taxonID	type	description	language	source
9954879CE7635603FC774105FBA0F84C.taxon	materials_examined	TYPE SPECIES. — Cyclostoma acutum Draparnaud, 1805 by subsequent designation of Gray (1847).	en	Esu, Daniela, Girotti, Odoardo (2019): Two new oligohaline Hydrobia (s. l.) (Caenogastropoda) from the transitional Upper Oligocene Galatone Fm (Apulia, Southern Italy). Geodiversitas 41 (1): 1-9, DOI: 10.5252/geodiversitas2019v41a1
9954879CE7625600FED04058FD9EF9C9.taxon	description	(Figs 2 A, B; 3 A-C)	en	Esu, Daniela, Girotti, Odoardo (2019): Two new oligohaline Hydrobia (s. l.) (Caenogastropoda) from the transitional Upper Oligocene Galatone Fm (Apulia, Southern Italy). Geodiversitas 41 (1): 1-9, DOI: 10.5252/geodiversitas2019v41a1
9954879CE7625600FED04058FD9EF9C9.taxon	materials_examined	TYPE MATERIAL. — Holotype: MPUR 7 - 3897 (Fig. 3 A), H = 4.9 mm, W = 2.3 mm. — Paratypes: MPUR 7 - 3898 / 2, 4198 / 1, 4211 / 90, 4200 / 40. DERIVATION OF NAME. — Named after the type locality. TYPE HORIZON. — Limy mudstone layers of Galatone Fm at Galatone (Esu et al. 1994; Bossio et al. 1998). TYPE LOCALITY. — Galatone (Lecce, Apulia, Italy) (40 ° 8 ’ 27.03 ” N, 18 ° 03 ’ 0.37 ” E). TYPE AGE. — Chattian (Late Oligocene).	en	Esu, Daniela, Girotti, Odoardo (2019): Two new oligohaline Hydrobia (s. l.) (Caenogastropoda) from the transitional Upper Oligocene Galatone Fm (Apulia, Southern Italy). Geodiversitas 41 (1): 1-9, DOI: 10.5252/geodiversitas2019v41a1
9954879CE7625600FED04058FD9EF9C9.taxon	diagnosis	DIAGNOSIS. — Small-sized shell, elongately conical, with up to 6 ¼ whorls, moderately convex to slightly flattened in the middle, often providing a faint angulation in the lower part of the whorl. Aperture ovate to quadrangular ovate, adapically rounded to barely angulate. Umbilicate.	en	Esu, Daniela, Girotti, Odoardo (2019): Two new oligohaline Hydrobia (s. l.) (Caenogastropoda) from the transitional Upper Oligocene Galatone Fm (Apulia, Southern Italy). Geodiversitas 41 (1): 1-9, DOI: 10.5252/geodiversitas2019v41a1
9954879CE7625600FED04058FD9EF9C9.taxon	description	DESCRIPTION Shell small-sized, solid, elongately conical, more or less turreted, with up to 6 ¼ regularly increasing whorls, moderately convex to slightly flattened in the middle, often providing a faint angulation in the abapical part and a step-like margin adapically, in proximity of the suture. Suture incised, deep. Last whorl high just under two thirds of shell height. Protoconch of c. 480 µm maximum diameter, smooth, consisting of one and a quarter slightly convex whorl (Fig. 2 B). Teleoconch glossy, with thin, prosocline to nearly orthocline and rather regularly spaced growth lines gradually appearing after the protoconch. Aperture ovate to quadrangular-ovate, adapically rounded to barely angulate, sometimes slightly thickened inside along the outer lip; peristome simple, continuous, columellar lip smoothly concave, adherent partially to the last whorl, outer lip slightly arched in lateral profile. Sometimes transverse, more or less raised wrinkles, irregularly spaced, occur on the last whorl just before the aperture (Figs 2 A; 3 Ac, Bc). Umbilicus open, narrow, and deep. Range dimensions: H = 4.1 - 5 mm, W = 2.1 - 2.4 mm, H / W = 1.95 - 2.08.	en	Esu, Daniela, Girotti, Odoardo (2019): Two new oligohaline Hydrobia (s. l.) (Caenogastropoda) from the transitional Upper Oligocene Galatone Fm (Apulia, Southern Italy). Geodiversitas 41 (1): 1-9, DOI: 10.5252/geodiversitas2019v41a1
9954879CE7625600FED04058FD9EF9C9.taxon	discussion	REMARKS The shell shape of Hydrobia (s. l.) galatoniana n. sp. shows a certain degree of variability, varying from more (Fig. 3 B) to less turreted (Fig. 3 C), with moderately convex to slightly flattened whorls, and ovate (Fig. 3 Ba) to quadrangularovate aperture (Fig. 3 Aa, Ca). Comparisons of the study material with several members of Hydrobiidae from the literature, and those stored in the rich malacological collection at the Senckenberg Museum of Frankfurt a. M., showed similarity mainly to two species: Hydrobia dubuissonii (Bouillet, 1834) and Hydrobia andreaei O. Boettger in Degrange-Touzin, 1892. Bouillet (1834: captions to pl. 18, fig. 14) gave a good description of H. dubuissonii from the Oligocene of the type locality (Aurillac, France), whilst the accompanying picture is unclear. Lozouet & Maestrati (2012: fig. 183.1 - 8) supplied good illustrations of French specimens of this species from the Oligocene (Stampian) of the Paris Basin. Hydrobia dubuissonii from Oligocene deposits of France (Aurillac and Paris Basin), and Middle Oligocene (Rupelian) of Vieux-Jone, Belgium, stored at the Senckenberg Museum (SMF 12.1594 c, ex coll. O. Semper), differs from H. (s. l.) galatoniana n. sp. in having a conical-pyramidal shape, a more acute spire with more prominent apex, whorls not stepped, narrower, more ovate and adapically more angled aperture, and a narrower umbilicus (cf. Esu & Girotti 2010: pl. 3, fig. 8; Lozouet & Maestrati 2012). Maximum dimensions (H = 4 mm, W = 2 mm) given by Bouillet (1834) for H. dubuissonii from the type-locality are on average smaller than those measured for the Galatone specimens (H = 4.1 - 5 mm, W = 2.1 - 2.4 mm). Hydrobia andreaei from Early Miocene localities of France, Cabanac (Pouquet), Aquitanian (SMF 245531 / 1 - 245532 / 1, coll. O. Boettger, ex Degrange-Touzin), Saucats (near Bordeaux), Aquitanian (SMF 245533 / 9, coll. O. Boettger), Lariey (Saucats), Aquitanian (SMF 245550 / 2, coll. O. Boettger), and Oligocene (Chattian, Cyrenenmergel) of ṙKolbsheim, Elsass (SMF 245534 / 24, coll. O. Boettger, ex A. Andreae) (cf. Degrange-Touzin 1892: 181), differs from H. (s. l.) galatoniana n. sp. in having a more cylindrical and elongate shell with more acute apical angle, a shallower suture, the last whorl slightly descending towards the aperture, restricted and provided with a faint ring-like varix close to the outer lip, a narrower and semiovate aperture, columellar margin slightly thickened and more oblique, outer margin well arched, a not deep umbilicus, and on average smaller size (H = 3.75 - 4.5, W = 1.62 - 1.75) (cf. Boettger in Degrange- Touzin 1892: pl. 5, figs 1 - 1 c; Lozouet et al. 2001: fig. 9). Hydrobia (s. l.) galatoniana n. sp. is distinct from the coeval Hydrobia dubuissonii hydruntina Esu & Girotti, 2010, from the Upper Oligocene marls of the bauxite quarry at Otranto (Apulia) (Esu & Girotti 2010: pl. 3, figs 1 - 7), in having a more cylindrical shell, more convex whorls, wider quadrangular-ovate aperture, and on average larger size (holotype of H. dubuissonii hydruntina: H = 3.8 mm, W = 2 mm; mean size: H = 4 mm, W = 2 mm). Quaggiotto & De Angeli (2016: fig. 3.1 - 3) illustrated some not well-preserved remains of Hydrobia cf. dubuissonii from the Lower Oligocene of Valle del Ponte (Vicenza, northeastern Italy). These specimens are clearly different from H. (s. l.) galatoniana n. sp. for their larger size (H = 7 mm), and more elongate and turreted shell shape.	en	Esu, Daniela, Girotti, Odoardo (2019): Two new oligohaline Hydrobia (s. l.) (Caenogastropoda) from the transitional Upper Oligocene Galatone Fm (Apulia, Southern Italy). Geodiversitas 41 (1): 1-9, DOI: 10.5252/geodiversitas2019v41a1
9954879CE760560FFEF140C5FD22FBF6.taxon	description	(Figs 2 C, D; 4 A-C)	en	Esu, Daniela, Girotti, Odoardo (2019): Two new oligohaline Hydrobia (s. l.) (Caenogastropoda) from the transitional Upper Oligocene Galatone Fm (Apulia, Southern Italy). Geodiversitas 41 (1): 1-9, DOI: 10.5252/geodiversitas2019v41a1
9954879CE760560FFEF140C5FD22FBF6.taxon	materials_examined	TYPE MATERIAL. — Holotype: MPUR 7 - 3899 (Fig. 4 A), H = 4.5 mm, W = 2.5 mm. — Paratypes: MPUR 7 - 4197 / 2, 4199 / 1, 3900 / 40. DERIVATION OF NAME. — Named after the Ionian Sea, whose shoreline is near the type locality. TYPE HORIZON. — Limy mudstone layers of Galatone Fm at Galatone (Esu et al. 1994; Bossio et al. 1998). TYPE LOCALITY. — Galatone (Lecce, Apulia, Italy) (40 ° 8 ’ 27.03 ” N, 18 ° 03 ’ 0.37 ” E). TYPE AGE. — Chattian (Late Oligocene).	en	Esu, Daniela, Girotti, Odoardo (2019): Two new oligohaline Hydrobia (s. l.) (Caenogastropoda) from the transitional Upper Oligocene Galatone Fm (Apulia, Southern Italy). Geodiversitas 41 (1): 1-9, DOI: 10.5252/geodiversitas2019v41a1
9954879CE760560FFEF140C5FD22FBF6.taxon	diagnosis	DIAGNOSIS. — Small-sized shell, ovately conical, with up to 5 slightly convex whorls, the last large and elongate. Aperture pear-shaped, relatively high and elongate, adapically barely angulate, somewhat flared and slightly thickened inside. Umbilicus slit-like.	en	Esu, Daniela, Girotti, Odoardo (2019): Two new oligohaline Hydrobia (s. l.) (Caenogastropoda) from the transitional Upper Oligocene Galatone Fm (Apulia, Southern Italy). Geodiversitas 41 (1): 1-9, DOI: 10.5252/geodiversitas2019v41a1
9954879CE760560FFEF140C5FD22FBF6.taxon	description	DESCRIPTION Shell small, solid, thick-walled, ovately conical, with up to 5 slightly convex whorls, the last large, elongate, exceeding two-thirds of shell height. Early whorls slowly increasing, the last faster and somewhat descending (Figs 2 C; 4 Ab, Ba). Suture shallow, oblique between the penultimate and ultimate whorl. Protoconch of c. 410 µm maximum diameter, smooth, consisting of one slightly convex whorl. Boundary to teleoconch distinct by the formation of closely spaced growth lines (Fig. 2 D). Teleoconch glossy, with very delicate and thin prosocline growth lines. An axial swelling occurs sometimes in the last whorl towards the aperture. Aperture pear-shaped, high, elongate, adapically barely angulate and abapically rounded, somewhat flared and slightly thickened inside; peristome continuous, columellar lip almost completely adherent to the last whorl, outer lip straight in lateral profile. Umbilicus slit-like. Range dimensions: H = 4 - 4.5 mm, W = 2.5 - 2.7 mm, H / W = 1.48 - 1.8. Provisionally, Esu et al. (1994) assigned this stout form to the subgenus Pseudamnicola (Staja) Brusina, 1897, currently considered a separate genus of the family Hydrobiidae (Harzhauser et al. 2015; MolluscaBase 2018 b). Examination of further material allowed distinguishing this species from Staja, which is a Miocene Eastern European taxon rich in several species (cf. MolluscaBase 2018 b), characterised by a very small size (dimensional range: H = 1.6 - 2.5 mm, W = 1 - 1.2 mm) and a mainly conical shell with regularly growing whorls (cf. Brusina 1897; Jekelius 1944). The assignment of ionica n. sp. to the same genus as galatoniana n. sp. is likely supported by similar conchological characters between the two species, such as hydrobioid shell shape with slightly convex whorls, last whorl higher, ovate-elongate and barely flared aperture, and smooth protoconch. Hydrobia (s. l.) ionica n. sp. differs from H. (s. l.) galatoniana n. sp. in having a more globose shell, a wider apical angle, a smaller protoconch, the last whorl proportionally wider and higher, the aperture more elongate, and a smaller H / W (max) ratio (1.8 vs 2.08). It differs from the coeval H. dubuissonii hydruntina from the Upper Oligocene marls of Otranto bauxite quarry (Esu & Girotti 2010) in being conical-ovate in shape, in having a less acute spire, a proportionally higher last whorl and wider and more elongate aperture. Hydrobia dubuissonii from Oligocene deposits of France and Belgium (see above) differs from H. (s. l.) ionica n. sp. in having a conical-pyramidal shape, a more acute spire and a smaller and more ovate aperture.	en	Esu, Daniela, Girotti, Odoardo (2019): Two new oligohaline Hydrobia (s. l.) (Caenogastropoda) from the transitional Upper Oligocene Galatone Fm (Apulia, Southern Italy). Geodiversitas 41 (1): 1-9, DOI: 10.5252/geodiversitas2019v41a1
