identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
217887A7990C80354BE75A39FBAF6572.text	217887A7990C80354BE75A39FBAF6572.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachystethus Laporte 1832	<div><p>Brachystethus Laporte, 1832</p> <p>Edessa (Brachystethus) LAPORTE, 1832:63; BURMEISTER, 1835:356.</p> <p>Brachystethus; SPINOLA, 1837:343; AMYOT &amp; SERVILLE, 1843:160; HERRICH- SCHÄFFER, 1845:1; 1853:310; DALLAS, 1851:336; SIGNORET, 1851:344; STÅL, 1867:532 (key); 1872:48; WALKER, 1868:454; DISTANT, 1880:84; BERG, 1892:11; LETHIERRY &amp; SEVERIN, 1893:187 (cat.); BERGROTH, 1908:178; KIRKALDY, 1909:152 (cat.); PIRÁN, 1955:337; ROLSTON et al., 1980:122 (key); FROESCHNER, 1981:69 (cat.); BRAILOVSKY, 1988:129 (diagnosis, distr.); RIDER, 1991:584 (key).</p> <p>Daimonocoris KORMILEV, 1951:37; PIRÁN, 1955:337 (syn.).</p> <p>Type species of Brachystethus, Cimex geniculatus Fabricius, 1787, by monotypy; of Daimonocoris, D. bolivianus Kormilev, 1951.</p> <p>Diagnosis. Medium-sized, broad oval species (fig. 1), predominantly dull, but some species brightly colored (figs. 2-13). Head small, short, jugae contiguous before tylus; first antennal segment surpassing apexof head. Rostrum short, notsurpassing mesocoxae. Metasternum elevated in an anteriorly partially bifurcated and posteriorly bifurcated carina, extended over part of mesosternum. Abdomen with medial tubercle apposed to the posterior margin of metasternal carina.</p> <p>Description. Mediumsize (totallengthinmm,, 13-20.2,, 14.9-22.4). Broadoval shape. Head about one and a half wider than long, with transversal wrinkles on jugae and tylus, and small sinuated wrinkles between oceli. Punctures concentrated in 1+1 narrow, semi-circular and divergent stripesthat arise from base of oceli, directedforward until base of jugae, and in a medianarea between oceli, variable in length. Jugae rounded at apex and contiguous before tylus. Eyes large and prominent. Antennae with each segment progressively longer than the preceding. First antennal segment clearly surpassing apex of head. Bucculae elevated, anteriorly truncated, evanescent posteriorly. First rostral segment clearly surpassing them, by more than half of segment length. Rostrum not surpassing mesocoxae. Pronotum trapezoidal, strongly declivent onanterior half. Anterior margin slightly concave, openly U-shaped. Anterolateral angles prominent, forming a small tooth. Anterolateral margins slightly sinuous, subretilineous. Humeri rounded, little projected. Posterolateral and posterior margins straight. Punctures fine, more concentrated before cicatrices and regularly distributed over remaining disc of pronotum; among them, coarse punctures less frequent and irregularly distributed. Scutellum with broad and rounded apex, surpassing an imaginary line across middle of fifth connexival segment. Corium with punctures fine, dense and regularly distributed. Frenum slightly surpassing apical half of scutellum. Posterolateral angles rounded, attaining about the middle of sixth tergite. Membrane with more than five longitudinal and bifurcated veins. Tibiae dorsally sulcated. Ventral surface of thorax more densely punctured on propleura, mesoepisternum and metaepimerum. Evaporatory surface covering mesoepimerum and metaepisternum. Ostiolar rugae variable in length. Prosternum flat. Mesosternum slightly carinated. Metasternum carinated, producedover mesosternum. Anterior margin of metasternal carina with a deep median depression, partially bifurcated; posterior margin bifurcated, apposed to medial abdominal tubercle. Third urosternite with astout medial tubercle. Posterolateral anglesof connexivum slightly prominent. Spiracles elliptical. Trichobothria over or laterad to spiracularline.</p> <p>Male genitalia. Pygophore ovoid to sub-quadrangular, shallowly open dorsally. Dorsal rim (dor) medially concave, forming or not 1+1 lateral lobes (ldor) variable in size and shape among the species. Tenthsegment (X) trapezoidal, wider basaly than apically. Parameres (pa) uniramous. Genital cup with dorsally visible, blade-like processes (gcp). Posterolateral angles of pygophore rounded and tumid. Phallus: basal plate (bp) wide; phallotheca (ph) tubular, elongated, curved dorsad and strongly esclerotized; vesica (v) short, about 1/5 of lengthof phallotheca, with 1+1 dorsal, hook-like processes, convergent at apex, and only one ventral membranous lobe; ductus seminis distalis (dsd) short, notsurpassing apex of vesica; conjunctiva absent.</p> <p>Female genitalia. Laterotergites 8 and 9 (la8, la9) without apical spine. Spiracles on laterotergites 8 present. Laterotergites 9 not surpassing transverse band uniting laterotergites 8. Gonocoxites 9 (gc9) broad, trapezoidal, anterolateral angles extended in arms variable in length and thickness; anterior margin medially concave. Gonapophyses 9 (g9) with a medial thickening (mt). Thickening of vaginal intima (tvi) with a ventral, beak-like projection, visible in profile, bearing the orificium receptaculi (or). Chitinellipsen (ch) present. Ductus receptaculi (dr) thin anteriorly to vesicular area, and variable after that. Capsula seminalis (cs) with three digitiform processes, directed toward annular crests (aac, pac).</p> <p>Distribution. Mexico, Centraland South America.</p> <p>Comments. Brachystethus can be easily distinguished from other edessines by the broad, oval outline and the metasternal carina partially bifurcated. These characters, combined with the presence of blade-like processes on genital cup and of a medial thickening on gonapophyses 9, are diagnostic for the genus. The exam of the holotype of Ochlerus discolor, transferred to Brachystethus by KIRKALDY (1909), allowed to conclude that B. discolor should be removed from Brachystethus, since it presents morphological characters different from the remaining species of the genus, as the body outline, narrowly ovate, the first rostral segment concealed by bucculae and laterotergites 9 with acute apex, surpassing band joining laterotergites 8 dorsally. As the holotype presents one of the gonocoxites 8 deformed, and the metasternum can not be clearly observed because it is crossed by the pin, the species is here considered as incertae sedis.</p> </div>	https://treatment.plazi.org/id/217887A7990C80354BE75A39FBAF6572	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barcellos, Aline;Grazia, Jocélia	Barcellos, Aline, Grazia, Jocélia (2003): Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Sér. Zool. 93 (4): 413-446, DOI: 10.1590/S0073-47212003000400008
217887A7990980354AFB5FC4FBAE60EC.text	217887A7990980354AFB5FC4FBAE60EC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachystethus Laporte 1832	<div><p>Key to the species of Brachystethus</p> <p>1. Metasternal carina, in profile, strongly elevated anteriorly.....................................................2 Metasternal carina, in profile, flat or little elevated anteriorly................................................ 7</p> <p>2. Pronotum with a yellowish sub-callous border....................................................................... 3 Pronotum without a yellowish sub-callous border..................................................................5</p> <p>3. Venter with longitudinal series of yellowish spots. Fourth antennal segment medially swollen............................................................................................................. B. cribrus (Fabricius)</p> <p>Venter uniformly castaneous to dark-castaneous, lacking yellowish spots. Fourth antennal segment cylindrical............................................................................................................ 4</p> <p>4. Lobes of dorsal rim of pygophore projected, well developed (fig. 22). Posterior borders of gonocoxites 8 sinuate, not projected over laterotergites 8 (fig. 63)............................................................................................................................................... B. improvisus Breddin</p> <p>Lobes of dorsal rim of pygophore scarcely projected (fig. 23). Posterior borders the gonocoxites 8 projected in acute angle, over laterotergites 8 (fig. 64)........................ B. vicinus Signoret</p> <p>5. Hemelytra with a small yellowish callus, near and entad to apex of radial vein........................................................................................................................................... B. schuhi, sp. nov.</p> <p>Hemelytra without callus........................................................................................................ 6</p> <p>6. Apex of femora and base of tibiae yellowish.................................................... B. signoreti Stål</p> <p>Legs uniformly castaneous to black.................................................. B. rubromaculatus Dallas</p> <p>7. Ostiolar rugae short, extending for 1/3 the distance between inner margin ostiolum and lateral margin of metapleura. Metasternum carina with a medial, transversal sulcus................... 8</p> <p>Ostiolar rugae long, extending for 2/3 of distance between inner margin of ostiolum and lateral margin of metapleura. Metasternal carina lacking sulcus.................................................... 9</p> <p>8. Dorsal surface black, with red spots on hemelytra, in variable extension (fig.8).......................................................................................................................................... B. tricolor Bolívar</p> <p>Dorsal surface black, with yellow spots on hemelytra, in variable extension (fig. 9)................................................................................................................................. B. vexillum Breddin</p> <p>9. Coxae, trochanters, base and apex of femora, base of tibiae and tarsi yellowish.......................................................................................................................................... B. coxalis Breddin</p> <p>Legs uniformly castaneous to dark-castaneous.................................. B. geniculatus (Fabricius)</p></div> 	https://treatment.plazi.org/id/217887A7990980354AFB5FC4FBAE60EC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barcellos, Aline;Grazia, Jocélia	Barcellos, Aline, Grazia, Jocélia (2003): Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Sér. Zool. 93 (4): 413-446, DOI: 10.1590/S0073-47212003000400008
217887A79909803A4B185B62FCB86659.text	217887A79909803A4B185B62FCB86659.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachystethus rubromaculatus Dallas 1851	<div><p>Brachystethus rubromaculatus Dallas, 1851</p> <p>(Figs. 2, 3, 14, 24, 34, 44, 53-56, 65, 66)</p> <p>Brachystethus rubromaculatus DALLAS, 1851:337; SIGNORET, 1851:345; STÅL, 1862:118; MAYR, 1866:68; WALKER, 1868:454; STÅL, 1872:48; DISTANT, 1880:85, pl. 8, fig. 6; LETHIERRY &amp; SEVERIN, 1893:187 (cat.); KIRKALDY, 1909:152 (cat.); BRAILOVSKY &amp; BARRERA, 1982:239 (redescription); BRAILOVSKY, 1988:129, fig. 118 (diagnosis, distr.); BRAILOVSKY et al., 1992:179, pl. 24, figs. 1-9 (nymphs, biology); MAES, 1994:8 (cat.).</p> <p>Brachystethus sexmaculatus HERRICH- SCHÄFFER, 1853:310, fig. 1010; STÅL, 1862:118 (syn.).</p> <p>Type material of B. rubromaculatus. Lectotype, here designated, labeled: a) Honduras/45123; b) “a”; c) Type; d) Dyson coll. (BMNH); paralectotype, here designated, labeled: a) Mexico; b) “b” (BMNH). Type locality: Honduras and Mexico.</p> <p>Description. Dorsal surface black, withred spots of variable extension on pronotum, scutellum andcorium (figs. 2, 3). Headentirely black, with conspicuous wrinkles. Punctures sparse; semi-circular stripes little conspicuous, moderately punctured. Antennae black; fourth antennal segment medially swollen, with longitudinal sulcus. Second antennal segment about one and a half as long as first; third slightly longer than second; fourth almost twice as longas thirdand slightly shorter than fifth. Pronotum black, with a large, rounded, red spot medially; in some specimens, extended laterally over most part of disc, except on a narrow black stripe on anterior and anterolateral margins. Punctures black, minute, regularly distributed on the disk of pronotum; among them, sparser and larger punctures. Scutellum black, with a rounded, red spot, which occupies the center of median-apical region; in some specimens, it extends anteriorly and posteriorly, covering the apical 2/3 of scutellum. Surfaceof basalhalf sub-rugose, with large punctures regularly distributed. Apical half with fine punctures, moreconcentrated lateraland apically. Corium without callus, color pattern variable, usually black, with a large red spot on basal third and another on apical third, except along lateral margins. In some specimens, the corium is predominantly red, with black portion restricted to a thin lateral stripe and one small spot near to apex of radial vein. Ventral surface of thorax dark reddish-brown to black, with a red spot near to outer margin of metaepimerum; punctures fine, concolor. Ostiolar rugae extending by 2/3 of distance between the inner margin of ostiolum and lateral margin of metapleura. Metasternum, in profile, strongly elevated anteriorly. Legs dark reddish-castaneous to black. Connexival segments entirely black; punctures fine, moderate and regularly distributed. Venter black to dark reddish-castaneous, with 1+1 oblong red spots of variable length, generally extendingfrom anterior margin of fourth sternite onto a little bit beyond the anterior margin of seventh. In some specimens, the spots can become coalescent, making the abdomen almost totally red, only with a lateral black stripe. Punctures fine, very sparse, if present restricted to medial area. Spiracles black.</p> <p>Male genitalia. Pygophore trapezoidal, wider apically. Lobes of dorsal rim conspicuous, slightly projected posterad, rounded at apex. Genital cup processes moderately visible dorsally (figs. 14, 24). Parameres with wide apex, anterolaterally directed, a little wider than half length of tenth segment (fig. 44). Ventral rim broadly Vshaped (fig. 34). Phallus as described for the genus (figs. 53-55).</p> <p>Measurements (n=10). Headlength 2.08 (1.92-2.44) ± 0.15, width 3.02 (2.88-3.28) ± 0.14; interoculardistance 1.59 (1.52-1.76) ± 0.08; anteocularlength 0.68 (0.6-0.8) ± 0.07; lengthof antennalsegments: I - 0.81 (0.72-0.88) ± 0.06; II - 1.24 (1.04-1.48) ± 0.14; III - 1.45 (1.28-1.68) ± 0.11; IV - 2.79 (2.4-3.28) ± 0.24; V - 3.28 (2.96-3.8) ±0.25; pronotallength 4.36 (4.01-4.84) ± 0.25, width 9.74 (8.85-10.69) ± 0.53; scutellarlength 6.66 (6.01-7.35) ± 0.43, width 6.16 (5.68-6.76) ± 0.35; coriumlength 9.16 (8.52-10.02) ± 0.45; abdominalwidth 10.99 (10.02-12.36) ± 0.79; totallength 14.69 (13.78-15.87) ± 0.70.</p> <p>Female genitalia. Gonocoxites 8 with posterior borders convex andsutural borders parallel and contiguous, except at apex (fig. 56). Gonocoxites 9 more than three times wider across arms apices than long medially; anterior border broadly U-shaped. Arms of gonocoxites 9 with length equal to 1/4 of plate width across apices of arms. Thickening of vaginal intima digitiform. Ductusreceptaculi thin, about one and a half longer before than after vesicular area. Capsula seminalis globose. Annular crests convergent (figs. 65, 66).</p> <p>Measurements (n=10). Headlength 2.19 (2-2.44) ± 0.16, width 3.25 (3.08-3.44) ± 0.14; interoculardistance 1.71 (1.56-1.84) ± 0.09; anteocularlength 0.74 (0.64-0.92) ± 0.10;</p> <p>lengthof antennalsegments: I - 0.88 (0.76-0.96) ± 0.06; II - 1.40 (1.28-1.6) ± 0.10; III - 1.60 (1.28-1.92) ± 0.18; IV - 3.01 (2.76-3.36) ± 0.21; V - 3.41 (3.28-3.56) ± 0.09; pronotallength 4.91 (4.51-5.34) ± 0.34, width 10.97 (10.10-11.77) ± 0.63; scutellarlength 7.46 (6.68-8.18) ± 0.57, width 7.02 (6.35-7.60) ± 0.47; coriumlength 10.31 (9.35-11.36) ± 0.68; abdominal width 12.46 (11.27-13.69) ± 0.88; totallength 16.15 (15.03-17.37) ± 0.90.</p> <p>Material examined. MEXICO, Veracruz: Atoyac (Paso del Macho, 5 mi. Southwest),, 24,25. VI.82, tropical forest 2500’, Rolf L. Aalbu col. (DARC); Santiago (7 mi. North Tuxtla),, 8.VII.63 (DARC); (Est. Biol. Los Tuxtlas),, 4.VII.1988, C. Mayorga (AMNH);, 22.VIII.1982, C. O’Brien, L. O’Brien &amp; Wibmer, 250’ (DARC); Las Cabañas,, 28.V.1953, L. Vazquez (AMNH); Fortin,, 23.VII. 1989 (DARC); La Buena Ventura,, 14.VII.09, M.Y.B. (AMNH); Catemaco (5 mi. East),,, 2-4.VII.1971, Murray, Hart &amp; Schaffner (DARC); Lake Catemaco,, 23.VII.1955, P. &amp; C. Vaurie, 1000’ (AMNH);, 16.VI.1969, Bright &amp; Campbell (DARC); Yucatán: Chichén-Itzá,, 10.VIII.1990, D.A. Rider (DARC); Oaxaca: Matias Romero (17 mi. North),, 27.VI.1971, C.W. O’Brien (DARC); Tehuantepec: Isth. of Tehuantepec,, Sumichrast, Collection Ashmead (USNM); Cinta Lapa Valley,, V.44, E. Hernandez (USNM); Quintana Roo: Klan Nuevo,, 6, 7.VI.1959, P. &amp; C. Vaurie (AMNH); Tabasco: Villahermosa (5 mi. South),, 13.VI.65, Burke, Meyer &amp; Schaffner (DARC); Chiapas: Ruins at Palenque,, 26-30.VI.1959, P. &amp; C. Vaurie (AMNH); Huatimac,, V.1958, Patiño col. (MZSP). GUATEMALA, Garcia (Dona Maria Zacapa),, 20.IX.81 (DARC); Retalhuleu: S. Sebastian, 25 o,, L. Thiel (USNM); Alta Verapaz: Chacoj,, Champion col., ex Godman &amp; Salvin (BMNH); Verapaz, Teleman,, Champion, ex. Godman &amp; Salvin (AMNH). HONDURAS, Yoro: Subirana,, II, Stadelmann (AMNH). NICARAGUA, Zelaya: El Recreo,, X.1984, J.M. Maes (SEAN). COSTA RICA,, (AMNH); Guanacaste: La Cruz (6 mi. North),, 19.VI.1974, C.W. O’Brien, L. O’Brien &amp; Marshall (DARC); Alajuela: Zapote de Upala (vic. Bijagua),, 8.XII.1972, R. Ortiz (DARC); Heredia: Puerto Viejo,, 8.VIII.1965, G.W. Frankie (DARC); (2 Km South, La Sielva Biol. Sta.), 2, 2, 3-5.VI.1984, Riley, Rider &amp; Le Doux (DARC); Cartago: Iica (Turrialba),, VIII.1963, C.C. Porter (AMNH); Port Parker (Elena Bay),, 13.I.1938, Zcca Exped. Acc. 37483 (AMNH); Puntarenas: Santa Elena (6 Km South),, 6, 7.VI. 1983, J.E. Wappes (DARC); Ricon de Osa (Osa Peninsula), 7, 2, 14-26.VII.1969, Toby Schuh &amp; Janet Crane (AMNH); San José:, C.P. Dodge (AMNH). PANAMA, Chiriquí: David,, Champion (AMNH);, ex Godman &amp; Salvin (AMNH); V. de Chiriquí,, ex Godman &amp; Salvin, 25-4000ft (AMNH); Panama: Summit,, XII.1953, N.L.H. Krauss (AMNH); Madden Forest,, 18.V.1980, Riley &amp; Le Doux (DARC); F.T. Kobbe,, 28.VI.1974, C.W. O’Brien, L. O’Brien &amp; Marshall, at night (DARC). No data:, F 1672 (AMNH);, Brachystethus rubromaculatus det. Dallas, H.G.B., F 1673 (AMNH);, F 1672;, 1955 (MNHN); (MZSP).</p> <p>Distribution. Mexico (Veracruz, Yucatán, Oaxaca, Tehuantepec, Quintana Roo, Tabasco, Chiapas), Guatemala (Retalhuleu, Alta Verapaz), Honduras, Nicaragua, Costa Rica (Guanacaste, Alajuela, Heredia, Cartago, Puntarenas, San José), Panama (Chiriquí, Panama).</p> <p>Comments. Brachystethus rubromaculatus is the sister group of the clade formed by B. signoreti, B. cribrus and B. schuhi, sharing with these species the fourth antennal segment medially swollen, the metasternal carina strongly elevated anteriorly and the little projected lobes of dorsal rim of pygophore (BARCELLOS &amp; GRAZIA, 2003). It can be easily distinguished by the most frequent color pattern, dorsally black, with rounded red spots and venter black, with 1 + 1 spots extending from the anterior margin of fourth sternite until anterior margin of seventh. However, in specimens in which the red spots are coalescent, B. rubromaculatus can be confounded with B. signoreti, but easily distinguished by the color pattern of legs, uniformly black, the finer punctuation of scutellum and by genitalia, especially the size of parameres in relation to X segment. Legs uniformly black also distinguish B. rubromaculatus from B. cribrus and B. schuhi. Brachystethus signoreti Stål, 1872</p> <p>(Figs. 4, 5, 15, 25, 35, 45, 57, 67)</p> <p>Brachystethus geniculatus SIGNORET, 1851:345 (preoccupied, not Fabricius, 1787); WALKER, 1868:455.</p> <p>Brachystethus signoretii STÅL, 1872:48 (new name); KIRKALDY, 1909:152 (cat.); GRAZIA, 1984:74.</p> <p>Brachystethus signoreti; LETHIERRY &amp; SEVERIN, 1893:187 (cat.).</p> <p>Typematerial. Lectotype, heredesignated, labeled: a) Laguaira, Coll. Signoret; b) geniculat. det. Signoret (NHMW); paralectotype, heredesignated, labeled: a) Laguaira, Coll. Signoret; b) geniculata det. Signoret (NHMW). Typelocality, Venezuela (“ LaGuaira ”).</p> <p>Description. Dorsal surface dark- to reddish-castaneous (figs. 4, 5). Head dark- to reddish-castaneous, with conspicuous wrinkles. Punctures of median stripe sparse; semicircular stripes moderately punctured. Antennae black; fourth antennal segment medially swollen, with a longitudinal sulcus. Second antennal segment about one and a half as long as first, third about one and a quarter the second, fourth almost twice the third and fifth slightly longer than fourth. Pronotum dark- to reddish-castaneous. Punctures fine regularly distributed, interposed with coarse punctures, more concentrated behind cicatrices; medium-sized punctures densely distributed before cicatrices. Scutellum concolor with pronotum; coarse punctures distributed on basal 3/4, among them, fine punctures, denserlaterally. Coriumwithout callus, dark- to reddish-castaneous; on the reddish-castaneous specimens, present two darker, grayish stripes, along outer margin and entad to radial vein. Ventral surface of thorax with black, medium-sized punctures. Ostiolar rugae extendingby 2/3 of distance between inner margin of ostiolum and lateral margin of metapleura. Metasternum, in profile, strongly elevated anteriorly. Legs dark-castaneous to black, yellowish at apexof femora and baseof tibiae. Connexival segments dark- to reddish-castaneous. Venter dark- to reddish-castaneous, in this case with reddish areas around and entadto spiracles andlittle less conspicuous sub-medially. Punctures fine, concolor, regularly distributed, but rare on medial fifth of abdomen. Spiracles black.</p> <p>Male genitalia. Pygophore trapezoidal, wider apically. Lobesof dorsal rim conspicuous, little projected posterad, rounded at apex. Genital cup processes moderately visible dorsally (figs. 15, 25). Parameres very wide apically; apexantero-laterad directed, slightly longerthan mediallengthof tenthsegment (fig. 45). Ventralrimopenly U-shaped (fig. 35).</p> <p>Measurements (n=10). Headlength 1.93 (1.84-2.04) ± 0.06, width 2.95 (2.88-3.12) ± 0.09; interoculardistance 1.55 (1.44-1.6) ± 0.06; anteocularlength 0.66 (0.60-0.72) ± 0.05; lengthof antennalsegments: I - 0.82 (0.80-0.88) ± 0.03; II - 1.22 (1.08-1.4) ± 0.08; III - 1.51 (1.24-1.68) ± 0.15; IV - 2.93 (2.72-3.2) ± 0.17; V- 3.35 (3.28-3.44) ± 0.07; pronotallength 4.23 (4.01-4.51) ± 0.18, width 9.44 (8.85-10.10) ± 0.39; scutellarlength 6.42 (6.01-6.93) ± 0.31, width 6.01 (5.68-6.35) ± 0.22; coriumlength 9.19 (8.85-9.69) ± 0.30; abdominalwidth 10.55 (10.02-11.02) ± 0.35; totallength 14.35 (13.78-15.03) ± 0.46.</p> <p>Female genitalia. Posterior borders of gonocoxites 8 convex over laterotergites 9. Sutural borders of gonocoxites 8 parallel and contiguous, except at apex (fig. 57). Gonocoxites 9 almost three and a half times wider across arms apices than long medially; anterior border broadly U-shaped. Length of arms equalto 1/6 of plate width. Thickening of vaginal intima digitiform. Ductus receptaculi thickafter vesiculararea, and about one and a half longer before than after this area. Capsula seminalis elongated, digitiform. Annular crestsparallel (fig. 67).</p> <p>Measurements (n=10). Headlength 2.09 (1.92-2.28) ± 0.12, width 3.11 (3.04-3.20) ± 0.06; interoculardistance 1.61 (1.52-1.68) ± 0.06; anteocularlength 0.70 (0.60-0.80) ± 0.06; lengthof antennalsegments: I - 0.94 (0.84-1.12) ± 0.08; II - 1.40 (1.28-1.52) ± 0.08; III - 1.68 (1.36-1.84) ± 0.15; IV - 3.18 (2.76-3.36) ± 0.18; V - 3.62 (3.56-3.68) ± 0.08; pronotallength 4.72 (4.51-5.01) ± 0.14, width 10.60 (10.19-11.02) ± 0.25; scutellarlength 7.18 (6.93-7.35) ± 0.13, width 6.75 (6.51-7.01) ± 0.20; coriumlength 10.27 (9.94-10.69) ± 0.22; abdominal width 11.87 (11.44-12.36) ± 0.28; totallength 16.30 (15.53-17.20) ± 0.56.</p> <p>Material examined. VENEZUELA,, pres. by Perth Museum (BMNH 1953-629);, Schauf. col., Coll. Mayr (NHMW); San Julian, 2, VII.1900, W. Robinson Bequest (USNM 1929); Rancho Grande, 3, 2, 19,26,27. IV.1945 (AMNH); Aragua: Villa de Cura (Est. Exp. Cataurito, (± 1000m),, 27.X.81, J.L. Garcia (IZAV); Maracay (450m, en luz), 2, 22.V.1963, E. Osuna (AMNH, IZAV); Mérida: Mérida, 4, 3 (AMNH); Tachira: San Cristobal (River Black),, 7.III.1980, Miguel G. (IZAV);, 5.I.1982, Gaban (IZAV); (Colon),, 1.IV.1982, Gaban (IZAV);, 10.X.1981, Ramirez (IZAV). ECUADOR, Pichincha: Santo Domingo de los Colorados (amazonian rainforest),,, X.1981, G. Onore (BMNH 1982-246); La Union, 2,, 12.VII.1984, R. Haswell (DARC); Cotopaxi: (1500 m, tropical rainforest general collecting),, X.1982, G. Onore, (BMNH, 1982-246); Paramba,, “ Coll. I.R. Sc. N.B. ” (DARC); 4, Rosemberg col. (USNM); (3500’, dry season),, II.97, Rosenberg (BMNH); Balrahamba,, F. Campos (USNM);, 18.V.1935, F. Campos; J.C. Lutz Collection (USNM 1961); Pastaza: Canelos,, Schouteden (DARC); Tungurahua: Ambato,, 2, Coll. Noualhier 1898 (MNHN);, E. de Ville &amp; M.R.Belg col., Coll. Noualhier 1898 (MNHN). No data: (BMNH);, Coll. Mayr (NHMW).</p> <p>Distribution. Venezuela (Aragua, Mérida, Tachira), Ecuador (Pichincha, Cotopaxi, Pastaza, Tungurahua).</p> <p>Comments. Brachystethus signoreti is the sister group of the clade composed by B. cribrus and B. schuhi, sharing the homoplastic character of color pattern of legs, apex of femoraand base of tibiae yellowish (BARCELLOS &amp; GRAZIA, 2003). Itcan bedistinguished from these species by the absence of calli on the corium and from B. cribrus, especially, by the color of venter, lacking yellow spots. Brachystethus signoreti is also close to B. rubromaculatus, distinguished by the color pattern of legs, the coarser punctures of scutellum and, on male genitalia, especially by the larger parameres.</p> </div>	https://treatment.plazi.org/id/217887A79909803A4B185B62FCB86659	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barcellos, Aline;Grazia, Jocélia	Barcellos, Aline, Grazia, Jocélia (2003): Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Sér. Zool. 93 (4): 413-446, DOI: 10.1590/S0073-47212003000400008
217887A79906803E4B2B5C28FC66655F.text	217887A79906803E4B2B5C28FC66655F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachystethus cribrus (Fabricius 1781)	<div><p>Brachystethus cribrus (Fabricius, 1781)</p> <p>(Figs. 6, 16, 26, 36, 46, 58, 68)</p> <p>Cimexcribrum FABRICIUS, 1781:357; 1794:117; GMELIN, 1793:2152.</p> <p>Edessa cribrum; FABRICIUS, 1803:154; BURMEISTER, 1835:356.</p> <p>Brachystethus cribrum; AMYOT &amp; SERVILLE, 1843:161; HERRICH- SCHÄFFER, 1845:1, fig. 788; DALLAS, 1851:337; HERRICH- SCHÄFFER, 1853:310; SIGNORET, 1851:344; WALKER, 1868:455; STÅL, 1872:48; LETHIERRY &amp; SEVERIN, 1893:187 (cat.); KIRKALDY, 1909:152 (cat.); PIRÁN, 1956:31.</p> <p>Brachystethus cribrus; FROESCHNER, 1981:69 (cat.).</p> <p>Typematerial. Holotype, labeled: Cimexcribrum F., det. P.H. V. Doesburg (ZMUC). The holotype of C. cribrum had been considered lost by the curator of ZMUC. ZIMSEN (1964) lists three syntypes of Cimex geniculatus and none of C. cribrum. The exam of one of the supposed syntypes of C. geniculatus showed that it had been identified as C. cribrum by Peter Von Doesburg, from Leiden Museum. In fact, this specimen presents the apex of femoraand the base of tibiae yellowish, aswell asyellowish spotson abdomen, fitting the Fabricius description of C. cribrum (“abdominis punctis genubusque flavis...”), that may indicate a mistake on the organization of Fabricius types, previously to ZIMSEN (1964) catalog. Thus, this specimen is here considered the holotype of Brachystethus cribrus. Typelocality: Suriname.</p> <p>Description. Dorsal surfacedark reddish-castaneous, witha thin marginal yellowish stripe (fig. 6). Head dark-castaneous, sometimes with yellow margin; wrinkles conspicuous. Punctures of median stripe sparse, more frequent on 1+1 semi-circular stripes and jugae. Antennae dark-castaneous to black; fourth antennal segment medially swollen, with longitudinal sulcus. Second antennal segment one and a half to almost twice as long as first; second and third subequal, fourth more than twice as long as third; fifth the longest, slightly longer than fourth. Pronotum dark reddish-castaneous, with yellowish stripe on anterolateral margins. Surface rugose; coarse punctures behind cicatrices, sparse over the disc. Punctures fine among the coarser ones, concentrated before cicatrices. Scutellum reddish-castaneous with coarse punctures more frequent on basal 2/3. Corium reddish-castaneous withyellowish callus lateral and entad theapex of radial vein. Ventral surface of thorax dark reddish-castaneous with medium-sized punctures. Ostiolar rugae extending by 2/3 of distance between inner margin of ostiolum and lateral margin of metapleura. Metasternum, in profile, strongly elevated anteriorly.</p> <p>Legs dark-castaneous, yellowish at apex of femora and base of tibiae. Connexival segments dark- to reddish-castaneous, narrowly bordered with yellow or light reddishcastaneous, over marginal sixth of each segment. Punctures fine, more concentrated entad to this stripe. Venter dark-castaneous, lateral margins with yellowish subcaloused stripe. Entad to spiracles, two longitudinal series of rounded yellow spots, one per segment, disposed in two rows: one lateral, on sternites III to VII and one median on sternites III to VI. This pattern varies intraspecifically; in some specimens, the external line presents two spots per segment on sternites III to VI. In some darker specimens, the spots may become less visible. Punctures fine, more concentrated along posteriormargin of sternites and on lateral thirds of venter. Spiracles yellowish-castaneous to black.</p> <p>Male genitalia. Pygophore trapezoidal, wider apically. Lobes of dorsal rim inconspicuous. Genital cup processes broadly visible dorsally (figs. 16, 26). Parameres distally wide (fig. 46). Ventral rim broadly V-shaped (fig. 36).</p> <p>Measurements (n=10). Headlength 1.90 (1.76-2.04) ± 0.09, width 2.98 (2.88-3.08) ± 0.07; interoculardistance 1.50 (1.44-1.60) ± 0.07; anteocularlength 0.65 (0.56-0.72) ± 0.05; lengthof antennalsegments: I - 0.90 (0.80-0.96) ± 0.05; II - 1.32 (1.04-1.52) ± 0.13; III - 1.44 (1.36-1.60) ± 0.09; IV - 3.07 (2.64-3.68) ± 0.32; V - 3.42 (2.96-4.08) ± 0.33; pronotallength 3.97 (3.67-4.26) ± 0.18, width 9.53 (9.19-10.02) ± 0.27; scutellarlength 6.45 (6.18-6.68) ± 0.20, width 6.04 (5.68-6.51) ± 0.22; coriumlength 8.98 (8.52-9.35) ± 0.32; abdominalwidth 10.30 (9.77-11.02) ± 0.39; totallength 14.14 (13.53-14.78) ± 0.44.</p> <p>Female genitalia. Posterior borders of gonocoxites 8 straight over laterotergites 9. Sutural borders of gonocoxites 8 parallel (fig. 58). Gonocoxites 9 about three timeswider across arms apices than long medially; anterior border broadly U-shaped. Arms of gonocoxites 9 short, length equal to 1/6 of plate width. Thickening of vaginal intima ogival. Ductus receptaculi thin after vesicular area; slightly more than one and a half longer before than after this area. Capsula seminalis elliptical. Annular crests convergent (fig. 68).</p> <p>Measurements (n=10). Headlength 2.11 (2.00-2.40) ± 0.12, width 3.20 (3.04-3.36) ± 0.09; interoculardistance 1.66 (1.60-1.76) ± 0.07; anteocularlength 0.68 (0.56-0.80) ± 0.07; lengthof antennalsegments: I - 0.85 (0.72-1.12) ± 0.11; II - 1.49 (1.28-1.68) ± 0.14; III - 1.60 (1.48-1.76) ± 0.09; IV - 3.39 (2.88-3.64) ± 0.25; V - 3.67 (3.20-3.92) ± 0.25; pronotallength 4.79 (4.18-6.18) ± 0.58, width 10.69 (9.85-11.61) ± 0.51; scutellarlength 7.37 (6.68-7.85) ± 0.35, width 6.81 (6.26-7.35) ± 0.31; coriumlength 10.14 (9.27-10.69) ± 0.57; abdominal width 11.92 (11.02-12.86) ± 0.62; totallength 16.22 (14.86-17.62) ± 0.83.</p> <p>Material examined. GUYANA,,, 1943, Bartlett col. (BMNH 1943-60, 1009);,, 20.II.1937, R. Pomaron &amp; A.S. Pinkus (AMNH); Kartabo (Distrito Bartica),, 1922 (BMNH 1954- 631); 3, 4, 13. VI, 16, 30.VII, 11.VIII.1922, M.D. Haviland, d.d. Collegium Newnhamense (BMNH 1954-631). SURINAME, Brokopondo: Brownsberg, 3, 3, 19-21.VIII.1975, L.H. Rolston (DARC); Brownsberg Nat. Pk.,, 19-22.VII.1975, D. Engleman (AMNH). VENEZUELA, Amazonas: Santa Lucía,, 15-21.XI.1982, A. Chacon &amp; G.Yepes (IZAV). BRAZIL, Amazonas: Tefé,, Raulin, Thayer Expedition (AMNH); Benjamin Constant (Rio Javari),, 3, 1.II-15.III.1942, August Rabaut (AMNH); Rio Caiary-Uaupes,, 1906, H. Schmidt (AMNH); Rio Negro (Tupurumara),, 9.II.1963, J. &amp; B. Bechyné (MPEG); Pará: Peixe-Boi,, 14.IV.1977, W. L. Overal. (MPEG);,, P. Waldir (MPEG); Jacareacanga,, X.69, R. Barbosa (AMNH); Itaituba (Santarensinho, Rio Tapajós), 2,, V.1962, Dirings (MZSP); Santa Maria do Guamá,, 2, 26.II.1987, P. Tadeu, J. Dias (MPEG);, 26.II.1987 (MPEG); Belém (Mocambo), 2, 21.XII.1978, A. Ramos (MPEG); Óbidos (bx/ Amazonas),, Dirings (MZSP); Bujaru,, 20.IV.1979, R.B. Neto (MPEG); Maranhão: Alto Turiaçu (Aldeia Gurupluna),, 19-22.II.1990, Helder Queiroz (MPEG); Mato Grosso: 10 o 25’S 59 o 28’W (300m),, 17-22.III.1977, D. Engleman (AMNH); Rondônia: Ariquemes (62 Km SW, Fazenda Rancho Grande),, 6-15.XII.1990, D.A. Rider &amp; J.E. Eger (DARC); (Linea C-20, 7 Km E. B-65),, 11-22.XI.1991, T.J. Riley (DARC); Ouro Preto do Oeste (Reserva INPA),, 26.III.1985, F.F. Ramos (MPEG); Vilhena,, 17.XII.1986, C. Elias (DZUP); Pernambuco: Cabo,, Distant Collection (BMNH 1911-383); Goiás: Goiânia,, I.1936, Borgmeier &amp; S. Lopes (UFRG). ECUADOR, Pichincha: Puerto Quito,, V.1982, G. Onore (BMNH 1982-246); Napo: Coca,,, XI, XII.1982, G. Onore (BMNH 1983-255); (tropical rainforest general collecting), 3,, XI.1982, G. Onore (BMNH 1982-246); (Pompeya on Napo R., 40 km from Coca),, V.1965, L.E. Pena (AMNH); Loja: S.E. Loja (Rio Jumboe, tributary Zamora R.),, 1-IV-1965, idem (AMNH); Zamora Chinchipe: Zamora (35 Mi Loja),,, 1-5.IV.1965, idem (AMNH). PERU, Ucayali: Pucallpa (Alt. 600 ft.),, 11.XI.1946, J.C. Pallister (AMNH); La Libertad: Cumpang (above Uctubamba, 2625m),, 13.X.1979, L. J. Barkley (DARC); Huánuco: 2, 23.XI.1946, Alt. 2200 ft., J.C. Pallister col. (AMNH); Tingo María (Alt. 2200 ft.),, 2, 23.IX, 21.X.1946, J.C. Pallister col. (AMNH); Cuzco: Cosñipata,, 6, 7.X.1966, Carrasco (UFRG); Pillcopata,, 11.X.68 (UFRG); Junín: Jauja Prov., Satipo,, V.1945, P. Paprzycki (AMNH); Rio Santiago,, 23.XI.1924, F. 6012, H. Bassler Collection Acc. 33591 (AMNH); Rio Morona,, 17.I.1925, F. 6194, H. Bassler Collection Acc. 33591 (AMNH); Upper Rio Maranon, 2, 26.VIII, 11.X.1924, F.6177, 6162, H. Bassler Collection Acc. 33591 (AMNH); Middle Rio Maranon,, 8.XI.1925, H. Bassler Collection Acc. 33591 (AMNH); Tarapoto Region,, 6.I.1926, F. 6042, H. Bassler Collection, Acc. 33591 (AMNH); Achinamiza,, 3, 14.X, 20.XI.1927, F. 6001, H. Bassler Collection Acc. 33591 (AMNH); Moyobamba Region,, 2, 12.I.1925, 8.I.1926, 5.XII.1945, F. 6097, F. 6149, F. 6072, H. Bassler Collection Acc. 33591 (AMNH). BOLIVIA, 3, 2, 1904, J. Steinbach col. (BMNH 1904 -311);, 1898, Coll. Noualhier (MNHN); Santa Cruz: Buena Vista Prov., Tohito, 400m, 2, 1956, Franz Steinbach (AMNH); Cochabamba: Rio Cristal Mavu (50 mi NE Cochabamba), 4, 4, 30.IX, 27.X, 12, 30.XI. 1949, L. Pena (AMNH, DARC). No data:,, (AMNH); San Jose, (MRSN).</p> <p>Distribution. Guyana, Suriname (Brokopondo), Venezuela (Amazonas), Brazil (Amazonas, Pará, Maranhão, Mato Grosso, Rondônia, Pernambuco, Goiás), Ecuador (Pichincha, Napo, Loja, Zamora Chinchipe), Peru (Ucayali, La Libertad, Huánuco, Cuzco, Junín), Bolivia (Santa Cruz, Cochabamba).</p> <p>Comments. Brachystethus cribrus is the sister species of B. schuhi, sharing a homoplasticcharacter, presence of yellowish calli oncorium (BARCELLOS &amp; GRAZIA, 2003). Brachystethus cribrus is separatedfrom all other congeners by autapomorphic characters: abdomen with yellow spots and genital cup processes widely visible dorsally. It can also be distinguished from B. schuhi by homoplastic characters – pronotum with yellowish border, yellowish coxae and the inconspicuous lobes of dorsal rim. The yellowish margin of pronotum and the color pattern of coxae, femora and tibiae of B. cribrus are similar to those of B. coxalis, B. improvisus and B. vicinus, but can be easily distinguished by the yellow spots on the ventral surface of abdomen, the fourth antennal segment medially swollen and, on male genitalia, by the genital cup processes widely visible dorsally.</p> </div>	https://treatment.plazi.org/id/217887A79906803E4B2B5C28FC66655F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barcellos, Aline;Grazia, Jocélia	Barcellos, Aline, Grazia, Jocélia (2003): Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Sér. Zool. 93 (4): 413-446, DOI: 10.1590/S0073-47212003000400008
217887A79902803E4BE65F2FFEF861C0.text	217887A79902803E4BE65F2FFEF861C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachystethus schuhi Jocélia Grazia 2003	<div><p>Brachystethus schuhi sp. nov.</p> <p>(Figs. 7, 17, 27, 37, 47)</p> <p>Etymology. Named in honor to Dr. Randall T. Schuh, for his great contribution to the knowledge of Heteroptera.</p> <p>Typematerial. Holotype, Guyana, nolocality, nodate, Babcock (AMNH). Type locality: Guyana.</p> <p>Description. Dorsal surface dark- to reddish-castaneous (reddish on apical 3/4 of corium) (fig. 7). Headentirely dark-castaneous, almost black; wrinkles conspicuous. Punctures densely distributed. Antennae black. Second antennal segment more than one and a half longer than first and as long as third; fourth and fifth lacking on the holotype. Pronotum dark-castaneous. Punctures coarse, moderately distributed, more concentrated behind cicatrices; finer punctures, densely distributed, before cicatrices. Scutellum dark-castaneous; coarse punctures on basal 2/3, more concentrated along lateral margins. Corium reddish, excepton basal fourth, dark-castaneous, almost black; a yellowish callus near and entad apex of radial vein. Ventral surface of thorax darkcastaneous, almost black; finepunctures regularly distributedon propleura, mesoepisternum and metaepimerum. Ostiolar rugae reaching about half the distance between inner margin of ostiolum and lateral margin of metapleura. Metasternal carina, in profile, strongly elevated anteriorly. Legs dark-castaneous, almost black, yellowish-castaneous at apex of femora and baseof tibiae. Connexivalsegments dark-castaneous, withfinepunctures regularly distributed. Venter uniformly dark-castaneous, almost black. Punctures fine, distributed more densely among spiracles, and a narrow, impuctatedmedial stripe. Spiracles black.</p> <p>Male genitalia. Pygophore subquadrangular, slightlywider apically. Lobes of dorsal rim little projected posterad, rounded at apex. Genital cup processes moderately visible dorsally (figs. 17, 27). Paramereswith wide apex, laterally directed (fig. 47). Ventral rim openly V-like (fig. 37).</p> <p>Measurements (n=1). Headlength 1.96, width 3.08; interoculardistance 1.52; anteocularlength 0.60; lengthof antennalsegments: I - 0.84; II - 1.36; III - 1.36; IV and V - lacking; pronotallength 4.34, width 10.19; scutellarlength 6.68, width 6.51; corium length 9.85; abdominalwidth 11.02; totallength 15.70.</p> <p>Distribution. Guyana.</p> <p>Comments. Brachystethus schuhi shares with its sister species B. cribrus the yellowish callus on corium, but can be distinguished by the following characters: venter uniformly castaneous, dark-castaneous coxae and genital cup processes moderately visible dorsally. Brachystethus schuhi is also closely related to B. signoreti and B. rubromaculatus (BARCELLOS &amp; GRAZIA, 2003), being distinguished from the the former by the callus on corium and male genitalia, and from the latter by the yellowish apex of femora and base of tibiae.</p> </div>	https://treatment.plazi.org/id/217887A79902803E4BE65F2FFEF861C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barcellos, Aline;Grazia, Jocélia	Barcellos, Aline, Grazia, Jocélia (2003): Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Sér. Zool. 93 (4): 413-446, DOI: 10.1590/S0073-47212003000400008
217887A7991D80234BE35EFCFD546630.text	217887A7991D80234BE35EFCFD546630.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachystethus tricolor Bolívar 1879	<div><p>Brachystethustricolor Bolívar, 1879</p> <p>(Figs. 8, 18, 28, 38, 48, 59)</p> <p>Brachystethustricolor BOLÍVAR, 1879:138; LETHIERRY &amp; SEVERIN, 1893:187 (cat.); KIRKALDY, 1909:152 (cat.); FROESCHNER, 1981:69 (cat.).</p> <p>Typematerial. Lectotype, heredesignated, labeled: a) Baeza (Ecuador), 4/mar - 20/abr/1865, F. Martinez; b) Expedición al Pacífico (1862-1865); c) M.N.C.N., Madrid; d) Brachystethustricolor Bol. ?; e) sintipo? (MNCS); paralectotypes,, both labeled: a) Martinez Baeza / 481; b) Baeza (Ecuador), 4/mar - 20/abr/1865, F. Martinez; c) Expedición al Pacífico (1862-1865); d) M.N.C.N., Madrid; e) Brachystethustricolor Bol. ?; f) sintipo? (MNCS). Typelocality: Baeza, Ecuador. (“Baeza Americae Equatorialis”).</p> <p>Description. Dorsal surface black, except corium and, in some specimens, part of pronotum, reddish (fig.8). Head entirely black, with conspicuous wrinkles. Mediumsized punctures moderate to sparsely distributed. Antennae black; fourth antennal segment cylindrical. Second antennal segment slightly longer than first and about 3/4 as long as third; third slightly morethan half of fourth; fifth the longest, a little less than one and a half as long as fourth. Pronotum entirely black, or, in some specimens, with about basal 2/3 reddish. Anterolateral margins entirely black or with a thin subcaloused, yellowish border. Medium-sized punctures concentrated before cicatrices. Some specimens with verycoarse punctures on basal 2/3, decreasing in size posterad. Scutellum black, sometimes slightly reddish in the middle. Coarse punctures, in variable density, decreasing in size toward apex. Remaining punctures medium-sized to fine, distributed allover surfaceof scutellum. Corium reddish, except along costal margins, with a narrow black stripe variable in length, and a little black spot, irregular-shaped, entad to apex of radial vein. Yellowishcallus nearand entad apex of radial vein, lackingin some specimens. Costal margins, in some specimens, with a thin, subcalousedyellowish border, evanescent posterad. Ventral surface of thorax black, sometimes reddish-castaneous on propleura and near coxae. Punctures concolor. Ostiolar rugae short, extending by about 1/3 of distance between inner margin of ostiolum and lateralmargin of metapleura. Metasternum, in profile, little elevated anteriorly; transversal sulcus in the middle. Legs black, coxae reddish-castaneous. Connexivum mostly black; in some specimens, a reddish stripe of variable width along inner margin. Venter red, with rounded black spots around each spiracle and one black continuous stripe along lateral margins, variable in width, occupying one fourth to half the distance between spiracles and lateral margin of abdomen. Some specimens present, on sternites III to VI, entad to spiracles, elliptical black spots, variable in size. Anterior and posterior margins of sternites occasionally with blackish stripes. Punctures sparse, fine and concolor with venter, more concentrated over black spots entad to spiracles. Spiracles black.</p> <p>Male genitalia. Pygophore quadrangular, equally wide at apex and base. Lobes of dorsal rim conspicuous, strongly projected posterad, rounded at apex. Genital cup processes little visibledorsally (figs. 18, 28). Parameres with acute apex, turned toward tenth segment (fig. 48). Ventral rim not so openly V-like as in other speciesof the genus (fig. 38).</p> <p>Measurements (n=4). Headlength 1.80 (1.72-1.87) ± 0.06, width 2.71 (2.61-2.77) ± 0.07; interoculardistance 1.46 (1.37-1.56) ± 0.08; anteocularlength 0.60 (0.56-0.62) ± 0.03; lengthof antennalsegments: I - 0.97 (0.94-1.01) ± 0.03; II - 1.12 (1.05-1.20) ± 0.06; III - 1.44 (1.25-1.56) ± 0.14; IV - 2.53 (2.50-2.56) ± 0.03; V - 3.44 (3.39-3.48) ± 0.06; pronotal length 3.86 (3.34-4.09) ± 0.35, width 9.14 (8.52-9.69) ± 0.50; scutellarlength 5.95 (5.52-6.35) ± 0.41, width 5.72 (5.36-6.01) ± 0.28; coriumlength 8.64 (8.52-8.77) ± 0.18; abdominalwidth 10.23 (9.69-10.69) ± 0.46; totallength 14.15 (13.53-14.86) ± 0.55.</p> <p>Female genitalia. Posterior borders of gonocoxites 8 in an obtuse angle near limit between laterotergites 9 and gonocoxites 9. Sutural borders of gonocoxites 8 parallel and contiguous, except at apex (fig. 59).</p> <p>Measurements (n=1). Headlength 1.87, width 2.89; interoculardistance 1.56; anteocularlength 0.62; lengthof antennalsegments: I - 1.17; II - 1.29; III - 1.40; IV - 2.77; V - 3.59; pronotallength 3.84, width 9.69; scutellarlength 6.24, width 6.00; coriumlength, not measured; abdominalwidth 11.19; totallength 15.20.</p> <p>Material examined. ECUADOR,, “71”, E. de Ville col., “Coll. R.I. Sc. N.B.” (DARC).</p> <p>PERU, Ucayali: Aguaytía (295m, low jungle),, 13.IX.1946, F. Woytkowski (USNM); Huánuco: Chinchao (25 Km below Carpish, 2500m),, 13.IX.1946, F. Woitkowski (AMNH).</p> <p>Distribution. Ecuador, Peru (Ucayali, Huánuco).</p> <p>Comments. Brachystethus tricolor is the sister species of B. vexillum, with which shares two synapomorphies and one homoplasy, respectively: venter predominantly red, metasternal carina transversely sulcated and ostiolar rugae short (BARCELLOS &amp; GRAZIA, 2003). These characters also separate both species from theremaining species of the genus. Brachystethus tricolor can be distinguished from B. vexillum by dorsal color with red spots, and quadrangular shape of pygophore.</p> </div>	https://treatment.plazi.org/id/217887A7991D80234BE35EFCFD546630	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barcellos, Aline;Grazia, Jocélia	Barcellos, Aline, Grazia, Jocélia (2003): Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Sér. Zool. 93 (4): 413-446, DOI: 10.1590/S0073-47212003000400008
217887A7991F80224BF85C0BFE906036.text	217887A7991F80224BF85C0BFE906036.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachystethus vexillum Breddin 1903	<div><p>Brachystethus vexillum Breddin, 1903</p> <p>(Figs. 9, 19, 29, 39, 49, 60, 69)</p> <p>Brachystethus vexillum BREDDIN, 1903:371; BERGROTH, 1908:178; KIRKALDY, 1909: 152; PIRÁN, 1955:337.</p> <p>Daimonocoris bolivianus KORMILEV, 1951:38; PIRÁN, 1955:337 (syn.).</p> <p>Type material of B. vexillum. Holotype, examined, labeled: a) Bolivia; b) Brachystethus vexillum Bredd; c) coll. Breddin; d) Holotypus; e) Dtsch. Entomol. Institut Berlin; f) HT vexillum (DEIC). Typelocality: Bolivia.</p> <p>Description. Dorsal surface black, with yellowish stripes on corium (fig. 9). Head black, with well-marked wrinkles. Punctures moderately distributed, restrict to base of jugae and tylus. Antennae black; fourth antennal segment cylindrical. First antennal segment about 3/4 longer than second, third slightly longer than second, fourth almost twice longer than third and about 3/4 the length of fifth, the longest. Pronotum black, with fine punctures regularly distributed and coarse punctures sparse behind cicatrices. Before cicatrices, medium-sized punctures densely distributed. Anterolateral angles yellowish at apex. Scutellum black, with coarse punctures sparse, more frequent beside lateral margins of basal half. Corium black, without callus, with two yellow stripes: one on clavus, adjacent to claval suture and another occupying 2/3 ectad to radial vein, becoming broader apically toward clavus. Legsdark reddish-castaneous to black. Ventral surface of thorax dark reddish-castaneous to black; medium-sized punctures, concolor. Ostiolar rugae short, reaching 1/3 of distance between inner margin of ostiolum and lateral margin of metapleura. Metasternal carina with conspicuous transversal sulcus; in profile, little elevatedanteriorly. Connexivum black, inner superior angle of each segment orange, segments finely punctured. In some specimens, the orange area extends along inner lateral margin of seventhsegment. Venter red, with black spots on lateral thirds, as follows: one near lateral margin of each segment, becoming broader posterad, other small, around spiracles. On sternites III to VI, also present, entad to others, a larger spot, sub-elliptical, close to spiracle. Coarse punctures more concentrated on lateral thirds, over these spots. Spiracles black.</p> <p>Male genitalia. Pygophore globose, apex and base equally wide. Lobes of dorsal rim conspicuous, strongly projected posterad, rounded at apex. Genital cup processes littlevisible dorsally (figs. 19, 29). Parameres withacute apex, anteriorly directed (fig. 49). Ventralrimopenly V-like (fig. 39).</p> <p>Measurements (n=3). Headlength 1.73 (1.68-1.76) ± 0.05, width 2.56 (2.52-2,60) ± 0.04; interoculardistance 1.49 (1.44-1.52) ± 0.05; anteocularlength 0.64; lengthof antennal segments: I - 0.85 (0.80-0.88) ± 0.05; II - 1.17 (1.12-1.20) ± 0.05; III - 1.44 (1.28-1.52) ± 0.14; IV - 2.59 (2.48-2.72) ± 0.12; V - 3.68; pronotallength 3.70 (3.59-3.84) ± 0.13, width 8.63 (8.52-8.85) ± 0.19; scutellarlength 5.87 (5.85-5.93) ± 0.05, width 5.29 (5.18-5.43) ± 0.13; coriumlength 8.24 (8.02-8.52) ± 0.26; abdominalwidth 10.08 (9.69-10.35) ± 0.35; total length 13.64 (13.03-14.28) ± 0.63.</p> <p>Female genitalia. Sutural borders of gonocoxites 8 parallel, except on apical fifth. Posterior borders convex on outer half and sub-retilineous on inner half (fig. 60). Gonocoxites 9 almost three times wider across apex of arms than longmedially; anterior border openly U-like, moderately excavate at middle. Length of arms of gonocoxites 9 equal to 1/6 of plate width. Thickening of vaginal intima digitiform, elongated. Ductus receptaculi thin, one and a half longer before than after vesicular area. Capsula seminalis digitiform. Annular crests sub-parallel, turned toward vesicular area (fig. 69).</p> <p>Measurements (n=9). Headlength 1.93 (1.80-2.08) ± 0.08, width 2.77 (2.72-2.84) ± 0.05; interoculardistance 1.60 (1.48-1.68) ± 0.06; anteocularlength 0.69 (0.64-0.72) ± 0.04; lengthof antennalsegments: I - 0.91 (0.84-0.96) ± 0.04; II - 1.24 (0.96-1.36) ± 0.13; III - 1.56 (1.36-1.88) ± 0.16; IV - 2.75 (2.60-2.96) ± 0.13; V - 3.58 (3.44-3.68) ± 0.09; pronotallength 4.12 (3.84-4.34) ± 0.15, width 9.65 (9.27-10.02) ± 0.25; scutellarlength 6.74 (6.51-7.01) ± 0.17, width 5.97 (5.68-6.43) ± 0.21; coriumlength 9.32 (8.93-9.85) ± 0.27; abdominalwidth 11.44 (11.19-11.52) ± 0.13; totallength 15.56 (14.95-16.70) ± 0.56.</p> <p>Material examined. PERU, Cuzco: Machu Picchu,, 28.XI.1982, E.R. Ross col., California Academy of Sciences (DARC); (over ruins, 2600-2800m), 2, 4, 1.VII.1964, B. Malkin (MZSP);, 7.V.1965 (arbusto) (MZSP);, 1942, Weyrauch (USNM); Prov. Paucartambo, Pillahuata (2800m),, 2, 19.XII.1952, F. Woytkowski (USNM); Marcapata,, “Coll. I.R.Sc. N.B.” (DARC). BOLIVIA, Yungas de Palmar (2000m),, R. Zischka (USNM); Cochabamba: Chapare prov., Alto Palmar (110m), 2, 2, XI.1961 (DARC).</p> <p>Distribution. Peru (Cuzco), Bolivia (Cochabamba).</p> <p>Comments. Bra chystethus vexillum is closely related to B. tricolor. The dorsal color pattern, with yellow stripes, and the globose pygophore distinguish this species from B. tricolor.</p> </div>	https://treatment.plazi.org/id/217887A7991F80224BF85C0BFE906036	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barcellos, Aline;Grazia, Jocélia	Barcellos, Aline, Grazia, Jocélia (2003): Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Sér. Zool. 93 (4): 413-446, DOI: 10.1590/S0073-47212003000400008
217887A7991E80264B315A6BFC8466E5.text	217887A7991E80264B315A6BFC8466E5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachystethus geniculatus (Fabricius 1787)	<div><p>Brachystethus geniculatus (Fabricius, 1787)</p> <p>(Figs. 1, 10, 20, 30, 40, 50, 61, 70)</p> <p>Cimex geniculatus FABRICIUS, 1787:293; 1794:115; GMELIN, 1793:2151.</p> <p>Edessa geniculata; FABRICIUS, 1803:153.</p> <p>Edessa (Brachystethus) marginatus LAPORTE, 1832:63, pl.54, fig.8.</p> <p>Edessa (Brachystethus) marginata; BURMEISTER, 1835:356 (syn.).</p> <p>Edessa circumscripta BURMEISTER, 1835:356.</p> <p>Brachystethus marginatus; SPINOLA, 1837:343; AMYOT &amp; SERVILLE, 1843:161; HERRICH- SCHÄFFER, 1845:2 (syn.); DALLAS, 1851:336; SIGNORET, 1851:344; HERRICH- SCHÄFFER, 1853:310; STÅL, 1860:28; 1872:48 (syn.); WALKER, 1868:455.</p> <p>Brachystethus circumscriptus; AMYOT &amp; SERVILLE, 1843:161 (syn.); HERRICH- SCHÄFFER, 1845:2, fig. 789; DALLAS, 1851:336; MAYR, 1866:68; WALKER, 1868:455; STÅL, 1872:48 (syn.).</p> <p>Brachystethus geniculatus; MAYR, 1866:68; STÅL, 1868:38 (redescription); 1872:48; BERG, 1892:11; LETHIERRY &amp; SEVERIN, 1893:187 (cat.); KIRKALDY, 1909:152 (cat.); PENNINGTON, 1920:10 (distr.); RUFINELLI &amp; PIRÁN, 1959:12 (distr.); BUCKUP, 1961:9; SILVA et al., 1968:54 (host plant); GRAZIA- VIEIRA &amp; CASINI, 1973:58 (distr.); GRAZIA, 1977:164-165; FROESCHNER, 1981:69 (cat.); LINK &amp; GRAZIA, 1987:118 (distr.).</p> <p>Type material. Holotype, labeled: geniculatus (ZMUC). This specimen, sent as one among three syntypes of C. geniculatus, is the only one whose characters fit with the original description, thus it is here designated as holotype. The second specimen fits the description of Cimex cribrum, asformerly commented, and the thirdfits the diagnostic characters of B. coxalis Breddin. Type locality: “Cajennae”. Most of the specimens studied were collected in eastern and southern Brazil, with Bahia State as its northern limit. Literaturealso indicates the occurrenceof B. geniculatus in Uruguay. As at Fabricius time the label data were quite vague, and Caiene was indeed a site of departure and arrival of the expeditions, we consider that Caiene does not actually correspond to the collecting locality of the specimen.</p> <p>Description. Dorsal surface castaneous to dark-castaneous (fig. 10). Head castaneous to dark-castaneous, with little conspicuous wrinkles. Punctures on medial region and semi-circular stripes densely distributed. Remainder of jugae moderately to sparsely punctured, depending on the specimen. Jugae with yellow stripe on outer lateral margins, evanescent anterad. Antennae reddish-castaneous; second antennal segment one and a half longer than first; third slightly longer than second; fourth one and a half longer than third and 3/4 of the length of fifth, the longest. Fourth antennal segment cylindrical. Pronotum castaneous to dark-castaneous, with yellowish stripe on anterolateral margins, and less conspicuous on theanterior margin. Punctures fine, darkcastaneous to black, regularly distributed, except on cicatrices. Among these punctures, present larger ones, black, more concentrated before cicatrices and behind them, where they achieve theirmaximum size. Scutellumwith colorsimilar to therest of dorsal surface, with coarserpunctures on anterior half (fig. 1). Remaining punctures fine, homogeneously and densely distributed. Corium castaneous, with a yellowish callus, internally and near to apex of radial vein. Legs uniformly reddish-castaneous. Ventral surface of thorax castaneous, punctured with dark-castaneous. Ostiolar rugae extending by about 2/3 of distance betweeninner margin of ostiolum and lateral margin of metapleura. Metasternal flat in profile. Connexivum castaneous, lateral outer 1/6 of each segment with subcaloused yellowish stripe. Entad to this border, punctures fine, densely and homogeneously distributed. Venter castaneous, bordered with a thin subcaloused, yellowish stripe. Medium-sized, concolor punctures densely distributed, except on a narrow medial stripe. Spiracles yellowish-castaneous, bordered with black.</p> <p>Male genitalia. Pygophore subquadrangular, slightly wider at apex. Lobesof dorsal rim conspicuous, moderately projected posterad, sub-acute at apex. Genital cup processes little visible in dorsal view (figs. 20, 30). Parameres with acute apex, turned anterad (fig. 50). Ventralrimopenly V-like (fig. 40).</p> <p>Measurements (n=10). Headlength 1.95 (1.76-2.12) ± 0.13, width 2.93 (2.72-3.16) ± 0.16; interoculardistance 1.61 (1.52-1.72) ± 0.07; anteocularlength 0.67 (0.56-0.80) ± 0.07; lengthof antennalsegments: I - 0.82 (0.76-0.88) ± 0.04; II - 1.33 (1.04-1.68) ± 0.22; III - 1.60 (1.36-1.84) ± 0.16; IV - 2.57 (2.32-3.04) ± 0.26; V - 3.28 (2.88-3.92) ± 0.37; pronotallength 4.08 (3.67-4.68) ± 0.37, width 9.82 (8.68-11.02) ± 0.81; scutellarlength 6.66 (6.01-7.52) ± 0.57, width 6.30 (5.68-7.01) ± 0.49; coriumlength 8.85 (8.18-9.69) ± 0.52; abdominalwidth 10.77 (9.52-12.02) ± 0.83; totallength 14.89 (13.36-16.37) ± 0.99.</p> <p>Female genitalia. Gonocoxites 8 rounded to sub-quadrangular. Sutural borders of gonocoxites 8 parallel and posterior borders bissinuated, in almost right angle over laterotergites 9 (fig. 61). Gonocoxites 9 more than threetimeswider across apexof armsthan long medially; anteriorborderopenly U-like, armslengthequal to 1/4 of platewidth. Thickening of vaginal intima digitiform. Ductus receptaculi thin after vesicular area and slightly shorter than before to this area. Capsula seminalis globose. Annular crests convergent (fig. 70).</p> <p>Measurements (n=10). Headlength 2.08 (1.92-2.24) ± 0.10, width 3.07 (2.84-3.40) ± 0.17; interoculardistance 1.72 (1.60-1.88)± 0.09; anteocularlength 0.70 (0.60-0.80) ± 0.07; lengthof antennalsegments: I- 0.90 (0.80-0.96) ±0.05; II - 1.40 (1.20-1.84) ± 0.20; III - 1.70 (1.44-2.08) ± 0.20; IV - 2.73 (2.36-3.36) ± 0.33; V - 3.38 (2.96-4.08) ± 0.43; pronotallength 4.43 (4.01-5.18) ± 0.37, width 10.48 (9.52-12.02) ± 0.86; scutellarlength 7.41 (6.85-8.68) ± 0.61, width 6.79 (6.35-7.77) ± 0.49; coriumlength 9.74 (8.85-11.36) ± 0.80; abdominalwidth 11.85 (10.69-13.28) ± 0.86; totallength 16.34 (15.11-18.54) ± 0.99.</p> <p>Material examined. BRAZIL,, Pres. by Perth Museum (BMNH 1953-629);, Blackmore col., 74, Pres. by Perth Museum (BMNH 1953-629); Chapada,, acc. 23739 (AMNH);, XII, acc. 23739 (AMNH); Bahia: Ilhéus, 3,, III.2000, C.A.L. Guimarães (in Attalea funifera) (UFRG); Minas Gerais: Poços de Caldas (Morro São Domingos),,, 19.XII.1967, 29.II.1968, J. Becker, O. Roppa &amp; O. Leoncini (MNRJ); Barbacena,, H. Ebert (AMNH); Goiás: Pirineus,, 2.II.1962, J. &amp; B. Bechyné (MPEG); Espírito Santo: Linhares,,, I.1962, Claudionor Elias (DZUP); Santa Tereza,, 24-31.IV.1967, C. &amp; C.T. Elias (DZUP);, 18-23.XII, C. Elias (DZUP); B. Guandu,, 17.V.1970, C. &amp; C.T. Elias (DZUP); Riode Janeiro:, IX, 23734 (AMNH); Riode Janeiro (Corcovado), 3, IX, X.1961, Seabra &amp; Alvarenga (DZUP);, 15.IX.1961, J.S. Moure, Alvarenga &amp; Seabra (DZUP); (Estr. Gávea),, 10.VII.1942, Pero M. Oliv. (UFRG); (Botafogo),, XI.1942, Pero M. Oliv. (UFRG); (Alto da Boa Vista),, X.1943, R. Más (UFRG); (Tijuca),, 13.IV.1943, R. Braga (UFRG); Itatiaia (816m),, 1.I.1933, N.B. Fagundes (UFRG); São Paulo:, 20.III.71 (DZUP);, 4.VIII.1974, C. Bordin (UFRG); Barueri, 6, 5, 02, 20.VII, 12.VIII, 12.XI.1954, 18.VII, 6.XII.1955, K. Lenko (MNRJ); Angatuba,, XII.1922, Azevedo Marques, Inst. Biologia Entomologia Agrícola Riode Janeiro 3.492 (UFRG); Paraná: Xaxim,, 1941, A. Souza (MNCE 13013); Imbaú,, IX.1942; (MNCE); Pinhão (Rio Iguaçu, próximo à Usina Hidrelétrica de Segredo), 2,, VII.1992, G. Skuk (MNCE); Alexandra,, 10.X.1970, Moure &amp; Mielke (DZUP); Ponta Grossa,, 3, XI.1945, 22.II, 23.IV.1946 (MNCE); (pedreira),, XII.1943 (DZUP); Piraquara,, 14.IV.1983, N. Gonçalves (MNCE); (mananciais da serra, (± 800m alt.),, S.L. Bonatto (UFRG); Curitiba,, IX.1945 (MNCE);, 16.II.1966, C. Ext. DZUFP (DZUP); (Fonte Ahú),, XII.1936, Claretiano (MNCE); Praia do Leste,, 26.V.1979, I.W. Forno (DZUP); Santa Catarina:, I.1946, A. Maller, Frank Johnson Donor;,, XI.1953, 2.VIII.1954, A. Maller (MNRJ); Nova Teutônia (27 o 11’N 52 o 23’L, 300- 500m),, II.1981, Fritz Plaumann (DZUP); Corupá (Hansa Humbolt),, II.1948, A. Maller col., Frank Johnson Donor (AMNH); Florianópolis (Canasvieiras),,, I.1946, C. Goffergé (MNCE 10258, 10257); Rio Grande do Sul:,, 6.XII.1973, R. Wagner Hanisch &amp; A. Ruszczyk (UFRG);, 17.XII.1973, E. Heinrichs (UFRG);, 19.III.1974, R.A. Ilana (UFRG); Derrubadas (Parque Florestal Estadual do Turvo), 3, 3, 3.IX.1982, 21-25.V.1984, S.L. Bonatto (UFRG); Porto Alegre,, V.1955, O.P.H. (em Ayrax l eprosum C. de vaca) (UFRG); (Morro Santana), 3, 10, 13.VIII. 1982, S. Bonatto (em quiescência) (UFRG);, 4, 15.IX.1983, M. Becker (em quiescência em bromeliáceas) (UFRG); (Reserva do Lami),, 12.II.1992, A.B.P. Santos (UFRG); (Petrópolis),, 13.XII.1982, G. Bond, L. Buckup (UFRG 271); (Vila Manresa),, 16.II.1965, Pe. Buck (UFRG 6306); (Belém Novo),, 2, 17.IV.1955, M.R. Oliveira (UFRG);, 11.IV.1981, R.P. Mocellin (UFRG); Santa Maria,, 10.IV.1981, A. Stringhini (UFRG); São Sepé,, 17.V.1981, C.R. Kelling (UFRG);, 27.IV.1989, Link &amp; Costa (UFRG); Guaíba (E.E. Guaíba),, 15.IV.1974, E. Heinrichs (UFRG); Viamão,, 18.IX.1993, L.A. Campos (UFRG); Pelotas,, 10.II.1941 (UFRG);, 14.IV.1961, C.M. Biezanko (BMNH); 2,, 16.XII.1961, 05.IV.1963, 7.IV.1966, C.M. Biezanko (BMNH 1965-175, 1963- 375, 1966-318); 2, 14.V.1967, C.M. Biezanko (AMNH); 2, 2, 14.V.1967, J. L. Mantovani &amp; Biezanko (BMNH 1967-265). BOLIVIA, Santa Cruz: San Francisco,, 1895, A. Borelli (MRSN). PARAGUAY, Itapúa: Hohenau,, III.1955, J.L. Stehlik (BMNH 1966-601). ARGENTINA, Buenos Aires: Buenos Aires,, Hoefel, Coll. J. Stehlik (BMNH 1964-449);, II.1953, Hoefel, Coll. J. Stehlik (BMNH 1966-601). No data: 2 (AMNH, UFRG);, P.R. Uhle col. (AMNH).</p> <p>Distribution. Brazil (Bahia, Minas Gerais, Goiás, Espírito Santo, Riode Janeiro, São Paulo to Rio Grande do Sul), Bolivia (Santa Cruz), Paraguay (Itapúa), Argentina (Buenos Aires).</p> <p>Comments. Brachystethus geniculatus is the sister group of the clade formed by B. coxalis, B. improvisus and B. vicinus, sharing with them the yellowish border of pronotum and the callus on corium (BARCELLOS &amp; GRAZIA, 2003). Brachystethus geniculatus is very similar to B. coxalis, by the parameres, with acute apices, and by the metasternal carina, flat in profile, contrasting with the carina strongly elevated of the other two species. Brachystethus geniculatus is differenciated from B. coxalis, B. improvisus and B. vicinus by the uniform color pattern of legs.</p> </div>	https://treatment.plazi.org/id/217887A7991E80264B315A6BFC8466E5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barcellos, Aline;Grazia, Jocélia	Barcellos, Aline, Grazia, Jocélia (2003): Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Sér. Zool. 93 (4): 413-446, DOI: 10.1590/S0073-47212003000400008
217887A7991A80284B1A5D51FC0666D2.text	217887A7991A80284B1A5D51FC0666D2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachystethus coxalis Breddin. 1904	<div><p>Brachystethus coxalis Breddin, 1904</p> <p>(Figs. 11, 21, 31, 41, 62)</p> <p>Brachystethus coxalis BREDDIN, 1904:178; BERGROTH, 1908:178; KIRKALDY, 1909:152 (cat.).</p> <p>Typematerial. Lectotype, heredesignated, labeled: a) Amazonas, Peru; b) Brachystethus coxalis Bredd.; coll. Breddin; d) Syntipus; e) DEI Eberswalde; f) 1 ST coxalis (DEIC). Breddin described the species based on two specimens, one male and one female. Only the male syntype was found. Type locality: Peru (Amazonas) and Panama (Chiriquí).</p> <p>Description. Dorsal surface dark-castaneous to black, with a marginal yellow stripe (fig. 11). Head dark-castaneous, with well-marked wrinkles. Puncturesdensely distributed. Outer margins of jugae with yellowish stripe, wider at base. Antennae dark-castaneous to black, except the first segment, yellowish. Fourth segment cylindrical. First antennal segment a little more than 3/4 as long as second, third almost twice as long as second, fourth slightly less than one and a half as long as third, fifth lacking. Pronotum darkcastaneous to black, with a broad yellow stripe along anterolateral margins. Punctures fine regularly distributed, with sparse coarse punctures restrict tothe area after cicatrices. Before cicatrices, smallerpunctures more concentrated alonganterior marginof pronotum. Scutellum dark-castaneous to black, with fine punctures regularly distributed. Coarse punctures on basal third, concentrated in 1+1 groups along lateral margins. Punctures slightly smaller, sparse, on basal half. Corium dark-castaneous, with yellow border on basal third of costal margin and with a small yellowish callus near and entad the apex of radial vein. Legs yellowish on coxae, trochanters, base and apex of femora, ventral face and apex of tibiae and tarsi; remaining segments dark-castaneous, almost black. Ventral surface of thorax dark-castaneous to black, with castaneous to black medium-sized punctures. Ostiolar rugae reaching 2/3 of distance between inner margin of ostiolumand lateral margin of metapleura. Metasternal carina flat in profile. Connexivum with a subcaloused yellow stripe, scarcely punctured, over outer lateral third of each segment; remaining of connexival segments dark-castaneous, almost black, with punctures minutely and densely distributed. Venter dark-castaneous, with a marginal, subcaloused yellowish stripe. Punctures fine regularly distributed, except on the middle, where they are less frequent.</p> <p>Male genitalia. Pygophore quadrangular. Lobes of dorsal rim conspicuous, moderately projected posterad, rounded at apex. Genital cup processes not visible in dorsal view. Parameres withacute apex, turned anterad (figs. 21, 31). Ventral rim openly V-like (fig. 41).</p> <p>Measurements (n=1). Headlength 2.03, width 3.00; interoculardistance 1.64; anteocularlength 0.74; antennalsegments - lacking; pronotallength 4.34, width 10.02; scutellarlength 6.40, width 6.24; corium length, not measured; abdominalwidth 11.19; totallength 15.87.</p> <p>Female genitalia. Posteriorborders of gonocoxites 8 forming an almost rightangle over laterotergites 9. Sutural borders of gonocoxites 8 parallel (fig. 62).</p> <p>Measurements (n=1). Headlength 2.00, width 3.04; interoculardistance 1.60; anteocularlength 0.72; lengthof antennalsegments: I - 0.88; II - 1.12; III - 2.00; IV - 2.80; V - lacking; pronotallength 4.18, width 10.02; scutellarlength 6.68, width 6.51; corium length 9.52; abdominalwidth 11.11; totallength 15.70.</p> <p>Material examined. VENEZUELA, Amazonas: Cerro de la Neblina (1880m),, 11.II.1984, Stupakoff (AMNH). No data, (ZMUC).</p> <p>Distribution. Panama (Chiriquí), Venezuela (Amazonas), Peru (Amazonas).</p> <p>Comments. Brachystethus coxalis is the sister group of B. vicinus and B. improvisus, sharing with them one synapomorphy, the yellowish tarsi, and two homoplastic characters, coxae, and apex of femora and base of tibiae yellowish (BARCELLOS &amp; GRAZIA, 2003). It can be distinguished from B. vicinus and B. improvisus by the flat metasternal carina, and from the latter also by the shape of parameres, with acute apex.</p> </div>	https://treatment.plazi.org/id/217887A7991A80284B1A5D51FC0666D2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barcellos, Aline;Grazia, Jocélia	Barcellos, Aline, Grazia, Jocélia (2003): Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Sér. Zool. 93 (4): 413-446, DOI: 10.1590/S0073-47212003000400008
217887A79914802A4B255CA9FE2D67E9.text	217887A79914802A4B255CA9FE2D67E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachystethus improvisus Breddin 1905	<div><p>Brachystethus improvisus Breddin, 1905</p> <p>(Figs. 12, 22, 32, 42, 51, 63, 71)</p> <p>Brachystethus improvisus BREDDIN, 1905:26; BERGROTH, 1908:178; KIRKALDY, 1909:152 (cat.).</p> <p>Typematerial. Holotype, examined, labeled: a) Esmerala; b) B. improvisus Bredd. / n. spec. genitalium!; c) coll. Breddin; d) Holotypus; e) DEI Eberswalde; f) 1 HT improvisus (DEIC). Typelocality, Venezuela.</p> <p>Description. Dorsal surface dark-castaneous, with yellow marginal stripe (fig. 12). Head with coarse punctures moderately distributed and conspicuous wrinkles. Outer margins of jugae with yellow stripe, broader at base. Second antennal segment about one and a halflonger than first, thirdslightly longer than second, fourth a little more than one and a half as long as third and about 3/4 as long as fifth, the longest. Fourth antennal segment cylindrical, lacking sulcus. Pronotum with fine punctures, concentrated before cicatrices, and regularly distributed on the remaining pronotum, interposed with a little coarser, sparse punctures. Scutellum with coarse punctures more frequent at basal 2/3. Corium with a yellowish callus near and entad apex of radial vein. Legs castaneous; coxae, base of trochanters and tibiae, apex of femora and tarsi reddish castaneous to yellowish, in extension variable among the specimens. Ventral surface of thorax with castaneous to black medium-sized punctures. Ostiolar rugae reaching about 2/3 the distance betweeninner margin of ostiolum and lateral margin of metapleura. Metasternal carina strongly elevated anteriorly. Connexivum castaneous to black, with subcaloused yellow stripe ocupying, at least, the outer lateral third of each segment. Punctures fine, more densely distributed entad to this stripe. Venter dark-castaneous, with yellow stripe along lateral margins. Punctures fine, inconspicuous at middle. Spiracles yellowish. Genitalia. Pygophore sub-quadrangular, slightly wider apically. Lobes of dorsal rim conspicuous, strongly projected, rounded at apex. Genital cup processes moderately visible dorsally (figs. 22, 32). Parameres with broad apex, anterolaterally directed (fig. 51). Ventralrimopenly U-like (fig. 42).</p> <p>Measurements (n=10). Headlength 2.09 (1.92-2.32) ± 0.12, width 3.34 (3.16-3.44) ± 0.11; interoculardistance 1.76 (1.68-1.84) ± 0.07; anteocularlength 0.76 (0.72-0.80) ± 0.04; lengthof antennalsegments: I - 1.02 (0.88-1.36) ± 0.13; II - 1.54 (1.36-1.76) ± 0.15; III - 2.06 (1.68-2.48) ± 0.25; IV - 3.31 (2.96-3.60) ± 0.19; V - 4.30 (4.16-4.40) ± 0.12; pronotallength 4.93 (4.51-5.26) ± 0.21, width 11.28 (10.69-12.02) ± 0.52; scutellarlength 7.57 (7.01-8.02) ± 0.35, width 7.13 (6.68-7.68) ± 0.35; coriumlength 10.71 (10.35-11.19) ± 0.26; abdominal width 12.49 (11.36-13.19) ± 0.61; totallength 17.48 (16.37-18.37) ± 0.71.</p> <p>Female genitalia. Posterior borders of gonocoxites 8 sinuate over laterotergites 9 (fig. 63); sutural borders of gonocoxites 8 parallel. Gonocoxites 9 less than three timesas wide acrossapex of armsthan long inthe middle; anterior border sinuate, little excavated medially. Arms of gonocoxites 9 a little more than 1/5 of plate width. Thickening of vaginal intima digitiform. Ductus receptaculi thin and longer after than before vesicular area. Capsula seminalis ovoid. Annular crests convergent (fig. 71).</p> <p>Measurements (n=10). Headlength 2.18 (2.00-2.36) ± 0.12, width 3.47 (3.36-3.60) ± 0.07; interoculardistance 1.82 (1.72-1.92) ± 0.06; anteocularlength 0.79 (0.68-0.88) ± 0.07; lengthof antennalsegments: I - 1.00 (0.80-1.20) ± 0.11; II - 1.67 (1.44-2.08) ± 0.19; III - 1.95</p> <p>(1.68-2.24) ± 0.18; IV - 3.37 (3.20-3.52) ± 0.14; V - 4.47 (4.08-4.72) ± 0.26; pronotallength 5.19 (4.84-5.51) ± 0.25, width 11.85 (10.86-12.86) ± 0.69; scutellarlength 8.26 (7.68-9.02) ± 0.47, width 7.54 (6.85-8.18) ± 0.42; coriumlength 11.51 (10.86-12.53) ± 0.52; abdominal width 13.20 (12.19-14.20) ± 0.79; totallength 18.72 (17.54-20.21) ± 0.89.</p> <p>Material examined. GUATEMALA, (AMNH). COSTA RICA, Peralta,, V.1935 (BMNH); Guanacaste: Tierras Morenas, Z. P. Tenorio (Rio San Lorenzo, 1050m),, 10-20.II.1992 (INBC LN 287800, 427600); Puntarenas: Monteverde,,, 4-6.VI.1980, J.E. Wappes col. (DARC); A.C. Arenal, Monteverde (San Luis, 1000-1350m),, I.1994, Z. Fuentes (INBC LN 449250, 250850, 2609); (San Luis, Buen Amigo, 1000-1350m), 2, V.1994, Grace Fuentes (INBC LN 250850, 449250, 2927); R.B. Monteverde,, 24.VIII-15.IX.1992, F. A. Quesada (INBC LN 250850, 449250); Coto Brus (Est. Biol. Las Alturas, 1500m),, VIII.1991, M. Ramirez (INBC LS 822500, 591300);, X.1991, M. A. Zumbado (INBC LS 822500, 591300);, I.1992, M. Ramirez, G. Mora, F. Quesada (INBC LS 822500, 591300);, 23.III-2.V.1992, F. Araya (INBC LS 822500, 591300). PANAMA, Chiriquí:,, 1898, Coll. Noualhier (MNHN); Dist. Renacimiento, Santa Clara (4000’),, 10.IX.82, R. Johnson (AMNH); Fortuna (82 o 15’W 8 o 44’N),, 3.IV.1977, Henk Wolda (AMNH); (1050m),, 18.V.1978, O’Brien &amp; Marshall (AMNH); N. Candelas (4000’),, 24.VIII.82, Stockwell (AMNH); Trail Fortuna to Soledad,, 20.V.1978 (s-facing slope) (AMNH); Repr. La Fortuna,, 17-21.IX.76, 3200’, D. Engleman (DARC); Cerro Azul: north of Tocumén,, 7.VI.1958, W.J. Hanson (DARC). COLOMBIA, Cundinamarca: Bogota,, Distant Coll. (BMNH 1911-383). BRAZIL, Amazonas: Tefé (Ega),, IV.78, I.79, M. de Mathan, Distant Coll. (BMNH 1911-383). BOLIVIA, Santa Cruz: Buena Vista Prov., Tohito (400m),, 1956, Franz Steinbach (AMNH). No data: 2 (BMNH).</p> <p>Distribution. Guatemala, Costa Rica (Guanacaste, Puntarenas), Panama (Chiriquí, Cerro Azul), Colombia (Cundinamarca), Brazil (Amazonas), Bolivia (Santa Cruz).</p> <p>Comments. Brachystethus improvisus is the sister group of B. vicinus (BARCELLOS &amp; GRAZIA, 2003), sharing with this species the metasternal carina anteriorly elevated and the anterior margin of gonocoxites 9 sinuate, little excavated, besides the general color and the color pattern of legs. The two species are very similar in general appearance, but can be distinguished by male and female genitalia. B. improvisus has the dorsal rim of pygophore with conspicuous lobes, genital cup processes moderately visible in dorsal view and parameres broad at apex; on the females, the posterior border of gonocoxites 8 is sinuate over laterotergites 9.</p> </div>	https://treatment.plazi.org/id/217887A79914802A4B255CA9FE2D67E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barcellos, Aline;Grazia, Jocélia	Barcellos, Aline, Grazia, Jocélia (2003): Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Sér. Zool. 93 (4): 413-446, DOI: 10.1590/S0073-47212003000400008
217887A79916802C4B175A62FD8861FC.text	217887A79916802C4B175A62FD8861FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachystethus vicinus Signoret 1851	<div><p>Brachystethus vicinus Signoret, 1851 (Figs. 13, 23, 33, 43, 52, 64, 72)</p> <p>Brachystethus vicinus SIGNORET,1851:344; WALKER, 1868:455; STÅL, 1872:48; DISTANT, 1880:85; LETHIERRY &amp; SEVERIN, 1893:187 (cat.); KIRKALDY, 1909:152 (cat.); FROESCHNER, 1981:69 (cat.); MAES, 1994:8 (cat.).</p> <p>Type material. Lectotype and paralectotype, here designated, both labeled: a) Brazil, Coll. Signoret, b) vicinus det. Signoret (NHMW). Typelocality: Brazil.</p> <p>Description. Dorsal surface castaneous to dark-castaneous, with a yellowish marginal stripe (fig. 13). Head castaneous to dark-castaneous, almost black, with wellmarked wrinkles. Punctures moderately distributed. Outer margins of jugae with yellow stripe, wider at base. Antennae dark-castaneous to black, except the first segment, yellowish at base. Fourth antennal segment cylindrical, lacking longitudinal sulcus. Second antennal segment one and a half longer than first and about 3/4 as long as third; third a little more than half as long as fourth, fifth the longest, a little longer than fourth. Pronotum castaneous to dark-castaneous; anterolateral margins with continuous yellow stripe. Medium-sized punctures concentrated before cicatrices and scarce after that, more frequent on anterior half of pronotum. Scutellum coarsely punctured, punctures more frequent on anterior half. Costal margins of corium with yellow stripe on anterior third, evanescent posterad. Yellowish callus near and entad apex of radial vein. Legs dark-castaneous, except coxae, basal 3/4 of trochanters, apex of femora and base of tibiae, yellowish. Ventral surface of thorax castaneous, with yellow stripe along lateral margins of propleura. Punctures fine, castaneous. Ostiolar rugae reaching 2/3 of distance between inner margin of ostiolum and lateral margin of metapleura. Metasternum, in profile, strongly elevated anteriorly. Connexivum with continuous yellow stripe along outer margin of segments, variable in width. Punctures fine, dense and homogeneously distributed on inner half of each segment. Venter castaneous; lateral margins with a continuous yellow stripe. Punctures fine and concolor, moderately distributed, less frequent in the middle. Spiracles yellowish.</p> <p>Male genitalia. Pygophore trapezoidal, much wider apically. Lobes of dorsal rim inconspicuous. Genital cup processeslittle visibledorsally (figs. 23, 33). Parameres long, withacute apex, turnedanterad (fig. 52). Ventral rim V-like, deeplyexcavatemedially (fig. 43).</p> <p>Measurements (n=4). Headlength 2.20 (2.00-2.34) ± 0.15, width 3.53 (3.36-3.67) ± 0.14; interoculardistance 1.85 (1.76-1.95) ± 0.09; anteocularlength 0.79 (0.74-0.86) ± 0.05; lengthof antennalsegments: I - 1.09 (0.96-1.25) ± 0.14; II - 1.68 (1.56-1.83) ± 0.11; III - 2.21 (1.84-2.42) ± 0.26; IV - 3.83 (3.24-4.29) ± 0.54; V - 4.78 (4.24-5.62) ± 0.73; pronotallength 5.30 (4.84-5.68) ± 0.37, width 12.11 (11.19-12.69) ± 0.67; scutellarlength 7.76 (7.28-8.24) ± 0.44, width 7.40 (7.04-7.76) ± 0.34; coriumlength 10.19; abdominalwidth 13.34 (12.27- 14.20) ± 0.93; totallength 18.83 (16.87-20.21) ± 1.59.</p> <p>Female genitalia. Posterior borders of gonocoxites 8 projected in acute angle, over the limit between laterotergites 8 and laterotergites 9; slightly convex on area adjacent to laterotergites 9 andgonocoxites 9 (fig. 64). Sutural borders of gonocoxites 8 parallel and contiguous. Gonocoxites 9 less than twice and a half wider across apices of arms than long medially; anterior border sinuate, little excavate medially. Arms of gonocoxites 9 about 1/6 as long as plate width. Thickening of vaginal intima digitiform. Ductus receptaculi more than one and a half longer before than after vesicular area, equally thickened in both regions. Capsula seminalis globose. Annular crests convergent (fig. 72).</p> <p>Measurements (n=6). Headlength 2.38 (2.16-2.50) ± 0.13, width 3.66 (3.36-3.90) ± 0.22; interoculardistance 1.95 (1.84-2.07) ± 0.12; anteocularlength 0.80 (0.74-0.88) ± 0.05; lengthof antennalsegments: I - 1.11 (1.04-1.21) ± 0.08; II - 1.72 (1.60-1.91) ± 0.13; III - 2.20 (1.84-2.42) ± 0.23; IV - 3.84 (3.40-4.10) ± 0.38; V - lacking; pronotallength 5.83 (5.18-6.35) ± 0.52, width 13.29 (12.02-14.53) ± 0.99; scutellarlength 8.72 (8.35-9.12) ± 0.33, width 8.24 (7.52-8.88) ± 0.50; coriumlength 12.08 (11.86-12.36) ±0.26; abdominalwidth 15.13 (14.03- 16.03) ± 0.88; totallength 20.61 (19.04-22.38) ± 1.53.</p> <p>Material examined. GUYANA, Essequibo: Kaieteur Fall (high forest),, 3.IX.1937, Richards &amp; Smart (BMNH 1937-776). BRAZIL, (NHMW); Pará: Mocajuba (Mangabeira),, XI.1952, Orlando Rego (MNRJ); Bahia:, 2, 1859, Dr. Felder (NHMW); Espírito Santo: Conceição da Barra,, 27.V.1969, C.T. &amp; C. Elias (DZUP); São Paulo: Araras (Us. Santa Lúcia),, 12.VII.1983, P.R. Cancian (UFRG); Campinas,, XI.1986, L.O.T.M. (UFRG).</p> <p>Distribution. Guyana (Essequibo), Brazil (Pará, Bahia, Espírito Santo, São Paulo).</p> <p>Comments. Brachystethus vicinus is the sister group of B. improvisus, distinguishing from this species, in the males, by the shape of parameres, with acute apex, by the inconspicuous lobes of dorsal rim of pygophore and by the genital cup processes little visible dorsally. In the females, the posterior borders of gonocoxites 8, projected in acute angle over the limit between laterotergites 8 and laterotergites 9, also separate B. vicinus from B. improvisus.</p> </div>	https://treatment.plazi.org/id/217887A79916802C4B175A62FD8861FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Barcellos, Aline;Grazia, Jocélia	Barcellos, Aline, Grazia, Jocélia (2003): Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae). Iheringia, Sér. Zool. 93 (4): 413-446, DOI: 10.1590/S0073-47212003000400008
