taxonID	type	description	language	source
4765755E2C50FFAEFCA054943FE2FCE1.taxon	diagnosis	Diagnosis: Presenting the characters of ‘ poneromorph’ subfamilies described by Bolton (2003: p. 40). Clypeus broadly inserted between frontal lobes (Bolton 2003); anterior clypeal margin with a narrow lamellar apron (Bolton 2003). Torulus not completely fused to frontal lobe (Bolton 2003). Antenna with 12 segments (Bolton 2003). Pronotum with the humeral corners angled, forming a distinct delimitation between the anterior and lateral margins (Lattke 2004). Antero-ventral angle of pronotum triangular (Feitosa 2015). Pretarsus without arolium (Lattke 2004). Petiole pedunculate (Keller 2011). Petiolar node as wide or wider than long (Feitosa 2015). Subpetiolar process very well developed, occupying more than onethird of the ventral portion of the petiolar sternite (Feitosa 2015). Helcium projecting from about midheight of the anterior face of abdominal segment III. Abdominal segment IV presclerites separated from the rest of segment by a constriction or slight thickening (Lattke 1994). Fourth abdominal tergite arched and larger than the sternite, giving the segment a curved appearance (Keller 2011).	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C51FFAEFF0057A43B0CFD4B.taxon	diagnosis	Diagnosis (Females): Head subquadrate; occipital lobe usually present; frontal carina broadly expanded laterad; row of stout setae on base of foretarsus opposite to strigil present; promesonotal suture absent to lightly impressed, never interrupting dorsal mesosomal sculpture; petiolar spiracle facing directly ventrad and sunken within a pit; second gastral (IV abdominal) sternite usually strongly reduced, so that the gaster is directed ventrally and anterad.	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C51FFAEFCA051523A17F9D1.taxon	materials_examined	Type Species: Ectatomma tuberculatum (Olivier) Diagnosis (Females): Occipital lobe absent. Antennal club absent. Palp formula 2,2. Pronotum usually with two or three tubercles. Mesonotum prominent and clearly differentiated from propodeum, separated by a deep transverse suture. Promesonotal suture well marked, interrupting or not the dorsal mesosomal sculpture. Propodeal spiracle elliptical or slit-shaped and separated from the declivous face of propodeum by a distance longer than its diameter. Apex of protibia with a stout seta close to the strigil base; dorsum of posterior coxa without projections. Species: brunneum, confine, edentatum, gibbum, goninion, † gracile, lugens, muticum, opaciventre, parasiticum, permagnum, planidens, ruidum, suzanae, tuberculatum, and vizottoi. Distribution: Exclusively found in the New World, from USA (Texas) to Argentina (Buenos Aires). Notes: Ectatomma are among the most conspicuous elements of the ant fauna in Neotropical ecosystems. Currently, the most comprehensive work including an identification key for the species in the genus is the revision by Kugler and Brown (1982). However, this work does not include the species Ectatomma parasiticum Feitosa and Fresneau, in Feitosa et al. (2008), E. suzannae Almeida (1986) and E. vizottoi Almeida (1987).	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C51FFADFCA054393CF7FC3A.taxon	diagnosis	Diagnosis (Females): Head subquadrate to elongate. Mandible subtriangular to subfalcate. Occipital lobe absent. Antennal club absent. Palp formula 2,2 to 3,2. Pronotum unarmed and without tubercles. Promesonotal suture feebly impressed to absent, never interrupting dorsal mesosomal sculpture, sometimes with a small pit frequently situated medially on a weakly impressed promesonotal suture. Mesonotum not prominent, forming a continuous line with the propodeum, separated by a transverse suture. Propodeal spiracle oval or rounded, separated from the declivous face of propodeum by a distance longer or shorter than its diameter. Apex of protibia without a stout seta close to the strigil base; dorsum of posterior coxae frequently with a lobe or spine.	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C53FFABFCA057F23FB7FECA.taxon	materials_examined	Type Species: Typhlomyrmex rogenhoferi Mayr Diagnosis (Females): Head subquadrate; antennal club sometimes well-defined and formed by 3 or 4 segments; cephalic vertex mostly smooth and shining, sometimes presenting faded striae or rugulae; eye absent or reduced, with less than 15 ommatidia; promesonotal suture well marked, totally interrupting dorsal mesosomal sculpture; propodeal spiracle separated from declivity margin by a distance longer than its diameter; metacoxal dorsum unarmed or at most with a small lobe or denticle; petiole pedunculate, sometimes with a prominent anteroventral process. Species: clavicornis, foreli, lavra new combination, lenis new combination, major, meire, prolatus, pusillus, reichenspergeri new combination, and rogenhoferi. Distribution: Exclusively Neotropical, occurring from Mexico to Argentina (Buenos Aires). Notes: Mostspecies of Typhlomyrmex can be identified usingthe key of Lacau et al. (2008), while T. lavra, T. lenis, and T. reichenspergeri (formerly included in the striatula group of Gnamptogenys) can be identified using Camacho et al. (2020).	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C55FFAAFF0053793C3FFBD4.taxon	materials_examined	Type Species: Aulacopone relicta Arnol'di Diagnosis (Queens): Ants of comparatively medium size. Mandibles subfalcate. Median portion of clypeus modified, raised as a short, blunt triangular point projecting from the antennal insertions to the mandible. Frontal lobe expanded, extending from the clypeal posterior margin to the vertex. Antennal scrobe wide and deep. Species: relicta. Distribution: The only known specimens were collected in Azerbaijan in mountainous forests. Notes: The genus is only known from two queens collected in Azerbaijan. The first specimen was collected in 1929 in Alazapin on the border with Iran, and later designated as the holotype by Arnol’di and deposited at the Zoological Institute of the Russian Academy of Sciences. The second specimen was collected in 1936, also by Arnol’di in the same country, in the region of Khachmaz, and later deposited in his personal collection at the Institute of Evolutionary Animal Morphology in Moscow. However, the holotype has been missing since 1979 and has not been examined for any study other than the original description. The second specimen was coated in gold-palladium for the study of its external morphology using scanning electron microscopy by Taylor (1980).	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C55FFAAFF0056243A48FD4C.taxon	diagnosis	Diagnosis (Females): Ants of comparatively small size (head width 0.42 - 0.53, head length 0.53 - 0.67). Mandible subfalcate. Palp formula 3,2. Frontal lobe expanded, completely covering antennal insertions. Antennal club with three antennomeres. Antennal scrobe absent. Eye drastically reduced, without conspicuous limits between ommatidia. Propodeum unarmed. Tarsal claw simple, without conspicuous preapical teeth nor a basal lobe. Petiole unarmed. Anterior face of abdominal segment III without an arched carina above the helcium.	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C55FFA9FCA0515A3CC0FEAC.taxon	diagnosis	Diagnosis (Females): Ants of comparatively small to medium size (head width 0.63 - 1.37; head length 0.72 - 1.54). Mandible triangular. Palp formula 3,2 to 4,3. Frontal lobe expanded, completely covering antennal insertions. Antennal club with three antennomeres. Antennal scrobe shallowly to deeply impressed. Eye well-developed to reduced, with clear limits between ommatidia. Propodeal spine absent to well-developed. Tarsal claw simple, without conspicuous preapical teeth (except in H. dolo and H. robusta) nor a basal lobe. Petiole with or without posterodorsal projections. Anterior face of abdominal segment III with an arched carina above the helcium.	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C51FFAEFF0051C93CA1FA54.taxon	diagnosis	Diagnosis (Females): Ectatommine ants of small to large size (head width 0.44 - 2.84 mm, head length 0.56 - 3.8 mm). Antennal scrobe usually absent. Eye absent to well-developed (Bolton 2003). Acetabulum of antennal socket apparatus spherical (Keller 2011); accessory chamber of antennal socket present (Keller 2011). Labial palp with two palpomeres (Keller 2011). Promesonotal suture fused and immobile to complete and flexible (Bolton 2003). Ventral flap on metapleural gland opening present (Keller 2011). Metacoxal cavity open (Bolton 2003). Petiolar sternite fused with tergite over its entire length (except in Rhytidoponera) (Bolton 2003, Keller 2011); laterotergites of petiole indistinct to absent.	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C52FFADFF0051823B63FBAA.taxon	diagnosis	Diagnosis (Females): Head wider posterad than anterad; mandible triangular with striae or rugulae on frontal surface; anterior clypeal margin convex; scape usually surpassing vertexal margin; eye slightly behind cephalic midlength; promesonotal suture frequently well marked, totally interrupting dorsal mesosomal sculpture; propodeal spiracle close to the declivous face of propodeum; propodeum unarmed; anterior prosternal process broadly concave medially; metacoxal dorsum always with a denticle or lobe; petiolar node high; anteroventral postpetiolar process relatively wide; second gastric segment only slightly arched ventrally.	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C52FFACFCA056243C4AFEE9.taxon	diagnosis	Diagnosis (Females): Head subquadrate or wider anterad than posterad in frontal view; anterior clypeal margin usually straight; frontal surface of mandible usually striate or rugulose; scape usually surpassing vertex; promesonotal suture feebly impressed to absent, never interrupting dorsal mesosomal sculpture; metanotal suture well impressed; propodeum usually armed with denticles or spines; petiolar node low; subpetiolar process shape variable, usually projecting anterad but sometimes subquadrate; metacoxal teeth generally present, usually acicular; second gastric segment slightly arched ventrally.	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C53FFACFF0053F13CB9FA74.taxon	diagnosis	Diagnosis (Females): Occipital lobe frequently present. Antennal club absent. Palp formula 2,2 to 3,2. Pronotum unarmed. Mesonotum not prominent, forming a continuous line with the propodeum, separated by a transverse suture. Promesonotal suture well marked, totally interrupting dorsal mesosomal sculpture. Propodeal spiracle oval or rounded, separated from the declivous face of propodeum by a distance longer than its diameter. Apex of protibia with a stout seta close to the strigil base; dorsum of posterior coxa without projections.	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C53FFACFF0054473DC1FAEC.taxon	diagnosis	Diagnosis (Females): Occipital lobe present. Antennal club absent. Palp formula 3,2. Pronotum usually unarmed, occasionally with humeral projections. Mesonotum not prominent, forming a continuous line with the propodeum, separated by a transverse suture. Promesonotal suture absent to feebly impressed, never interrupting the dorsal mesosomal sculpture. Propodeal spiracle oval to rounded and separated from the declivous face of propodeum by a distance longer than its diameter. Apex of protibia without a stout seta close to the strigil base; apex of meso- and metatibia with two spurs; dorsum of posterior coxae frequently with a lobe or spine.	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C54FFAAFCA052A43E8DFE91.taxon	materials_examined	Type Species Acanthoponera mucronata (Roger) Diagnosis (Females): Ants of comparatively medium size (head width 0.90 – 1.61, head length 1.00 – 1.75). Mandible triangular. Palp formula 6,4. Frontal lobe reduced, only partially covering antennal insertions. Antennal club with four antennomeres. Antennal scrobe deeply impressed. Eye well-developed, with clear limits between ommatidia. Propodeum with a pair of welldeveloped spines. Tarsal claw with conspicuous preapical teeth and a basal lobe. Petiole with a long posterodorsal projection. Anterior face of abdominal segment III with an arched carina above the helcium. Tribe Heteroponerini Bolton new combination Diagnosis: Ectatommine ants of small to medium size (head width 0.42 – 1.61 mm, head length 0.53 – 1.75 mm); cephalic dorsum with a longitudinal carina extending from anterior margin of clypeus to posterior margin of head (Bolton 2003); antennal scrobe present (Bolton 2003); eye present; acetabulum of antennal socket apparatus hemispherical (Keller 2011); accessory chamber of antennal socket absent (Keller 2011); labial palp with three or four palpomeres (Bolton 2003, Keller 2011); promesonotal suture complete and flexible (Bolton 2003); ventral flap on metapleural gland opening absent (Keller 2011); metacoxal cavity closed (Bolton 2003); petiolar sternite articulated with tergite over its entire length (Bolton 2003, Keller 2011); laterotergites of petiole present. Genera: Acanthoponera, Aulacopone, Boltonia new genus, Heteroponera. Species: goeldii, minor, mucronata, and peruviana. Distribution: Exclusively Neotropical, from southern Mexico to northern Argentina. Fig. 3. Worker of Boltonia microps in A) frontal view; B) dorsal view; and C) lateral view. Images by April Nobile (CASENT 0173544); available from www. antweb. org (Antweb 2021). Notes: Acanthoponera represents a lineage of arboreal and nocturnal Neotropical ants. Most species can be identified using the work of Feitosa and Prada-Achiardi (2019) for the Colombian fauna.	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C54FFAAFCA052A43E8DFE91.taxon	diagnosis	Diagnosis: Ectatommine ants of small to medium size (head width 0.42 – 1.61 mm, head length 0.53 – 1.75 mm); cephalic dorsum with a longitudinal carina extending from anterior margin of clypeus to posterior margin of head (Bolton 2003); antennal scrobe present (Bolton 2003); eye present; acetabulum of antennal socket apparatus hemispherical (Keller 2011); accessory chamber of antennal socket absent (Keller 2011); labial palp with three or four palpomeres (Bolton 2003, Keller 2011); promesonotal suture complete and flexible (Bolton 2003); ventral flap on metapleural gland opening absent (Keller 2011); metacoxal cavity closed (Bolton 2003); petiolar sternite articulated with tergite over its entire length (Bolton 2003, Keller 2011); laterotergites of petiole present.	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
4765755E2C54FFABFF0053D23DF7FF54.taxon	diagnosis	Diagnosis: Ectatommine ants of small to medium size (head width 0.42 – 1.61 mm, head length 0.53 – 1.75 mm); cephalic dorsum with a longitudinal carina extending from anterior margin of clypeus to posterior margin of head (Bolton 2003); antennal scrobe present (Bolton 2003); eye present; acetabulum of antennal socket apparatus hemispherical (Keller 2011); accessory chamber of antennal socket absent (Keller 2011); labial palp with three or four palpomeres (Bolton 2003, Keller 2011); promesonotal suture complete and flexible (Bolton 2003); ventral flap on metapleural gland opening absent (Keller 2011); metacoxal cavity closed (Bolton 2003); petiolar sternite articulated with tergite over its entire length (Bolton 2003, Keller 2011); laterotergites of petiole present.	en	Camacho, G P, Franco, W, Branstetter, M G, Pie, M R, Longino, J T, Schultz, T R, Feitosa, R M (2022): UCE Phylogenomics Resolves Major Relationships Among Ectaheteromorph Ants (Hymenoptera: Formicidae: Ectatomminae, Heteroponerinae): A New Classification For the Subfamilies and the Description of a New Genus. Insect Systematics and Diversity 6 (1): 6, DOI: 10.1093/isd/ixab026, URL: http://dx.doi.org/10.1093/isd/ixab026
