taxonID	type	description	language	source
8710D84ACC2F850DFF43FC71FE2AFD58.taxon	description	(Figs. 1, 3 – 4, 6 – 7, 11) urn: lsid: zoobank. org: act: 35 B 1590 C- 6 AF 4 - 4 DD 8 - 9 AF 7 - 20 FF 977 C 3 EB 4	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2F850DFF43FC71FE2AFD58.taxon	materials_examined	Type material. Holotype, here designated, male, with labels: “ PERU, Madre de Dios Dept. CICRA field station, ~ 2 km NW of cafeteria, research plot, 12.55212 ° S 70.10921 ° W, 295 m, 11 – 13. vi. 2011, Chaboo team, flight intercept trap, PER- 11 - FIT- 023 ” / “ [barcode label] SEMC 0985242 ”. In the collection of MUSM. Paratypes. Four: same label as holotype except 9 – 11. vi. 2011, PER- 11 - FIT- 022 and barcode label SEMC 0955098 (1 ♀ SEMC); same label as holotype except 7 – 9. vi. 2011, PER- 11 - FIT- 021 and barcode labels SEMC 0956709, SEMC 0956710 (1 ♀ MUSM, 1 ♀ UTCI); same label as holotype except 12.55261 ° S 70.11008 ° W, 11 – 13. vii. 2010, PER- 10 - 07 - FIT- 004, barcode label SEMC 0955348 (1 ♀ SEMC). All paratypes with label “ PARATYPE Philothalpus anapodos Chatzimanolis, des. Chatzimanolis 2019 ”.	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2F850DFF43FC71FE2AFD58.taxon	diagnosis	Diagnosis. Among Philothalpus species with glossy integument of head and pronotum, P. chotaenus, P. ecuadorensis, P. juanecoi, P. porphyros, P. portokalis, P. loksos, P. stravos, and P. anapodos all have large, numerous, contiguous to virtually contiguous punctures on pronotum. Philothalpus anapodos and P. portokalis are the only two species in that group that have reddish-brown to orange head and pronotum and punctures on pronotum that are round and not elongated longitudinally. In P. portokalis the head and pronotum are bright orange, the pronotum is strongly concave posteriorly and the apex of the paramere is emarginate, while in P. anapodos the head and pronotum are reddish-brown, the pronotum is weakly concave posteriorly and the apex of the paramere is rounded; P. anapodos also does not have any peg setae on the paramere.	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2F850DFF43FC71FE2AFD58.taxon	description	Description. Forebody length 5.9 – 6.7 mm. Coloration of head and pronotum shinning reddish brown; legs, mouthparts, antennomeres 1 – 3 reddish brown; antennomeres 4 – 11 brown; elytra shinning metallic purple-brown or green, with golden overtones; elytra epipleuron lighter in coloration, reddish brown; mesoscutellum brown; abdominal segments 3 – 5 reddish brown with dark brown areas medially and on paratergites; segment 6 brown except posterior 1 / 6 reddish brown; segment 7 brown on anterior half and reddish brown on posterior half; segment 8 reddish brown except anterior 1 / 3 brown. Head with 4 – 5 rows of large punctures on each side of impunctate middle epicranium; head with 4 – 5 rows of smaller punctures posteriorly; punctures contiguous; head appearing glossy due to lack of closely spaced micropunctures and microsculpture. Head transverse, width / length ratio = 1.25; antennomeres 1 – 6 longer than wide; antennomeres 7 – 8 subquadrate; antennomeres 9 – 10 slightly transverse. Neck with 4 – 5 irregular rows of small punctures. Pronotum longer than wide, width / length ratio = 0.83; with large, numerous, contiguous to virtually contiguous umbilicate punctures over most of pronotal surface; with impunctate midline, midline wider anteriorly than posteriorly; punctures round, not longitudinally elongate; punctures contiguous but not confluent; pronotum with polygon-shaped microsculpture on anterolateral corners. Pronotum weakly concave posteriorly. Pronotum / elytra length ratio = 0.96; elytra with large, dense punctures. Abdominal tergites with sparse, small punctures; sternites with denser larger punctures. Males with broad shallow emargination on sternite 7; small, round porose structure present positioned anteriorly on sternite 7; sternite 8 with broad shallow emargination; females with no secondary sexual structures. Aedeagus as in Figs. 6 – 7; paramere asymmetrical, in dorsal view converging to narrow rounded apex; in lateral view paramere becoming narrower medially to apex; paramere with two small apical teeth; paramere without peg setae; paramere shorter and narrower than median lobe; median lobe asymmetrical, in dorsal view, converging to narrow rounded tip; in lateral view median lobe becoming narrower medially, converging to upwards apex; medial lobe ventrally with to small teeth, almost beneath the parameral teeth.	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2F850DFF43FC71FE2AFD58.taxon	distribution	Distribution. Known from the department of Madre de Dios in Peru. Habitat. Philothalpus anapodos was collected in flight intercept traps in lowland tropical rainforest.	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2F850DFF43FC71FE2AFD58.taxon	etymology	Etymology. The specific epithet is derived from the Greek word ανάποδος (upside down) and refers to the shape of the median lobe.	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2D850BFF43FF3DFD70F80F.taxon	description	(Figs. 2, 5, 8 – 11) urn: lsid: zoobank. org: act: 62 C 36 DDB- 542 B- 4131 - 94 DF-F 3 FEB 2 B 26044	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2D850BFF43FF3DFD70F80F.taxon	materials_examined	Type material. Holotype, here designated, male, with labels: “ GUATEMALA: Retalhuleu, Finca los Chicharros, SW of Zunil, 1270 m, 14.69460 - 91.53792, 3. vi. 2015. R. Anderson, ex: sifted mixed hardwood litter, GUAT 1 A 15 102 ” / “ [barcode label] SEMC 1538104 ” / “ HOLOTYPE Philothalpus boreios Chatzimanolis, des. Chatzimanolis 2018 ”. In the collection of SEMC. Paratypes. Four; “ El. Sal., Los Chorros, 4 km S. Santa Tecla, 10. v. 1971, S. Peck ” / “ Ber. 200 ravine leaf & fis litter ” (1 ♀ FMNH); “ Est. La Pailas, 800 m, P. N. Rincón de la Vieja, Prov. Guanacaste, Costa Rica, 27 jul [July] — 15 ago [August] 1992, D. Garcia, L-N 306300 388600 ” / “ [barcode label] INBIO CRI 000842173 ” (1 ♀ MNCR- A); “ Costa Rica, Prov. San Jose, Platanares, Vista de Mar, Finca Bosque W. Villalobas, 1100 m, 16 Jul [July] — 2 ago [August] 2001, R. González, Malaise, L _ S 348200 _ 499690 # 64209 ” / “ [barcode label] INB 0003356842 ” (1 ♀ MNCR-A); “ Boruca (Caudau de Osa), i. 1943, (S. Dengo), Costa Rica ” / “ Field Mus. Nat. Hist. 1966, A. Bierig Colln, Acc. Z- 13812 ” (1 ♀ FMNH). All paratypes with label “ PARATYPE Philothalpus boreios Chatzimanolis, des. Chatzimanolis 2019 ”.	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2D850BFF43FF3DFD70F80F.taxon	diagnosis	Diagnosis. Among Philothalpus species with glossy integument of head and pronotum, P. boreios is more similar to P. chloropennis, P. fabiolae, P. fervidus, and P. rufus. Philothalpus boreios can be easily distinguished from these species based on the coloration of head and pronotum that is dark metallic purple-green.	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2D850BFF43FF3DFD70F80F.taxon	description	Description. Forebody length 5.3 - 6.2 mm. Coloration of head, pronotum and elytra dark metallic purple-green; mouthparts, legs and antennomere 1 reddish brown, antennomeres 2 – 11 brown; mesoscutellum dark brown to black; abdominal segments 3 – 4 reddish brown to brown; segments 5 – 8 reddish brown. Head with 1 – 2 rows of large punctures on each side of impunctate middle epicranium; head with 2 – 3 irregular rows of small punctures posteriorly; punctures not contiguous; impunctate area on middle of epicranium much larger than punctuate areas in lateral regions; head appearing glossy due to lack of closely spaced micropunctures and microsculpture. Head transverse, width / length ratio = 1.5; antennomeres 1 – 4 longer than wide; antennomeres 5 – 6 subquadrate; antennomeres 7 – 10 transverse. Neck with 2 – 3 irregular rows of small punctures. Pronotum longer than wide, width / length ratio = 0.9; with less numerous punctures; punctures not contiguous, separated by width of 0.5 – 1 punctures; with impunctate midline having width of 4 – 5 punctures; punctures round, not longitudinally elongate; pronotum without polygon-shaped microsculpture on anterolateral corners. Pronotum weakly concave posteriorly. Pronotum / elytra length ratio = 0.95; elytra with large punctures. Abdominal tergites with medium-sized punctures; sternites with denser larger punctures. Males with small, round porose structure present positioned anteriorly on sternite 7; sternite 8 with broad shallow emargination; females with no secondary sexual structures. Aedeagus as in Figs. 8 – 10; paramere in dorsal view gradually converging to narrowly rounded apex; in lateral view paramere converging to pointed apex; paramere with peg setae as in Fig. 10; paramere shorter and narrower than median lobe; median lobe in dorsal view converging to subacute apex; in lateral view median lobe becoming narrower from middle to apex; with small dorsal subapical tooth.	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2D850BFF43FF3DFD70F80F.taxon	distribution	Distribution. Known from the department of La Libertad in El Salvador, the department of Retalhuleu in Guatemala and the provinces of Guanacaste, San Jose and Puntarenas in Costa Rica. Habitat. The holotype was sifted from hardwood litter at mid-elevation in a cloud forest fragment on the Pacific slope of Guatemala (Robert Anderson per. comm.) and most paratypes were also collected at mid-elevation in cloud forest.	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2D850BFF43FF3DFD70F80F.taxon	etymology	Etymology. The specific epithet is derived from the Greek word βóρειος (northern) and refers to this being the northern most species of Philothalpus known to date.	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2B850AFF43FB11FC14F9F7.taxon	materials_examined	PERU: Cusco, Villa Carmen field station, ~ 1.7 km west cafeteria, research transect, 12.89250 ° S 71.41917 ° W, 555 m, 28 – 30. v. 2011, D. J. Bennett & E. Razuri, flight intercept trap, PER- 11 - FIT- 015, SEMC 0956743 (1 ♂ SEMC); Madre de Dios, CICRA field station, ~ 2 km NW of cafeteria, research plot, 12.55212 ° S 70.10921 ° W, 295 m, 9 – 11. vi. 2011, Chaboo team, malaise trap, PER- 11 - MAT- 025, SEMC 0955091 (1 ♂ SEMC); CICRA field station, ~ 2 km NW of cafeteria, research plot, 12.55236 ° S 70.10989 ° W, 295 m, 7 – 9. vi. 2011, Chaboo team, flight intercept trap, PER- 11 - FIT- 017, SEMC 1465573 (1 ♂ SEMC); CICRA field station, ~ 2 km NW of cafeteria, re- search plot, 12.55212 ° S 70.10921 ° W, 295 m, 7 – 9. vi. 2011, Chaboo team, flight intercept trap, PER- 11 - FIT- 021, SEMC 0956720 (1 ♂ SEMC); CICRA field station, ~ 2 km NW of cafeteria, research plot, 12.55212 ° S 70.10921 ° W, 295 m, 11 – 13. vi. 2011, Chaboo team, flight intercept trap, PER- 11 - FIT- 023, SEMC 0985253 (1 ♂ SEMC); CICRA field station, ~ 2 km NW of cafeteria, research plot, 12.55236 ° S 70.10989 ° W, 295 m, 11 – 13. vi. 2011, Chaboo team, flight intercept trap, PER- 11 - FIT- 019, SEMC 1465573 (1 ♂ UTCI).	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2B850AFF43F92DFADEF89F.taxon	materials_examined	SURINAME: Sipaliwini, 2.362933 ° N 56.697682 ° W, 237 m, camp 3, Wehepal SE Kwamala, FIT, Larson & Short leg, 3 – 7. ix. 2010, SR 10 - 0903 - TN 3, 2010 CI-RAP Survey, Chatzimanolis DNA Voucher SC- 234, UTCI 000023102 (1 ♂ UTCI); 2.182883 ° N 56.78725 ° W, on Sipaliwini River, Camp 2, 210 m, 27. viii – 1. ix. 2010, FIT, Larsen & Short coll., Chatzimanolis DNA Voucher SC- 240, UTCI 000023116 (1 sex unknown UTCI).	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2B850AFF43F8C5FE30F86F.taxon	materials_examined	COLOMBIA: Magdalena, Finca San Josè, 8 km SE of Socorpa Mission, 1450 – 1500 m, 27 – 31. vii. 1968, B. Malkin leg (1 ♀ FMNH).	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC288509FF43FF3CFDD6FEE8.taxon	materials_examined	ECUADOR: Napo, Campana Cocha, E Puerto Misahualli, 2. vi. 1982, 1100 ’, leaf litter, ridge, H. Frania (1 ♂ CNC); Tena-Baeza Rd., km 24, N. Cotundo, 36 – 4000 ’, 29. iv. 1982, litter, H. Frania (1 ♂, 1 ♀ CNC). Note: the female specimen was previously identified incorrectly by Chatzimanolis & Ashe (2005) as a paratype of P. loksos Chatzimanolis & Ashe, 2005.	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC288509FF43FED4FC05FE5C.taxon	materials_examined	PERU: Madre de Dios, Amazonas Lodge, N. Atalaya, 12 ° 52.2 ’ S 71 ° 22.6 ’ W, 480 m, 10 – 13. xi. 2007, D. Brzoska, FIT, PER 1 B 07 002, SEMC 0869838 (1 ♀ UTCI); Tambopata Wildlife Reserve, 30 km SW Puerto Maldonado, 12 ° 50 ’ S 69 ° 20 ’ W, 18 – 20. xi. 1982, J. J. Anderson coll. (1 ♀ CMNH).	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC288509FF43FD80FA83FD2C.taxon	materials_examined	COSTA RICA: Cartago: Turrialba, M. N. Guayabo, Seud. Monuculus, 1100 – 1200 m, 8 – 12. v. 2007, Moraga, Azofeifa, Gonzalez, Navarro. Tp. Foso. L _ N _ 217200 _ 570300 # 91255, INB 0004082351 (1 ♀ NHMD).	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC288509FF43FD10FCB7FCBC.taxon	materials_examined	COLOMBIA: Valle del Cauca, PNN Farallones de Cali, Anchicaya, 3 ° 26 ’ N 76 ° 48 ’ W, 650 m, pitfall, 2 – 4. i. 2001, S. Sarria Leg. M 1555, SM 0548893 (1 ♀ SEMC).	en	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
