970 Ent. Div. \; 4 males and 2 females labeled: \ NEW ZEALAND NN Whangamoa Sdle 700 m Slaters Rd 4 May 1997 J S Dugsdale N menziesii N truncata kamahi litter \ NZ PB \; 5 males and 2 females labeled: \ Nelson, N. Z. Whangamoa Saddle 14.9.74 J. I. Townsend leafmould \ Coll. J. I. Townsend \; 2 females labeled: \ Titirangi Marlborough 2 Oct 69 F. Alack \ Litter 69 / 174 \ NZ PB \; 1 male and 2 females labeled: \ Titirangi Marlb. 22. x. 69 F. A. Alack \ Litter 69 / 174 \ DSIR \ No \ NZ PB \. Additional material: \ Wairoa Gorge Nelson 5 - 4 - 60 J. I. Townsend \ NZ PB \ (4 females).	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6753C5BA1DCFDFF91EFFCCD.taxon	etymology	Etymology. The specific epithet is a Latinized eponym in the genitive case and is based on the surname of Christopher E. Carlton, Professor and former Director of Louisiana State Arthropod Museum, Baton Rouge, Louisiana, United States, who triggered my interests in research of the members of the tribe Anillini in the USA as well as in New Zealand.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6753C5BA1DCFDFF91EFFCCD.taxon	materials_examined	Type locality. New Zealand, South Island, Nelson, Richmond Range, Dun Mountain. Recognition. Adults of this species (Fig. 2 B) are virtually similar to the adults of many Nesamblyops species (cf. with Figs. 2 A, 3 B), however can be easily distinguished from them by the shape of posterior angle of pronotum, formed by tiny blunt denticle, and the structure of the male genitalia.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6753C5BA1DCFDFF91EFFCCD.taxon	description	Description. Large for genus (SBL range 1.75 – 2.03 mm, mean 1.89 ± 0.078 mm, n = 29). Habitus. Body form (Fig. 2 B) markedly convex, elongate ovoid, general proportions wide (WE / SBL 0.40 ± 0.010), head narrow relative to pronotum (WH / WPm 0.67 ± 0.016), proportions of pronotum in comparison to elytra average for genus (WPm / WE 0.76 ± 0.019). Color. Body color brunneopiceous, appendages testaceous. Prothorax. Pronotum (Fig. 4 B) moderately long in comparison to elytra (LP / LE 0.43 ± 0.011) and moderately transverse (WPm / LP 1.23 ± 0.026), with lateral margins arcuately constricted posteriorly (WPm / WPp 1.26 ± 0.022). Anterior angles indistinct, posterior angles obtuse (106 – 115 °), in form of protruding blunt denticles with faint sinuation anteriorly. Width between posterior angles greater than between anterior angles (WPa / WPp 0.85 ± 0.032). Basal margin almost rectilinear, slightly convex at middle. Elytra. Ovoid, narrowly depressed along suture, comparatively long (LE / SBL 0.58 ± 0.007) and moderately wide (WE / LE 0.69 ± 0.020). Humeri completely rounded. Lateral margins slightly divergent at basal half, subparallel at middle and evenly rounded to apex in apical third. Male genitalia. Median lobe of aedeagus (Fig. 5 F) moderately arcuate and moderately twisted. Shaft subparallel, tapering in apical third. Apex slightly curved dorsally with narrowly rounded tip. Apical orifice of moderate length, reaches one third of the shaft length. Ventral margin of median lobe straight. Walls of shaft with numerous poriferous canals. Dorsal copulatory sclerites in form of characteristic “ bird’s head ” plate, with apically attenuated dorsal part. Scaled membraneous fields lacking. Spines of internal sac absent. Left paramere (Fig. 5 D) comparatively narrow, with shortly attenuate apex, bearing three long setae. Right paramere (Fig. 5 E) elongated, bearing three long setae, which are slightly shorter the length of paramere. Ring sclerite as in Fig. 7 B. Female internal genitalia. Not examined. Geographical distribution. The range of this species stretches along the eastern coast of Tasman Bay from the Wairoa George in the south to Titirangi Bay of the Marlborough Sound region (Crosby et al. 1998) to the north (Fig. 10, black circles). Habitat. Specimens were collected from leaf litter, leaf mold, moss in bush and Nothofagus forests (labels mention kamahi shrub Weinmannia racemosa, silver N. menziesii, and hard N. truncata beeches). Relationships. Based on the structure of the male genitalia N. carltoni occupies an isolated position among examined species of the genus.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6743C58A1DCFC17920EFD89.taxon	description	Figs. 2 C, 4 C, 5 G – I, 10	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6743C58A1DCFC17920EFD89.taxon	materials_examined	Type material. HOLOTYPE, male, in NZAC, dissected, labeled: \ Mt Stokes 1160 m Marlborough 12 Oct 67 \ J. I. Townsend \ Litter 67 / 244 \ DSIR \ No \ NZ PB \. PARATYPES (2 specimens, in NZAC, dissected 1 male), 1 female labeled same as holotype; 1 male labeled: \ Mt Stokes Marlb. 11. x. 67 J. I. Townsend \ moss 67 / 247 \ DSIR \ Ns \ NZ PB \.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6743C58A1DCFC17920EFD89.taxon	etymology	Etymology. The specific epithet is a Latin adjective, confusus, in the masculine form, meaning “ having been mixed ”, and refers to the similarity between the external appearance of the new species and its closest relatives.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6743C58A1DCFC17920EFD89.taxon	materials_examined	Type locality. New Zealand, South Island, Marlborough Sounds, Mount Stokes. Recognition. Adults of this species (Fig. 2 C) are practically indistinguishable from the adults of many Nesamblyops species based on external characters (cf. with Figs. 2 A – B, 2 F) and are distinguished from them by the structure of the male genitalia.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6743C58A1DCFC17920EFD89.taxon	description	Description. Average size for genus (SBL range 1.64 – 1.67 mm, mean 1.66 ± 0.017 mm, n = 3). Habitus. Body form (Fig. 2 C) markedly convex, elongate ovoid, general proportions wide (WE / SBL 0.42 ± 0.012), head narrow relative to pronotum (WH / WPm 0.68 ± 0.038), in comparison with other species pronotum less bulky in comparison to elytra (WPm / WE 0.72 ± 0.027). Color. Body color dark brunneous, appendages testaceous. Prothorax. Pronotum (Fig. 4 C) moderately long in comparison to elytra (LP / LE 0.42 ± 0.024) and moderately transverse (WPm / LP 1.24 ± 0.013), with lateral margins arcuately and strongly constricted posteriorly (WPm / WPp 1.35 ± 0.026). Anterior angles indistinct, posterior angles strongly obtuse (123 – 127 °), widely rounded. Width between posterior angles slightly greater than between anterior angles (WPa / WPp 0.93 ± 0.054). Basal margin slightly convex. Elytra. Ovoid, narrowly depressed along suture, comparatively long (LE / SBL 0.59 ± 0.015) and moderately wide (WE / LE 0.71 ± 0.034). Humeri completely rounded. Lateral margins slightly divergent at basal third, subparallel at middle and evenly rounded to apex in apical third. Male genitalia. Median lobe of aedeagus (Fig. 5 I) moderately slightly arcuate and moderately twisted. Shaft arcuate, tapering in apical half. Apex slightly enlarged and straight with rounded tip. Apical orifice long, occupies almost half of the shaft length. Ventral margin of median lobe slightly concave in apical part. Walls of shaft with few poriferous canals apically. Dorsal copulatory sclerites with distinct V-shaped structure, directed with its sharp end apically. Scaled membraneous fields lacking. Spines of internal sac absent. Left paramere (Fig. 5 G) comparatively wide, with shortly attenuate apex, bearing three long setae. Right paramere (Fig. 5 H) short and wide, bearing three long setae, which are slightly longer the length of paramere. Female internal genitalia. Not examined. Geographical distribution. This species is known only from Mount Stokes in the Marlborough Sounds area (Crosby et al. 1998) (Fig. 10, green circle). Habitat. According to label data s pecimens were collected from moss without more precise data about plant communities. Relationships. The structure of the male genitalia suggests the relationship of N. confusus to other species having a similar V-shaped structure of dorsal copulatory sclerites in the internal sac of the median lobe, such as N. oreobius (Broun) and N. lescheni.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6773C59A1DCFDD39267F869.taxon	description	Figs. 2 D, 4 D, 5 J – L, 7 C, 9	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6773C59A1DCFDD39267F869.taxon	materials_examined	Type material. HOLOTYPE, male, in NZAC, dissected, labeled: \ Head of Fabians Valley 920 m Marlborough 23 Oct 63 \ J. I. Townsend \ Litter 63 / 28 \ DSIR \ No \ NZ PB \. PARATYPES (3 specimens, in NZAC, dissected 1 male), 1 male and 2 females labeled same as holotype. Additional material: \ Mt. Riley 2400 ’ Blenheim Dist. 24. vii. 66 A. K. Walker \ litter from dry ridge under rata, beech and cedar trees 66 / 217 \ DSIR \ No \ (1 female).	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6773C59A1DCFDD39267F869.taxon	etymology	Etymology. The specific epithet is a Latin adjective, distinctus, in the masculine form, meaning “ distinguished ”, and refers to the distinctive shape of the sclerites in the internal sac of male genitalia, allowing to distinguish this species from his closest relatives.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6773C59A1DCFDD39267F869.taxon	materials_examined	Type locality. New Zealand, South Island, Marlborough, Richmond Range, Fabians Valley. Recognition. Adults of this species (Fig. 2 D) can be distinguished from the adults of majority species of Nesamblyops by the distinctive shape of pronotum (Fig. 4 D), however are practically indistinguishable from the adults of some Nesamblyops species (cf. with Figs. 3 C) and are distinguished from all of them by the structure of the male genitalia.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6773C59A1DCFDD39267F869.taxon	description	Description. Large for genus (SBL range 1.93 – 2.26 mm, mean 2.06 ± 0.130 mm, n = 5). Habitus. Body form (Fig. 2 D) markedly convex, elongate ovoid, general proportions wide (WE / SBL 0.42 ± 0.011), head narrow relative to pronotum (WH / WPm 0.66 ± 0.008, proportions of pronotum in comparison to elytra average for genus (WPm / WE 0.76 ± 0.011). Color. Body color brunneopiceous, appendages testaceous. Prothorax. Pronotum (Fig. 4 D) moderately long in comparison to elytra (LP / LE 0.43 ± 0.022) and moderately transverse (WPm / LP 1.28 ± 0.055), with lateral margins almost rectilinear and slightly constricted posteriorly (WPm / WPp 1.23 ± 0.016). Anterior angles indistinct, posterior angles obtuse (105 – 115 °), dented, and slightly protruding backwards. Width between posterior angles greater than between anterior angles (WPa / WPp 0.80 ± 0.030). Basal margin slightly concave. Elytra. Ovoid, narrowly depressed along suture, comparatively long (LE / SBL 0.58 ± 0.008) and wide (WE / LE 0.72 ± 0.026). Humeri completely rounded. Lateral margins slightly divergent at basal half, shortly subparallel at middle and evenly rounded to apex in apical half. Male genitalia. Median lobe of aedeagus (Fig. 5 L) moderately arcuate and moderately twisted. Shaft subparallel basally, tapering in apical half. Apex moderately curved dorsally with narrowly narrowly rounded tip. Apical orifice long, occupies almost half of the shaft length. Ventral margin of median lobe straight. Walls of shaft without poriferous canals. Dorsal copulatory sclerites in form of characteristic “ bird’s head ” plate with apically attenuated dorsal part, and surrounded by distinct scaled membraneous field. Spines of internal sac absent. Left paramere (Fig. 5 J) comparatively wide, with shortly attenuate apex, bearing three long setae. Right paramere (Fig. 5 K) of moderate length, bearing three long setae, which are slightly shorter the length of paramere. Ring sclerite as in Fig. 7 C. Female internal genitalia. Not examined. Geographical distribution. This species is known from two localities on the Richmond Range in the Marlborough area (Crosby et al. 1998) (Fig. 9, red circles). Habitat. Specimens were collected from litter “ under rata, beech and cedar trees ”. Relationships. Based on the structure of the male genitalia N. distinctus forms a natural group with other species having a similar dorsal copulatory sclerite in the internal sac of the median lobe, such as N. brouni, N. tararua, and N. townsendi. Based on the details of the sclerite configuration and the state of the scaled membraneous field, it is postulated that N. distinctus is a sister taxon to N. brouni.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6793C57A1DCFBD69153FD61.taxon	description	Figs. 2 E, 4 E, 5 M – O, 7 E, 8 B, 10	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6793C57A1DCFBD69153FD61.taxon	materials_examined	Type material. HOLOTYPE, male, in NZAC, labeled: \ E. Attempt Hill, D’Urville Isd. 28 - 9 - 63 Litter J. I. Townsend \ J. I. Townsend Collection \ Nesamblyops oreobius (Broun) Det. I. Townsend \. PARATYPES (13 specimens, in NZAC, dissected 1 male and 1 female), 1 male and 11 females labeled same as holotype; 1 female labeled: \ Coll. A. C. O’Connor, October, 1942 \ D’Urville Is. Cook Strait, N. Z. \ A. E. Brookes Collection \ DSIR \ No \ NZ PB \.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6793C57A1DCFBD69153FD61.taxon	etymology	Etymology. The specific epithet is a Latinized eponym in the genitive case and is based on the surname of Richard A. B. Leschen, New Zealand coleopterist (New Zealand Arthropod Collection, Maanaki Whenua — Landcare Research, Auckland, New Zealand), who made material of New Zealand Anillini available for this study.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6793C57A1DCFBD69153FD61.taxon	materials_examined	Type locality. New Zealand, South Island, Marlborough Sounds, D’Urville Island. Recognition. Adults of this species (Fig. 2 E) can be distinguished from adults of many Nesamblyops species (cf. with Figs. 2 A – C, 3 A – C) by their smaller size and characteristic body proportions, as well by the structure of the male genitalia.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6793C57A1DCFBD69153FD61.taxon	description	Description. Of moderate size for genus (SBL range 1.43 – 1.68 mm, mean 1.54 ± 0.091 mm, n = 12). Habitus. Body form (Fig. 2 E) markedly convex, elongate ovoid, general proportions wide (WE / SBL 0.42 ± 0.013), head narrow relative to pronotum (WH / WPm 0.69 ± 0.013), proportions of pronotum in comparison to elytra average for genus (WPm / WE 0.77 ± 0.034). Color. Body color brunneorufous, appendages testaceous. Prothorax. Pronotum (Fig. 4 E) moderately long in comparison to elytra (LP / LE 0.43 ± 0.024) and moderately transverse (WPm / LP 1.28 ± 0.037), with lateral margins arcuately constricted posteriorly (WPm / WPp 1.29 ± 0.027). Anterior angles indistinct, posterior angles obtuse (106 – 115 °), but distinct. Width between posterior angles greater than between anterior angles (WPa / WPp 0.89 ± 0.015). Basal margin almost rectilinear, slightly convex at middle. Elytra. Ovoid, narrowly depressed along suture, comparatively long (LE / SBL 0.58 ± 0.012) and wide (WE / LE 0.72 ± 0.021). Humeri completely rounded. Lateral margins slightly divergent at basal third, subparallel at middle and evenly rounded to apex in apical third. Male genitalia. Median lobe of aedeagus (Fig. 5 O) slightly arcuate and moderately twisted. Shaft subparallel, tapering in apical third. Apex short and semicircular. Apical orifice short, occupies about one third of the shaft length. Ventral margin of median lobe widely concave. Walls of shaft with numerous poriferous canals. Dorsal copulatory sclerites with distinct V-shaped structure, directed with its sharp end apically. Scaled membraneous fields lacking. Spines of internal sac absent. Left paramere (Fig. 5 M) comparatively wide, with shortly attenuate apex, bearing three long setae. Right paramere (Fig. 5 N) short and wide, bearing three long setae, which are slightly longer the length of paramere. Ring sclerite as in Fig. 7 E. Female internal genitalia. Spermatheca (Fig. 8 B) sclerotized, of bulb-like shape with wider rounded apical part and narrower rounded basal part. Length of spermatheca shorter than the lengths of either spermathecal gland or spermathecal duct. Spermathecal duct short without coils. Attachments of spermathecal duct and gland to spermatheca close together. Geographical distribution. This species is known only from the D’Urville Island in the Marlborough Sounds area (Crosby et al. 1998) (Fig. 10, red circles). Possibly, N. lescheni is an endemic of this island. Habitat. Labels do not include specific information about habitats. Relationships. The structure of the male genitalia suggests relationship of N. lescheni to other species with similar V-shaped structure of dorsal copulatory sclerites in the internal sac of the median lobe, such as N. confusus and N. oreobius.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6783C52A1DCFCFB91C8FCA9.taxon	description	Figs. 2 F, 4 F, 5 P – R, 7 D, 8 A, 11	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6783C52A1DCFCFB91C8FCA9.taxon	materials_examined	Examined material (more than 280 specimens): WO: Mt. Karioi, nr base, nr summt, 11 Oct 1981, C. F. Butcher, litter 81 / 95 (5 ex.); Tangarakau Gorge, 450 m, 18 - 4 - 65, N. A. Walker, litter (1 ex.); Mt. Pirongia, 962 m, 16 Jan 1977, A. K. Walker, litter 77 / 2 (2 ex.), Mt. Pirongia, 2500 ’, 23 - 9 - 64, litter, G. Kushel (1 ex.); TeKauri Scenic Reserve, nr Kawhia, 20 Sep 1981, C. F. Butcher, litter 81 / 89 (6 ex.); Waitomo, Maori L., Tumutumu Rd., 24 May 1983, J. C. Watt, litter and soil 83 / 60 (5 ex.); Waipuna Reserve near Waitomo, 28 Oct 1977, A. R. Ferguson, litter 77 / 134 (4 ex.); Herangi Range, 500 m, above Mangatoa Saddle, 9 Dec 1982, J. S. Dugsdale, mixed swards and bryophytes 82 / 134 (8 ex.); Mohoenui, Gribbons Road, 26 Jun 1977, N. H. Mancer or G. W. Ramsay, litter 77 / 78 (118 ex.); Mt. Messenger, 610 ’, Taranaki, 24. xi. 68, J. S. Dugsdale, litter 68 / 196 (1 ex.); TK: Pouakai Range, 1006 m, Ahuwakawa Track, 11 Jan 1978, J. C. Watt, litter 78 / 11 (30 ex.), same but, 1067 m, litter and moss 78 / 10 (5 ex.), same but, 1220 m, 9 Jan 1978, litter 78 / 13 (4 ex.); Mt. Egmont, 3500 ’, 2 - 4.1.82, G. Kushel, leafmould (9 ex.); Mt. Egmont, 823 m, Taranaki, 29 Nov 70, A. W. Don, litter (3 ex.); Mt. Egmont, 4600 ’, Stratford side, 25. x. 68, J. S. Dugsdale, mat plants 68 / 198 (2 ex.); Mt. Egmont, 1000 m, Stratford Rd., 22 Feb 1984, B. M. May, litter 84 / 52 (1 ex.); Egmont NP, Ngatoro TK, 650 m, 19. III. 1998, Larochelle, Larivière, wet broadleaf forest along mud flats, in litter & rotten wood on & along big logs (1 ex.); N Egmont, 23 Feb 1984, B. M. May, litter 84 / 51 (1 ex.); 853 m E Egmont, Nov 76, K. E. Fox, moss beside Kamahi Track (5 ex.); Summit track, 4500 ’, Mt. Egmont, 15.6.65, J. I. Townsend, moss sample from wet bank above Hooker Hut (1 ex.); Stratford Plat., 4000 ’, 15 - 6 - 65, moss, J. I. Townsend (5 ex.); Stratford Plt. Rd., Mt. Egmont, 1800 ’, 17 - 4 - 65, litter, N. A. Walker (2 ex.), same but 2150 ’ (3 ex.), same but 2550 ’ (1 ex.), same but 2600 ’ (1 ex.), same but 3250 ’ (1 ex.), same but 3500 ’ (2 ex.); Mt Egmont NP, Stratford Mtn. House, 846 m, 23 May 1986, C. L. Lyal, Leaf litter, CL 474 (2 ex.); Wilkies Pools, track litter, Dawson Falls, 3200 ’, 16 - 6 - 65, J. I. Townsend (2 ex.); Dawson Falls, 975 m, Mt. Egmont, Taranaki, 16 Jun 65, J. I. Townsend, litter 65 / 387 (1 ex.); Pouakai Hump, 1220 m, 3 Dec 1975, A. K. Walker, dry moss 75 / 213 (4 ex.); W edge Hump, Plat. Pouakai Ra., 2 Dec 1975, J. S. Dugsdale, litter 75 / 200 (1 ex.); Pouakai Ra, 1372 m, Summit Ridge, 10 Jan 1978, J. C. Watt, litter 78 / 9 (2 ex.), same but 1400 m, litter 78 / 8 (1 ex.); Pouakai Trig, Pouaki Range, 3 Dec 1975, litter 75 / 206 (4 ex.), same but moss 75 / 208 (12 ex.), same but litter 75 / 209 (3 ex.); Pouakai Saddle, 1220 m, Pouakai Ra., 3 Dec 1975, J. S. Dugsdale, litter 75 / 202 (2 ex.), same but Poa colensoi swards 75 / 203 (10 ex.), same but litter 75 / 204 (9 ex.), same but litter 75 / 205 (6 ex.); Pouakai Ra. Hut, 1280 m, 2 Dec 1975, A. K. Walker, litter 75 / 210 (1 ex.); Tangarakau Gorge, 31 - 12 - 78 (1 ex.). The description of N. oreobius contains little diagnostic information that allows correct identification. Below, I redescribe the species to make comparison of N. oreobius with other species easier. Type locality. New Zealand, North Island, Waikato, Mount Pirongia. Recognition. Adults of this species (Fig. 2 F) are practically indistinguishable from the adults of many Nesamblyops species based on external characters (cf. with Figs. 2 A – C) and are distinguished from them by the structure of the male genitalia.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6783C52A1DCFCFB91C8FCA9.taxon	description	Description. Medium sized for genus (SBL range 1.51 – 1.83 mm, mean 1.68 ± 0.100 mm, n = 23). Habitus. Body form (Fig. 2 F) markedly convex, elongate ovoid, general proportions wide (WE / SBL 0.40 ± 0.008), head narrow relative to pronotum (WH / WPm 0.70 ± 0.014), proportions of pronotum in comparison to elytra average for genus (WPm / WE 0.76 ± 0.027). Color. Body color brunneorufous, appendages testaceous. Prothorax. Pronotum (Fig. 4 F) moderately long in comparison to elytra (LP / LE 0.41 ± 0.016) and moderately transverse (WPm / LP 1.26 ± 0.021), with lateral margins arcuately and strongly constricted posteriorly (WPm / WPp 1.34 ± 0.032). Anterior angles indistinct, posterior angles obtuse (111 – 124 °), rounded. Width between posterior angles greater than between anterior angles (WPa / WPp 0.93 ± 0.029). Basal margin almost rectilinear, slightly convex at middle. Elytra. Ovoid, narrowly depressed along suture, comparatively long (LE / SBL 0.58 ± 0.008) and wide (WE / LE 0.72 ± 0.026). Humeri completely rounded. Lateral margins slightly divergent at basal half, shortly subparallel at middle and evenly rounded to apex in apical half. Male genitalia. Median lobe of aedeagus (Fig. 5 R) strongly arcuate and moderately twisted. Shaft subparallel in basal half, tapering in apical half. Apex slightly curved dorsally with narrowly rounded tip. Apical orifice occupies about one third of the shaft length. Ventral margin of median lobe straight. Walls of shaft with poriferous canals at basal half. Dorsal copulatory sclerites with V-shaped plate, directed with its sharp end apically. Scaled membraneous fields lacking. Spines of internal sac absent. Left paramere (Fig. 5 P) comparatively wide, with shortly attenuate apex, bearing three long setae. Right paramere (Fig. 5 Q) moderately long and wide, bearing three long setae, which are shorter the length of paramere. Ring sclerite as in Fig. 7 D. Female internal genitalia. Spermatheca (Fig. 8 A) sclerotized, of bulb-like shape with wide circular apical part and narrow basal part. Length of spermatheca shorter than the lengths of either spermathecal gland or spermathecal duct. Spermathecal duct short without coils. Attachments of spermathecal duct and gland to spermatheca close together. Geographical distribution. This species is distributed only on the North Island, where it is known from the Taranaki and Waikato areas (Crosby et al. 1998) from Mount Taranaki (former Mount Egmont) in the south, north to Mount Pirongia (Fig. 11). Habitat. According to label data, specimens were collected from leafmold, litter, and moss without more precise data about plant communities. Relationships. The structure of the male genitalia of N. oreobius, especially the V-shaped sclerite of the internal sac suggests a possible relationship to the other species with similar configuration of copulatory sclerites, such as N. confusus and N. lescheni.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC67D3C53A1DCFC339478F9E4.taxon	description	Figs. 3 A, 4 G, 6 A – C, 7 F, 10	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC67D3C53A1DCFC339478F9E4.taxon	materials_examined	Type material. HOLOTYPE, male, in NZAC, labeled: \ Mt Stokes 1172 m Kenepuru Snds Marlb. 11. x. 67 J. McBurney \ moss 67 / 243 \ DSIR \ Ns \ NZ PB \. PARATYPES (24 specimens, in NZAC, dissected 4 exx.), 3 males and 6 females labeled same as holotype; 3 males and 8 females labeled: \ Mt Stokes 1160 m Marlborough 12 Oct 67 \ J. I. Townsend \ Litter 67 / 244 \ DSIR \ Ns \ NZ PB \; 1 male: \ Mt Stokes Marlb. 3850 ’ 12. x. 67 J. I. Townsend \ Litter 67 / 244 \ DSIR \ Ns \ NZ PB \; 1 male and 1 female labeled: \ Mt Stokes 3850 ’ Marlb. 13. x. 67 J. I. Townsend \ Litter 67 / 245 \ DSIR \ Ns \ NZ PB \; 1 male labeled: \ Mt. Stokes 3850 ’ 11. x. 67 \ Kenepuru Sounds J. I. Townsend \ moss sample 67 / 243 \.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC67D3C53A1DCFC339478F9E4.taxon	etymology	Etymology. The specific epithet is a Latin adjective, parvulus, in the masculine form, diminutive of Latin parvus meaning “ small ”, and refers to the small size of the new species.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC67D3C53A1DCFC339478F9E4.taxon	materials_examined	Type locality. New Zealand, South Island, Marlborough Sounds, Mount Stokes. Recognition. Adults of this species (Fig. 3 A) can be distinguished from adults of many Nesamblyops species (cf. with Figs. 2 A – C, 3 B – C) by the small size, elongated habitus, diagnostic shape of pronotum (Fig. 4 G) as well by the structure of the male genitalia.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC67D3C53A1DCFC339478F9E4.taxon	description	Description. Small for genus (SBL range 1.26 – 1.49 mm, mean 1.37 ± 0.081 mm, n = 15). Habitus. Body form (Fig. 3 A) moderately convex, elongate ovoid, general proportions slightly wide (WE / SBL 0.39 ± 0.025), head wide relative to pronotum (WH / WPm 0.75 ± 0.023), proportions of pronotum in comparison to elytra average for genus (WPm / WE 0.74 ± 0.063). Color. Body color rufotestaceous, appendages testaceous. Prothorax. Pronotum (Fig. 4 G) moderately long in comparison to elytra (LP / LE 0.40 ± 0.021) and moderately transverse (WPm / LP 1.23 ± 0.034), with lateral margins arcuately and strongly constricted posteriorly (WPm / WPp 1.37 ± 0.047). Anterior angles indistinct, posterior angles very obtuse (126 – 141 °), almost effaced. Width between anterior angles equals the width between posterior angles (WPa / WPp 1.00 ± 0.047). Basal margin slightly concave. Elytra. Elongate-ovoid, moderately depressed along suture, comparatively long (LE / SBL 0.60 ± 0.013) and slightly narrow for genus (WE / LE 0.66 ± 0.032). Humeri completely rounded. Lateral margins slightly divergent at basal fourth, subparallel at middle and evenly rounded to apex in apical third. Male genitalia. Median lobe of aedeagus (Fig. 6 C) slightly arcuate and slightly twisted. Shaft diverging basally, tapering in apical half. Apex enlarged with straight rounded tip. Apical orifice long, occupies almost half of the shaft length. Ventral margin of median lobe straight basally, moderately enlarged in apical third. Walls of shaft with few poriferous canals apically. Dorsal copulatory sclerites in form of characteristic U-shaped plate, directed with its open end apically. Scaled membraneous fields lacking. Spines of internal sac absent. Left paramere (Fig. 6 A) wide, apex blunt, bearing three setae of moderate length. Right paramere (Fig. 6 B) short and wide, bearing three long setae, which are slightly longer the length of paramere. Ring sclerite as in Fig. 7 F. Female internal genitalia. Not examined. Geographical distribution. This species is known only from Mount Stokes in the Marlborough Sounds area (Crosby et al. 1998) (Fig. 10, yellow circle). Habitat. According to label data s pecimens were collected from moss without more precise data about plant communities. Relationships. Based on the structure of the male genitalia it is postulated that N. parvulus occupies an isolated position among the examined species of the genus. At Mount Stokes N. parvulus lives in sympatry and syntopically with N. confusus: according to the label data at least one sample included both species.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC67C3C4EA1DCF9009503F87A.taxon	description	Figs. 3 B, 4 H, 6 D – F, 7 G, 9	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC67C3C4EA1DCF9009503F87A.taxon	materials_examined	Type material. HOLOTYPE, male, in NZAC, labeled: \ NEW ZEALAND WN Tararua Range Judd Ridge Otaki Forks 800 m \ 5 Nov 1980 C. F. Butcher litter 80 / 101 \ NZ PB \. PARATYPES (129 specimens glued on boards and triangles plus more than 20 specimens in plastic capsules, in NZAC, dissected 16 exx.), 9 males and 6 females labeled same as holotype; 5 males and 4 females labeled: \ NEW ZEALAND WN Tararua Ra Dundas Hut Ridge 950 m 3 Dec 1984 \ R. C. Craw Sifted litter 84 / 88 \ Duplicate specimens in alcohol \ NZ PB \; 1 female labeled: \ NEW ZEALAND WN Tararua Ra Dundas Hut Ridge 1350 m, E side 6 Dec 1984 \ R. C. Craw Sifted litter 84 / 88 \ Duplicate specimens in alcohol \ NZ PB \; 2 females labeled: \ NEW ZEALAND WN Tararua Ra Dundas Hut Ridge 990 m 3 Dec 1984 \ B. G. Bennet & T. K. Crosby Sifted litter 84 / 83 \ NZ PB \; 1 female labeled: \ Tararua Ra Dundas Hut Ridge River Forks 900 m 11 Feb 1985, G. W. Ramsay, litter 85 / 12 \ NZ PB \; 2 females labeled: \ Tararua Ra., Dundas Hut Ridge, 800 m, 13 Feb 1985, G. W. Ramsay, Sifted litter 85 / 16 \ NZ PB \; 2 females labeled: \ Tararua Ra., Dundas Hut Ridge, 950 m, 29 Nov 1984, R. C. Crow, in Blechnum discolour \ NZ PB \; 1 female labeled: \ Tararua Ra., Dundas Hut Ridge, 990 m, 13 Feb 1985 \ C. F. Butcher Litter 85 / 19 \ NZ PB \; 1 male and 1 female labeled: \ 10 km S of Levin, Waikawa, 150 m, 8 Mar 1978 S. B. & J. Peck \ litter \ S. & B. Peck Collection \ NZ PB \; 1 male labeled: \ Nth. Manakau Rd 12.5.91 J. A. & J. I. Townsend \ J. I. Townsend Collection \; 1 male labeled: \ NEW ZEALAND WN Levin Kimberley Reserve 6 Nov 1980 C. F. Butcher \ litter 80 / 103 \ NZ PB \; 2 females labeled: \ NEW ZEALAND WN Mt Holdsworth Start of Interior Track 30 Jan 1985 \ H. P. McColl Sifted litter 2 / 85 \ NZ PB \; 4 males labeled: \ Mt. Holdsworth 2000 ’ Tararuas 19. xi. 68 S. Edridge \ moss 68 / 202 \ DSIR \ No \ NZ PB \; 2 females labeled: \ NEW ZEALAND WN Tararua Range Judd Ridge Otaki Forks, 272 m 5 Nov 1980 \ C. F. Butcher litter 80 / 102 \ NZ PB \; 1 male and 1 female labeled: \ Tararua Ra. 17 Nov 68 S. Edridge \ N. D 3 \ 2000 ’ litter 68 / 200 \ DSIR \ No \ NZ PB \; 1 female labeled: \ NEW ZEALAND WN W. Peak Tararua Ra 1280 m 14 Apr 1974 J. C. Watt \ Litter under Olearia colensoi & Chionochloa 74 / 31 \ NZ PB \; 1 female labeled: \ Nr. Field Hut 3000 ’ 6.9.65 J. I. Townsend moss in forest \; 1 male labeled: \ litter under grass track N. of Wharite 3000 ’ 13.11.89 J. I. Townsend \ J. I. Townsend Collection \; 1 male labeled: \ Waiotauru Rd Akatarawa Range litter 16.11.91 J. Nunn \ J. I. Townsend Collection \; more than 10 specimens in plastic capsule labeled: \ Waiotauru Rd. 22 - 10 - 91 Akatarawa Range litter under silver beech J. I. Townsend \ J. I. Townsend Collection \; more than 10 specimens in plastic capsule labeled: \ Akatarawa Range Waiotauru Rd litter 16.11.91 J. I. Townsend \ Nesamblyops oreobius \ J. I. Townsend Collection \; 1 male labeled: \ leaf litter silver beech forest Waiotauru 830 m 26.4.88 J. Cooksley Tararua Forest Park \ J. I. Townsend Collection \; 1 male labeled: \ NEW ZEALAND WN Tararua Forest Park Waiotauru Rd 20 Jul 1999 E. Spurr \ 7 / S \ NZ PB \; 1 male labeled: \ NEW ZEALAND WN Tararua Forest Park Waiotauru Rd 11 May 1999 E. Spurr \ 10 / W \ NZ PB \; 8 males and 4 females labeled: \ NEW ZEALAND WN Akatarawa Sdle 600 m, 7 Mar 1978 S. B. Peck Litter \ S. & B. Peck Collection \ NZ PB \; 3 males and 8 females labeled: \ NEW ZEALAND WN Akatarawa Sdle Tararua FP Tk 17 Jan 1984 H. P. McColl \ 8 / 84 litter \ Duplicate specimens in alcohol \ NZ PB \; 1 female labeled: \ Akatarawa Range S. of Saddle 1700 ’ Leaf litter 8 - 9 - 85 A. J. Townsend \ Anillini Nesamblyops oreobius (Broun) \ J. I. Townsend Collection \; 1 male labeled: \ NEW ZEALAND WN Kaitoke Waterworks Swingbridge Tk 200 m, 29 Jan 1985 \ C. Welch Nothofagus truncata litter \ NZ PB \; 1 female labeled: \ NEW ZEALAND WN Kaitoke Waterworks Bush Walk, 200 m 29 Jan 1985 \ C. Welch Mixed forest litter \ NZ PB \; 1 male labeled: \ NEW ZEALAND WN Orongorongo V, W. N. 15 Jan 1995 \ Hard beech forest litter Tullgren # 61 \ Nesamblyops oreobius Broun det. A. Larochelle 1995 \; 1 male and 1 female labeled: \ NEW ZEALAND WN Orongorongo V, W. N. 17 Dec 1993 \ Hard beech forest litter Tullgren # 34 \ Nesamblyops oreobius Broun det. A. Larochelle 1995 \ NZ PB \; 1 male labeled: \ NEW ZEALAND WN Orongorongo V, W. N. 17 Dec 1993 \ Hard beech forest litter Tullgren # 35 \ Nesamblyops oreobius Broun det. A. Larochelle 1995 \ NZ PB \; 3 females labeled: \ NEW ZEALAND WN Orongorongo V Stn Ridge 100 m 27 Jan 1992 J. S. Dugdale \ Litter Nothophagus solandri emergence trap \ NZ PB \; 1 female labeled: \ Orongorongo V. Well. 152 m \ 23 Sep 69 J. S. Dugdale \ litter \ DSIR \ No \ NZ PB \; 1 male labeled: \ Orongorongo Res. Stn. Well. \ 21 May 69 J. S. Dugdale \ litter \ DSIR \ No \ NZ PB \; 1 male labeled: \ Orongorongo F. S. Wellington \ 18 Sep 69 J. C. Watt J. S. Dugdale \ litter \ DSIR \ No \; 1 female labeled: \ Orongorongo Res. Sta Wellington 21 May 69 \ J. S. Dugdale \ litter \ DSIR \ No \ NZ PB \; 1 female labeled: \ NEW ZEALAND WN Wainuiomata Whakanui Tr 10 Jan 1993 GR Grehan \ Moss, 700 m beech / kamahi \ NZ PB \; 1 male labeled: \ New Zealand WN Table Top Tararua FP 10.2.85 \ in leatherwood litter \ Nesamblyops oreobius det. J. Nunn \; 4 males and 1 female labeled: \ New Zealand WN 4 km along Waiotauru Rd. 7 - Nov- 06 \ Washed soil sample. Taruarua Rng. \; 1 male labeled: \ New Zealand WN Waiotauru Rd. Tararua FP, 23.4.95 \ in leaf litter \. 7 males labeled: \ Port Ligar 304 m Marlborough 26 Oct 69 F. Alack \ Litter 69 / 175 \ DSIR \ No \ NZ PB \; 14 males and 8 females labeled: \ Port Ligar 1000 ’ Marlb. 26. x. 69 F. A. Alack \ Litter 69 / 175 \ DSIR \ No \ NZ PB \; 1 male labeled: \ Canoe Bay Pelorus Snd. 2 - 9 - 62 J. I. Townsend \ J. I. Townsend Collection \ Nesamblyops oreobius (Broun) Det. I. Townsend 1973 \.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC67C3C4EA1DCF9009503F87A.taxon	etymology	Etymology. The specific epithet is a noun in apposition and refers to the name of the Mountain Range, where the type series was collected.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC67C3C4EA1DCF9009503F87A.taxon	materials_examined	Type locality. New Zealand, North Island, Wellington, Tararua Range. Recognition. Adults of this species (Fig. 3 B) are practically indistinguishable from the adults of many Nesamblyops species based on external characters (cf. with Figs. 2 A – B, 2 D) and are distinguished from them by the structure of the male genitalia.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC67C3C4EA1DCF9009503F87A.taxon	description	Description. Large for genus (SBL range 1.80 – 2.33 mm, mean 2.05 ± 0.162 mm, n = 25). Habitus. Body form (Fig. 3 B) markedly convex, elongate ovoid, general proportions wide (WE / SBL 0.40 ± 0.010), head narrow relative to pronotum (WH / WPm 0.68 ± 0.017), proportions of pronotum in comparison to elytra average for genus (WPm / WE 0.75 ± 0.015). Color. Body color brunneopiceous, appendages brunneorufous. Prothorax. Pronotum (Fig. 4 H) moderately long in comparison to elytra (LP / LE 0.41 ± 0.016) and moderately transverse (WPm / LP 1.21 ± 0.021), with lateral margins rectilinear constricted posteriorly (WPm / WPp 1.27 ± 0.038). Anterior angles indistinct, posterior angles slightly obtuse (98 – 113 °), rounded. Width between posterior angles greater than between anterior angles (WPa / WPp 0.86 ± 0.033). Basal margin almost straight. Elytra. Ovoid, narrowly depressed along suture, comparatively long (LE / SBL 0.60 ± 0.012) and moderately wide (WE / LE 0.67 ± 0.024). Humeri completely rounded. Lateral margins slightly divergent at basal half, shortly subparallel at middle and evenly rounded to apex in apical half. Male genitalia. Median lobe of aedeagus (Fig. 6 F) strongly arcuate and moderately twisted. Shaft subparallel, tapering in apical third. Apex moderately curved dorsally with narrowly rounded tip. Apical orifice occupies almost third of the shaft length. Ventral margin of median lobe almost straight, slightly concave before apex. Walls of shaft with poriferous canals basally. Dorsal copulatory sclerites in form of characteristic “ bird’s head ” plate, surrounded by scaled membraneous field. Spines of internal sac absent. Left paramere (Fig. 6 D) of normal proportions, with moderately attenuate apex, bearing three long setae. Right paramere (Fig. 6 E) long and narrow, bearing three long setae, slightly shorter the length of paramere. Ring sclerite as on Fig. 7 G. Female internal genitalia. Not examined. Geographical distribution. The range of this species splits into two parts. The large northern area of the range occupies the southern parts of the North Island and stretches along the Rimutaka and Tararua Ranges north to Wharite Peak in the Ruahine Range at least, while the small southern part occupies the outer parts of Pelorus Sound to the north of Tennison Inlet in the Marlborough Sounds area (Crosby et al. 1998) of the South Island (Fig. 9, black circles). So, the range of this species includes adjacent territories of both the North and South Islands. Habitat. Specimens were collected from litter and moss in forest, shrub, fern, and grass communities (labels mention silver N. menziesii, mountain N. solandri, and hard N. truncata beeches, leatherwood Olearia colensoi, and kamahi Weinmannia racemosa shrubs, fern Lomaria (= Blechnum) discolor, and tussock grass Chionochloa). Relationships. The structure of the male genitalia of N. tararua suggests relationship to other species having dorsal copulatory sclerite in a shape of a “ bird’s head, ” such as N. brouni, N. distinctus, and N. townsendi. The details of the copulatory sclerite configuration suggest that N. tararua is close to N. brouni. The reduced scaled membraneous field and the presence of an area with poriferous canals at the base of the shaft suggest that N. tararua may share common ancestry with N. townsendi. Significantly, males of N. tararua are the only members of the genus that possess unexpectedly great variation in the size and to a lesser extent in the shape of the median lobe (see Fig. 8). However, among different populations the dorsal sclerite is almost identical, and I was unable to find distinct and stable characters of the median lobe allowing consistent arrangement of this variability geographically. Within-group relationships of N. tararua are unclear.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6603C4AA1DCFA049597FF69.taxon	description	Figs. 3 C, 4 I, 6 G – I, 7 H, 9	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6603C4AA1DCFA049597FF69.taxon	materials_examined	Type material. HOLOTYPE, male, in NZAC, labeled: Tennyson Inlet Marlb. 22.5.64 J. I. Townsend \ Forest litter \ DSIR \ No \ NZ PB \. PARATYPES (12 specimens, in NZAC, dissected 4 exx.), 1 female labeled same as holotype; 1 male labeled: \ Opouri Sdle. Tennyson Inlet Rd. 4. viii. 67 \ F. Alack Litter 67 / 225 \ DSIR \ No \; 1 female labeled: \ Tennyson Inlet 22.5.64 litter J. I. Townsend \ NZ PB \; 2 males and 2 females labeled: \ M Robinson Ridge Kenepuru Sd. 500 m Marlborough \ 13 Mar 1970 J. I. Townsend \ litter 70 / 142 \ DSIR \ No \ NZ PB \; 3 males labeled: \ Ship Cove 365 m. SD 30 Nov. 72 J. S. Dugdale \ Litter 72 / 273 \ DSIR \ No \ NZ PB \; 1 female labeled: \ Shakespeare Bay, Picton \ 11 Aug 69 J. McBurney \ DSIR \ No \; 1 female labeled: \ Marlborough Mt. Robertson 1,600 16 - 5 - 60 I. Mannering \ Leafmould \ NZ PB \.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6603C4AA1DCFA049597FF69.taxon	etymology	Etymology. The specific epithet is a Latinized eponym in the genitive case, and is based on the surname of James Ian Townsend, a famous New Zealand carabidologist and the collector of the type series.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6603C4AA1DCFA049597FF69.taxon	materials_examined	Type locality. New Zealand, South Island, Marlborough Sounds, Tennyson Inlet. Recognition. Adults of this species (Fig. 3 C) can be distinguished from the adults of majority species of Nesamblyops by the distinctive shape of pronotum (Fig. 4 I), however are practically indistinguishable from the adults of some Nesamblyops species (cf. with Figs. 2 D) and are distinguished from all of them by the structure of the male genitalia.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
1F070A1EC6603C4AA1DCFA049597FF69.taxon	description	Description. Large for genus (SBL range 1.85 – 2.05 mm, mean 1.95 ± 0.071 mm, n = 6). Habitus. Body form (Fig. 3 C) markedly convex, elongate ovoid, general proportions wide (WE / SBL 0.41 ± 0.010), head narrow relative to pronotum (WH / WPm 0.66 ± 0.014), proportions of pronotum in comparison to elytra average for genus (WPm / WE 0.77 ± 0.013). Color. Body color brunneopiceous, appendages brunneorufuos. Prothorax. Pronotum (Fig. 4 I) moderately long in comparison to elytra (LP / LE 0.44 ± 0.009) and moderately transverse (WPm / LP 1.22 ± 0.016), with lateral margins rectilinear and slightly constricted posteriorly (WPm / WPp 1.18 ± 0.019). Anterior angles indistinct, posterior angles obtuse (94 – 100 °), dented, and slightly protruding backward. Width between posterior angles much greater than between anterior angles (WPa / WPp 0.77 ± 0.009). Basal margin concave. Elytra. Ovoid, narrowly depressed along suture, comparatively long (LE / SBL 0.59 ± 0.010) and moderately wide (WE / LE 0.69 ± 0.008). Humeri completely rounded. Lateral margins slightly divergent at basal half, shortly subparallel at middle and evenly rounded to apex in apical half. Male genitalia. Median lobe of aedeagus (Fig. 6 I) strongly arcuate and moderately twisted. Shaft diverging basally, tapering in apical half. Apex moderately curved dorsally with narrowly rounded tip. Apical orifice long, occupies almost half of the shaft length. Ventral margin of median lobe straight. Walls of shaft with scattered poriferous canals. Dorsal copulatory sclerites in form of specific “ bird’s head ” plate, surrounded by small scaled membraneous field. Spines of internal sac absent. Left paramere (Fig. 6 G) of normal proportions, with moderately attenuate apex, bearing three long setae. Right paramere (Fig. 6 H) long and narrow, bearing three long setae, slightly shorter the length of paramere. Ring sclerite as on Fig. 7 H. Female internal genitalia. Not examined. Geographical distribution. This species is distributed within the Marlborough Sounds area (Crosby et al. 1998) (Fig. 9, yellow circles). Habitat. According to label data specimens were collected from leafmold and litter without more precise data about plant communities. Relationships. The structure of the male genitalia of N. townsendi suggests relationship to other species having dorsal copulatory sclerite in a shape of “ bird’s head, ” such as N. brouni, N. distinctus, and N. tararua. The state of the scaled membraneous field and the presence of the area with poriferous canals suggest that N. townsendi may share the common ancestry with N. tararua. Within-group relationships of N. townsendi remains to be examined.	en	Sokolov, Igor M. (2023): Eight new species of the genus Nesamblyops Jeannel (Anillini: Carabidae: Coleoptera) from New Zealand with notes about dispersal of the genus to the North Island. Zootaxa 5230 (2): 179-201, DOI: https://doi.org/10.11646/zootaxa.5230.2.3
