identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A37718FF83356EFDC6F9B9FD37F971.text	03A37718FF83356EFDC6F9B9FD37F971.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allokoenenia Silvestri 1913	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Genus  Allokoenenia Silvestri, 1913</p>
            <p>Type species</p>
            <p> Allokoenenia afra Silvestri, 1913 by monotypy. </p>
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	https://treatment.plazi.org/id/03A37718FF83356EFDC6F9B9FD37F971	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Souza, Maysa Fernanda Villela Rezende;Ferreira, Rodrigo Lopes	Souza, Maysa Fernanda Villela Rezende, Ferreira, Rodrigo Lopes (2022): Two extraordinary troglobitic species of Allokoenenia (Eukoeneniidae: Palpigradi) from Brazil: first records of this initially monotypic genus more than a century after its description. European Journal of Taxonomy 789: 11-48, DOI: 10.5852/ejt.2022.789.1627, URL: http://dx.doi.org/10.5852/ejt.2022.789.1627
03A37718FF83356BFDD1F8E8FC8DFAD2.text	03A37718FF83356BFDD1F8E8FC8DFAD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allokoenenia afra Silvestri 1913	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Allokoenenia afra Silvestri, 1913</p>
            <p>Figs 1–2</p>
            <p>Material examined</p>
            <p> Typus (♀) and cotypus (immature)</p>
            <p> GUINEA •  Mamou . </p>
            <p>Description</p>
            <p>MEASUREMENTS AND RATIO. See Tables 1–2.</p>
            <p>ADULT FEMALE (“ typus ”). Relative width of opisthosomal segments VIII–XI as described in the diagnosis of the genus by Rowland &amp; Sissom (1980) (Fig. 1A); last opisthosomal segment with dorsal row of 2 +2 setae inserted around its half and two pairs of ventral setae, one inserted around middle of segment and other inserted in its distal half (Fig. 2); flagellum formed by 14 short and rounded segments (Fig. 1A– B), each with whorl of setae of similar shape (at least one setae smaller than others in each segment); flagellar segments 1–3, 5, 7 and 9 with apical cuticular spines.</p>
            <p>IMMATURE SPECIMEN (“cotypus”). Frontal organ formed by two branches distally pointed and basally expanded, with fine reticulation (Fig. 1C); cheliceral fingers with 8 teeth each; 6 setae (grt, r, gla, esp and 2 esd) on basitarsus of leg IV; relative width of last opisthosomal segments and chaetotaxy of segment XI as in adult.</p>
            <p>Remarks</p>
            <p>As previously mentioned, the poor condition of the type material allowed us to observe and describe only a small number of morphological features. From the simple and short description of the female adult of this species provided by Silvestri (1913), which includes a few drawings, it is possible to recognize the following additional characters: lateral organ with a single blade, deutotritosternum with a single seta, and opisthosomal sternites IV–VI with three pairs of a setae flanked by one s seta on each side (1 s +3 a +3 a +1 s). The author also described and illustrated the chaetotaxy of the genital lobes (10 +10 and 2+2 setae on first and second lobes respectively), and illustrated the articles of the palp, leg I and leg IV. However, it is not clear whether he represented the complete chaetotaxy of these body parts. According to the drawing of the opisthosoma presented by Silvestri (1913: 217, fig. VII, 2), the immature already has the primordia of genital lobes, but it was not possible to confirm whether it is a male or a female juvenile.</p>
            <p> Due to the poor preservation of the type specimens, the collection of new specimens from the type locality is essential for its redescription according to the modern taxonomy of  Palpigradi , thus allowing a more detailed comparison with the congeneric species. </p>
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	https://treatment.plazi.org/id/03A37718FF83356BFDD1F8E8FC8DFAD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Souza, Maysa Fernanda Villela Rezende;Ferreira, Rodrigo Lopes	Souza, Maysa Fernanda Villela Rezende, Ferreira, Rodrigo Lopes (2022): Two extraordinary troglobitic species of Allokoenenia (Eukoeneniidae: Palpigradi) from Brazil: first records of this initially monotypic genus more than a century after its description. European Journal of Taxonomy 789: 11-48, DOI: 10.5852/ejt.2022.789.1627, URL: http://dx.doi.org/10.5852/ejt.2022.789.1627
03A37718FF86357DFDC7FA89FEF0FCC1.text	03A37718FF86357DFDC7FA89FEF0FCC1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allokoenenia canhembora Souza & Ferreira 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Allokoenenia canhembora sp. nov.</p>
            <p>urn:lsid:zoobank.org:act: F6B9B9E0-6B19-4FCB-82CF-A8B4F754F492</p>
            <p>Figs 3–9</p>
            <p>Diagnosis</p>
            <p>Frontal organ formed by 2 reticulated and expanded branches; 5 blades finely reticulated in the lateral organs; 5 setae on deutotritosternum; 7 pairs of short setae on propeltidium; 3 pairs of setae with similar length on metapeltidium; cheliceral fingers with 9 teeth each; coxae II–IV with 3, 3, 1 thick setae, respectively; 5 setae (grt, r, esp and 2 esd) on basitarsus of leg IV; opisthosomal tergites II–VI with two pairs of setae t between one setae s on each side; opisthosomal sternites IV–VI with three pairs of a setae (a 1 shorter than a 2 and a 3) flanked by one s seta on each side; opisthosomal segment XI with 2 pairs of ventral setae arranged in 2 rows; first lobe of female genitalia with 10 pairs of setae; flagellum formed by 14 moniliform segments with setae of different lengths, including very long setae inserted in a V arrangement from the sixth flagellar segment onwards.</p>
            <p>Etymology</p>
            <p>The species ephitet is derived from a word of the Tupi language meaning ʻfugitiveʼ. The species received this name due to the difficulty of capturing the specimens, which quickly ran away and hid under the calcite plates on the cave floor as soon as they were discovered (see: Habitats and threats). The name is to be treated as a noun in apposition.</p>
            <p>Material examined</p>
            <p>
                  Holotype BRAZIL • ♀; Bahia, Campo Formoso,  
                <a title="Search Plazi for locations around (long -40.89503/lat -10.511373)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-40.89503&amp;materialsCitation.latitude=-10.511373">Toca do Gonçalo Cave</a>
                 ; 10°30′40.94″ S, 40°53′42.10″ W; 527 m a.s.l.; 11 Jun. 2012; R.L. Ferreira leg.; ISLA 50394. 
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            <p>Other material</p>
            <p> BRAZIL • 1 ♀; same collection data as for holotype; 16 Mar. 2014; ISLA 50395.</p>
            <p>Description</p>
            <p>Adult female</p>
            <p>MEASUREMENTS AND RATIOS. See Table 2.</p>
            <p>BODY LENGTH. Without flagellum: 1420 μm.</p>
            <p>PROSOMA. Frontal organ formed by two reticulated and expanded branches (42 long), with blunt tips (Fig. 3A). Lateral organ with 5 blades pointed-lanceolate (32 long) and finely reticulated (Fig. 3B). Propeltidium with 7 +7 short setae (Fig. 4B). Setae t 1, t 2 and t 3 of metapeltidium 77, 75 and 77 long, respectively. Deuto-tritosternum with 5 setae in U-shaped arrangement (Fig. 4A). Labrum with 5+5 short setae. Basal segment of chelicera 195 long (dorsal length), with 6 proximal setae (p 4 and p 6 thickened and densely barbed; p 1 slightly thinner and barbed) (Fig. 4C), and 3 distal setae: d 3 (107) longer than d 1 (60) and d 2 (62); d 3 smooth near base and barbed in its distal half, d 1 and d 2 thin, flexible and with tiny projections in apex; and one apical seta. Hand of chelicera with 7 setae: 4 dorsal setae, 2 setae in its outer portion (1 close to articulation of movable finger and 1 on a tubercle close to teeth of fixed finger) and 1 seta inserted in its inner portion. Fingers with 9 teeth each.</p>
            <p>COXAL CHAETOTAXY. Pedipalp coxa with 18 setae (Fig. 3C); coxa I with 13 ordinary setae and two microsetae (Fig. 3D); coxa II with 3 thick setae, 2 macrosetae and 8 ordinary setae (Fig. 3E); coxa III with 3 thick setae, 1 macroseta, and 8 ordinary setae (including 1 small seta adjacent to macroseta) (Fig. 3F) and coxa IV with 1 thick and 8 ordinary setae (Fig. 3G).</p>
            <p>PEDIPALP. tc with 8 setae (2 considerably smaller than others); fe with 8 setae (Fig. 5A); ti with 8 setae; bta1 with 2 m and 1 normal seta; bta2 with 2 normal setae and 4 m; ta1 with 2 m; ta2 with 6 m (Fig. 5B); ta3 with 1 long fs, 1 cs with a conspicuous spine, 2 r, 10 m (one macroseta with basal denticle and conspicuous spine) and 10 normal setae (Fig. 7A).</p>
            <p>LEG I. tc with 12 normal setae (1 considerably smaller than others); fe with 9 normal setae; pa with 9 normal setae and 1 tb (Fig. 6A); ti with 9 normal setae; bta1 with 1 m, 1 normal seta, 2 tb and 1 fs (with inner branch shorter than outer branch); bta2 with 4 m, 2 tb and 1 long fs (Fig. 6B); bta3 with 1 r, 1 grt and 1 short normal seta; bta4 with 5 m, 1 tb and 1 long fs; ta1 with 5 normal setae (2 considerably smaller than others); ta2 with 5 m, 1 tb and 1 long fs (Fig. 6C); ta3 with 5 fs (with subequal branches) arranged as fs 1 / fs 2 / fs 3 / fs, rs (rs / fs 1 =2.2), 2 r, 1 cs, 13 m and 5 normal setae (Fig. 6D).</p>
            <p>4+5</p>
            <p>LEG II. tc with 3 normal setae; fe with 1 m and 4 normal setae; pa with 2 m, 1 thick seta, and 2 normal setae; ti with 1 thick seta and 4 normal setae; bta with 6 normal setae; ta with 1 r, 1 m and 9 normal setae.</p>
            <p>LEG III. tc with 2 normal setae; fe, pa, and ti with 1 thick seta and 4 normal setae each; bta with 6 normal setae; ta with 1 r, 1 m and 9 normal setae.</p>
            <p>LEG IV. tc with 3 normal setae; fe with 1 m and 2 normal setae; pa and ti with 1 thick and 4 normal setae each; bta with grt, r, esp, and 2 esd; ta1 with 4 normal setae; ta2 with 8 normal setae.</p>
            <p>IVBTA. 7.2 × longer than wide and with 5 setae (grt, r, esp and 2 esd). Seta r inserted in distal half (dr / IVbta =0.64) and grt inserted in proximal half of segment (Fig. 7B).</p>
            <p>OPISTHOSOMA. Tergites II–VI with 3 +3 dorsal setae, two pairs of t setae (t 1 =35‒41 μm, t 2 = 37‒42 μm) between a pair of slender setae (s =27‒35 μm) (Fig. 8A). Sternite III with 2+ 2 setae. Sternites IV–VI each with 3+ 3 thickened setae (a 1 =30 μm, a 2 =41‒45 μm, a 3 =48‒53 μm) between a pair of slender setae (s = 36‒40 μm) (inserted caudal to thick setae); pair of pores present between a 1 setae on sternites IV‒VI (Fig. 8B). Presence of inconspicuous paired cavities in intersegmental furrows between sternites III–IV, IV–V, V–VI and VI–VII, one at each side of opisthosoma. Segments VII–X with 8 setae each (4 dorsal and 4 ventral). Segment XI elongated (1.15 × as long as wide), with dorsal row of 2+2 long setae (inner pair=160 μm; outer pair =117‒125 μm) inserted in its distal half and 2 pairs of ventral setae, 1 inserted around middle of segment (60 μm) and other inserted in apical region (75 μm) (Fig. 8C). Intermediate ring of flagellum reduced in size (11 μm) and bears 2 tiny setae (8 μm).</p>
            <p>FEMALE GENITALIA. First lobe with 10 +10 setae in 5 transverse rows: 2+ 2 sternal setae (st 1, st 2) followed by 2+2, 1+1, 1 +1 and 4 +4 distal setae (a 1 =11 μm; a 2 = 12 μm; a 3 =20 μm; a 4 =30 μm); interior surface of the first lobe with a group of 3 orifices on either side and a medial pair of small orifices (Fig. 7C). Second lobe with 3+2 setae, asymmetry caused by lack of 1 y seta (x = 15 μm; y = 29 μm; z = 27 μm), and with cuticular spines; presence of group of 4 orifices on each half (Fig. 7D).</p>
            <p>FLAGELLUM. Arched and formed by 14 short and moniliform segments. First flagellar segment with 4 short setae (21–44 μm); second with 2 short setae (27–30 μm); third with 2 long setae (75 μm) and 4 short setae (10 – 30 μm); fourth with 3 short setae (15–31 μm); fifth with 5 short setae (25–33 μm); sixth with 2 very long setae; seventh with 1 short seta (15 μm) and 4 very long setae; eighth with 3 short setae (10–15 μm) and 2 very long setae; remaining segments bear 1 short seta (10–23 μm) and 4 very long setae each (Fig. 8D). Very long setae inserted laterally on segments, being arranged in V along flagellum (Fig. 9H). Length (μm) and total number of setae (N) of flagellar articles available in Table 1.</p>
            <p>Male and immatures</p>
            <p>Unkown.</p>
            <p>Distribution</p>
            <p>Known only from the type locality.</p>
            <p>Remarks</p>
            <p> Allokoenenia canhembora sp. nov. belongs to the genus  Allokoenenia mainly because of the shape of the last opisthosomal segments and flagellum (Figs 8C–D, 9F–H). It shares with  A. afra the presence of a short and arched flagellum formed by 14 moniliform segments, the chaetotaxy of opisthosomal sternites IV–VI (1 s +3 a +3 a +1 s), and the arrangement of the ventral setae of the last opisthosomal segment (in two rows) (Figs 2, 8C). However, they can be distinguished by the shape of the frontal organ (Figs 1C, 3A), and by the number of blades on lateral organs (5 vs 1) and deutotritosternal setae (5 vs 1) (Silvestri, 1913). Furthermore, we observed that the immature specimen of  A. afra has 6 setae on basitarsus IV (grt, r, gla, esp and 2 esd). This feature was not visible in the adults and was not described by Silvestri (1913), but they probably have at least 6 setae, since in general the setation of this article is already complete in immatures of this stage or they have fewer setae than the adults. In the new species, the basitarsus IV bears only 5 setae (grt, r, esp and 2 esd). Some flagellar characters can also be useful to distinguish these two species: the flagellar segments 1–3, 5, 7 and 9 of  A. afra bear the typical apical cuticular spines present in the flagellum of other palpigrades, as well as setae of uniform shape despite showing some variation in their lengths, as represented in Silvestri (1913: 216, fig. VI, 5) and confirmed by the study of the specimens (Fig. 1B); in  A. canhembora sp. nov. , the cuticular spines are absent and the flagellar segments carry setae of very different lengths and shapes, including very short (around 10–15 μm) to very long setae (longer than 100 μm). </p>
            <p>Habitat and threats</p>
            <p>The Toca do Gonçalo Cave is associated to carbonatic rocks from the Caatinga geological group, which comprises relatively young rocks deposited around three million years ago under a lacustrine condition. Such rocks are closely inserted to old carbonatic rocks from the Una geological group, from the Neoproterozoic. The cave is located in a semiarid biome (the Caatinga Formation), in the Campo Formoso Municipality, Bahia State, Northeastern Brazil (Fig. 9A). It presents around 500 m of topographed conduits and a single horizontal lenticular-shaped entrance (Fig. 9B), which connects to a descending conduit. This cave presents two distinct levels: the upper level (which corresponds to the left branch of the cave), is predominantly dry, and the lower level (corresponding to the right branch) is quite humid (Fig. 9C) (Souza-Silva &amp; Ferreira 2016). The lower level used to be mostly submerged by the water table, but it is currently accessible, since the water table reduced dramatically, as will be discussed further on. The main sources of organic matter for both terrestrial and aquatic species is the dissolved and particulate organic matter imported from the external habitats during strong rains, but also the small guano deposits and some root mats that can be used as food by invertebrates. The Toca do Gonçalo Cave is considered a hotspot of subterranean biodiversity (only three hotspots are currently known to South America), presenting 22 troglobitic species (Souza-Silva &amp; Ferreira 2016). Hostpots of subterranean biodiversity are those caves (or cave systems) with 20 or more cave-restricted species (Culver &amp; Sket 2000). However, most troglobitic species from this cave remain undescribed.</p>
            <p> All individuals of  A. canhembora sp. nov. were found at the lower cave level, always in very damp areas. In 2012, several specimens were observed, but only a single exemplar was captured since the cave substrata in the area prevented us to collect most of them. The water in karst systems usually presents high contents of dissolved calcium carbonates, which sometimes can form conspicuous deposits in the water surface inside caves. Such formations are usually called “calcite rafts” and are considered to be a kind of speleothem (Fig. 9D). Since the water table in Toca do Gonçalo Cave used to fluctuate in time, the areas devoid of water usually present a muddy substrate covered by many layers of calcite rafts (Fig. 9E). Most specimens of  A. canhembora sp. nov. were found below those calcite rafts, so in most cases, as soon as we turned a piece of calcite, the specimen used to run away and hide in the uncountable small spaces between the layers, thus making their capture unfeasible. This is the reason why we decided to name the species “canhembora”, meaning ʻfugitiveʼ as explained in the etymology. A visit in 2013 revealed specimens (although in smaler number in comparison to 2012), but none could be captured. Finally, in 2014, we could find a single specimen that was successfully sampled (Fig. 9F–H). </p>
            <p> All the cave visits from 2015 onward resulted in no further observations of specimens. Since the lower level of the cave was extremely dry, we believe that the specimens had probably migrated to inaccessible lower compartments. However, it is important to emphasize the fact that this species might be seriously threatened, as are other cave-restricted species from the Toca do Gonçalo Cave. This cave has been the target of several human impacts, which were highly intensified over the last years. In 1997, one of the authors (RLF) could attest that most part of the lower level was inaccessible due to the high level of the water table. In that period, the villagers of Gonçalo settlement used to draw water from the cave for their subsistence and this practice occurred during decades, by means of a diesel pump installed inside the cave. The villagers used to remove water from the cave once a week, thus the water table was persisting (although varying along the time), since the amount of water pumped out from the cave was negligible. However, an electric pump used by a local farmer for irrigation was installed in the cave in 2010 (Prevorčnik et al. 2012; Souza-Silva &amp; Ferreira 2016). According to the villagers, this pump remained active during the whole day. In one of our surveys in 2010, a pronounced decrease of the water table (at least two meters) was observed. In 2012, the equipments were removed from the cave since the National Cave Research and Conservation Center intervened and asked the municipal government to dig an artesian well for the villagers. In December 2018, a cave visit revealed that the water table was no longer observed. Hence, previously inaccessible areas could be explored, revealing the cave to be much longer than thought. However, from the 22 troglobitic species occurring in this cave, only two (an isopod and a springtail) were observed in this visit, in some moist areas located deep inside the cave. No specimens of  A. canhembora sp. nov. were observed, despite the intense efforts employed by the team in searching for them. The villagers mentioned that the building of a reservoir some dozens of kilometers from the cave, has altered the water balance, so that eventual flooding that used to reach the area never occurred again. Therefore, the Toca do Gonçalo Cave is seriously threatened and it is crucial that the Brazilian government intervene, in order to ensure the protection of this unique habitat in South America. </p>
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	https://treatment.plazi.org/id/03A37718FF86357DFDC7FA89FEF0FCC1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Souza, Maysa Fernanda Villela Rezende;Ferreira, Rodrigo Lopes	Souza, Maysa Fernanda Villela Rezende, Ferreira, Rodrigo Lopes (2022): Two extraordinary troglobitic species of Allokoenenia (Eukoeneniidae: Palpigradi) from Brazil: first records of this initially monotypic genus more than a century after its description. European Journal of Taxonomy 789: 11-48, DOI: 10.5852/ejt.2022.789.1627, URL: http://dx.doi.org/10.5852/ejt.2022.789.1627
03A37718FF903577FDF0FC98FC0FFCA9.text	03A37718FF903577FDF0FC98FC0FFCA9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allokoenenia stygia Souza & Ferreira 2022	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Allokoenenia stygia sp. nov.</p>
            <p>urn:lsid:zoobank.org:act: C37DF62D-1144-4E02-9076-4BF344AEB140</p>
            <p>Figs 10–19</p>
            <p>Diagnosis</p>
            <p>Frontal organ formed by 2 reticulated and lanceolate branches; 3 blades finely reticulated in the lateral organs; 5 setae on deutotritosternum; 7 pairs of short setae on propeltidium; 3 pairs of setae with similar lengths on metapeltidium; cheliceral fingers with 9 teeth each; 2 thickened dorsal setae proximally inserted in the cheliceral hand; coxae II–IV with 3, 3, 0 thick setae, respectively; 6 setae (grt, r, gla, esp and 2 esd) on basitarsus of leg IV; opisthosomal tergites II–VI with 2 pairs of robust setae (one pair of setae t between one setae s on each side); opisthosomal sternites IV–VI with 2 pairs of a setae flanked by 1 s seta on each side; opisthosomal segment XI with 2 pairs of ventral setae arranged in 2 rows; first lobe of female genitalia with 11 pairs of setae.</p>
            <p>Etymology</p>
            <p>The species epithet is derived from Styx, a Titan goddess and also one of the five rivers that form the boundary between Earth and the Underworld according to Greek mythology. From the Latin ‘stygius’ derived from the Ancient Greek ‘Στυγιος’ (‘Stúgios’, ‘relating to Styx’), from ‘Στυγξ’ (‘Stúx’, ‘Styx, chief river of underworld’). The name is a noun in apposition.</p>
            <p>Material examined</p>
            <p>
                  Holotype BRAZIL • ♀; Pará,  
                <a title="Search Plazi for locations around (long -49.97728/lat -6.352839)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.97728&amp;materialsCitation.latitude=-6.352839">Canaã dos Carajás</a>
                 , SB_0114 cave; 6°21′10.22″ S, 49°58′38.20″ W; 326 m a.s.l.; 10–31 Jan. 2013; F.F. Pellegatti et al. leg.; MZSP 54247. 
            </p>
            <p>
                  Paratype BRAZIL • 1 ♀; Pará,  
                <a title="Search Plazi for locations around (long -49.97728/lat -6.3535166)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-49.97728&amp;materialsCitation.latitude=-6.3535166">Canaã dos Carajás</a>
                 , SB_0112 cave; 6°21′12.66″ S, 49°58′38.20″ W; 331 m a.s.l.; 10–31 Jan. 2013; F.F. Pellegatti et al. leg.; MZSP 54249  . 
            </p>
            <p>Description</p>
            <p>Adult female</p>
            <p>MEASUREMENTS AND RATIOS. See Table 2.</p>
            <p>BODY LENGTH. Without flagellum: 1030‒1295 μm.</p>
            <p>PROSOMA. Frontal organ formed by 2 reticulated and lanceolate branches (40 long) (Fig. 10A). Lateral organ with 3 blades pointed-lanceolate (24‒27 long) and finely reticulated (Fig. 10B). Propeltidium with 7 +7 short setae (Fig. 10D). Setae t 1, t 2 and t 3 of metapeltidium 72‒75, 67‒80 and 80‒82 long, respectively (Fig. 10C). Deuto-tritosternum with 5 setae in U-shaped arrangement (Fig. 11A). Labrum with 5+ 5 short setae. Basal segment of chelicera 280‒310 long (dorsal length), with 6 proximal setae (p 4 and p 6 thickened and densely barbed; remaining setae sparsely barbed, with p 5 more conspicuous than usual) (Fig. 11B), and 3 distal setae: d 3 (142) longer than d 1 (55‒60) and d 2 (62); d 3 smooth near base and barbed in distal half, d 1 and d 2 thin, flexible and with tiny projections at apex (Fig. 10E); and 1 apical seta. Hand of chelicera with 7 setae: 4 dorsal setae (2 inserted close to base of article and considerably thicker than others) (Fig. 11D), 2 setae in outer portion (1 close to articulation of movable finger and 1 on tubercle close to teeth of fixed finger) and 1 seta inserted in inner portion. Fingers with 9 teeth each.</p>
            <p>COXAL CHAETOTAXY. Pedipalp coxa with 18 setae (Fig. 12A); coxa I with 11 ordinary setae and 2 microsetae (Fig. 12B); coxa II with 3 thick setae, 2 macrosetae, and 8 ordinary setae (Fig. 12C); coxa III with 3 thick setae, 1 macroseta, and 8 ordinary setae (including 1 small seta adjacent to macroseta) (Fig. 12D) and coxa IV with 8 ordinary setae (thick seta absent) (Fig. 12E).</p>
            <p>PEDIPALP. tc with 8 (9 in paratype) setae (2 considerably smaller than others); fe with 8 setae; ti with 8 setae; bta1 with 2 m and 1 normal seta; bta2 with 2 normal setae and 4 m; ta1 with 2 m; ta2 with 6 m (Fig. 13); ta3 with 1 long fs, 1 cs with conspicuous spine, 2 r, 8 m (1 with basal denticle and conspicuous spine) and 12 normal setae (Fig. 16A).</p>
            <p>LEG I. tc with 11 normal setae (2 considerably smaller than others); fe with 9 normal setae; pa with 9 normal setae and 1 tb (Fig. 14A); ti with 9 normal setae; bta1 with 2 normal setae, 2 tb and 1 fs (with subequal branches); bta2 with 2 m, 2 normal setae, 2 tb and 1 fs (with subequal branches) (Fig. 14B); bta3 with 1 r, 1 grt and 1 short normal seta; bta4 with 5 m, 1 tb and 1 long fs; ta1 with 5 normal setae (2 considerably smaller than others); ta2 with 5 m, 1 tb and 1 long fs (Fig. 15A); ta3 with 5 fs (with subequal branches) arranged as fs 1 / fs 2 + 3 / fs 4+5, rs (rs / fs 1 = 2.4), 2 r, 1 cs with conspicuous spine, 9 m and 9 normal setae (Fig. 15B).</p>
            <p>LEG II. tc with 3 normal setae; fe with 5 normal setae; pa and ti with 1 thick seta and 4 normal setae each; bta with 6 normal setae; ta with 1 r, 1 m and 9 normal setae.</p>
            <p>LEG III. tc with 2 normal setae; fe, pa, and ti with 1 thick seta and 4 normal setae each; bta with 6 normal setae; ta with 1 r, 1 m and 9 normal setae.</p>
            <p>LEG IV. tc with 3 normal setae; fe with 1 m and 2 normal setae; pa and ti with 1 thick and 4 normal setae each; bta with grt, r, gla, esp, and 2 esd; ta1 with 4 normal setae; ta2 with 8 normal setae.</p>
            <p>IVBTA. 7 × longer than wide and with 6 setae (grt, r, gla, esp and 2 esd). Seta r inserted around middle of article (dr / IVbta =0.55); gla inserted close to r and grt inserted in proximal half (Fig. 16B).</p>
            <p>OPISTHOSOMA. Tergites II–VI with 2+2 dorsal robust setae, one pair of t setae (t 1 = 37‒50 μm) between pair of s setae (s =25‒35 μm) (Fig. 11E). Sternite III with 2+ 2 setae. Sternites IV–VI each with 2 +2 thickened setae (a 1 =50‒62 μm, a 2 = 55‒65 μm) between pair of slender setae (s =45‒52 μm) (inserted caudal to thick setae); pair of pores present between a 1 setae on sternites IV‒VI (Fig. 11F). In paratype, presence of inconspicuous paired cavities in intersegmental furrows between sternites III–IV, IV–V, and V–VI, 1 at each side of opisthosoma; these cavities not observed in holotype. Segments VII–X with 8 setae each (4 dorsal and 4 ventral). Segment XI elongated (1.08‒1.14 × as long as wide), with dorsal row of 2+ 2 long setae (inner pair =122 μm; outer pair=112 μm) inserted around half and two pairs of ventral setae, one inserted in proximal half (62 μm) and other around half (77–82 μm). Intermediate ring of flagellum lost or absent in both specimens (Figs 11C, 17).</p>
            <p>FEMALE GENITALIA. First lobe with 11 +11 setae in 5 transverse rows: 2 +2 sternal setae (st 1, st 2) followed by 3+3, 1+1, 1 +1 and 4+ 4 distal setae (a 1 =20 μm; a 2 =25 μm; a 3 = 30 μm; a 4 =37 μm); interior surface of first lobe with group of 3 orifices on either side and medial pair of small orifices (Fig. 18A). Second lobe with 3+ 3 setae (x = 10–11 μm; y = 35–37 μm; z = 25–34 μm), and with cuticular spines; presence of group of 4 orifices on each half (Fig. 18B).</p>
            <p>FLAGELLUM. Not preserved.</p>
            <p>Male and immatures</p>
            <p>Unkown.</p>
            <p>Distribution</p>
            <p>Known only from SB_0114 and SB_0112 caves (municipality of Canaã dos Carajás, Pará State, Brazil).</p>
            <p>Remarks</p>
            <p> Allokoenenia stygia sp. nov. fits the diagnosis of  Allokoenenia by the relative width of the last opisthosomal segments. The flagellum was lost during the collection and therefore its morphology remains unknown. As with  A. afra ,  A. canhembora sp. nov. , and  Allokoenenia sp. , the ventral setae on the last opisthosomal segment of this new species is arranged in two rows. </p>
            <p> Allokoenenia stygia sp. nov. differs from  A. afra by the shape of frontal organ (Figs 1C, 10A), and by the number of blades on lateral organs (3 vs 1) and deutotritosternal setae (5 vs 1) (Silvestri 1913). Considering the new congeneric species herein described,  A. stygia sp. nov. differs from both by presenting the two most proximal setae on cheliceral hand thickened, the cs seta of leg I with a conspicuous spine, trochanter of leg I with 11 setae (vs 12 setae), coxa I with 13 setae (vs 15), coxa IV without thick seta (vs one seta), a single pair of t setae on opisthosomal tergites II–VI (vs two pairs), 6 setae on bta IV (vs 5 setae) and 3 blades on lateral organs (vs 4 in  Allokoenenia sp. and 5 in  A. canhembora sp. nov. ). Additionally, the new species can be distinguished from  A. canhembora sp. nov. by having 2 pairs of thickened setae (a) on opisthosomal sternites IV–VI (vs 3 pairs) and 11 pairs of setae on the first lobe of female genitalia (vs 10 pairs). </p>
            <p>Habitat and threats</p>
            <p> Individuals of  Allokoenenia stygia sp. nov. (Fig. 19E) were found in two adjacent iron ore caves (about 90 m bee-line from each other) located in a ferruginous plateau known as Serra da Bocaina (Fig. 19D), inserted in the domain of Amazon Forest (Fig. 19A), Northern Brazil. Other caves in this region were sampled, but no specimens of this species were found at other localities. These two caves, named SB_0114 (Figs 19B–C) and SB_0112, have horizontal projections of 89 and 16.1 m, respectively, and are formed by a single spongiform gallery (Fig. 19G). In 2017, a National Park was created (Parque Nacional dos Campos Ferruginosos) incorporating the Serra da Bocaina region. This conservation unit has 790.9 km 2 and protects two important iron ore formations within the Carajás geological province: the Serra da Bocaina and the Serra do Tarzan. However, although the top and the slopes of those plateaus are preserved, most of the area (especially in the surroundings of the Serra da Bocaina) is very altered by human activities (especially pastures) (Fig. 19D, F). Fortunately, the area where the SB_0112 and SB_0114 caves are located is preserved. At the same time, despite the recently created conservation unit, frequent criminal fires hit the region, often advancing over the forest. Such fires usually come from conflicts between the local farmers and landless people, and such impacts may represent a threat for the species, especially when considering its very restricted distribution. </p>
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	https://treatment.plazi.org/id/03A37718FF903577FDF0FC98FC0FFCA9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Souza, Maysa Fernanda Villela Rezende;Ferreira, Rodrigo Lopes	Souza, Maysa Fernanda Villela Rezende, Ferreira, Rodrigo Lopes (2022): Two extraordinary troglobitic species of Allokoenenia (Eukoeneniidae: Palpigradi) from Brazil: first records of this initially monotypic genus more than a century after its description. European Journal of Taxonomy 789: 11-48, DOI: 10.5852/ejt.2022.789.1627, URL: http://dx.doi.org/10.5852/ejt.2022.789.1627
