identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5C11879EFFB5FFDEFD84EF7DFDF610CB.text	5C11879EFFB5FFDEFD84EF7DFDF610CB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aquattuor Frederiksen 2013	<div><p>Genus Aquattuor Frederiksen, 2013</p> <p>Diagnosis (modified after Enghoff &amp; Frederiksen 2015)</p> <p>Differs from all other Odontopygidae by the structure of the limbus, which consists of large, rectangular, easily detachable flaps. Gonopods of the Prionopetalini type, i.e., basomere of telopodite with a torsate flexible zone followed by a posttorsal narrowing; solenomere elongated. Gonopodal proplica apically expanded into a distal palette; metaplica with an oblique flange; solenomere simple, thin and whip-like; telomere terminating in a long, curved, gutter-like to tubular part.</p> <p>Descriptive notes</p> <p>For characters not mentioned here, see Enghoff &amp; Frederiksen (2015).</p> <p>Mandibular stipes (Fig. 2): in males, but not in females with a disto-ventral expansion.</p> <p>First pair of male legs: in the genus Aquattuor the first pair of legs is remarkably diverse. Figs 3–7 illustrate the first pair of legs in all known species of the genus, with a focus on the prefemur. In all of them, the prefemur carries a more or less finger-shaped process (pfp) which projects orad from the anterior surface of the prefemur (Figs 3–7). The distal (ventral) surface of the process is smooth, the basal (dorsal) surface is more or less scaly-rugose. The shape of the process varies from slender fingershaped to relatively broad/triangular, and in one species (A. nguruensis sp. nov.) there is a clearly delimited depression (pfd) on the distal surface of the process (Fig. 4 A–C). In A. major Enghoff, 2015, A. longipala Enghoff, 2015 and A. mollilobus sp. nov. there is in addition a second, more distal, more irregularly shaped prefemoral lobe (dpl, Figs 5–6).</p> <p>Another type of variation is seen in the setation of the telopodites of the first pair of legs (Fig. 8). Here, A. mollilobus sp. nov. (Fig. 8C) stands out by having very short setae on all podomeres. Aquattuor major and A. longipala have long setae on the tarsus, but very short ones on the femur (Fig. 8B). The remaining species have long setae (same length as on the following leg-pairs) on all podomeres (Fig. 8A). These differences in setation do not seem to reflect the setation of normal walking legs.</p> <p>In one species (A. udzungwensis), the podomeres distal to the prefemur of the first leg pair seem to be invariably missing (several specimens examined) (Fig. 7 D–E).</p> <p>Adhesive pads on walking legs are absent.</p> <p>Included species (alphabetically)</p> <p>Aquattuor claudiahempae Enghoff &amp; Frederiksen, 2015</p> <p>A. denticulatus Frederiksen, 2013 (type species)</p> <p>A. fasciatus (Attems, 1896)</p> <p>A. longipala Enghoff, 2015</p> <p>A. major Enghoff, 2015</p> <p>A. mollilobus sp. nov.</p> <p>A. nguruensis sp. nov.</p> <p>A. stereosathe Enghoff, 2015</p> <p>A. submajor Enghoff, 2015</p> <p>A. udzungwensis Enghoff, 2015</p></div> 	http://treatment.plazi.org/id/5C11879EFFB5FFDEFD84EF7DFDF610CB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Enghoff, Henrik	Enghoff, Henrik (2020): A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1-32, DOI: https://doi.org/10.5852/ejt.2020.626
5C11879EFFB2FFD0FE0DE875FAB01676.text	5C11879EFFB2FFD0FE0DE875FAB01676.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aquattuor claudiahempae Enghoff 2015	<div><p>Aquattuor claudiahempae Enghoff &amp; Frederiksen, 2015</p> <p>Figs 1, 3 D–I, 9–10</p> <p>Aquattuor claudiahempae Enghoff &amp; Frederiksen, 2015: 17.</p> <p>Diagnosis</p> <p>Diameter 1.41–1.58 mm, 50–53 podous rings (Fig. 9). Shares a short (as broad as long, or slightly longer) gonopodal palette with A. denticulatus, A. fasciatus, A. stereosathe, A. submajor, and A. udzungwensis. Differs from these species, except A. fasciatus, by the combination of fully developed telopodites on the first pair of male legs, the gonopodal telomere describing a ca 270° curve, curving almost in one plane only, by lacking a spinose lid-like flap on the besal telomeral lamella, and by having a distinct mesobasal lobe of the gonopod palette. Very similar to, and perhaps not distinct from A. fasciatus, differs by being mostly smaller (male diameter &lt;1.7 mm) and by having the prefemoral processes of the first pair of male legs more slender, finger-shaped.</p> <p>Material studied (total: 25 ♂♂, 29 ♀♀) TANZANIA • 1 ♂, paratype; Kilimanjaro Region, Mt Kilimanjaro, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=37.269245&amp;materialsCitation.latitude=-3.2331583" title="Search Plazi for locations around (long 37.269245/lat -3.2331583)">Hai District</a>; 03°13′59.37″ S, <p>37°16′09.28″ E; 1345 m a.s.l.; 20 Nov. 2013; S.B. Frederiksen leg.; coffee plantation, Plot 2; NHMD 621638 • 24 ♂♂, 29 ♀♀; Morogoro Region, Udzungwa Mts National Park, Kidatu, Plot 2; <p>07°41′14.9″ S, 36°56′24.7″ E; 650 m a.s.l.; 24 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from eight traps); NHMD 621639.</p></p> </p> <p>37°16′09.28″ E; 1345 m a.s.l.; 20 Nov. 2013; S.B. Frederiksen leg.; coffee plantation, Plot 2; NHMD 621638 • 24 ♂♂, 29 ♀♀; Morogoro Region, Udzungwa Mts National Park, Kidatu, Plot 2; <p>07°41′14.9″ S, 36°56′24.7″ E; 650 m a.s.l.; 24 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from eight traps); NHMD 621639.</p></p> <p>07°41′14.9″ S, 36°56′24.7″ E; 650 m a.s.l.; 24 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from eight traps); NHMD 621639.</p> <p>Descriptive notes</p> <p>The specimens from Udzungwa Mts National Park very much resemble those from Mt Kilimanjaro (type locality of A. claudiahempae), in size (Fig. 9) as well as in gonopod structure (Fig. 10, cf. Enghoff &amp; Frederiksen 2015: fig. 10). In particular, the two samples agree in having the distal part of the telomere compressed, i.e., instead of forming a gutter, the lamella constituting the telomere is folded flat such that the inside of the gutter becomes invisible; the outside is strongly microspinose on one side and the free margin of the other side is characteristically wavy-spinose (Fig. 10D, Enghoff &amp; Frederiksen 2015: fig. 10d).</p> <p>First pair of male legs (Fig. 3 D–I) each with a single, finger-shaped prefemoral process; prefemoral processes more slender than those of the otherwise very similar A. fasciatus (Fig. 3 A–C); setae of telopodite distal to prefemur similar in length to those of normal walking legs.</p> <p>Distribution</p> <p>Described from Mt Kilimanjaro (1169–1345 m a.s.l., in habitats disturbed by human activities; Enghoff &amp; Frederiksen 2015); now also known from one site at 650 m a.s.l. in the Udzungwa Mts (Fig. 1).</p> <p>Remarks</p> <p>Enghoff (2016 b) discussed A. fasciatus (Attems, 1896) (see below), emphasizing its great similarity with A. claudiahempae and suggesting that the latter might be a synonym of A. fasciatus. The similarity between both nominal species also extends to the compressed distal part of the telomere (Enghoff 2016 b: fig. 7c, f). The only differences between the two seem to be size (cf. the identification key below, couplet 9) and the shape of the prefemoral process of the first pair of legs (Fig. 3). The new specimens of A. claudiahempae from Kidatu, Udzungwa Mts are of the same size as A. claudiahempae from Mt Kilimanjaro (Fig. 9), and their prefemoral processes strongly resemble those of Kilimanjaro specimens. Aquattuor claudiahempae and A. fasciatus are, for the time being, upheld as separate species.</p> </div>	http://treatment.plazi.org/id/5C11879EFFB2FFD0FE0DE875FAB01676	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Enghoff, Henrik	Enghoff, Henrik (2020): A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1-32, DOI: https://doi.org/10.5852/ejt.2020.626
5C11879EFFBCFFD1FDE1EFC8FD091685.text	5C11879EFFBCFFD1FDE1EFC8FD091685.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aquattuor denticulatus Frederiksen 2013	<div><p>Aquattuor denticulatus Frederiksen, 2013</p> <p>Figs 1, 4 D–F</p> <p>Aquattuor denticulatus Frederiksen, 2013: 64.</p> <p>Diagnosis</p> <p>Diameter 1.58–1.77 mm, 50–52 podous rings. Shares a short (as broad as long, or slightly longer) gonopodal palette with A. claudiahempae, A. fasciatus, A. stereosathe, A. submajor and A. udzungwensis. Differs from these species by having the subdistal part of the gonopodal telomere almost straight, parallel to the coxa, the telomere tip being bent abruptly laterad.</p> <p>Material studied (total: 1 ♂) TANZANIA • 1 ♂, paratype; Tanga Region, Muheza District, East Usambara Mts, Amani, at Sigi<p>River; 500 m a.s.l.; 7 Feb. 1977; H. Enghoff, O. Lomhodlt and O. Martin leg.; NHMD 621640 [= ZMUC 00020528].</p></p> <p>River; 500 m a.s.l.; 7 Feb. 1977; H. Enghoff, O. Lomhodlt and O. Martin leg.; NHMD 621640 [= ZMUC 00020528].</p> <p>Descriptive notes</p> <p>First pair of male legs (Fig. 4 D–F) each with a single, finger-shaped prefemoral process; setae of telopodite distal to prefemur similar in length to those of normal walking legs.</p> <p>Distribution</p> <p>Known only from the type locality, Tanga Region, Muheza District, E Usambara Mts, Amani.Altitudinal range: 500– 1000 m a.s.l. (Frederiksen 2013).</p></div> 	http://treatment.plazi.org/id/5C11879EFFBCFFD1FDE1EFC8FD091685	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Enghoff, Henrik	Enghoff, Henrik (2020): A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1-32, DOI: https://doi.org/10.5852/ejt.2020.626
5C11879EFFBDFFD7FD87EFB3FB281065.text	5C11879EFFBDFFD7FD87EFB3FB281065.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aquattuor fasciatus (Attems 1896)	<div><p>Aquattuor fasciatus (Attems, 1896)</p> <p>Figs 1, 3 A–C</p> <p>Odontopyge fasciata Attems, 1896: 40.</p> <p>Prionopetalum fasciatum – Attems 1914: 210. — Brolemann 1920: 123. — Kraus 1960: 86. — VandenSpiegel &amp; Pierrard 2009: 152. — Enghoff et al. 2016: 69.</p> <p>Aquattuor aff. claudiahempae – Enghoff &amp; Frederiksen 2015: 19.</p> <p>Aquattuor fasciatus – Enghoff 2016b: 14.</p> <p>Diagnosis</p> <p>Diameter 1.55–1.91 mm, 51–53 (55?) podous rings. Shares a short (as broad as long, or slightly longer) gonopodal palette with A. claudiahempae, A. fasciatus, A. stereosathe, A. submajor and A. udzungwensis. Differs from these species, except A. claudiahempae, by the combination of fully developed telopodites on the first pair of male legs, the gonopodal telomere describing a ca 270° curve, curving almost in one plane only, by lacking a spinose lid-like flap on the basal telomeral lamella, and by having a distinct meso-basal lobe of the gonopod palette. Very similar to, and perhaps not distinct from A. claudiahempae, differs by being mostly larger (male diameter&gt; 1.5 mm) and by having the prefemoral processes of the first pair of male legs stouter, subtriangular.</p> <p>Material studied (total: 1 ♂)</p> <p>TANZANIA • 1 ♂; Zanzibar, the sultan’s palace; 16 Jun. 1979; M. Stoltze leg.; NHMD 621641.</p> <p>Descriptive notes</p> <p>First pair of male legs (Fig. 3 A–C) each with a single, relatively stout, triangular prefemoral process; prefemoral processes stouter than those of the otherwise very similar A. claudiahempae; setae of telopodite distal to prefemur similar in length to those of normal walking legs.</p> <p>See remarks under A. claudiahempae.</p> <p>Distribution</p> <p>Recorded from Zanzibar (type locality), Pwani Region, Bagamoyo Distr., Vula Mountain, Pongwe (VandenSpiegel &amp; Pierrard, 2009, as Prionopetalum fasciatum) and from Morogoro Region, Kilosa District, Rubeho Mts, 900 m a.s.l. (Enghoff &amp; Frederiksen 2015, as A. aff. claudiahempae).</p> </div>	http://treatment.plazi.org/id/5C11879EFFBDFFD7FD87EFB3FB281065	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Enghoff, Henrik	Enghoff, Henrik (2020): A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1-32, DOI: https://doi.org/10.5852/ejt.2020.626
5C11879EFFBBFFD5FD86E9D7FC7F1642.text	5C11879EFFBBFFD5FD86E9D7FC7F1642.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aquattuor longipala Enghoff, in Enghoff & Frederiksen 2015	<div><p>Aquattuor longipala Enghoff, 2015</p> <p>Figs 1, 5 D–F, 8B, 9, 11</p> <p>Aquattuor longipala Enghoff, in Enghoff &amp; Frederiksen, 2015: 12.</p> <p>Diagnosis</p> <p>Diameter 1.78–2.02 mm, 50–52 podous rings (Fig. 9). Differs from congeners, except A. longipala and A. major, by having the gonopodal palette considerably longer than broad, and by having a distal prefemoral lobe on the first pair of male legs, in addition to the normal prefemoral process. Differs from A. mollilobus sp. nov. by having distinct lateral and mesal incisions on the gonopod coxa and by having the tarsal setae of the first pair of male legs of normal length; differs from A. major in lacking a triangular tooth on the anterior margin of the gonopod telomere.</p> <p>Material studied (total: 3 ♂♂, 3 ♀♀, 3 juvs)</p> <p>TANZANIA • 2 ♂♂, 3 ♀♀, 3 juvs; Morogoro Region, Udzungwa Mts National Park, Mito Mitatu, Plot 15; 07°49′39.8″ S, 36°50′26.0″ E; 1552 m a.s.l.; 27 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from two traps); NHMD 621642 • 1 ♂; Morogoro Region, Udzungwa Mts National Park, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.846333&amp;materialsCitation.latitude=-7.8373055" title="Search Plazi for locations around (long 36.846333/lat -7.8373055)">Mito Mitatu</a>; 07°50′14.3″ S, 36°50′46.8″ E; 1207 m a.s.l.; 25 Nov. 2013; T. Pape and N. Scharff leg.; pitfall trapping; NHMD 621643.</p> <p>Descriptive notes</p> <p>The new specimens perfectly fit the original description of A. longipala, except that they are marginally larger (51–52 podous rings, diameter 1.84–2.02 mm, vs 50–52 podous rings, diameter 1.78–1.90 mm for type specimens) (Fig. 9).</p> <p>First pair of male legs (Fig. 5 D–F) each with a single slender, finger-shaped prefemoral process (pfp); in addition, each prefemur has a smaller, irregular lobe (dpl) on the oral side of the distal margin; setae of femur much shorter than corresponding setae on normal walking legs (Fig. 8B).</p> <p>Fig. 11 shows a gonopod of one of the newly collected specimens.</p> <p>Aquattuor longipala is very similar to A. major (see Discussion).</p> <p>Distribution</p> <p>Known from two sites in the Udzungwa Mts, viz., West Kilombero FR and Mito Mitatu in Udzungwa Mts National Park (Fig. 1). Altitudinal range: 1207–1552 m a.s.l.</p></div> 	http://treatment.plazi.org/id/5C11879EFFBBFFD5FD86E9D7FC7F1642	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Enghoff, Henrik	Enghoff, Henrik (2020): A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1-32, DOI: https://doi.org/10.5852/ejt.2020.626
5C11879EFFB9FFD5FD9DEFF5FAD21272.text	5C11879EFFB9FFD5FD9DEFF5FAD21272.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aquattuor major Enghoff, in Enghoff & Frederiksen 2015	<div><p>Aquattuor major Enghoff, 2015</p> <p>Figs 1, 5 A–C, 9</p> <p>Aquattuor major Enghoff, in Enghoff &amp; Frederiksen, 2015: 10.</p> <p>Diagnosis</p> <p>Diameter 1.97–2.11 mm, 53–54 podous rings (Fig. 9). Differs from congeners, except A. longipala and A. mollilobus sp. nov., by having the gonopodal palette considerably longer than broad, and by having a distal prefemoral lobe on the first pair of male legs, in addition to the normal prefemoral process. Differs from A. longipala and A. mollilobus sp. nov. by having a triangular tooth on the anterior margin of the gonopod telomere, from A. mollilobus sp. nov. further by having distinct lateral and mesal incisions on the gonopod coxa and by having the tarsal setae of the first pair of male legs of normal length.</p> <p>Material studied (total: 1 ♂)</p> <p>TANZANIA • 1 ♂, paratype; Morogoro Region, Udzungwa Mts, Mwanihana FR, above Sanje; 1650 m a.s.l.; 18 Aug. 1982; M. Stoltze and N. Scharff leg.; pitfall trap; NHMD 621644.</p> <p>Descriptive notes</p> <p>First pair of male legs (Fig. 5 A–C) each with a single, finger-shaped prefemoral process (pfp); in addition each prefemur has a smaller, irregular lobe (dpl) on the oral side of the distal margin; setae of femur much shorter than corresponding setae on normal walking legs, as in A. longipala (see Fig. 8B).</p> <p>Aquattuor major is very similar to A. longipala (see Discussion).</p> <p>Distribution</p> <p>Known only from Mwanihana FR in the Udzungwa Mts. Altitudinal range: 1650–1850 m a.s.l.</p></div> 	http://treatment.plazi.org/id/5C11879EFFB9FFD5FD9DEFF5FAD21272	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Enghoff, Henrik	Enghoff, Henrik (2020): A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1-32, DOI: https://doi.org/10.5852/ejt.2020.626
5C11879EFFB9FFC8FDA9EBCCFF331274.text	5C11879EFFB9FFC8FDA9EBCCFF331274.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aquattuor mollilobus Enghoff 2020	<div><p>Aquattuor mollilobus sp. nov.</p> <p>urn:lsid:zoobank.org:act: A453CBE6-5FF7-4327-9B45-FA0C6A9AE84F</p> <p>Figs 1, 6 A–C, 8C, 9, 12–13</p> <p>Diagnosis</p> <p>Diameter 1.80–1.96 mm, 48–53 podous rings (Fig. 9). Differs from congeners, except A. longipala and A. major, by having the gonopodal palette considerably longer than broad, and by having a distal prefemoral lobe on the first pair of male legs, in addition to the normal prefemoral process. Differs from A. longipala and A. major by having the lateral and mesal incisions of the gonopod coxa indistinct, and by having the tarsal setae of the first pair of male legs much shorter than tarsal setae of normal walking legs.</p> <p>Etymology</p> <p>The name is an adjective meaning soft (Latin: mollis) lobe and refers to the distal modification of the first male prefemora.</p> <p>Material studied (total: 21 ♂♂, 11 ♀♀, 1 juv.)</p> <p>Holotype</p> <p>TANZANIA • ♂; Morogoro Region, Udzungwa Mts National Park, Kidatu, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.918556&amp;materialsCitation.latitude=-7.678361" title="Search Plazi for locations around (long 36.918556/lat -7.678361)">Plot</a> 6; 07°40′42.1″ S, 36°55′06.8″ E; 1482 m a.s.l.; 21 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping; NHMD 621645. Paratypes</p> <p>TANZANIA • 8 ♂♂; Morogoro Region, Udzungwa Mts National Park, Kidatu, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.918583&amp;materialsCitation.latitude=-7.6793056" title="Search Plazi for locations around (long 36.918583/lat -7.6793056)">Plot</a> 5; 07°40′45.5″ S, 36°55′06.9″ E; 1448 m a.s.l.; 21 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from three traps); NHMD 621646 • 7 ♂♂; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.918556&amp;materialsCitation.latitude=-7.678361" title="Search Plazi for locations around (long 36.918556/lat -7.678361)">Udzungwa Mts National Park</a>, Kidatu, Plot 6; 07°40′42.1″ S, 36°55′06.8″ E; 1482 m a.s.l.; 21 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from six traps); NHMD 621647 • 5 ♂♂; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.918556&amp;materialsCitation.latitude=-7.6850557" title="Search Plazi for locations around (long 36.918556/lat -7.6850557)">Udzungwa Mts National Park</a>, Kidatu, Plot 9; 07°41′06.2″ S, 36°55′06.8″ E, 1527 m a.s.l.; 23 Oct. 2014; J. Malumbres- Olarte leg.; pitfall trapping (summed catch from three traps); NHMD 621648.</p> <p>Referred non-type material</p> <p>TANZANIA • 11 ♀♀, 1 juv.; same collection data as for paratypes (plots 5, 6, 9); NHMD.</p> <p>Description (male)</p> <p>MEASUREMENTS. Body length 28–31 mm; vertical diameter 1.82–1.91 mm; 48–53 podous rings, no apodous rings in front of telson.</p> <p>COLOUR. Similar to that of A. claudiahempae (Enghoff &amp; Frederiksen 2015: fig. 1). A light mid-dorsal stripe present.</p> <p>LIMBUS (Fig. 12E). With rectangular, detachable flaps, as typical of the genus.</p> <p>FIRST PAIR OF MALE LEGS (Figs 6 A–C, 8C). Prefemoral processes (pfp) of first pair of legs slender, fingershaped, slightly curved laterad; in addition an irregular, rugose lobe (dpl) on oral-distal margin of prefemur; setae of all podomeres much shorter than femoral setae of normal walking legs.</p> <p>GONOPOD COXA (Fig. 12 A–D). In anterior or posterior view ca 3½ × as long as broad. Margins of proplica (pp) parallel in basal ca ¾. Mesal (mi) and lateral (li) incisions shallow and poorly developed; apical palette (pa) narrower than basal part of proplica, lateral margin regularly rounded, mesal margin straight.</p> <p>GONOPOD TELOPODITE (Fig. 13). Solenomere (slm) thin, whiplike. Main part of telomere (tm) a rather broad gutter describing a 270° curve in almost one plane; tip of telomere broad rounded, densely beset with minute spicules on external surface.</p> <p>In all other characters conforming to the general description of Aquattuor in Enghoff &amp; Frederiksen (2015).</p> <p>Distribution</p> <p>Known only from a single site in Udzungwa Mts National Park (Fig. 1). Altitudinal range: 1448–1527 m a.s.l.</p></div> 	http://treatment.plazi.org/id/5C11879EFFB9FFC8FDA9EBCCFF331274	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Enghoff, Henrik	Enghoff, Henrik (2020): A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1-32, DOI: https://doi.org/10.5852/ejt.2020.626
5C11879EFFA4FFCFFDA5EBC7FB431711.text	5C11879EFFA4FFCFFDA5EBC7FB431711.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aquattuor nguruensis Enghoff 2020	<div><p>Aquattuor nguruensis sp. nov.</p> <p>urn:lsid:zoobank.org:act: B691ECD6-3E7E-4731-AD33-1DD9D39381DB</p> <p>Figs 1, 4 A–C, 14–15</p> <p>Aquattuor sp. – Enghoff &amp; Frederiksen 2015: 21 (?).</p> <p>Diagnosis</p> <p>Diameter 2.0 mm. Differs from congeners by lacking lateral and mesal incisions on the gonopod coxa and by having the telomere tip flattened and smooth. Differs from A. mollilobus sp. nov., in which the lateral and mesal incisions are indistinct, by lacking a distal prefemoral lobe on the first pair of male legs.</p> <p>Etymology</p> <p>The name is an adjective referring to the type locality.</p> <p>Material studied (total: 1 ♂, 2 ♀♀)</p> <p>Holotype</p> <p>TANZANIA • ♂; Morogoro Region and District, west side of Nguru Mountains, near Maskati mission, 20 km W of Turisani; 1900 m a.s.l.; unknown date; Jan Kielland leg.;moist evergreen forest; VMNH110617.</p> <p>Referred non-type material</p> <p>TANZANIA • 1 ♀; same collection data as for holotype; VMNH110618 • 1 ♀; Nguru Mts, Mhonda Mission at Turiani; Oct. 1992; M. Andersen leg.; NHMD 621649.</p> <p>Description (male)</p> <p>MEASUREMENTS. Vertical diameter 2.0 mm; posterior part of body missing.</p> <p>COLOUR. Faded, but apparently similar to that of A. claudiahempae (Enghoff &amp; Frederiksen 2015: fig. 1). Traces of a light mid-dorsal stripe present.</p> <p>LIMBUS (Fig. 14F). Quite worn, but still shows rectangular, detachable flaps as typical of the genus. FIRST PAIR OF MALE LEGS (Figs 4 A–C). Prefemoral processes relatively stout, triangular; distal (ventral) surface of processes with a large, sharply delimited depression (pfd); setae of telopodite distal to prefemur same length as setae of normal walking legs.</p> <p>GONOPOD COXA (Fig. 14). In anterior or posterior view ca 3 × as long as broad. Margins of proplica subparallel, but lateral margin regularly convex, mesal margin correspondingly concave. No mesal and lateral incisions, apical palette (pa) hence poorly delimited, triangular.</p> <p>GONOPOD TELOPODITE (Fig. 15). Solenomere (slm) thin, whip-like. Main part of telomere a rather broad gutter describing a 270° curve in almost one plane; tip of telomere flattened, smooth.</p> <p>In all other characters conforming to the general description of Aquattuor in Enghoff &amp; Frederiksen (2015).</p> <p>Distribution</p> <p>Known only from one, possibly two sites in the Nguru Mts. Altitude: 1900 m a.s.l.</p></div> 	http://treatment.plazi.org/id/5C11879EFFA4FFCFFDA5EBC7FB431711	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Enghoff, Henrik	Enghoff, Henrik (2020): A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1-32, DOI: https://doi.org/10.5852/ejt.2020.626
5C11879EFFA3FFCFFDFDEE2BFAB61241.text	5C11879EFFA3FFCFFDFDEE2BFAB61241.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aquattuor stereosathe Enghoff, in Enghoff & Frederiksen 2015	<div><p>Aquattuor stereosathe Enghoff, 2015</p> <p>Figs 1, 6 D–F, 8A, 9</p> <p>Aquattuor stereosathe Enghoff, in Enghoff &amp; Frederiksen, 2015: 15.</p> <p>Diagnosis</p> <p>Diameter 1.68–1.75 mm, 46–51 podous rings (Fig. 9). Shares a short (as broad as long, or slightly longer) gonopodal palette with A. claudiahempae, A. denticulatus, A. fasciatus, A. submajor and A. udzungwensis. Differs from these species by having the gonopodal telomere describing a ca 360° curve, curving in three dimensions, and by having the telomere tip with coarsely laciniate margins and a surface that is not microspiculate.</p> <p>Material studied (total: 1 ♂)</p> <p>TANZANIA • 1 ♂, paratype; Iringa Region, Udzungwa Mts, Udzungwa Scarp FR, above Chita Village; 1050 m a.s.l.; 26–29 Oct. 1984; N. Scharff leg.; pitfall trap in intermediate rain forest; NHMD 621650.</p> <p>Descriptive notes</p> <p>First pair of male legs (Figs 6 D–F): prefemoral processes relatively stout, triangular; setae of telopodite distal to prefemur same length as setae of normal walking legs.</p> <p>Distribution</p> <p>Known only from the Udzungwa Scarp FR in the Udzungwa Mts. Altitudinal range: 900–1500 m a.s.l.</p></div> 	http://treatment.plazi.org/id/5C11879EFFA3FFCFFDFDEE2BFAB61241	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Enghoff, Henrik	Enghoff, Henrik (2020): A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1-32, DOI: https://doi.org/10.5852/ejt.2020.626
5C11879EFFA3FFCDFD86EBFBFED512DE.text	5C11879EFFA3FFCDFD86EBFBFED512DE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aquattuor submajor Enghoff, in Enghoff & Frederiksen 2015	<div><p>Aquattuor submajor Enghoff, 2015</p> <p>Figs 1, 7 A–C, 9, 16, 18</p> <p>Aquattuor submajor Enghoff, in Enghoff &amp; Frederiksen, 2015: 7.</p> <p>Diagnosis</p> <p>Diameter 1.41–1.87 mm, 41–48 podous rings (Fig. 9). Shares a short (as broad as long, or slightly longer) gonopodal palette with A. claudiahempae, A. denticulatus, A. fasciatus, A. stereosathe and A. udzungwensis. Differs from these species by the combination of fully developed telopodites on the first pair of male legs, the gonopodal telomere describing a ca 270° curve, curving almost in one plane only, by lacking a spinose lid-like flap on the besal telomeral lamella and by lacking a distinct mesobasal lobe of the gonopod palette.</p> <p>Material studied (total: 8 ♂♂)</p> <p>TANZANIA • 1 ♂, paratype; Udzungwa Mts, Mwanihana Forest above Sanje; 1000 m a.s.l.; 1 Aug. 1981; N. Scharff leg.; pitfall trap; NHMD 261651 • 1 ♂; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.918583&amp;materialsCitation.latitude=-7.6793056" title="Search Plazi for locations around (long 36.918583/lat -7.6793056)">Udzungwa Mts National Park</a>, Kidatu, Plot 5; 07°40′45.5″ S, 36°55′06.9″ E; 1448 m a.s.l.; 21 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping; NHMD 621652 • 1 ♂; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.918556&amp;materialsCitation.latitude=-7.678361" title="Search Plazi for locations around (long 36.918556/lat -7.678361)">Udzungwa Mts National Park</a>, Kidatu, Plot 6; 07°40′42.1″ S, 36°55′06.8″ E; 1482 m a.s.l.; 21 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping; NHMD 621653 • 4 ♂♂; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.91456&amp;materialsCitation.latitude=-7.6850557" title="Search Plazi for locations around (long 36.91456/lat -7.6850557)">Udzungwa Mts National Park</a>, Kidatu, Plot 9; 07°41′06.2″ S, 36°54′52.4″ E; 1527 m a.s.l.; 23 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from four traps); NHMD 621654 • 1 ♂; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.840805&amp;materialsCitation.latitude=-7.827722" title="Search Plazi for locations around (long 36.840805/lat -7.827722)">Udzungwa Mts National Park</a>, Mito Mitatu, Plot 15; 07°49′39.8″ S, 36°50′26.9″ E; 1552 m a.s.l.; 27 Oct. 2014; J. Malumbres- Olarte leg.; pitfall trapping; NHMD 621655.</p> <p>Descriptive notes</p> <p>The newly studied specimens from Kidatu are considerably smaller than those from Mwanihana FR; see Table 1.</p> <p>First pair of male legs (Figs 7 A–C): prefemoral processes finger-shaped; setae of telopodite distal to prefemur same length as setae of normal walking legs.</p> <p>Gonopod telopodite (Fig. 16): basal telomeral lamella (btl) forming a subspherical chamber but without a spinose lid-like flap, cf. A. udzungwensis. In the specimen illustrated in Fig. 16 the tip has remarkably long marginal laciniae, approaching the condition typical of A. stereosathe (Enghoff &amp; Frederiksen 2015: fig. 9c).</p> <p>Remarks</p> <p>The distinction between A. submajor and A. udzungwensis is discussed under the latter species.</p> <p>Distribution</p> <p>Known from Mwanihana FR and Kidatu in Udzungwa Mts National Park. Altitudinal range: 1000–1552 m a.s.l.</p></div> 	http://treatment.plazi.org/id/5C11879EFFA3FFCDFD86EBFBFED512DE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Enghoff, Henrik	Enghoff, Henrik (2020): A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1-32, DOI: https://doi.org/10.5852/ejt.2020.626
5C11879EFFA1FFC1FDEAEA60FAFE156F.text	5C11879EFFA1FFC1FDEAEA60FAFE156F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aquattuor udzungwensis Enghoff, in Enghoff & Frederiksen 2015	<div><p>Aquattuor udzungwensis Enghoff, 2015</p> <p>Figs 1, 2, 7 D–E, 9, 17–18</p> <p>Aquattuor udzungwensis Enghoff, in Enghoff &amp; Frederiksen, 2015: 13.</p> <p>Diagnosis</p> <p>Diameter 1.44–1.75 mm, 44–54 podous rings (Fig. 9). Shares a short (as broad as long, or slightly longer) gonopodal palette with A. claudiahempae, A. denticulatus, A. fasciatus, A. stereosathe and A. submajor. Differs from these species by having the telopodite of the first pair of male legs represented by the prefemur only, and by having the basal telomeral lamella forming a subspherical chamber with a spinose, lid-like flap.</p> <p>Material studied (total: 302 ♂♂, 120 ♀♀)</p> <p>TANZANIA • 4 ♂♂, paratypes; Iringa Region, Udzungwa Mts, Udzungwa Scarp FR, above Chita Village; 730 m a.s.l.; 25–29 Oct. 1984; N. Scharff leg.; pitfall traps in lowland rain forest; NHMD 621656 • 16 ♂♂, 13 ♀♀; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.94128&amp;materialsCitation.latitude=-7.6870832" title="Search Plazi for locations around (long 36.94128/lat -7.6870832)">Udzungwa Mts National Park</a>, Kidatu, Plot 1; 07°41′13.5″ S, 36°56′28.6″ E; 650 m a.s.l.; 24 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from seven traps); NHMD 621657 • 1 ♂, 4 ♀♀; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.940197&amp;materialsCitation.latitude=-7.6874723" title="Search Plazi for locations around (long 36.940197/lat -7.6874723)">Udzungwa Mts National Park</a>, Kidatu, Plot 2; 07°41′14.9″ S, 36°56′24.7″ E; 650 m a.s.l.; 24 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping; NHMD 621658 • 45 ♂♂, 12 ♀♀; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.930363&amp;materialsCitation.latitude=-7.684" title="Search Plazi for locations around (long 36.930363/lat -7.684)">Udzungwa Mts National Park</a>, Kidatu, Plot 3; 07°41′02.4″ S, 36°55′49.3″ E; 1005 m a.s.l.; 14 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from twelve traps); NHMD 621659 • 24 ♂♂, 2 ♀♀; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.93011&amp;materialsCitation.latitude=-7.6847777" title="Search Plazi for locations around (long 36.93011/lat -7.6847777)">Udzungwa Mts National Park</a>, Kidatu, Plot 4; 07°41′05.2″ S, 36°55′48.4″ E; 993 m a.s.l.; 15 Sep. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from eight traps); NHMD 621660 • 8 ♂♂, 5 ♀♀; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.93353&amp;materialsCitation.latitude=-7.689833" title="Search Plazi for locations around (long 36.93353/lat -7.689833)">Udzungwa Mts National Park</a>, Kidatu, Plot 7; 07°41′23.4″ S, 36°56′00.7″ E; 708 m a.s.l.; 24 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from six traps); NHMD 621661 • 167 ♂♂, 68 ♀♀; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.927555&amp;materialsCitation.latitude=-7.686917" title="Search Plazi for locations around (long 36.927555/lat -7.686917)">Udzungwa Mts National Park</a>, Kidatu, Plot 8; 07°41′12.9″ S, 36°55′39.2″ E; 978 m a.s.l.; 14 Oct.2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from eleven traps); NHMD 621662 • 2 ♂♂; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.867027&amp;materialsCitation.latitude=-7.8415556" title="Search Plazi for locations around (long 36.867027/lat -7.8415556)">Udzungwa Mts National Park</a>, Mito Mitatu, Plot 13; 07°50′29.6″ S, 36°52′01.3″ E; 674 m a.s.l.; 26 Oct. 2014; J. Malumbres- Olarte leg.; pitfall trapping (summed catch from two traps); NHMD 621663 • 5 ♂♂, 2 ♀♀; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.859165&amp;materialsCitation.latitude=-7.8405833" title="Search Plazi for locations around (long 36.859165/lat -7.8405833)">Udzungwa Mts National Park</a>, Mito Mitatu, Plot 14; 07°50′26.1″ S, 36°51′33.0″ E; 1006 m a.s.l.; 26 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from four traps); NHMD 621664 • 1 ♂; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=36.847195&amp;materialsCitation.latitude=-7.8375278" title="Search Plazi for locations around (long 36.847195/lat -7.8375278)">Udzungwa Mts National Park</a>, Mito Mitatu, Plot MM2; 07°50′15.1″ S, 36°50′49.9″ E; 1109 m a.s.l.; T. Pape and N. Scharff leg.; pitfall trapping; NHMD 621665 • 13 ♂♂, 4 ♀♀; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.91914&amp;materialsCitation.latitude=-8.503722" title="Search Plazi for locations around (long 35.91914/lat -8.503722)">Udzungwa Scarp Catchment Forest Reserve</a>, Chita, Plot 16; 08°30′13.4″ S, 35°55′08.9″ E; 659 m a.s.l.; 31 Oct. 2014; J. Malumbres-Olarte leg.; pitfall trapping (summed catch from seven traps); NHMD 621666 • 19 ♂♂, 10 ♀♀; Morogoro Region, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=35.91653&amp;materialsCitation.latitude=-8.499473" title="Search Plazi for locations around (long 35.91653/lat -8.499473)">Udzungwa Scarp Catchment Forest Reserve</a>, Chita, Plot 17; 08°29′58.1″ S, 35°54′59.5″ E; 908 m a.s.l.; 30 Oct. 2014; J. Malumbres- Olarte leg.; pitfall trapping (summed catch from six traps); NHMD 621667.</p> <p>Descriptive notes</p> <p>The newly studied specimens on average have more podous rings than those studied by Enghoff &amp; Frederiksen (2015) (Fig. 9). In particular, the specimens from Kidatu and Mito Mitatu have more podous rings (49–54, most specimens 53), whereas the new specimens from Chita (49–51 podous rings) overlap with the previously studied ones; see also Discussion.</p> <p>First pair of male legs (Fig. 7 D–E): prefemoral processes finger-shaped to triangular; telopodital podomeres distal to prefemur missing in all studied specimens; see Discussion.</p> <p>Gonopod telopodite (Fig. 17): basal telomeral lamella (btl) forming a subspherical chamber with a spinose, lid-like flap, cf. A. submajor and remarks below.</p> <p>Remarks – The distinction between A. submajor and A. udzungwensis</p> <p>The distinction between A. udzungwensis and A. submajor according to Enghoff &amp; Frederiksen (2015) is quite subtle. The newly studied material has contributed to a better understanding of the differences between these two species. Among the differences stated by Enghoff &amp; Frederiksen (2015), the shape of the meso-posterior lamella of the telomere seems to be the only reliable one (Figs 16–17). The mesoposterior lamella is high and basally angled in A. udzungwensis, and towards the tip of the telomere, the lamella narrows gradually. In A. submajor, the lamella is lower, not angled, and not narrowing so strongly towards the tip.</p> <p>However, two new clear differences can now be added:</p> <p>• The basal telomeral lamella (btl) in A. udzungwensis has a small, spinose, lid-like flap (spl) (Fig. 17); such a flap is absent in A. submajor (Fig. 16).</p> <p>• In all dissected males of A. udzungwensis (&gt; 10), collected in several parts of the Udzungwa Mts, the telopodites of the first leg-pair are strongly reduced distal to the prefemur (Fig. 7 D–E). Only tiny, somewhat irregular and darkened remnants are present, suggesting that their absence may be due to some physical damage inflicted on the male. In all examined males of A. submajor, the telopodites are normal (Fig. 7 A–C).</p> <p>For both species, the newly studied material considerably widens their size range: newly studied specimens of A. submajor have fewer podous rings and a smaller body diameter than those studied by Enghoff &amp; Frederiksen (2015); in A. udzungwensis the newly studied specimens tend to have more podous rings than those previously studied, while there is a marked difference in diameter. These differences may, at least in part, be correlated with altitude: both species show a clear decrease of numbers of podous rings with increasing altitude from mostly&gt; 50 podous rings at 600–900 m altitude to &lt;46 rings at 1400–1550 m altitude (Fig. 18).</p> <p>Distribution</p> <p>Known from several localities in the Udzungwa Mts (Fig. 1). Altitudinal range: 650–1410 m a.s.l.</p></div> 	http://treatment.plazi.org/id/5C11879EFFA1FFC1FDEAEA60FAFE156F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Enghoff, Henrik	Enghoff, Henrik (2020): A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1-32, DOI: https://doi.org/10.5852/ejt.2020.626
5C11879EFFADFFC6FD06ECD2FAB01735.text	5C11879EFFADFFC6FD06ECD2FAB01735.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aquattuor sp. in Enghoff & Frederiksen 2015	<div><p>Aquattuor sp.</p> <p>Material studied</p> <p>TANZANIA • Several ♀♀ /juveniles; Morogoro, Sokoine University Campus, NORAD guest house; 28 Mar. 1989; Zicsi leg.; HMNH.</p> <p>Remarks</p> <p>These specimens represent a range extension of the genus Aquattuor, but a closer identification is not possible.</p> <p>Key to the species of Aquattuor (males)</p> <p>1. Palette (pa) of gonopods elongate, considerably to much longer than broad (Figs 11–12; Enghoff &amp; Frederiksen 2015: figs 6–7); first pair of legs with a distal prefemoral lobe (Figs 5, 6 A–C); 48+ podous rings, diameter&gt; 1.7 mm..................................................................................................... 2</p> <p>– Palette as long as broad or slightly longer; first pair of legs without a distal prefemoral lobe (as in Fig. 3); diameter &lt;1.9 mm............................................................................................................... 4</p> <p>2. Palette (pa) not clearly delimited by lateral (li) and mesal (mi) incisions (Fig. 12A); tarsal setae of first pair of legs much shorter than tarsal setae of normal walking legs (Fig. 8C)................................................................................................................................. A. mollilobus sp. nov. (Udzungwa)</p> <p>– Palette delimited by lateral and mesal incisions (e.g., Fig. 11A); tarsal setae of first pair of legs ca of same length as tarsal setae of normal walking legs (Fig. 8B).......................................................... 3</p> <p>3. 53–54 podous rings, diameter&gt; 1.9 mm; anterior margin of telomere with a triangular tooth (Enghoff &amp; Frederiksen 2015: fig. 6A, tt)............................... A. major Enghoff, 2015 (Udzungwa)</p> <p>– 50–52 podous rings; anterior margin of telomere without a triangular tooth................................................................................................................................. A. longipala Enghoff, 2015 (Udzungwa)</p> <p>4. Subdistal part of telomere almost straight, parallel to coxa; telomere tip bent abruptly laterad (Enghoff &amp; Frederiksen 2015: fig. 11); 50–52 podous rings, diameter 1.6–1.8 mm......................................................................................................... A. denticulatus Frederiksen, 2013 (E Usambara)</p> <p>– Telomere describing a more complex curve..................................................................................... 5</p> <p>5. Gonopod coxa without lateral and mesal incisions; telomere tip flattened, smooth (Figs 14–15)................................................................................................................ A. nguruensis sp. nov. (Nguru)</p> <p>– Gonopod coxa with lateral (li) and mesal (mi) incisions; telomere tip microspiculate or with serrated margins.............................................................................................................................................. 6</p> <p>6. Telomere describing a ca 360° curve, curving in three dimensions; telomere tip with coarsely laciniate margins, but surface not microspiculate (Enghoff &amp; Frederiksen 2015: fig. 9)............................................................................................................ A. stereosathe Enghoff, 2015 (Udzungwa)</p> <p>– Telomere describing a ca 270° curve, curving almost in one plane only; telomere tip more or less microspiculate................................................................................................................................... 7</p> <p>7. Telopodites of first pair of legs only represented by prefemur; basal telomeral lamella (btl) forming a subspherical chamber with a spinose, lid-like flap (Fig. 17)................................................................................................................................................ A. udzungwensis Enghoff, 2015 (Udzungwa)</p> <p>– Telopodites of first pair of legs fully developed; basal telomeral lamella without a spinose, lid-like flap.................................................................................................................................................... 8</p> <p>8. &lt;49 podous rings; meso-basal lobe of gonopod palette indistinct (Enghoff &amp; Frederiksen 2015: fig. 5)................................................................................... A. submajor Enghoff, 2015 (Udzungwa)</p> <p>–&gt; 48 podous rings; meso-basal lobe (mbl) of gonopod palette well-developed (Enghoff &amp; Frederiksen 2015: fig. 10; Enghoff 2016: fig. 7).................................................................................................. 9</p> <p>9. Diameter&gt; 1.5 mm. Prefemoral processes of first pair of legs subtriangular (Fig. 3 A–C).......................................................................... A. fasciatus (Attems, 1896) (Zanzibar, Rubeho Mts, Mt Vula)</p> <p>– Diameter &lt;1.7 mm. Prefemoral processes of first pair of legs finger-shaped (Fig. 3 D–I)..................................................... A. claudiahempae Enghoff &amp; Frederiksen, 2015 (Udzungwa, Kilimanjaro)</p> </div>	http://treatment.plazi.org/id/5C11879EFFADFFC6FD06ECD2FAB01735	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Enghoff, Henrik	Enghoff, Henrik (2020): A mountain of millipedes VIII. The genus Aquattuor Frederiksen, 2013 revisited - a new species from the Udzungwa Mts, Tanzania, another from the Nguru Mts, and introduction of the first pair of male legs as a source of taxonomic characters (Diplopoda, Spirostreptida, Odontopygidae). European Journal of Taxonomy 626: 1-32, DOI: https://doi.org/10.5852/ejt.2020.626
