taxonID	type	description	language	source
03B087C8070444053ACF18CCDFAECD3F.taxon	description	Figs 1 – 4	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070444053ACF18CCDFAECD3F.taxon	materials_examined	Type materials. Holotype male: China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Linkouzi, 31 ° 28 ’ 19.47 ” N, 109 ° 52 ’ 58.34 ” E, alt. 1680 m, 17 August 2022, X. K. Jiang & H. M. Chen leg. Paratypes: 6 males and 2 females, same data as holotype; 1 female, same locality, 18 August 2022, X. K. Jiang & H. M. Chen leg.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070444053ACF18CCDFAECD3F.taxon	diagnosis	Diagnosis. This species is very similar to Zephronia hui Liu & Wesener, 2022 from Jiangkou County, Guizhou Province, China (Zhao et al. 2022), but it differs from the latter by the antennae slender (Fig. 2 A) (short and stout in Z. hui), the process of telopoditomere 2 of the anterior telopod obviously shorter than the length of telopoditomeres 3 and 4 (Figs 3 C, 4 C) (subequal in Z. hui).	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070444053ACF18CCDFAECD3F.taxon	etymology	Etymology. This species is named after the type locality, Linkouzi, noun in apposition.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070444053ACF18CCDFAECD3F.taxon	description	Description. Male body length ca. 19 – 31 mm. Width of thoracic shield = 8.0 – 11.0 mm, of tergite 8 = 8.8 – 12.0 mm. Height of thoracic shield = 4.3 – 6.0 mm, of tergite 8 = 6.0 – 8.2. Holotype 31 mm long, 11.0 mm (thoracic shield) wide, 6.0 mm (thoracic shield) high, 12.0 (tergite 8) wide, 8.2 (tergite 8) high. Female body length ca. 32 – 34 mm long. Width, of thoracic shield = 10.8 – 11.0 mm, of tergite 8 = 11.9 – 12.0 mm. Height, of thoracic shield = 6.4 – 6.9 mm, of tergite 8 = 8.3 – 8.9 mm. Body generally brown with irregular dark marks scattered on body surface (Fig. 1). Eyes with ca. 55 ocelli. Antennae short and thick. Last antennomere obviously longer than other antennomeres, lengths of antennomeres: 1 = 2 = 3 = 4 = 5 << 6 (Fig. 2 A). Apical disc with 39 – 55 apical cones (male), 24 – 27 (female) (Fig. 2 B, C). Organ of Tömösváry located inside antennal groove. Palpi sensory cones located in a single field. Head surface setose. Structure of gnathochilarium typical. Sensory cones of palpi all located in single field. Mandibles not examined. First stigmatic plate widely rounded, apex slightly curved anteriorly. Posterior-lateral margin of laterotergite 1 strongly projecting into a sharp tip. Laterotergite 2 with a broad, stout, much shorter projection. Collum glabrous except for marginal setae. Thoracic shield grooves deep, anterior margin thickened. Surface glabrous like tergites, setae only in grooves. Tergites surface glabrous. Tips of paratergites of midbody tergites projecting posteriorly. Inner section of endotergum lacking any spines or setae. Middle area with a single row of small, elliptical, cuticular impressions. Distance between impressions twice as wide as their diameter. Apically, 2 dense rows of short marginal bristles, tips of longest setae barely protruding beyond midpoint towards tergal margin. Anal shield massive, well-rounded, shiny and glabrous. Locking carina weakly developed. Leg 1 and 2 with 3 or 4 ventral spines. Leg pairs 4 – 21 (Fig. 2 D) with 8 – 10 ventral spines and single apical spine. Coxal process weakly developed and well-rounded (Fig. 2 D), absent from leg pairs 1 and 2. Tarsus 3.3 times longer than wide. Femur with toothed ridge of medium length. Male gonopore covered with a small, inconspicuous plate. Telopoditomere 1 of anterior telopods (Figs 3 A, B; 4 A, B) stout, slightly wider than long. Telopoditomere 2 as long as telopoditomere 3 in anterior view. Process of telopoditomere 2 originated posteriorly, broader than telopoditomere 3, tapering apically, curved, and protruding as high as basal part of telopoditomere 4. Inner margin towards movable finger with one large, membranous lobe. Posterior surface with a row of 5 crenulate and sclerotized teeth (Figs 3 C, 4 C). Telopoditomere 4 short, with at inner margin two long spines (Figs 3 A – C, 4 A – C). Mesial and lateral parts of telopoditomere 1 and anterior and lateral parts of telopoditomere 2 covered by setae. Other parts glabrous. Telopoditomere 1 of posterior telopods cylindrical, twice as long as wide. telopoditomere 2 stout (Figs 3 D, E; 4 D, E). Immovable finger (process of telopoditomere 2) slightly shorter than movable finger, consisting of telopoditomeres 3 and 4. Margin towards movable finger with two large, membranous lobes. Tip of immovable finger with a white spot in posterior view. Telopoditomere 3 elongated, 3 times longer than telopoditomere 4. Margin near immovable finger with a large membranous lobe and two slender spine, posterior surface with a row of 15 crenulate and sclerotized teeth. Telopoditomere 4 with at inner margin two long spines and a single membranous lobe. Entire telopod with few setae, mostly at anterior margins of telopoditomere 1 and inner margins of telopoditomere 2 (Figs 3 D, E; 4 D, E). Female vulva (Fig. 2 E) large, covering 2 / 3 of coxa, located at mesal margin, extending mesally to basal third of prefemur length. Operculum rounded. Subanal plate large and wide, not subdivided.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070444053ACF18CCDFAECD3F.taxon	distribution	Distribution. Known only from the type locality.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070044033ACF1A6BDC60C8F8.taxon	description	Figs 5, 6	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070044033ACF1A6BDC60C8F8.taxon	materials_examined	Materials examined: 1 male, China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Linkouzi, 31 ° 28 ’ 19.47 ” N, 109 ° 52 ’ 58.34 ” E, alt. 1680 m, 17 August 2022, X. K. Jiang & H. M. Chen leg; 1 female and 3 juveniles, same locality, 18 August 2022, X. K. Jiang & H. M. Chen leg.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070044033ACF1A6BDC60C8F8.taxon	distribution	Distribution. This species was reported from Sichuan, Guizhou and Hubei Provinces. This is the first record of this species from Chongqing.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070044033ACF1A6BDC60C8F8.taxon	discussion	Remarks. The characters of the habitus and the gonopods of this species are shown in Figs 5 – 6. All the collected locations of this species from China are near the Yangtze River, implying the species may disperse through the water flow.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80700440E3ACF1EF8DED0CB80.taxon	description	Figs 7 – 10	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80700440E3ACF1EF8DED0CB80.taxon	materials_examined	Type materials. Holotype male: China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Hongqi, Longtan Cave, 31 ° 30 ’ 40.73 ” N, 109 ° 49 ’ 40.93 ” E, alt. 1677 m, 14 August 2022, X. K. Jiang & H. M. Chen leg. Paratypes: 5 males and 9 females, same data as holotype; 1 male and 2 females, Yintiaoling National Nature Reserve, Hongqi, a manual tunnel under the Shuangtong Dam, 31 ° 31 ’ 16.83 ” N, 109 ° 49 ’ 51.84 ” E, alt. 1458 m, 14 August 2022, X. K. Jiang & H. M. Chen leg.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80700440E3ACF1EF8DED0CB80.taxon	etymology	Etymology. This specific name is an adjective Latin word, meaning ‘ inflated’ and refers to the shape of the prefemoroidal lateral processes of the gonopod.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80700440E3ACF1EF8DED0CB80.taxon	diagnosis	Diagnosis. This species can be distinguished from other congeneric species by the large and inflated prefemoroidal lateral processes of the gonopod (Figs 9 A, C; 10 A, C) (vs. clavate or stick-like in other species) and the particular shape of the apex of the gonopod (Figs 9 D, 10 D).	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80700440E3ACF1EF8DED0CB80.taxon	description	Description. Adult males body length 46 – 50 mm, width 2.8 – 3.0 mm, 56 – 58 pleurotergites + telson. Adult females body length 48 – 52 mm, width 3.1 – 3.4 mm, 57 – 60 pleurotergites + telson. Body coloration uniformly light brownish (Figs 7, 8). Head convex, as broad as pleurotergite 5, covered with minute setae. Eyes with ca. 55 pigmented ocelli in 8 or 9 horizontal rows (Fig. 8 A). Organ of Tömösváry small, about 2 – 3 times larger than an ocellus, situated close to anteroventral side of eye. Antennae slender, reaching the posterior edge of PT 6; length of antennomeres: 2 = 3> 4 = 5> 6> 7> 1; tip of antennomere 7 with four cones (Fig. 8 A). Collum much narrower than head (Fig. 7). pleurotergites 6 and 7 in males strongly enlarged (Fig. 8 A). Crests on collum and metazona moderately developed, 5 + 5 primary crests distinct, some secondary crests inconspicuous, 5 – 8 + 5 – 8 (Figs 7, 8 B), and ca. 12 small crests down to ventral pleurotergal edge. 3 rd primary crest strongly enlarged (Fig. 8 B). Ozopores starting with pleurotergite 6, present until penultimate pleurotergite, placed on tip of 3 rd primary crest. Setal pattern as in Table 1. Axial line rather distinct (Fig. 8 B). Epiproct simple, rear part with ca. 30 long setae in 3 – 4 transverse rows. Anal valves smooth, each divided into a small triangle and a large sclerite, both with a pair of macrosetae. Spinnerets comparatively thin and long, ending with a long macroseta. Hypoproct tripartite, medial sclerite largest, subrectangular, bearing two paramedian macrosetae; each lateral sclerite with a single macroseta (Fig. 8 C). Male leg-pairs 1 and 2 evidently shorter, leg-pair 3 slightly shorter than midbody legs. Midbody legs about twice as long as pleurotergite height. Coxal sacs present from 3 rd at least to leg-pair 18, but most abraded. Coxae 2 with a gonopore posteriorly, opening from a small cone. Coxae 7 normal without mesal spine; prefemora slightly incrassate. Tarsal pads small, present from 3 rd to about leg pair 21. Gonopod coxae freely connected through a medial membranous lamina. Coxa strongly broadened, bearing a large, broad anterior process, tapering apically, near the length of telopodite (Figs 9 A – C, 10 A – C). Prefemur with two processes. Lateral process inflated and drop-shaped, densely setose, about half the length of telopodite (Figs 9 A, C; 10 A, C). Inner process small, base with several setae (Figs 9 C, 10 C). Femoroidal stem long, slender, directed cephalad. Subterminal part of femoroidal stem broadened, forming a complex tip with ten apices (Figs 9 A – C, 10 A – C). One dorsal and one ventral process located subterminal part directed basally. Terminal part trifid (Figs 9 D, 10 D). Dorsal branch with two apices, basal one short and straight, directed mesally and distal one strongly bent, with a sharp tip. Ventral branch with three tips and a small apical lobe. A broad and rounded lobe located below the two branches. Seminal groove ending on the ventral branch. Female. Slightly larger than males, pleurotergites 2 and 3 strongly enlarged. Leg pairs 1 and 3 with tarsal pads. Leg pair 1 strongly reduced in size. Leg pair 2 normal. All legs with unmodified coxae. Cyphopods small, densely setose (Fig. 9 E).	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80700440E3ACF1EF8DED0CB80.taxon	distribution	Distribution. Known only from the type locality.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80700440E3ACF1EF8DED0CB80.taxon	discussion	Remarks. The family Paracortinidae contains two genera and 14 species, distributed in Southwest China (Yunnan and Sichuan) and Vietnam (Liu & Tian 2015). The new species described here is the most northeast one on record, significantly expanding the distribution range of this family.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070C440D3ACF1A6BD996CB4C.taxon	description	Figs 11, 12	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070C440D3ACF1A6BD996CB4C.taxon	materials_examined	Materials examined: 20 males and 38 females, China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Guanshan, Yanzi Cave, 31 ° 28 ’ 52.67 ” N, 109 ° 43 ’ 59.70 ” E, alt. 2230 m, 12 August 2022, X. K. Jiang & H. M. Chen leg; 1 male and 4 females, Yintiaoling National Nature Reserve, Guanshan, Gangkou Cave, 31 ° 30 ’ 32.22 ” N, 109 ° 41 ’ 33.65 ” E, alt. 2070 m, 13 August 2022, X. K. Jiang & H. M. Chen leg.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070C440D3ACF1A6BD996CB4C.taxon	distribution	Distribution. Shennongjia (Hubei) and Yintiaoling (Chongqing).	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070C440D3ACF1A6BD996CB4C.taxon	discussion	Remarks. The characters of the habitus and the gonopods of this species are shown in Figs 11 – 12. The type locality of this species is in Shennongjia National Natural Reserve, Hubei Province (Mauriès & Nguyen Duy-Jacquemin, 1997), bordering on Yintiaoling. It is not surprising to collect this species in the region.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070E440D3ACF1AB3D878C90C.taxon	description	Figs 13 – 15	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070E440D3ACF1AB3D878C90C.taxon	materials_examined	Material examined. 1 male, China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Linkouzi, 31 ° 28 ′ 30.0 ″ N, 109 ° 52 ′ 40.8 ″ E, alt. 1213 m, 17 August 2022, X. K. Jiang & H. M. Chen leg.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070E440D3ACF1AB3D878C90C.taxon	discussion	Remarks. The characters of the habitus and the gonopods of this species are shown in Figs 13 – 15. This species is easily distinguished by the coxite with two strong setae ventrolaterally, the prefemoral part less than half as long as the acropodite, and the latter distinctly excavate, forming a mesal cavity surrounded by fringed or microspiculate lamellae. It has hitherto been throughout Taiwan and Jiangsu, China, as well as in Japan and Vietnam (Golovatch et al. 2018). The above samples thus represent the first formal record of N. nodulosa in Chongqing, China.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070844083ACF1B95D846CAA8.taxon	description	Figs 16, 17	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070844083ACF1B95D846CAA8.taxon	materials_examined	Materials examined. 5 males and 7 females, China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Hongqi, 31 ° 30 ’ 25.43 ” N, 109 ° 49 ’ 28.42 ” E, alt. 1166 m, 13 August 2022, X. K. Jiang & H. M. Chen leg; 13 males and 11 females, Yintiaoling National Nature Reserve, Linkouzi, 31 ° 28 ’ 19.47 ” N, 109 ° 52 ’ 58.34 ” E, alt. 1680 m, 16 August 2022, X. K. Jiang & H. M. Chen leg.; 6 males and 8 females, same locality, 17 August 2022, X. K. Jiang & H. M. Chen leg; 2 males and 4 females, Yintiaoling National Nature Reserve, Guanshan, Yanzi Cave, 31 ° 28 ’ 52.67 ” N, 109 ° 43 ’ 59.70 ” E, alt. 2230 m, 12 August 2022, X. K. Jiang & H. M. Chen leg; 1 female, Yintiaoling National Nature Reserve, Shizhuzi, 31 ° 32 ’ 4.68 ” N, 109 ° 42 ’ 16.26 ” E, alt. 2210 m, 11 August 2022, X. K. Jiang & H. M. Chen leg.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070844083ACF1B95D846CAA8.taxon	distribution	Distribution. This species is widespread in central and southern China (Qinghai, Gansu, Shaanxi, Sichuan, Guizhou, Yunnan and Zhejiang) (Golovatch 2019).	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070844083ACF1B95D846CAA8.taxon	discussion	Remarks. The characters of the habitus and the gonopods of this species are shown in Figs 16 – 17.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070B44163ACF1A06D85BC8A0.taxon	description	Figs 18 – 20	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070B44163ACF1A06D85BC8A0.taxon	materials_examined	Type materials. Holotype male: China, Chongqing, Wuxi County, Tongcheng Town, Zengjiayan Cave, 31 ° 22 ’ 30.23 ” N, 109 ° 46 ’ 11.59 ” E, alt. 1301 m, 15 August 2022, X. K. Jiang & H. M. Chen leg. Paratypes: 3 males, same data as holotype; 2 males and 2 females, Yintiaoling National Nature Reserve, Hongqi, Longtan Cave, 31 ° 30 ’ 40.73 ” N, 109 ° 49 ’ 40.93 ” E, alt. 1677 m, 14 August 2022, X. K. Jiang & H. M. Chen leg.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070B44163ACF1A06D85BC8A0.taxon	diagnosis	Diagnosis. This species can be distinguished from other species in this genus by the femorite of the gonopods extremely bent, lamellar processes present apically, and without an oblique lateral sulcus (Figs 19, 20).	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070B44163ACF1A06D85BC8A0.taxon	etymology	Etymology. This species is named by the lamellar processes of the tip of femorite of the gonopods, adjective.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070B44163ACF1A06D85BC8A0.taxon	description	Description. Length ca. 43 – 50 mm (male), 45 – 48 mm (female), width of midbody pro- and metazona 3.7 – 3.8 and 5.2 – 5.3 mm (male), 3.9 – 4.7 and 5.2 – 6.0 mm (female), respectively. Head and dorsum of the body black, paraterga light yellow, venter and legs reddish brown (Fig. 18 A). Clypeolabral region sparsely setose, vertex bare, epicranial suture distinct. Antennae slender, antennomeres 2 – 7 clavate. Lengths of antennomeres: 3 ≈ 2 ≈ 4 ≈ 5 ≈ 6> 1> 7. Collum glabrous (Fig. 18 A). Paraterga only slightly declivous, broadly rounded, and narrowly bordered. Postcollum constriction obvious. Tegument of metaterga shining, rugulose, prozona and metazona below paraterga smooth. Axial line barely visible both on pro- and metazona. Paraterga strongly developed, broad, subhorizontal, always lying below dorsum, thick in lateral view (Fig. 18 A, B). Ozopores evident, lateral, lying in an ovoid groove at about 1 / 3 in front of caudal corner. Pore formula normal. Transverse sulcus evident, narrow, rather shallow and only slightly incomplete on metaterga 2 and 3, reaching base of paraterga on metaterga 4 – 18. Stricture between pro- and metazona narrow and shallow, evidently beaded at bottom down to base of paraterga. Sterna sparsely setose, without modifications, but with a small, rounded cone between male coxae 4. No conspicuous ridge in front of gonopod aperture. Legs long and slender, obviously longer than body height. Prefemora without modifications. Epiproct conical, flattened dorsoventrally, apical papillae well-developed (Fig. 18 C). Hypoproct arch shaped. Gonopods (Figs 19, 20) simple. Coxa long and slender, with several setae distodorsally (Figs 19, 20). Prefemora densely setose, about 1 / 3 as long as acropodite (Figs 19, 20). Femorite slender, evidently curved and not enlarged distad, without a lateral sulcus (Figs 19, 20). Tip of femorite present two lamellar processes (Figs 19 B, 20 B). One situated at the inner margin, with a sharp tip. Another one situated at the base of solenophore. Solenophore bent as semicircle. Tip of solenophore slightly bifid, upper denticle slender and longer than lower lobule. Lower lobule broad (Figs 19 B, 20 B). Solenomere about as long as solenophore, flagelliform (Figs 19 A, B; 20 A, B).	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070B44163ACF1A06D85BC8A0.taxon	distribution	Distribution. Known only from the type locality.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8070B44163ACF1A06D85BC8A0.taxon	discussion	Remarks. Orthomorpha is distributed in southeast Asia and it is very similar to the sympatric genus Antheromorpha (Likhitrakarn et al. 2011, 2016). Their taxonomic boundary is vague. Orthomorpha can only be distinguished from the latter by the poorly differentiated gonopod tip, usually feebly bi- or trifid, whereas Antheromorpha shows a deeply bifid gonopod tip (Likhitrakarn et al. 2011, 2016). Considering the structure of the gonopod tip of the new species, we chose to place it in Orthomorpha. Orthomorpha is a large genus comprising 50 species, and has been reviewed recently (Likhitrakarn et al. 2011; Nguyen & Sierwald 2013). However, no species of this genus have been reported from China. Therefore, the new species described here is the first Orthomorpha recorded in this country. Two individuals from deep inside Zengjiayan Cave show the distinct pale coloration of the body (Fig. 18 B), but their gonopods are identical to the epigean congeners indicating that they belong to the same species.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071544163ACF180ED82ACEF0.taxon	description	Figs 21, 22	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071544163ACF180ED82ACEF0.taxon	materials_examined	Materials examined: 2 males, China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Hongqi, 31 ° 30 ’ 25.43 ” N, 109 ° 49 ’ 28.42 ” E, alt. 1166 m, 16 August 2022, X. K. Jiang & H. M. Chen leg; 11 males and 15 females, Yintiaoling National Nature Reserve, Guanshan, 31 ° 31 ’ 0.78 ” N, 109 ° 43 ’ 18.33 ” E, alt. 2034 m, 12 August 2022, X. K. Jiang & H. M. Chen leg.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071544163ACF180ED82ACEF0.taxon	distribution	Distribution. Cosmopolitan.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071544163ACF180ED82ACEF0.taxon	discussion	Remarks. The characters of the habitus and the gonopods of this species are shown in Figs 21 – 22. This species is distributed nearly world-wide in tropical and temperate environments and closely relates to human activities.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071744153ACF1BDBDFAECDA6.taxon	description	Figs 23 – 25	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071744153ACF1BDBDFAECDA6.taxon	materials_examined	Type materials. Holotype male: China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Huangcaoping, 31 ° 25 ’ 7.20 ” N, 109 ° 55 ’ 59.33 ” E, alt. 2098 m, 20 August 2022, X. K. Jiang & H. M. Chen leg. Paratypes: 2 females, same data as holotype; 1 male, Yintiaoling National Nature Reserve, Linkouzi, 31 ° 28 ’ 19.47 ” N, 109 ° 52 ’ 58.34 ” E, alt. 1680 m, 17 August 2022, X. K. Jiang & H. M. Chen leg; 2 females, Yintiaoling National Nature Reserve, Shizhuzi, 31 ° 32 ’ 4.68 ” N, 109 ° 42 ’ 16.26 ” E, alt. 2210 m, 11 August 2022, X. K. Jiang & H. M. Chen leg.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071744153ACF1BDBDFAECDA6.taxon	diagnosis	Diagnosis. This species is very similar to Polylobosoma panda (Golovtach, 2009) from Foping Nature Reserve, Shaanxi Province, China. It can be distinguished from the latter by the tips of the solenomere and the solenophore pointed laterally (Figs 24, 25), not caudally as in P. panda.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071744153ACF1BDBDFAECDA6.taxon	etymology	Etymology. This specific name is an adjective Latin word, meaning ‘ coronal’ and refers to the shape of the solenophore of the gonopod.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071744153ACF1BDBDFAECDA6.taxon	description	Description. Length ca. 43 – 46 mm (male), 44 – 45 mm (female), width of midbody pro- and metazona 4.0 – 4.1 mm and 5.8 – 6.1 mm (male), 5.0 – 5.2 mm and 6.1 – 6.3 mm (female), respectively. Head and dorsum of the body black, paraterga whitish or light yellow, venter and legs reddish brown (Fig. 23). Clypeolabral region poorly setose, vertex glabrous, epicranial suture distinct. Antennae slender, antennomeres 2 – 7 clavate. Lengths of antennomeres: 3 ≥ 2 ≥ 4 ≥ 5 ≥ 6> 1> 7. Collum glabrous (Fig. 23 A). Paraterga only slightly declivous, broadly rounded, and narrowly bordered (Fig. 23 A, B). Postcollum constriction moderate (Fig. 23 A). Tegument of metaterga shining, rough, and leather-like; prozona and metazona below paraterga smooth. Axial line barely visible both on pro- and metazona. Paraterga strongly developed, broad, subhorizontal, located in the middle of body segment in lateral view (Fig. 23 A, B). Ozopores evident, lateral, lying in an ovoid groove at about 1 / 3 in front of caudal corner. Pore formula normal. Transverse sulcus evident reaching base of paraterga on metaterga 5 – 17, absent in metaterga 2 – 4. Stricture between pro- and metazona narrow and shallow, evidently beaded at bottom down to base of paraterga. Sterna sparsely setose, without modifications, but with a small, rounded cone between male coxae 4 (Fig. 23 C). No conspicuous ridge in front of gonopod aperture. Legs long and slender, obviously longer than body height. Prefemora without modifications. Epiproct conical, flattened dorsoventrally, apical papillae well-developed (Fig. 23 D). Hypoproct arch shaped. Gonopods simple (Figs 24, 25). Coxa subcylindrical, poorly setose distodorsally (Figs 24 A, C; 25 A, C). Prefemora densely setose, about 1 / 3 as long as acropodite (Figs 24, 25). Femorite slender, slightly curved and not enlarged distad, without a lateral sulcus (Figs 24, 25). Tip of femorite present one small lobe (Figs 24 A, B; 25 A, B). Solenophore obviously bent. Anterior margin of solenophore expanded anteriorly, corona-shaped (Figs 24, 25). Tip of solenophore deeply trifid, all branches with a sharp tip. Solenomere about as long as solenophore, flagelliform (Figs 24 B, C; 25 B, C).	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071744153ACF1BDBDFAECDA6.taxon	distribution	Distribution. Known only from the type locality.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071144123ACF1BDBD8D9C848.taxon	description	Figs 26, 27	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071144123ACF1BDBD8D9C848.taxon	materials_examined	Materials examined: 2 males and 1 female, China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Shizhuzi, 31 ° 32 ’ 4.68 ” N, 109 ° 42 ’ 16.26 ” E, alt. 2210 m, 11 August 2022, X. K. Jiang & H. M. Chen leg.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071144123ACF1BDBD8D9C848.taxon	discussion	Remarks. The characters of the habitus and the gonopods of this species are shown in Figs 26 – 27. Sellanucheza is a small genus containing six species distributed in China (three species in Sichuan, Chongqing and Shaanxi) and northern Vietnam. The generic name has been changed from Szechuanella Hoffman, 1961 to the anagram Sellanucheza because of preoccupation (Enghoff et al. 2004), and the original name refers to Sichuan Province, China. The type locality of S. jaegeri is in Houzhenzi, Shaanxi, about 400 km from Yintiaoling. Further investigation of the actual distribution range of this species is needed, especially on some areas of south slope of Qinling Mountains.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071144133ACF1C01DF25CAA9.taxon	description	Figs 28 – 31	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071144133ACF1C01DF25CAA9.taxon	materials_examined	Materials examined. 1 Male and 1 female, China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Yanping, 31 ° 31 ′ 9.1 ″ N, 109 ° 43 ′ 1.2 ″ E, alt. 1802 m, 11 August 2022, X. K. Jiang & H. M. Chen leg.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071144133ACF1C01DF25CAA9.taxon	discussion	Remarks. This species is very common in southern China, and has been recorded from Gansu, Sichuan and Yunnan Provinces. The new samples almost fully agree with the original description. The habitus and gonopodal structures are illustrated here (Figs 28 – 31).	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80719441A3ACF1BDBD948CC00.taxon	description	Figs 32 – 34	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80719441A3ACF1BDBD948CC00.taxon	materials_examined	Type materials. Holotype Male: China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Hongqi, Longtan Cave, 31 ° 30 ’ 40.73 ” N, 109 ° 49 ’ 40.93 ” E, alt. 1677 m, 14 August 2022, X. K. Jiang & H. M. Chen leg. Paratypes: 12 females, same data as holotype; 1 male, a manual tunnel near the Longtandong Cave, 31 ° 31 ’ 16.83 ” N, 109 ° 49 ’ 51.84 ” E, alt. 1458 m, same time and collectors as holotype; 1 female, same cave as holotype, 11 August 2022, Q. D. Zhu leg.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80719441A3ACF1BDBD948CC00.taxon	etymology	Etymology. The specific name is a noun in apposition, derived from the type locality, Wuxi County.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80719441A3ACF1BDBD948CC00.taxon	diagnosis	Diagnosis. The new species is similar to Epanerchodus latus Liu & Golovatch, 2018 in having the gonopodal femorite with a finger-shaped process distally and a process medially, the endomere carrying a subapical lobe, but distinguished by the broad subapical endomere with a spiny process and a row of apical denticles (Fig. 34).	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80719441A3ACF1BDBD948CC00.taxon	description	Description: Length of holotype 22 mm, male paratypes 22 mm, female paratypes 20 – 37 mm; width of midbody pro- and metazona in holotype 1.6 mm and 3.1 mm, respectively, another male paratype 1.5 mm and 3.0 mm, in female paratypes up to 1.2 – 1.9 mm and 2.4 – 3.5 mm. Live specimens whitish to light yellow and semitransparent (Fig. 32 A), in alcohol light yellow to light brown (Figs 32 B, C, 33). Adults with 20 segments. Width of holotype: head (1.8) << collum (2.7) <segment 2 (2.8) = 3 = 4 = 5 <6 (2.9) – 8 <9 (3.1) – 16, thereafter segments 17 (2.9) – 19 (1.7). Body gradually tapering posteriorly towards telson (Figs 32, 33). Head covered with sparse pubescence throughout, vertigial sulcus apparent (Figs 32 A, B; 33 A, C). Antennae long and slender, clavate, in situ reaching behind segment 4. Collum fan-shaped, with irregular sparse setae, and a faint lateral incision / denticle on each side (Figs 32 A, B; 33 C). Paraterga distinct and broad (Figs 32, 33), midbody metaterga about 1.9 times as wide as prozonae. All paraterga slightly upturned dorsally above a faintly convex dorsum (Fig. 32). Anterior edge of metaterga forming a shoulder (Figs 32, 33). Caudolateral corners of paraterga from rings 2 protruded toward back-inner past tergal margin, subsequent ones acuminate, especially on rings 18 – 19 (Figs 32, 33). Integument shining, translucent, prozonae very delicately alveolate (Figs 32, 33 C). Three or four notches at lateral margins of paraterga (Figs 32, 33). Constriction between pro- and metazonae narrow, shallow and smooth (Fig. 32). Metatergal sculpture faint, with three irregular transverse rows of setigerous polygonal bosses (Fig. 32). Sulcus between front and middle rows of setae a little deeper than that between middle and caudal rows (Fig. 32). Tergal setae very short, sparse. Pore formula normal: 5, 7, 9, 10, 12, 13, 15 – 19. Ozopores large and evident, located between last and penultimate marginal incisions. Epiproct short, conical, slightly surpassing paraprocts caudally, pre-apical lateral papillae small, with two long setae (Fig. 32 C). Hypoproct lunate, with two setigerous conical papillae at caudal margin (Fig. 33 B, C). Sterna sparsely setose, cross-shaped impressions shallow (Fig. 33 A, B). Legs long and slender, about 2.0 times as long as body ring height. Coxa of gonopod (Fig. 34) large and squarish, as usual. Prefemur densely setose and nearly half the length of telopodite. Femorite with a clivus (cl), a process (p 2) medially and a finger-shaped process (p 1) distally. Endomere (en) strongly curved, broad subapical, with a small subapical curved lobe (l), a row of apical denticles (d) and a spiny protuberance medially (sp). Seminal groove starting mesally, distally recurved laterad near base of p 1, then run into an accessory seminal chamber, the latter opening on a hairy pulvillus, which makes a clear-cut distolateral loop. Exomere absent.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C80719441A3ACF1BDBD948CC00.taxon	distribution	Distribution. Known only from the type locality, Wuxi, Chongqing, China.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071944193ACF1DA7D861CAA8.taxon	description	Figs 35 – 37	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071944193ACF1DA7D861CAA8.taxon	materials_examined	Type materials. Holotype male: China, Chongqing, Wuxi County, Yintiaoling National Nature Reserve, Linkouzi, 31 ° 28 ’ 19.47 ” N, 109 ° 52 ’ 58.34 ” E, alt. 1680 m, 17 August 2022, X. K. Jiang & H. M. Chen leg. Paratypes: 2 males and 1 female, same data as holotype.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071944193ACF1DA7D861CAA8.taxon	diagnosis	Diagnosis. This species is very similar to R. davidiani Golovatch, 2014 from Sichuan in the slender and bent prefemoral process and solenomere of the gonopods, but differs from the latter by the prefemoral process of the gonopods only slightly bending mesially (Figs 36 A – C, 37 A – C) not forming semicircle as in R. davidiani.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071944193ACF1DA7D861CAA8.taxon	etymology	Etymology. The species name is derived from the shape of the prefemoral process of the gonopods, meaning ‘ spine’, noun.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071944193ACF1DA7D861CAA8.taxon	description	Description. Length ca. 48 – 52 mm, midbody paranotal width 9.7 – 10.6 mm, metatergal length 4.1 – 4.8 mm, collum width 8.3 – 8.5 mm, length 3.3 – 3.6 mm. Body uniformly light yellow (Fig. 35 A). Fluorescence in UV light strong (Fig. 35 B). Head smooth, epicranial suture distinct, several setae scattered above clypeus, with 2 dense rows at its margin and on labrum. Antennae slender. Antennomeres 1 and 7 short and small, slightly longer than wide. Antennomeres 2 – 6 clavate. Lengths of antennomeres: 6> 5 ≈ 4 ≈ 3 ≈ 2> 1> 7. Gnathochilarium densely setose. Collum convex, smooth, shiny, without marginal ridge, tapering laterally. Pro- and metaterga smooth without any traces of tubercles or punctuation, not even wrinkles (Fig. 35 A). Posteriolateral edge of paranota strongly pointed caudad. Pore formula normal, pores on paranota 5,7,9,10,12,13,15,16,17, and 18, in median excavation of paranota (in lateral view). Epiproct in dorsal view subtriangular, in lateral view protruding over paraprocts, parallel-sided, slightly curved ventrad, with 7 + 7 setae, 3 + 3 of them sitting on knobs. Paraprocts strongly marginate with 2 + 2 setae, hypoproct with 1 + 1 setae on knobs. Midbody legs well separated, sterna wide and smooth. Postgonopodal legs (Fig. 36 E) with moderately developed ventral spine on prefemur, increasingly stronger towards body end. Femur as long as tarsus. Postfemur stout, about half-length of femur. Tibia straight, both subequal in length of postfemur. Tarsus slender. Claw curved. Male 2 nd leg pair coxa with strong median projections about half as long as length of coxa, densely setose, without membraneous tubules apically (Fig. 36 D). Male gonopodal aperture on segment 7 wide, elliptical, about twice as wide as long, gonopods in situ usually deeply embedded, with acropodites crossing each other. Coxa of gonopod (Figs 36 A – C, 37) stout, approximately as long as wide, glabrous, without proximal apophysis. Cannula normal, situated on mesal side (Figs 36 B, C; 37 B, C). Telopodite consists of two slender and simple processes, prefemoral process and acropodite (solenomere), typical for Riukiaria. Prefemur densely covered with long hairs. Base of prefemoral process developed, originated from the base of prefemur. Prefemoral process extremely slender, spine-like bending mesially, as long as acropodite, with no setae, hairs, or processes. Acropodite long, scythe-shaped with a sharp tip, arched proximally towards prefemoral process. Prostatic groove running along mesal side, and ending on the pointed tip (Fig. 37 B).	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071944193ACF1DA7D861CAA8.taxon	distribution	Distribution. Known only from the type locality.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
03B087C8071944193ACF1DA7D861CAA8.taxon	discussion	Remarks. Riukiaria contains 36 species distributed in China and Japan (Marek et al. 2014). In China, thirteen species have been reported from Shaanxi (1 species), Sichuan (2 species), Zhejiang (2 species), Fujian (1 species) and Taiwan (7 species). Riukiaria spina sp. nov. is the first species of Riukiaria reported from Chongqing.	en	Chen, Hui-Ming, Zheng, Chang-Bin, Jiang, Xuan-Kong (2023): The millipedes (Diplopoda) in Yintiaoling National Natural Reserve, Southwest China. Zootaxa 5257 (1): 49-81, DOI: 10.11646/zootaxa.5257.1.6, URL: http://dx.doi.org/10.11646/zootaxa.5257.1.6
