identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B287EAFFE0FFE922B5F9EEFB767306.text	03B287EAFFE0FFE922B5F9EEFB767306.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucania mopan Adams and McCabe 2023	<div><p>Leucania mopan Adams and McCabe new species</p> <p>Figs. 2 (imago), 15 (valvae), 16 (endophallus), 35 (bursa copulatrix)</p> <p>Material examined. Dissections examined (5♁♁, 2♀♀). Type material: Holotype male. BELIZE: Cayo: Mountain Pine Ridge, 1000’ Falls, 17.060529,- 88. 848458, 623m, 26–30 May 1990, L.C. Dow, dissection TLM♁6326 (deposited in NYSM). Paratypes. (28♁♁, 8♀♀) BELIZE: Cayo: Mountain Pine Ridge, 1000’ Falls, 17.060529,- 88. 848458, 623m, 15 May 1990, L.C. Dow, 1♁, dissection MSA♁3406, 22–24 May 1990, L.C. Dow 10♁♁, 3♀♀, dissections MSA♁2408, MSA ♀ 2409, 26-30 May 1990, L.C. Dow, 3♁♁, 2♀♀, 13–16 Jun 1990, L.C. Dow, 1♁, 2-3 Jul 1990, L.C. Dow, 1♀, dissection TLM ♀ 6329 (NYSM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-88.86409&amp;materialsCitation.latitude=17.043169" title="Search Plazi for locations around (long -88.86409/lat 17.043169)">Granite Cairn</a>, 17.043169, -88.864088, 702m, 15 Jun 1990, M.S. Adams, 1♁, 1 Jul 1990 L.C. Dow, 3♁♁ (NYSM); no specific locality, 16.956938, -88.979678, 625m, M.S. Adams 6♁♁, 1♀ (NYSM); Stann Creek, Cockscomb Basin Sanctuary, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-88.4589&amp;materialsCitation.latitude=16.780802" title="Search Plazi for locations around (long -88.4589/lat 16.780802)">Jaguar Reserve</a>, 16.780801, -88.458902, 71m, 10–12 May 1990, M.S. Adams, 1♁, 1♀, 4-9 Jun 1990, L.C. Dow, 1♁, 18–19 Jun 1990, M.S. Adams, 1♁, dissection TLM♁6317 (NYSM). GUATEMALA: Alto Verapez, Chejel, 15.283333, -90.050000, 287m, 1♁, dissection MSA US♁180 (USNM).</p> <p>Diagnosis. The Belizean endemic L. mopan bears a close resemblance to its sibling species, the widespread Central American L. albifasciata (Hampson, 1905) (Fig. 1), from which the habitus differs in having a generally paler rufous forewing. The hind wing of both sexes of L. mopan are infuscated with brown scales contrasting with the pure white hind wing of L. albifasciata. Both species have basal abdominal eversible tubular structures and a lappet (Fig. 13) (an inflatable flap on the outside of the valva). The ornamentation of the everted endophallus separates L. mopan from L. albifasciata (Fig. 14). Both species have a small cluster of short, stout cornuti at the proximal end of the endophallus, which is a simple straight tube bent medially at a right angle, at which point there are three long cornuti. Beyond this point, on L. mopan, the endophallus is unadorned except for an array of four or five retrorse cornuti proximal to the primary gonopore. In contrast, L. albifasciata has the endophallus adorned by a single, long row of cornuti, which continues to the primary gonopore. The appendix bursae of L. mopan is straight (Fig. 35) whereas that of L. albifasciata has.a sharp angle at midpoint (Fig. 36).</p> <p>Description. (Fig. 2) Wingspan 35–38 mm. Head, palpi, frons, and thorax pale rufous. Tegulae and patagia pale rufous with dusting of black scales. No scale tufts on male fore- and mid-tibiae. Abdomen paler ventrally with dusting of black scales. Forewing pale rufous with dusting of black scales. White-scaled cubital vein bisects forewing and broadens at distal end of cell where it curves around inferior distal margin of reniform (represented by black dot). Cubital vein a narrow white line subtended by dark shade along its entire length. Forewing with faint darker stripe adjacent to white-scaled second anal vein. Postmedial (p.m.) line a curving row of small black dots on veins. Pale oblique fascia subtended by dark wedge from vein M3 to wing apex. Forewing with row of fine black terminal points. Fringe dark. Hind wing off-white, infuscated with dark scales on veins and along costal margin. A field of black specks on sub-costal margin of underside of both fore and hind wing. Basal abdominal eversible tubular structures present.</p> <p>Male genitalia. (Figs. 15 &amp; 16). Uncus slightly dilated, terminating in well-defined, claw-like tip; tegumen and vinculum unmodified; valva broad with few non-deciduous setae on margin and with an inflatable flap (lappet) on outside of valva; cucullus distinctively short and broad with non-deciduous setae on its outer margin; pore plate present at valvulus (base of cucullus); ampulla short and thin; editum conspicuous below ampulla; digitus short and sharp pointed; clasper an elongate, acutely curved projection, extending almost to shoulder of valva margin; claval area of sacculus smooth and unmodified; phallus a simple straight tube. Everted endophallus with small cluster of short, stout cornuti at proximal section followed by a simple straight tube bent medially at a right angle, three long cornuti near bend; distal portion of endophallus mostly unadorned except for an array of four or five retrorse cornuti at the primary gonopore.</p> <p>Female genitalia. (Fig. 35) Ductus bursae extending from ostium bursae as a straight tube until an angle at junction with appendix bursae. Appendix bursae sclerotized and extending beyond ductus bursae. Corpus bursae sac-like and thin-walled, arising at juncture of ductus bursae and appendix bursae.</p> <p>Global Distribution. Belize (type locality), Guatemala.</p> <p>Etymology. The specific epithet mopan is in recognition of the dialect of the Yucatanian Branch of the Mayan language.</p> <p>Food plant. Unknown.</p> <p>Larva. Unknown.</p> <p>Remarks. Leucania mopan appears to be a narrow endemic to the pre-montane [650–700m] pine savanna of the Mountain Pine Ridge of Belize and Guatemala. This small (less than 3000 km 2) ecoregion is recognized by the World Wildlife Fund as Critical/Endangered (Dinerstein, et al., 1995). The principal physical factors of this habitat include xeric, acidic, nutrient-poor soil and periodic low intensity burns (Johnson &amp; Chaffey, 1973). The predominant tree is the endemic variety of Caribbean pine (Pinus caribaea Morelet var. hondurensis (Sénéciauze) W.H. Barrett &amp; Golfari). Other characteristic plants include: Crescentia cujete L.(Calabash tree), Quercus spp., Curatella americana L. (Chaparro), Byrsonima crassifolia H.B.K. (Manteco), and Acoelorraphe wrightii (Griseb. &amp; H. Wendl.) H. Wendl. ex Becc. (Paurotis palm) (Corrales, et al., 2015; Hampshire, 1988; Laughlin, 2002). The canopy is open with many low shrubs and open savanna areas with grasses, sedges and a variety of herbs.</p> </div>	https://treatment.plazi.org/id/03B287EAFFE0FFE922B5F9EEFB767306	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mccabe, Timothy L.;Adams, Morton S.	Mccabe, Timothy L., Adams, Morton S. (2023): Five new species of the genus Leucania Ochsenheimer in Central America (Lepidoptera: Noctuidae). Zootaxa 5256 (3): 250-266, DOI: 10.11646/zootaxa.5256.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5256.3.2
03B287EAFFE2FFEE22B5F93EFD8A7513.text	03B287EAFFE2FFEE22B5F93EFD8A7513.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucania merga Adams and McCabe 2023	<div><p>Leucania merga Adams and McCabe new species</p> <p>Figs. 4 (imago), 17 (valvae), 18 (endophallus), 37 (bursa copulatrix)</p> <p>Material examined. Dissections examined (13♁♁, 5♀♀). Type material: Holotype male. GUATEMALA: Alta Verapaz: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-90.21246&amp;materialsCitation.latitude=15.191822" title="Search Plazi for locations around (long -90.21246/lat 15.191822)">Biotopo del Quetzal</a>, 15.191822, -90.212461, 1700m, 1♁, dissection TLM♁4385 (deposited in NYSM); Paratypes. (36♁♁, 11♀♀) COSTA RICA Guanacaste, 11.01602, -85.38053, 380m, 10-SRNP-114591, D. Janzen, 1♁ dissection TLM♁6020 (USNM), 1♀, dissection TLM ♀ 6021 (USNM); Puntarenas: Monteverde, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-84.82202&amp;materialsCitation.latitude=10.316877" title="Search Plazi for locations around (long -84.82202/lat 10.316877)">Pension Quetzal</a>, 10.316877, -84.822019, 13880m, 14–21 Apr 1990, T. McCabe, 10–21 Feb 2007, T. McCabe, 10♁♁, dissection TLM♁1818 (TLM). GUATEMALA:: Alta Verapaz: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-90.21246&amp;materialsCitation.latitude=15.191822" title="Search Plazi for locations around (long -90.21246/lat 15.191822)">Biotopo del Quetzal</a>, 15.191822, -90.212461, 1700m, 3♁ (TLM); Suchitepéquez: Patulul, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.136246&amp;materialsCitation.latitude=14.522925" title="Search Plazi for locations around (long -91.136246/lat 14.522925)">Los Tarrales Natural Res.</a>, 14.522925, -91.136243, 758m, 22 Jul 2009, T. Mc-Cabe, 1♁, 23–24 May 2014, T. McCabe, 1♀ (TLM); Quetzaltenango: Fuentes Georgina, Volcan Zunil, 14.748972,- 91.48031, 2420m, 4–5 Oct 2012, T. McCabe, 1♁, dissection TLM♁4827, 13-21 Feb 2007, T. McCabe, 4♀♀, (TLM) 1 dissection TLM ♀ 6484; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-92.36025&amp;materialsCitation.latitude=15.54411" title="Search Plazi for locations around (long -92.36025/lat 15.54411)">Bosqueren de Majades</a>, 15.54411, -92.36025, 2933m, 6 Oct 2012, T. McCabe, 1♁ (TLM); Sacatapequez, Antigua, 25–29 Feb 1992, P.J. Landolt, 1♁, dissection MSA♁2900 (NYSM). NICARAGUA: Matagalpa: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.91395&amp;materialsCitation.latitude=12.99698" title="Search Plazi for locations around (long -85.91395/lat 12.99698)">Selva Negra</a>, 12.99698, -85.91395, 1300m, 10 Nov 2010, T. McCabe, 1♀ (TLM). MEXICO: Oaxaca: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.54298&amp;materialsCitation.latitude=16.24104" title="Search Plazi for locations around (long -96.54298/lat 16.24104)">7 mi S Miahuatian</a>, 16.241039, -96.542979, 1672m, 19 Aug 1992, H. Romack, 1♁ (TLM); Loxicha, 20 km N Candelaria, 1585m, 22–23 Jul 1993, P.J. Landolt, 1♁, dissection MSA♁3149 (NYSM); Mitla, 19 Aug 1969, L.A. Kelton, 1♀, dissection MSA ♀ CNC22; (CNC); Querétaro: 15 mi W Xilitia, 1585m, 31 Jul 1992, P.J. Landolt, 1♁, dissection MSA♁3148; Vera Cruz: Las Minas, near Permet, 1200m, 18 Jul 1993, P.J. Landolt,1♁, dissection MSA♁3190, 18 Jul 1993, P.J. Landolt, 1♀, dissection, MSA ♀ 3191 (NYSM); Chiapas: El Bosque, 1♁, dissection MSA♁CNC19; Bochii, 24 Jul 1969, L.A. Kelton, 2♁♁; San Christóbal, Las Casas, 2295m, 12 May 1969, J.E.H. Martin,1♁, 6 May 1969, J.E.H. Martin, 1♁, dissection MSA♁CNC3; no specific locality, 18 Jul 1969, D. Kritsch, 1♁; Tapilulu, 21 May 1969, A. Mutuura, 3♁♁; 9 mi SE Tropisco, 16 May 1969, J.E.H. Martin, 1♁, 1♀; Durango, 10 mi W El Salto, 2743m, 3 Aug 1964, J.E.H. Martin, 1♀, dissection MSA ♀ CNC4 (CNC). ECUADOR: Zamora, Valladolid, 1♁, dissection MSA♁3593; Chinchipe, 3 km E Sabaanilla, Rio Zamora, 1610m, 1♀ dissection MSA ♀ 3896 (NYSM). VENEZUELA: no specific locality or date, 1♁ MSA♁2785; Aragua: Henri Pittier National Park, Rancho Grande, 22-31 Aug 1967, R. Poole, 1♁, dissection MSA♁ US197 (USNM).</p> <p>Diagnosis. Leucania merga is compared to the widespread L. dorsalis Walker, 1856 (Fig. 12). Leucania merga typically has a black spot similar to p.m. dots, but below the mid portion of the cell. This black spot is lacking in L. dorsalis. Diagnostic genitalia characters are as follows: In L. merga the combination of a “pitchfork-like” digitus and basal sclerite of the clasper contrasts with the “spade-like” structure of these two elements in L. dorsalis (Fig. 19). The everted endophalli of the two species is also strikingly different (compare Fig. 18 with Fig. 20). In L. merga, after the basal straight portion the endophallus, makes a right angle capped by a diverticulum with a cluster of long, stiff cornuti. In L. dorsalis this structure is reduced to two small diverticula, one of which has a single hair-like cornutus. The terminal segment of the endophallus in L. merga has a row of robust, retrorse cornuti, which diminishes to a sparse row as it continues to the gonopore. In L. dorsalis this segment of the endophallus is completely unadorned. The bursae copulatrix are also distinct. In L. merga (Fig. 37) the ductus bursae is short and thick whereas in L. dorsalis (Fig. 38) the ductus bursae is long, narrow, and twisted.</p> <p>Description. (Fig. 4) Wingspan 34–36.5 mm. Head, palpi, frons, thorax, and tegulae light tan. Patagia with three bands, most anterior with brown scales, second less distinct, third with distinct, black-tipped scales; sex tufts present on male fore- and mid-tibia. Forewing light tan, veins with white interspaces with brown between veins, cubital vein white with faint brown shade along entire length, a black dot (not reniform) present at middle of cell below origin of vein Cu2; p.m. line not produced, apical shade faint, no terminal dots present. Hind wings of both sexes infuscated with some dark scaling on veins. Ventral forewing light tan, darker in subcostal area. Ventral hind wing with light infuscation. Abdomen light tan, shaggy. Sexes similar, except females somewhat darker. Males with basal abdominal eversible tubular structures.</p> <p>Male genitalia. (Figs. 17 &amp; 18) Uncus slightly dilated before terminating in well-defined, claw-like tip; tegumen and vinculum unmodified; cucullus elongate and somewhat rectangular with row of marginal setae in sockets, pore plate present at valvulus; ampulla long and thin; digitus long and sharp-pointed; editum a conspicuous protuberance; basal sclerite of clasper produced into a sharp-pointed projection; claval area of sacculus unmodified. Phallus short and straight; proximal portion of everted endophallus unadorned followed by a right angle, then a diverticulum adorned by clump of long, sharp spines, then the endophallus balloons out, followed by a portion without cornuti; distal portion of endophallus with short single row of heavy retrorse spines leading to somewhat irregular row of shorter cornuti that extends to a narrower terminal portion.</p> <p>Female genitalia. (Fig. 37) Ductus bursae moderately long and sclerotized. Appendix bursae sclerotized and striate for proximal half, directed to left before overlapping ductus bursae and leading to membranous sac that terminates at the ductus seminalis. Corpus bursae sac-like and thin-walled, arising at juncture of ductus bursae and appendix bursae.</p> <p>Global distribution. Mexico, Guatemala (type locality), Belize, Costa Rica, Ecuador.</p> <p>Etymology. The specific epithet merga, a noun in apposition (Latin merga for hayfork) refers to the sharp, tinelike digitus and pointed basal sclerite of the clasper.</p> <p>Food plant. Unknown.</p> <p>Larva. Unknown.</p> <p>Remarks. A Guatemala specimen of L. merga with Janzen code 10-SRNP-114591 was sequenced in BOLD under the name Leucania Poole 11.</p> </div>	https://treatment.plazi.org/id/03B287EAFFE2FFEE22B5F93EFD8A7513	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mccabe, Timothy L.;Adams, Morton S.	Mccabe, Timothy L., Adams, Morton S. (2023): Five new species of the genus Leucania Ochsenheimer in Central America (Lepidoptera: Noctuidae). Zootaxa 5256 (3): 250-266, DOI: 10.11646/zootaxa.5256.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5256.3.2
03B287EAFFE4FFEE22B5FE6FFA2873AB.text	03B287EAFFE4FFEE22B5FE6FFA2873AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucania colorada McCabe & Adams 2023	<div><p>Leucania colorada McCabe &amp; Adams new species</p> <p>Figs. 7 (imago), 21 (valvae), 22 (endophallus), 39 (bursa copulatrix)</p> <p>Material examined. Dissections examined (2♁♁, 2♀♀). Type material: Holotype female. MEXICO: 3 mi. N. Tlaxiaco, Tierra Azul, 7 August 1992, H. Romack, 7500’, dissection TLM ♀ 6362 (deposited in NYSM). Paratypes. (3♁♁, 1♀) MEXICO, Oaxaca: 5 mi. S. Tlaxiaco, Tierra Azul, 7 August 1992, H. Romack, 7000’, 3♁♁, dissections TLM♁6260 and TLM♁6361; 7 mi. S. Miahuitlan, 19 August 1992, H. Romack, 7000’, 1♀, dissection TLM ♀ 6276 (NYSM).</p> <p>Diagnosis. Leucania colorada bears a close resemblance to L. sororia. Leucania colorada is red-brown versus the gray-brown of L. sororia. The white scaling on the cubital vein typically does not curve around the reniform in L. colorada, as it does in L. sororia. The valvae are similar, but the tip of the clasper in L. colorada is blunt and bent ventrally, whereas in L. sororia there is a pointed tip that is bent dorsally (Figs. 21 vs. 23). In habitus, L. colorada is like L. complicata Strecker, 1898 (Figs. 5 &amp; 6) and L. oaxacana Schaus, 1898 (Fig. 3). The endophalli of L. colorada and L. sororia have elongate fingerlike diverticula lacking in L. oaxacana (Fig. 34) and L. complicata (Fig. 28). Leucania complicata lacks the white scaling on the cubital vein; it also has a unique spur at the apex of the phallus (Fig. 28). Leucania oaxacana has a much smaller costal lobe of the sacculus (Fig. 33). Leucania colorada has an appendix bursae that is barely upturned at its tip whereas L. sororia (Fig. 40) has an appendix bursae that is longer and doubles back on itself.</p> <p>Description. (Fig. 7) Wingspan 32–36 mm. Head, palpi, thorax, patagia, tegula, and forewings reddish-brown. No scale tufts on male fore- and mid-tibiae. Forewing cubital vein clearly marked with white scales from base to reniform, after which vein thickens distally. White scales on cubital vein typically not extending above bottom of reniform. Post-medial line a curving row of small black dots on veins. Hind wing pale at base with brown infuscation towards margin. Abdomen concolorous with hind wing margins. Ventral surface of both fore and hind wing duller than dorsal surface and terminal dots more distinct. Pair of eversible tubular structures present on ventrolateral aspect of male second abdominal segment.</p> <p>Male genitalia. (Figs. 21 &amp; 22) Uncus with spear-shaped spiny terminus, only slightly thickened and attenuated distally, lacks a narrow claw-like tip. Tegumen and vinculum unmodified. Lappet inflates beyond the margin of valva; cucullus short and broad with row of non-deciduous setae on outer margin; valvular pore plate large; ampulla short and thin, editum conspicuous below ampulla, digitus short, sharp-pointed, and widely splayed in respect to clasper; clasper an elongated projection extending almost to valva margin and terminating in a blunt knob. Claval area of sacculus wide, producing a flat-topped bulge. Phallus a simple straight tube. Everted endophallus with a basal patch of spines followed by a finger-like diverticulum at about one-third of its length, at base of diverticulum a similar patch of a few spines. A long, somewhat irregular double row of spines covering distal half of endophallus, most proximal of these spines much longer than the others.</p> <p>Female genitalia. (Fig. 39) Ductus bursae extending from ostium bursae as a straight tube before sharp turn into appendix bursae. At base of the turn a spherical, membranous, corpus bursae arises after a narrow constriction. Appendix bursae striated, folded back upon itself and splayed in relation to ductus bursae. A small portion of the appendix bursae overlaps the ductus bursae in ventral view, terminating in a short, upturned prominence.</p> <p>Global Distribution. Mexico (type locality).</p> <p>Etymology. The specific epithet colorada refers to the muddy reddish color of the moth.</p> <p>Food plant. Unknown.</p> <p>Larva. Unknown.</p> <p>Remarks. Leucania colorada and L. sororia can be sympatric and synchronic at high elevation in the Mexican state of Oaxaca. A female was chosen as holotype as the tip of the appendix bursa is the most diagnostic feature.</p> </div>	https://treatment.plazi.org/id/03B287EAFFE4FFEE22B5FE6FFA2873AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mccabe, Timothy L.;Adams, Morton S.	Mccabe, Timothy L., Adams, Morton S. (2023): Five new species of the genus Leucania Ochsenheimer in Central America (Lepidoptera: Noctuidae). Zootaxa 5256 (3): 250-266, DOI: 10.11646/zootaxa.5256.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5256.3.2
03B287EAFFE7FFE322B5F931FBC3709E.text	03B287EAFFE7FFE322B5F931FBC3709E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucania sororia McCabe & Adams 2023	<div><p>Leucania sororia McCabe &amp; Adams, new species</p> <p>Figs. 8 (imago), 23 (valvae), 24 (endophallus), 40 (bursa copulatrix)</p> <p>Material examined. Dissections examined (3♁♁, 3♀♀). Type material: Holotype female. MEXICO, Oaxaca, 7 mi. S. Miahuitlan, 19 August 1992, 7,000’, H. Romack, dissection TLM ♀ 6308 (deposited in NYSM). Paratypes. (4♁♁, 3♀♀). MEXICO: Durango, 10 mi. W. of El Salto, 18 June 1964, J.E.H. Martin, 9000’, 4♁♁, 2♀♀, dissections TLM ♀ 6440, TLM ♁6436, MSA♁CNC1, MSA ♀ CNC2 (CNC); Oaxaca, 5 mi. S. Tlaxiaco, 5 August 1992, 7,000’, 1♁, H. Romack, dissection TLM♁6067 [NYSM].</p> <p>Diagnosis. Leucania sororia bears a close resemblance in both habitus and genitalia to L. colorada (Fig. 7). Leucania sororia is gray-brown versus the red-brown of L. colorada. Another reliable diagnostic character is the configuration of the appendix bursae. The upward bent tip of the appendix bursae in L. sororia is much shorter and smaller than in L. colorada (Fig. 39). In the male genitalia, the digitus and clasper run parallel in L. sororia but are splayed apart in L. colorada (Fig. 21). Leucania sororia may be confused with L. oaxacana and L. complicata (see diagnosis of L. colorada). Based solely on habitus, L. oaxacana is not reliably separated from L. sororia. Leucania oaxacana has the white of the cubital vein thickened beneath the reniform whereas L. sororia typically has no thickening of the white at this location. The forewing color and pattern of L. sororia is also close to that of L. imperfecta Smith, 1894 (Fig. 9), but L. imperfecta has much lighter hind wings.</p> <p>Description. Wingspan 34–36 mm. Head, palpi, frons, thorax, patagia, tegula, and forewing gray-brown. No scale tufts on male fore- and mid-tibiae. Forewing with cubital vein clearly marked with white scales from base to reniform where it thickens (expands). Reniform above and adjacent to white-scaled cubital vein. White scales present below or rarely beyond the reniform. White scaling of cubital vein not thickened near reniform. Post-median line a curving row of small black dots on veins. Hind wings pale at base with brown marginal infuscation. Abdomen concolorous with hind wing margin. Ventral surface of hind wings duller and terminal dots more distinct than on dorsum. Eversible tubular structures present on ventrolateral aspect of male second abdominal segment.</p> <p>Male genitalia. (Figs. 23 &amp; 24) Uncus with distal end spear-shaped and spiny, slightly thickened distally, and attenuated, but lacking a narrow claw-like tip. Tegumen and vinculum unmodified. Valva broad with inflatable flap (lappet) on the outside. Lappet, when fully inflated, extends well beyond valva margin. Cucullus short and broad with a row of numerous non-deciduous setae on outer margin. Pore plate well-developed. Ampulla short and thin. Editum conspicuous. Digitus short, sharp pointed, and typically parallel to clasper (see right valva in Fig. 23). Clasper an elongated projection extending almost to valva margin and terminating in upturned knob with acute point. Claval area of the sacculus wide, producing flat-topped bulge. Phallus a simple straight tube. Everted endophallus with finger-like diverticulum at one-third of its length from base, a patch of a few spines at base of diverticulum, another similar patch of spines between the diverticulum and the phallus. Distal half of endophallus with a long, somewhat irregular, double row of spines, proximal spines much longer than the others.</p> <p>Female genitalia. (Fig. 40) Ductus bursae extends from ostium bursae as a straight tube, sharply bent at appendix bursae. Corpus bursae arising following a narrow constriction, spherical, membranous. Striate appendix bursae folding back upon itself, overlapping ductus bursae for most of its length before terminating in a long, upturned prominence.</p> <p>Global distribution. Mexico (type locality).</p> <p>Etymology. The epithet “sororia” (soror Latin for sister) refers to the relationship to L. colorada.</p> <p>Food plant. Unknown.</p> <p>Larva. Unknown.</p> <p>Remarks. Leucania sororia and L. colorada are sympatric and synchronic at high elevations in the Mexican state of Oaxaca. Leucania oaxacana Schaus, 1898 (syntype illustrated by Poole, 2022) occurs in Mexico and Central America and is easily confused with two of the newly described species (L. sororia and L. colorada) and with L. complicata.</p> <p>We have also examined L. complicata Strecker, 1898, Restored Status, from northern Mexico and southwestern USA (photograph of type examined). Previously treated as a synonym of L. oaxacana (Franclemont &amp; Todd, 1983). Leucania complicata was misidentified as L. oaxacana by Poole (accessed 16 June 2022). Leucania complicata is a member of an informal “oaxacana group” that includes: Leucania oaxacana, L. albifasciata, L. colorada, L. sororia, L. mopan, and L. merga. Leucania imperfecta Smith, 1894, is sympatric with L. complicata in southwestern USA and northern Mexico and may be confused with species of the “ oaxacana ” group.</p> </div>	https://treatment.plazi.org/id/03B287EAFFE7FFE322B5F931FBC3709E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mccabe, Timothy L.;Adams, Morton S.	Mccabe, Timothy L., Adams, Morton S. (2023): Five new species of the genus Leucania Ochsenheimer in Central America (Lepidoptera: Noctuidae). Zootaxa 5256 (3): 250-266, DOI: 10.11646/zootaxa.5256.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5256.3.2
03B287EAFFE9FFE422B5FAE2FD537512.text	03B287EAFFE9FFE422B5FAE2FD537512.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Leucania championi Adams and McCabe 2023	<div><p>Leucania championi Adams and McCabe new species</p> <p>Figs. 10 (imago), 31 (valvae), 32 (endophallus), 44 (bursa copulatrix)</p> <p>Leucania humidicola (not Guenée, 1852: 90). Troubridge, 2020: Fig. 102 [Misidentification].</p> <p>Leucania februalis (not Hill, 1924: 186, Fig.17). Troubridge, 2020: Fig. 101 [Misidentification].</p> <p>Mythimna solita (not Walker, 1856: 99). Hayes, 1975, page 168–169, Fig. 43. [Misidentification].</p> <p>Material examined. Dissections examined (7♁♁, 9♀♀). Type material: Holotype male. GUATEMALA: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.48031&amp;materialsCitation.latitude=14.748972" title="Search Plazi for locations around (long -91.48031/lat 14.748972)">Fuentes Georginas</a>, 14.748972, -91.480310, 2455m, 13–16 Feb 2007, 1♁, T. McCabe, dissection TLM♁6056 (deposited in NYSM); Paratypes. (21♁♁, 14♀♀). MEXICO: Chiapas; Tapilula, 21 May, 1969, A. Matuura, 1♁, dissection MSA♁CNC11 (CNC); Durango; Pueblo Nuevo, 10 km W El Salto, 02 Aug 1969, J.E.H. Martin, 1♀, dissection MSA ♀ 3907 (CNC); Chiapas: San Cristóbal, 13 May 1969. J.E.H. Martin, 1♀, (CNC); Oaxaca: Sola de Vega, Llano Verde, 16.5652,- 973639, 825m, 07 Jul 1977, J.E. Rawlins, 2♁♁, 1♀, dissection MSA ♀ 251 (CMNH); 55 km N Putia, 1♁ (CMNH); 15 Aug 1986, 1♁(CMHH), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-97.68153&amp;materialsCitation.latitude=17.312765" title="Search Plazi for locations around (long -97.68153/lat 17.312765)">Tlaxiaco</a>, 3 mi N Tlaxiaco, 17.312765, -97.681532, 2438m, H. Romack, 1♀, dissection TLM ♀ 6065 (TLM); Ixtia, Guelatao de Juárez, 18 Aug 1969, L.A. Kelton, 1♁ (CMNH); Puebla: Puebla, no specific locality, 20 Jun 1920, C.C. Hoffmann, 1♁ (CMNH); Michoacan: Chupicuara, 13 Jul 1977, J.E. Rawlins, 1♁ (CMNH); San Luis Petosi, Querótaro, 24–25 Jul 1982, J.E. Rawlins, 1♁ (CMNH); Baja California Sur: La Paz, Guaycura Hotel, 05 Dec 1961, 1♁ (CMNH); Sierra Madre Oriental: nr. Bajan, 26.569722,- 101.221389, 960m, 16 Jul 1992, T. McCabe, 1♁ (TLM); Cuatrocienagos, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-102.18389&amp;materialsCitation.latitude=26.843332" title="Search Plazi for locations around (long -102.18389/lat 26.843332)">Dunes</a>, 26.843333, -102.183889, 16 Jul 1992, T. McCabe, 1♁, dissection TLM♁5280 (TLM). COSTA RICA: Guanacaste, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-85.38053&amp;materialsCitation.latitude=11.01602" title="Search Plazi for locations around (long -85.38053/lat 11.01602)">Santa Rosa National Park</a>, 11.01602, -85.38053, 380m, 10–12 Jul 1979, D. Janzen, 3♁♁, 1♀, dissections MSA♁297, MSA ♀ 296; 9-12 Aug 1977, 1♁, 1♀ dissections MSA ♁301, MSA ♀ 302; 14 Jan 1978, 1♀, dissection MSA ♀ 298, (NYSM), Guanacaste, 11.01602, -85.38053, 380m, 1♁ 07-SRNP-102951, D. Janzen, dissection TLM ♁6028 (USNM), 1♀, 07-SRNP-104287, dissection TLM ♀ 6029 (USNM); no locality or date specified, Cooper, 1♁ (CMNH). GUATEMALA: [Quetzaltenango]: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.48031&amp;materialsCitation.latitude=14.748972" title="Search Plazi for locations around (long -91.48031/lat 14.748972)">Fuentes Georginas</a>, 14.748972, -91.480310, 2455m, 13–16 Feb 2007, 1♁, T. McCabe; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.47972&amp;materialsCitation.latitude=14.748972" title="Search Plazi for locations around (long -91.47972/lat 14.748972)">Fuentes Georginas</a>, 8 km SE Zunil, 14.748972, -91.479722, 2313m, 14–15 Feb 2007, M.S. Adams, 1♁ (NYSM); 26 Feb 2007, 1♀ (TLM); [Suchitepéquez: Paulul], <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.136246&amp;materialsCitation.latitude=14.522925" title="Search Plazi for locations around (long -91.136246/lat 14.522925)">Los Tarrales Natural Reserve</a>, 14.522925, -91.136243, 1400m, 22 Jul 2009, T. McCabe 2♀ (TLM), 1♀ 11 Feb 2007 (TLM), 22 July 2009, 1♀ dissection TLM ♀ 6479; NICARAGUA: Matagalpa, Fuente Pura, 12 km N Matagalpa, 8–9 Jan 1994, E. van den Berghe, 1♁ (CMNH); Selva Nigra, 29 Dec 1993, E. van den Berghe, 1♀ (CMNH); ECUADOR: Imbabura, Valle de la Chota, 16 km W Ambuqui, 17 Nov 1987, R. Davidson, 1♀, dissection MSA ♀ 3903 (CMNH); VENEZEULA: Aragua: Maracay, 5–11 Jul 1981, B. LaLanne-Cassou, 1♁, dissection MSA♁2027 (NYSM).</p> <p>Diagnosis. Solely on the basis of habitus, it is probably impossible to distinguish L. championi from L. humidicola Guenée, 1852 (Fig. 11) consistently. The forewing of L. championi is usually not as bright and contrasting as L. humidicola but otherwise similar. Males of both species have distinctive heavily tufted fore- and mid-tibia. The range of L. championi extends from Mexico to northwestern South America, and potentially overlaps L. humidicola in northern Mexico. The genitalia of both sexes are distinctive. In L. championi the extended basal sclerite of the clasper is attenuated into a sharp point, which reaches beyond the margin of the valva. In L. humidicola this structure is shorter and bluntly upturned to the mid-margin (Fig. 29). In L. championi the everted endophallus initially is a simple tube but has an unadorned pyramid-shaped diverticulum at approximately one-third of its length from base, followed by a single row of robust retrorse cornuti, which extends to the gonopore. In L. humidicola the everted endophallus (Fig. 30) is distinguished by a long narrow diverticulum, which arises near the base and ends in a long pointed cornutus. In both L. championi and L. humidicola (Fig. 43) the appendix bursae arises near the ostium bursae, however the ductus bursae in L. championi is a short straight tube ending in a sac-like corpus bursae whereas in L. humidicola this structure is long and ends in a twisted loop before entering the corpus bursae.</p> <p>Description. (Fig. 10) Wingspan 35–38 mm. Male palpi with dark scales, female palpi tan without dark scales; frons light tan. Thorax tan with three patagia bands, first band with brown scales, middle band less distinct, posterior band with distinct black-tipped scales; tegula and thorax tan; scale tufts on male fore- and mid-tibiae. Forewing ground light brown; cubital vein white scaled with brown shade for entire length; reniform reduced to a black dot at end of cell; p.m. line indicated by dots at veins, strongest at veins M1 and Cu2; veins white; terminal dots present. Hind wing of male pearly white, the female hind wing white, infuscated near margin. Ventral forewing light tan, darker in subcostal area. Ventral hind wing pearly white, costal margin tan. Abdomen light tan, shaggy; male basal abdominal eversible tubular structures present. Sexes similar, except female hind wing slightly darker near margin.</p> <p>Male genitalia. (Figs. 31 &amp; 32) Uncus, tegumen, and vinculum unmodified; cucullus short and rounded with a single row of fine non-deciduous setae in sockets on the lateral margin of both cucullus and valva; a large pore plate present at valvulus; ampulla thick and hook-like; digitus short and thick; editum inconspicuous; top of clasper sharp and hooked, whereas the basal sclerite produced into a long, straight, attenuated, sharp-pointed projection reaching or exceeding margin of valva; claval area of the sacculus with a slight prominence. Phallus long, thin and straight; everted endophallus with a short, pointed diverticulum at one-third length from base followed by a row of stout, retrorse cornuti, which extends to the gonopore.</p> <p>Female genitalia. (Fig. 44) Ductus bursae basally short and thick. Sclerotized appendix bursae branching to left and becoming sac-like, ending in ductus seminalis; ductus bursae continuing beyond the origin of the appendix bursae with a short, straight portion before entering thin walled, sac-like corpus bursae.</p> <p>Global distribution. Mexico, Guatemala (type locality), Costa Rica, Panama, Venezuela, Ecuador including Galápagos Islands (vide Hayes, 1975, as Mythimna solita).</p> <p>Food plant. Seaside Dropseed Sporobolus virginicus (Linnaeus) Kunth was reported as food plant of L. championi (as M. solita) on the Galápagos Islands (vide Hayes, 1975). Seaside Dropseed is a perennial tussock grass of coastal marshes, dunes, and beaches of tropical and subtropical countries worldwide.</p> <p>Larva. Hayes (1975) described the larva as follows: “Head gray with brown reticulation. Body reddish brown with darker markings and diffuse white lines.”</p> <p>Etymology. The specific epithet “championi ” honors George Charles Champion FLS (b.1851–d.1927) who collected specimens in Guatemala in 1879 for the “Biologia Centrali-Americana” (Selander &amp; Vaurie, 1962). Today we are faced with increasingly limited access to collecting at a time of precipitous declines in biodiversity. We wish to call attention to the often overlooked contribution of collectors who document our natural heritage.</p> <p>Remarks. We have not verified L. humidicola from Mexico, however as it occurs in California and Texas it undoubtedly also occurs in Mexico. We recognize two new junior synonyms of L. humidicola: Leucania februalis Hill, 1924, New Synonym, and Leucania elephas Troubridge, 2020, New Synonym. Leucania championi has been misidentified as L. humidicola in a recent publication (Troubridge, 2020). We reiterate the necessity of examination of types of all available names. The practice of wantonly ignoring primary types and junior synonyms leads to the generation of more synonyms hindering biodiversity research.</p> <p>Guatemala specimens of L. championi with Janzen codes 07-SRNP-102951 and 07-SRNP-10487 were sequenced in BOLD under the name Leucania februalis.</p> </div>	https://treatment.plazi.org/id/03B287EAFFE9FFE422B5FAE2FD537512	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Mccabe, Timothy L.;Adams, Morton S.	Mccabe, Timothy L., Adams, Morton S. (2023): Five new species of the genus Leucania Ochsenheimer in Central America (Lepidoptera: Noctuidae). Zootaxa 5256 (3): 250-266, DOI: 10.11646/zootaxa.5256.3.2, URL: http://dx.doi.org/10.11646/zootaxa.5256.3.2
