identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
8710D84ACC2F850DFF43FC71FE2AFD58.text	8710D84ACC2F850DFF43FC71FE2AFD58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothalpus anapodos Chatzimanolis 2020	<div><p>Philothalpus anapodos Chatzimanolis, new species</p> <p>(Figs. 1, 3–4, 6–7, 11)</p> <p>urn:lsid:zoobank.org:act: 35B1590C-6AF4-4DD8-9AF7-20FF977C3EB4</p> <p>Type material. Holotype, here designated, male, with labels: “ PERU, Madre de Dios Dept. CICRA field station, ~ 2 km NW of cafeteria, research plot, 12.55212°S 70.10921°W, 295 m, 11–13.vi.2011, Chaboo team, flight intercept trap, PER-11-FIT-023” / “[barcode label] SEMC0985242 ”. In the collection of MUSM.</p> <p>Paratypes. Four:same label as holotype except 9–11.vi.2011, PER-11-FIT-022and barcode label SEMC0955098 (1 ♀ SEMC); same label as holotype except 7–9.vi.2011, PER-11-FIT-021 and barcode labels SEMC0956709, SEMC0956710 (1 ♀ MUSM, 1 ♀ UTCI); same label as holotype except 12.55261°S 70.11008°W, 11–13.vii.2010, PER-10-07-FIT-004, barcode label SEMC0955348 (1 ♀ SEMC). All paratypes with label “ PARATYPE Philothalpus anapodos Chatzimanolis, des. Chatzimanolis 2019 ”.</p> <p>Diagnosis. Among Philothalpus species with glossy integument of head and pronotum, P. chotaenus, P. ecuadorensis, P. juanecoi, P. porphyros, P. portokalis, P. loksos, P. stravos, and P. anapodos all have large, numerous, contiguous to virtually contiguous punctures on pronotum. Philothalpus anapodos and P. portokalis are the only two species in that group that have reddish-brown to orange head and pronotum and punctures on pronotum that are round and not elongated longitudinally. In P. portokalis the head and pronotum are bright orange, the pronotum is strongly concave posteriorly and the apex of the paramere is emarginate, while in P. anapodos the head and pronotum are reddish-brown, the pronotum is weakly concave posteriorly and the apex of the paramere is rounded; P. anapodos also does not have any peg setae on the paramere.</p> <p>Description. Forebody length 5.9–6.7 mm. Coloration of head and pronotum shinning reddish brown; legs, mouthparts, antennomeres 1–3 reddish brown; antennomeres 4–11 brown; elytra shinning metallic purple-brown or green, with golden overtones; elytra epipleuron lighter in coloration, reddish brown; mesoscutellum brown; abdominal segments 3–5 reddish brown with dark brown areas medially and on paratergites; segment 6 brown except posterior 1/6 reddish brown; segment 7 brown on anterior half and reddish brown on posterior half; segment 8 reddish brown except anterior 1/3 brown.</p> <p>Head with 4–5 rows of large punctures on each side of impunctate middle epicranium; head with 4–5 rows of smaller punctures posteriorly; punctures contiguous; head appearing glossy due to lack of closely spaced micropunctures and microsculpture. Head transverse, width/length ratio = 1.25; antennomeres 1–6 longer than wide; antennomeres 7–8 subquadrate; antennomeres 9–10 slightly transverse. Neck with 4–5 irregular rows of small punctures. Pronotum longer than wide, width/length ratio = 0.83; with large, numerous, contiguous to virtually contiguous umbilicate punctures over most of pronotal surface; with impunctate midline, midline wider anteriorly than posteriorly; punctures round, not longitudinally elongate; punctures contiguous but not confluent; pronotum with polygon-shaped microsculpture on anterolateral corners. Pronotum weakly concave posteriorly. Pronotum/elytra length ratio = 0.96; elytra with large, dense punctures. Abdominal tergites with sparse, small punctures; sternites with denser larger punctures.</p> <p>Males with broad shallow emargination on sternite 7; small, round porose structure present positioned anteriorly on sternite 7; sternite 8 with broad shallow emargination; females with no secondary sexual structures.</p> <p>Aedeagus as in Figs. 6–7; paramere asymmetrical, in dorsal view converging to narrow rounded apex; in lateral view paramere becoming narrower medially to apex; paramere with two small apical teeth; paramere without peg setae; paramere shorter and narrower than median lobe; median lobe asymmetrical, in dorsal view, converging to narrow rounded tip; in lateral view median lobe becoming narrower medially, converging to upwards apex; medial lobe ventrally with to small teeth, almost beneath the parameral teeth.</p> <p>Distribution. Known from the department of Madre de Dios in Peru.</p> <p>Habitat. Philothalpus anapodos was collected in flight intercept traps in lowland tropical rainforest.</p> <p>Etymology. The specific epithet is derived from the Greek word ανάποδος (upside down) and refers to the shape of the median lobe.</p></div> 	http://treatment.plazi.org/id/8710D84ACC2F850DFF43FC71FE2AFD58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chatzimanolis, Stylianos	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2D850BFF43FF3DFD70F80F.text	8710D84ACC2D850BFF43FF3DFD70F80F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothalpus boreios Chatzimanolis. Not 2020	<div><p>Philothalpus boreios Chatzimanolis, new species</p> <p>(Figs. 2, 5, 8–11)</p> <p>urn:lsid:zoobank.org:act: 62C36DDB-542B-4131-94DF-F3FEB2B26044</p> <p>Type material. Holotype, here designated, male, with labels: “ GUATEMALA: Retalhuleu, Finca los <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.53792&amp;materialsCitation.latitude=14.6946" title="Search Plazi for locations around (long -91.53792/lat 14.6946)">Chicharros</a>, SW of Zunil, 1270m, 14.69460 -91.53792, 3.vi.2015. R. Anderson, ex: sifted mixed hardwood litter, GUAT1A15 102” / “[barcode label] SEMC1538104 ” / “ HOLOTYPE Philothalpus boreios Chatzimanolis, des. Chatzimanolis 2018”. In the collection of SEMC.</p> <p>Paratypes. Four; “ El. Sal., Los Chorros, 4 km S. Santa Tecla, 10.v.1971, S. Peck ” / “Ber. 200 ravine leaf &amp; fis litter” (1 ♀ FMNH); “Est. La Pailas, 800 m, P. N. Rincón de la Vieja, Prov. Guanacaste, Costa Rica, 27 jul [July]— 15 ago [August] 1992, D. Garcia, L-N 306300 388600” / “[barcode label] INBIO CRI000842173” (1 ♀ MNCR- A); “ Costa Rica, Prov. San Jose, Platanares, Vista de Mar, Finca Bosque W. Villalobas, 1100 m, 16 Jul [July]—2 ago [August] 2001, R. González, Malaise, L_S 348200_499690 #64209” / “[barcode label] INB0003356842” (1 ♀ MNCR-A); “Boruca (Caudau de Osa), i. 1943, (S. Dengo), Costa Rica ” / “Field Mus. Nat. Hist. 1966, A. Bierig Colln, Acc. Z-13812” (1 ♀ FMNH). All paratypes with label “ PARATYPE Philothalpus boreios Chatzimanolis, des. Chatzimanolis 2019 ”.</p> <p>Diagnosis. Among Philothalpus species with glossy integument of head and pronotum, P. boreios is more similar to P. chloropennis, P. fabiolae, P. fervidus, and P. rufus. Philothalpus boreios can be easily distinguished from these species based on the coloration of head and pronotum that is dark metallic purple-green.</p> <p>Description. Forebody length 5.3-6.2 mm. Coloration of head, pronotum and elytra dark metallic purple-green; mouthparts, legs and antennomere 1 reddish brown, antennomeres 2–11 brown; mesoscutellum dark brown to black; abdominal segments 3–4 reddish brown to brown; segments 5–8 reddish brown.</p> <p>Head with 1–2 rows of large punctures on each side of impunctate middle epicranium; head with 2–3 irregular rows of small punctures posteriorly; punctures not contiguous; impunctate area on middle of epicranium much larger than punctuate areas in lateral regions; head appearing glossy due to lack of closely spaced micropunctures and microsculpture. Head transverse, width/length ratio = 1.5; antennomeres 1–4 longer than wide; antennomeres 5–6 subquadrate; antennomeres 7–10 transverse. Neck with 2–3 irregular rows of small punctures. Pronotum longer than wide, width/length ratio = 0.9; with less numerous punctures; punctures not contiguous, separated by width of 0.5–1 punctures; with impunctate midline having width of 4–5 punctures; punctures round, not longitudinally elongate; pronotum without polygon-shaped microsculpture on anterolateral corners. Pronotum weakly concave posteriorly. Pronotum/elytra length ratio = 0.95; elytra with large punctures. Abdominal tergites with medium-sized punctures; sternites with denser larger punctures.</p> <p>Males with small, round porose structure present positioned anteriorly on sternite 7; sternite 8 with broad shallow emargination; females with no secondary sexual structures.</p> <p>Aedeagus as in Figs. 8–10; paramere in dorsal view gradually converging to narrowly rounded apex; in lateral view paramere converging to pointed apex; paramere with peg setae as in Fig. 10; paramere shorter and narrower than median lobe; median lobe in dorsal view converging to subacute apex; in lateral view median lobe becoming narrower from middle to apex; with small dorsal subapical tooth.</p> <p>Distribution. Known from the department of La Libertad in El Salvador, the department of Retalhuleu in Guatemala and the provinces of Guanacaste, San Jose and Puntarenas in Costa Rica.</p> <p>Habitat. The holotype was sifted from hardwood litter at mid-elevation in a cloud forest fragment on the Pacific slope of Guatemala (Robert Anderson per. comm.) and most paratypes were also collected at mid-elevation in cloud forest.</p> <p>Etymology. The specific epithet is derived from the Greek word βóρειος (northern) and refers to this being the northern most species of Philothalpus known to date.</p> </div>	http://treatment.plazi.org/id/8710D84ACC2D850BFF43FF3DFD70F80F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chatzimanolis, Stylianos	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2B850AFF43FB11FC14F9F7.text	8710D84ACC2B850AFF43FB11FC14F9F7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothalpus asymmetros Chatzimanolis & Ashe 2005	<div><p>Philothalpus asymmetros Chatzimanolis &amp; Ashe, 2005</p> <p>PERU: Cusco, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.41917&amp;materialsCitation.latitude=-12.8925" title="Search Plazi for locations around (long -71.41917/lat -12.8925)">Villa Carmen</a> field station, ~ 1.7 km west cafeteria, research transect, 12.89250°S 71.41917°W, 555 m, 28–30.v.2011, D.J. Bennett &amp; E. Razuri, flight intercept trap, PER-11-FIT-015, SEMC0956743 (1 ♂ SEMC); Madre de Dios, CICRA field station, ~ 2 km NW of cafeteria, research plot, 12.55212°S 70.10921°W, 295 m, 9–11.vi.2011, Chaboo team, malaise trap, PER-11-MAT-025, SEMC0955091 (1 ♂ SEMC); CICRA field station, ~ 2 km NW of cafeteria, research plot, 12.55236°S 70.10989°W, 295 m, 7–9.vi.2011, Chaboo team, flight intercept trap, PER-11-FIT-017, SEMC1465573 (1 ♂ SEMC); CICRA field station, ~ 2 km NW of cafeteria, re- search plot, 12.55212°S 70.10921°W, 295 m, 7–9.vi.2011, Chaboo team, flight intercept trap, PER-11-FIT-021, SEMC0956720 (1 ♂ SEMC); CICRA field station, ~ 2 km NW of cafeteria, research plot, 12.55212°S 70.10921°W, 295 m, 11–13.vi.2011, Chaboo team, flight intercept trap, PER-11-FIT-023, SEMC0985253 (1 ♂ SEMC); CICRA field station, ~ 2 km NW of cafeteria, research plot, 12.55236°S 70.10989°W, 295 m, 11–13.vi.2011, Chaboo team, flight intercept trap, PER-11-FIT-019, SEMC1465573 (1 ♂ UTCI).</p> </div>	http://treatment.plazi.org/id/8710D84ACC2B850AFF43FB11FC14F9F7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chatzimanolis, Stylianos	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2B850AFF43F92DFADEF89F.text	8710D84ACC2B850AFF43F92DFADEF89F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothalpus bilobus Chatzimanolis & Ashe 2005	<div><p>Philothalpus bilobus Chatzimanolis &amp; Ashe, 2005</p> <p>SURINAME: Sipaliwini, 2.362933°N 56.697682°W, 237 m, camp 3, Wehepal SE Kwamala, FIT, Larson &amp; Short leg, 3–7.ix.2010, SR10-0903-TN3, 2010 CI-RAP Survey, Chatzimanolis DNA Voucher SC-234, UTCI000023102 (1 ♂ UTCI); 2.182883°N 56.78725°W, on Sipaliwini River, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-56.78725&amp;materialsCitation.latitude=2.182883" title="Search Plazi for locations around (long -56.78725/lat 2.182883)">Camp</a> 2, 210 m, 27.viii–1.ix.2010, FIT, Larsen &amp; Short coll., Chatzimanolis DNA Voucher SC-240, UTCI000023116 (1 sex unknown UTCI).</p> </div>	http://treatment.plazi.org/id/8710D84ACC2B850AFF43F92DFADEF89F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chatzimanolis, Stylianos	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC2B850AFF43F8C5FE30F86F.text	8710D84ACC2B850AFF43F8C5FE30F86F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothalpus fervidus (Erichson 1840)	<div><p>Philothalpus fervidus (Erichson, 1840)</p> <p>COLOMBIA: Magdalena, Finca San Josè, 8 km SE of Socorpa Mission, 1450–1500 m, 27–31.vii.1968, B. Malkin leg (1♀ FMNH).</p> </div>	http://treatment.plazi.org/id/8710D84ACC2B850AFF43F8C5FE30F86F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chatzimanolis, Stylianos	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC288509FF43FF3CFDD6FEE8.text	8710D84ACC288509FF43FF3CFDD6FEE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothalpus juanecoi Asenjo & Ribeiro-Costa 2009	<div><p>Philothalpus juanecoi Asenjo &amp; Ribeiro-Costa, 2009</p> <p>ECUADOR: Napo, Campana Cocha, E Puerto Misahualli, 2.vi.1982, 1100’, leaf litter, ridge, H. Frania (1 ♂ CNC); Tena-Baeza Rd., km24, N. Cotundo, 36–4000’, 29.iv.1982, litter, H. Frania (1 ♂, 1 ♀ CNC).</p> <p>Note: the female specimen was previously identified incorrectly by Chatzimanolis &amp; Ashe (2005) as a paratype of P. loksos Chatzimanolis &amp; Ashe, 2005.</p> </div>	http://treatment.plazi.org/id/8710D84ACC288509FF43FF3CFDD6FEE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chatzimanolis, Stylianos	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC288509FF43FED4FC05FE5C.text	8710D84ACC288509FF43FED4FC05FE5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothalpus mundus (Sharp 1876)	<div><p>Philothalpus mundus (Sharp, 1876)</p> <p>PERU: Madre de Dios, Amazonas Lodge, N. Atalaya, 12°52.2’S 71°22.6’W, 480 m, 10–13.xi.2007, D. Brzoska, FIT, PER1B07 002, SEMC 0869838 (1 ♀ UTCI); <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.333336&amp;materialsCitation.latitude=-12.833333" title="Search Plazi for locations around (long -69.333336/lat -12.833333)">Tambopata Wildlife Reserve</a>, 30 km SW Puerto Maldonado, 12°50’S 69°20’W, 18–20.xi.1982, J. J. Anderson coll. (1 ♀ CMNH).</p> </div>	http://treatment.plazi.org/id/8710D84ACC288509FF43FED4FC05FE5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chatzimanolis, Stylianos	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC288509FF43FD80FA83FD2C.text	8710D84ACC288509FF43FD80FA83FD2C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothalpus rufus Chatzimanolis & Ashe 2005	<div><p>Philothalpus rufus Chatzimanolis &amp; Ashe 2005</p> <p>COSTA RICA: Cartago: Turrialba, M.N. Guayabo, Seud. Monuculus, 1100–1200 m, 8–12.v.2007, Moraga, Azofeifa, Gonzalez, Navarro. Tp. Foso. L _N_217200_570300 #91255, INB0004082351 (1 ♀ NHMD).</p> </div>	http://treatment.plazi.org/id/8710D84ACC288509FF43FD80FA83FD2C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chatzimanolis, Stylianos	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
8710D84ACC288509FF43FD10FCB7FCBC.text	8710D84ACC288509FF43FD10FCB7FCBC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philothalpus rugosus Chatzimanolis & Ashe. A 2005	<div><p>Philothalpus rugosus Chatzimanolis &amp; Ashe 2005</p> <p>COLOMBIA: Valle del Cauca, PNN Farallones de Cali, <a href="http://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.8&amp;materialsCitation.latitude=3.4333334" title="Search Plazi for locations around (long -76.8/lat 3.4333334)">Anchicaya</a>, 3°26’N 76°48’W, 650 m, pitfall, 2– 4.i.2001, S. Sarria Leg. M1555, SM0548893 (1 ♀ SEMC).</p> </div>	http://treatment.plazi.org/id/8710D84ACC288509FF43FD10FCB7FCBC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chatzimanolis, Stylianos	Chatzimanolis, Stylianos (2020): Even more new species, additional records, and a revised key for Philothalpus Kraatz (Coleoptera: Staphylinidae: Staphylininae). Zootaxa 4742 (1): 129-138, DOI: https://doi.org/10.11646/zootaxa.4742.1.8
