taxonID	type	description	language	source
894087A5FFA7FFEBF5E1FA10FB5F0393.taxon	description	(Figs. 1 A, 2 A, E, 3 A; Table 1)	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA7FFEBF5E1FA10FB5F0393.taxon	materials_examined	Holotype. — KU 168849, adult male from the Río Calima, 1.5 km (by road) west of Lago Calima, 1230 - m elevation (3 ° 53 ′ N, 76 ° 34 ′ W; datum ¼ WGS 84), Departamento del Valle del Cauca, Colombia, one of a series collected by William E. Duellman and Linda Trueb on 31 May and 1 June 1975. Paratypes. — KU 168846 – 48, 168851, adult females collected with the holotype.	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA7FFEBF5E1FA10FB5F0393.taxon	diagnosis	Diagnosis. — Pristimantis calima is diagnosed by the following combination of characters: (1) skin on dorsum smooth, becoming finely shagreen laterally and coarse on the flanks; ventral skin areolate; discoidal fold present, well anterior to groin; dorsolateral folds and ridges absent in scapula region; (2) tympanic membrane differentiated, round; its length 36.3 % of eye length in one male and 26.4 – 38.6 % in four females; tympanic annulus prominent, not covered by supratympanic fold, which extends from posterior corner of orbit along upper edge of temporal region toward insertion of arm; (3) snout moderately long, subacuminate in dorsal view, rounded in profile, with papilla at tip; canthus rostralis straight; (4) upper eyelid bearing 3 – 6 tubercles small, narrower than IOD (53.5 – 73.3 % IOD); small interocular tubercle present; cranial crests absent; (5) choanae small, ovoid; not concealed by palatal shelf of maxillary arch; dentigerous processes of vomers prominent, triangular in outline, separated medially by a distance equal to the width of the visible dentigerous process, located well behind posterior edges of choanae, each odontophore bearing 4 – 5 teeth; (6) male with vocal slits and subgular vocal sac evident externally; nuptial pads present on thumbs; (7) finger I shorter than finger II; discs and circumferential grooves present on all fingers; discs truncate, except on finger I, which is round; disc of finger I smaller than that of finger II and this in turn smaller than discs on fingers III and IV; (8) fingers with lateral fringes; palmar tubercle not divided; thenar tubercle oval, slightly smaller than palmar tubercle; supernumerary tubercles low, distributed on all fleshy parts of palm; subarticular tubercles round, two on thumb and second finger and three on third and fourth fingers, and larger than supernumerary tubercles; (9) two or three ulnar tubercles subconical, not coalesced (Fig. 2 A); (10) heel and outer edge of tarsus lacking tubercles; inner tarsal tubercle indistinct; (11) inner metatarsal tubercle elongate, its length twice its width; low, conical outer metatarsal tubercle one-fourth size of inner metatarsal tubercle; subarticular tubercles round, two on toes I and II, three on toes III and V, and four on toe IV; supernumerary plantar tubercles low, rounded, on proximal segments of toes; (12) toes bearing prominent lateral fringes; basal webbing not encompassing basal subarticular tubercles, except on outer border of toe IV (Fig. 2 E); toe III shorter than toe V; toe III extending to distal edge of the antepenultimate subarticular tubercle of toe IV; toe V reaching distal edge of penultimate subarticular tubercle of toe IV; discs and circumferential grooves present on all toes; discs of toes III – V equal to discs on fingers III – IV and larger than discs of toes I – II; (13) color in life: dorsum dull green, reddish brown, or olive brown; ventral surfaces of hind limbs pinkish lavender; groin and anterior surfaces of thighs cream brown; throat and belly surfaces yellow; iris pale creamy bronze with median red streak; (14) SVL in the only known adult male 24.0 mm; in adult females 37.7 – 40.0 mm (mean ± 1 SD ¼ 38.8 ± 0.90 mm; n ¼ 4). 3 CORRESPONDENCE: e-mail, sarriajhon @ gmail. com Comparisons. — Pristimantis calima differs from the other species in the genus by having basal webbing on outer edge of toe IV, subarticular tubercles beneath the joint between distal phalanges, papilla at the tip of snout, and uniform cream-brown coloration on the anterior surfaces of thighs and in the groin. It is most similar to three Ecuadorian species — P. eugeniae, P. nŋctophŋlax, and P. subsigillatus (Fig. 1) by having subarticular tubercles beneath the joint between distal phalanges, dorsal skin smooth, ventral skin areolate, discoidal fold well anterior to groin, snout subacuminate in dorsal view and rounded in profile, papilla at tip of snout, vocal slits and nuptial pads present, dentigerous processes of vomers prominent, tympanic membrane differentiated, tympanic annulus not covered by supratympanic fold, and by lacking of tubercles on medial surface of tarsus. However, P. calima has more webbing on outer edge of toe IV that encompasses the basal subarticular tubercle (not encompassing basal subarticular tubercle in those species; Fig. 2 E – H). Furthermore, females of P. calima are larger (SVL 37.7 – 40.0 mm) than those of P. eugeniae (SVL 30.5 – 37.6 mm; Lynch and Duellman 1997) and P. subsigillatus (SVL 30.0 – 33.4 mm; Lynch and Duellman 1997). Also, P. calima differs from P. eugeniae by having a row of three or four subconical tubercles on the postaxial surface of the forearm (only the antebrachial tubercle is present in P. eugeniae; Fig. 2 A, B). Furthermore, P. calima differs from P. nŋctophŋlax and P. subsigillatus by having uniform cream-brown coloration on anterior surfaces of thighs and groin area (black bars or gray reticulations of the anterior surfaces of the thighs and groin in P. nŋctophŋlax and P. subsigillatus; Fig. 3). Finally, P. calima could be considered similar in appearance to P. mindo Arteaga et al. (2013); however, P. calima has basal webbing on the outer edge of toe IV, papilla at the tip of snout, nuptial pads present on thumbs, and females having uniform creambrown coloration on the anterior surfaces of thighs and in the groin (instead of flash marks on the thighs and groin in P. mindo). Among species of Pristimantis in western Colombia, few have webbing between toes; these are P. albericoi (Lynch and Ruiz-Carranza 1996), P. bernali (Lynch 1986), P. diaphonus (Lynch 1986), P. diogenes (Lynch and RuizCarranza 1996), P. hŋbotragus (Lynch 1992), P. jaimei (Lynch 1992), and P. loustes (Lynch 1979). In comparison with P. calima, P. albericoi is smaller, with a SVL of 20.6 – 20.7 mm in adult males and 25.0 – 29.3 mm in adult females (Lynch and Ruiz-Carranza 1996) and has low postocular folds (SVL 24.0 mm in adult males and 37.7 – 40.0 mm of SVL in adult females, and postocular folds absent in P. calima). Likewise, P. bernali lacks vocal slits, nuptial pads on thumb, and papilla on the tip of the snout (characters present in P. calima), whereas P. diaphonus and P. loustes differ by having toes half webbed and no tympanic membrane (basal webbing not encompassing basal subarticular tubercles except on outer border of toe IV and tympanic membrane evident in P. calima). Webbing encloses the basal subarticular tubercles on the toes and an inner tarsal fold characterizes P. diogenes and P. jaimei (basal webbing not encompassing basal subarticular tubercles except on outer border of toe IV and tarsal fold absent in P. calima). Finally, P. hŋbotragus is distinguished by having a flaplike inner tarsal fold (inner tarsal fold absent in P. calima).	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA7FFEBF5E1FA10FB5F0393.taxon	description	Description of the holotype. — An adult male with head wider than body; head width 42.5 % of SVL; head length 37.5 % of SVL; snout moderately long, subacuminate in dorsal view, rounded in profile; snout having one papilla at its tip; eye – nostril distance 69.6 % of diameter of eye; nostrils not protuberant, directly lateral at level of lower jaw. Canthus rostralis straight, not elevated; loreal region concave lacking tubercles; lips rounded; internarial region slightly depressed; top of head flat with small conical tubercle between upper eyelids; upper eyelid with three small tubercles; its width 66.6 % of IOD; supratympanic fold slightly curved downward just posterior to orbit; tympanic membrane and tympanic annulus present, not covered by the supratympanic fold; two enlarged postrictal tubercles present. Choanae small, nearly round, not obscured by palatal shelf; dentigerous processes of vomers prominent, each process bearing four teeth; tongue as long as wide, its posterior border not notched, posterior third not adherent to floor of mouth; paired vocal slits present, longitudinal, lateral to base of tongue; median subgular vocal sac present. Skin on dorsum smooth, becoming finely shagreen on the flanks; skin on belly, throat, and ventral surfaces of the thighs areolate; discoidal fold present well anterior to groin; dorsolateral folds absent; cloacal sheath short; no tubercles in cloacal region. Three ulnar tubercles forming a row along the ventrolateral edge of the forearm; palmar tubercle not divided; thenar tubercle oval, slightly smaller than palmar tubercle; subarticular tubercles round, prominent, two on thumb and second finger and three on third and fourth fingers, and larger than supernumerary tubercles; supernumerary tubercles low, on all fleshy parts of palm; fingers having lateral fringes; relative lengths of fingers I <II <IV <III, all fingers having terminal ventral pads well defined by circumferential grooves; disc on thumb narrow, that on finger II slightly smaller than tympanic annulus; discs on fingers III and IV as wide as tympanic annulus; white, nonspinous nuptial pads are present on the dorsomedial surface of the base of the thumb. Hind limbs moderately robust; when hind limbs flexed perpendicular to axis of body, heels overlap; tibia length 50.4 % of SVL; foot length 46.2 % of SVL; heel and outer edge of tarsus lacking of tubercles; inner tarsal tubercle indistinct; inner metatarsal tubercle elongate, its length twice its width; oval outer metatarsal tubercle one-fourth size of inner; toes bearing lateral fringes and discs (about as wide as long) on expanded pads; basal webbing not encompassing basal subarticular tubercles, except on outer border of toe IV; relative lengths of toes I <II <III <V <IV; toe III much shorter than toe V; toe III extending to distal edge of the antepenultimate subarticular tubercle of toe IV; toe V reaching distal edge of penultimate subarticular tubercle of toe IV; all toes having terminal ventral pads well defined by circumferential grooves; discs of toes III – V equal to discs on fingers III – IV and larger than discs of toes I – II; subarticular tubercles prominent, rounded, two on toes I and II, three on toes III and V, and four on toe IV; supernumerary tubercles low, rounded, numerous on proximal segments of digits. In life, dorsum dull green, reddish brown, or olive brown (Fig. 1 A). Ventral surfaces of hind limbs pinkish lavender, throat and belly surfaces yellow. Iris pale creamy bronze with medial red streak (WED, field catalog, 31 May 1974). Measurements of holotype (mm). — SVL 24.0, tibia length 12.1, foot length 11.1, HL 9.0, head width 10.2, IOD 3.0, internarial distance 2.0, width of upper eyelid 2.0, diameter of eye 3.3, eye – nostril distance 2.3, diameter of tympanum 1.2.	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA7FFEBF5E1FA10FB5F0393.taxon	distribution	Distribution and ecology. — Pristimantis calima is known only from the type locality in lower montane rain forest in a deep valley with small, cascading streams (Fig. 4). The frogs were found on leaves, on the ground, and on tree trunks 2 – 3 m above the ground at night. During the 2 d and nights spent at the type locality on 31 May and 1 June 1975, temperature varied from 14 to 25 ° C. The observed daily rainfall was 19 mm and 4 mm, respectively.	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA7FFEBF5E1FA10FB5F0393.taxon	etymology	Etymology. — The specific name is a noun in apposition. It is the name of the type locality, the Río Calima. This locality is recognized as one of the most important archaeological reserves in Colombia because it contains the prints of an ancient Colombian civilization that was identified as ‘‘ Calima. ’’	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA7FFEBF5E1FA10FB5F0393.taxon	discussion	Remarks. — The humid Pacific slopes of the Cordillera Occidental and adjacent Chocoan lowlands harbor a great diversity of amphibians, especially centrolenid, dendrobatid, and strabomantid frogs. WED (with L. Trueb and J. E. Simmons) spent 3 d at the Río Calima site in September 1974 and 2 d there in May and June 1975. During these 5 d and nights they found five new species: Nŋmphargus prasinus (Duellman 1981), Gastrotheca dendronastes Duellman 1983, Hŋloscirtus simmonsi (Duellman 1989), Pristimantis calima, and P. diaphonus (Lynch 1986). Other sympatric species found were: P. achatinus (Boulenger 1898), P. erŋthropleura (Boulenger 1896), P. orpacobates (Lynch et al. 1994), N. grandisonae (Cochran and Goin 1970), and N. griffithsi (Goin 1961).	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA3FFE7F5F1FB3DFDDB01D7.taxon	description	(Figs. 5 A, B, 6 A, B; Table 2)	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA3FFE7F5F1FB3DFDDB01D7.taxon	materials_examined	Holotype. — KU 145021, adult female from the Laguna San Rafael, 3450 - m elevation (2 ° 23 ′ N, 76 ° 19 ′ W; datum ¼ WGS 84), Departamento del Cauca, Colombia, one of a series collected by Kjell von Sneidern in December 1938. Paratypes. — KU 145019 – 20, 145022, adult males collect-ed with the holotype.	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA3FFE7F5F1FB3DFDDB01D7.taxon	diagnosis	Diagnosis. — Pristimantis sneiderni is diagnosed by the following combination of characters: (1) skin on dorsum bearing low warts (most obvious on flanks); ventral skin areolate; discoidal fold present, well anterior to groin; dorsolateral folds and ridges absent in scapula region; (2) tympanic membrane not evident; tympanic annulus present, its upper edge covered by supratympanic fold, extending only just posterior to tympanic annulus; (3) snout round in dorsal view and in profile, with papilla at the tip (most obvious in males); canthus rostralis concave; (4) upper eyelid warty, narrower than IOD (50.0 – 80.0 % IOD); interocular tubercle absent; cranial crests absent; (5) choanae small, ovoid; not concealed by palatal shelf of maxillary arch; dentigerous processes of vomers absent; (6) males lacking vocal slits and nuptial pads; (7) finger I shorter than finger II; discs and circumferential grooves present on all fingers; discs truncate, except on finger I, which is round; disc of finger I smaller than that of finger II and this in turn smaller than discs on fingers III and IV; (8) fingers with narrow lateral fringes; palmar tubercle divided; thenar tubercle oval, same size as palmar tubercle; supernumerary tubercles low, distributed on all fleshy parts of palm; subarticular tubercles round, prominent, two on thumb and second finger and three on third and fourth fingers, and larger than supernumerary tubercles; (9) antebrachial tubercle present; other ulnar tubercles absent; (10) heel bearing one small tubercle; outer edge of tarsus bearing low tubercles; inner tarsal fold absent; (11) inner metatarsal tubercle elongate, its length twice its width; conical outer metatarsal tubercle onefourth the size of inner metatarsal tubercle; subarticular tubercles round, prominent, two on toes I and II, three on toes III and V, and four on toe IV; supernumerary plantar tubercles low, rounded, on proximal segments of toes; (12) toes bearing narrow lateral fringes; webbing absent; toe III shorter than toe V; toe III extending to distal edge of the antepenultimate subarticular tubercle of toe IV; toe V reaching the edge distal to the penultimate subarticular tubercle of toe IV; discs and circumferential grooves present on all toes; discs of toes III – V equal to discs on fingers II – IV and larger than discs of toe I; (13) color in preservative: dorsum and flanks tan with small dark brown markings; venter tan with brown mottling, mainly on chest, canthal stripe dark brown, throat tan with fine brown spots; posterior surfaces of thighs uniformly tan; (14) males smaller than females; SVL in three adult males 21.0, 22.0, and 23.0 mm, in one adult female 28.0 mm. Comparisons. — Pristimantis sneiderni differs from the other species in the genus by having subarticular tubercles beneath the joint between distal phalanges, dorsal skin with low warts, papilla at the tip of snout in adult males, and by lacking a tympanic membrane, dentigerous processes of vomers, and vocal slits. It is most similar to P. boulengeri and two Peruvian species, P. deŋi and P. schultei (Figs. 5, 6), by having tubercles beneath the joint between distal phalanges, dorsal skin with low scattered tubercles or warts, canthal stripe darker, papilla at tip of snout, tubercles or warts on upper eyelids, heel with one or more conical tubercles, low tubercles on medial surface of tarsus, dorsal coloration in preservative tan with darker brown markings (except P. schultei), and by being relatively large (except P. deŋi). From P. boulengeri, P. sneiderni is distinguished in having a round snout in profile (rather acutely round; Fig. 5 B, D), dorsal skin with low warts (rather than finely granular), and in lacking dentigerous processes of vomers and vocal slits. Pristimantis sneiderni differs from P. schultei by having a round snout in profile (rather than inclined posteroventrally; Fig. 5 B, F), and in lacking ulnar tubercles, dentigerous processes of vomers, and vocal slits. Pristimantis sneiderni is distinguished from P. deŋi by having a snout round in profile (rather than protruding and inclined posteroventrally) and in lacking weakly defined dorsolateral folds, dentigerous processes of vomers, and subgular vocal sac (Fig. 6). From the genus Pristimantis, other species with similar appearance to P. sneiderni are P. acuminatus (Shreve 1935), P. breƲifrons (Lynch 1981), P. enigmaticus Ortega-Andrade et al. (2015), P. limoncochensis Ortega-Andrade et al. (2015), P. omeƲiridis Ortega-Andrade et al. (2015), and P. pseudoacuminatus (Shreve 1935). However, P. sneiderni is distinguished from those species by lacking the dentigerous processes of vomers (dentigerous processes of vomers are present in those species). In addition, P. sneiderni differs from P. enigmaticus, P. limoncochensis, and P. omeƲiridis by lacking vocal slits (character state present in those species) and by having a dorsal skin with low warts (rather than the dorsal skin shagreened as in P. enigmaticus, or smooth as in P. limoncochensis and P. omeƲiridis). Additional characters that distinguish P. sneiderni from P. acuminatus and P. breƲifrons are the round snout in dorsal view (rather acuminate or subacuminate in those species) and the absence of vocal slits (present in P. breƲifrons). Finally, P. pseudoacuminatus is distinctively smaller (12.7 – 17.6 mm in males, 18.1 – 22.4 mm in females; Lynch 1980 a) when compared with P. sneiderni (21.0 – 23.0 mm in males, 28.0 mm in the only known female). Among species distributed on the southern portions of the Cordillera Central of Colombia, Pristimantis sneiderni could be confused with P. petersi (Lynch and Duellman 1980). However, P. sneiderni is larger (SVL 21.0 – 23.0 mm in adult males and 28.0 in one adult female) than P. petersi (SVL 14.5 – 19.9 in adult males and 20.3 – 23.1 mm in adult females; Lynch and Duellman 1980), and it lacks dentigerous processes of vomers and vocal slits (character states present in P. petersi).	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA3FFE7F5F1FB3DFDDB01D7.taxon	description	Description of the holotype. — An adult female with head slightly wider than body; head width 39.2 % of SVL; HL 32.1 % of SVL; snout rounded in dorsal view and in profile; eye – nostril distance 66.6 % of diameter of eye; nostrils not protuberant, directed slightly anterolaterally at level of lower jaw. Canthus rostralis curved, not elevated; canthal stripe dark brown; loreal region concave lacking tubercles; lips rounded; internarial region not depressed; top of head flat; upper eyelid warty, its width 50.0 % of IOD; supratympanic fold slightly curved downward only just posterior to tympanic annulus, tympanic membrane not evident, tympanic annulus present, upper fifth covered by the supratympanic fold; two nonenlarged postrictal tubercles. Choanae small, nearly round, not obscured by palatal shelf; dentigerous processes of vomers absent; tongue as long as wide, its posterior border not notched, posterior third not adherent to floor of mouth. Skin on dorsum bearing low warts (most obvious on flanks); skin on belly and throat areolate; skin on the ventral surface of the thighs, just below the cloaca bearing low warts; skin on posteroventral surfaces of thighs smooth; discoidal fold present, well anterior to groin; dorsolateral folds absent; cloacal sheath short. Antebrachial tubercle present; other ulnar tubercles absent; palmar tubercle deeply bifid; thenar tubercle oval, as wide as palmar tubercle; subarticular tubercles round, prominent, larger than supernumerary tubercles, two on thumb and second finger and three on fingers III and IV; supernumerary tubercles low, on all the fleshy parts of the palm; fingers bearing lateral fringes; relative lengths of fingers I <II <IV <III, all fingers having terminal ventral pads well defined by circumferential grooves; disc on thumb narrow, that on finger II slightly smaller than tympanic annulus; discs on fingers III and IV slightly larger than tympanic annulus. Hind limbs moderately robust; when hind limbs flexed perpendicular to axis of body, heels overlap; tibia length 50.0 % of SVL; foot length 50.0 % of SVL; heel bearing one low tubercle; outer edge and medial surface of tarsus bearing low tubercles; tarsal fold absent; inner metatarsal tubercle elongate, its length twice its width; conical outer metatarsal tubercle one-fourth size of inner; toes slender, bearing lateral fringes and discs (about as wide as long) on expanded pads; webbing absent; relative lengths of toes I <II <III <V <IV; toe III much shorter than toe V; toe III extending to distal edge of the antepenultimate subarticular tubercle on toe IV; toe V extending to edge distal of the penultimate subarticular tubercle on toe IV; all toes having terminal ventral pads well defined by circumferential grooves; discs of toes III – V equal to discs on fingers II – IV and larger than discs of toe I; subarticular tubercles round, prominent, two on toes I and II, three on toes III and V, and four on toe IV; supernumerary plantar tubercles low, rounded, numerous on proximal segments of digits. Measurements of holotype (mm). — SVL 28.0, tibia length 14.0, foot length 14.0, HL 9.0, head width 11.0, IOD 4.0, internarial distance 2.0, width of upper eyelid 2.0, diameter of eye 3.0, eye – nostril distance 2.0.	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA3FFE7F5F1FB3DFDDB01D7.taxon	distribution	Distribution. — Pristimantis sneiderni is known only from the type locality, the vicinity the Laguna de San Rafael, Páramo de Puracé, Cauca, Colombia (Fig. 4). The Laguna de San Rafael is located within the Puracé National Park, Cordillera Central, Colombia.	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA3FFE7F5F1FB3DFDDB01D7.taxon	etymology	Etymology. — The specific name is a noun in the genitive case and is a patronym for the naturalist, taxidermist, and collector Kjell von Sneidern (1910 – 1999), who first found the species during his explorations of Puracé, Cauca. Originally from Sweden, von Sneidern was director of the Museo de Historia Natural de la Universidad del Cauca from 1940 – 1960, and founder in 1954 of NoƲedades Colombianas, the journal published by the Museo de Historia Natural de la Universidad del Cauca.	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
894087A5FFA3FFE7F5F1FB3DFDDB01D7.taxon	discussion	Remarks. — Kjell von Sneidern visited the Laguna de San Rafael (southeast of Popayán) at the end of 1938 and the beginning of 1939. Together with the new species, von Sneidern collected Atelopus ebenoides Rivero 1963, Osornophrŋne bufoniformis (Peracca 1904), Hŋloxalus pinguis (Rivero and Granado-Díaz 1990), Pristimantis buckleŋi (Boulenger 1882), and P. obmutescens (Lynch 1980 b). WED (with L. Trueb and J. E. Simmons) in September 1974 visited several sites in Puracé, vicinity of the type locality. They found two species of bufonids (A. ebenoides and O. bufoniformis) and six species of Pristimantis — P. boulengeri, P. buckleŋi, P. leptolophus (Lynch 1980 b), P. mŋersi (Goin and Cochran 1963), P. obmutescens, and P. Ʋicarius (Lynch and Ruiz-Carranza 1983).	en	Ospina-Sarria, Jhon Jairo (2019): Two New Species of Pristimantis (Amphibia: Anura: Strabomantidae) from Southwestern Colombia. Herpetologica 75 (1): 85-95, DOI: 10.1655/d-18-00019
