taxonID	type	description	language	source
03B187B7FFBDFFF8FF3AFC46C439A1E8.taxon	diagnosis	Diagnosis. Black or yellow flies with one pair of setulae on lunule; palpus usually short and clavate, but sometimes elongate; proboscis long to very long; crossvein dm-cu absent; comb-like row of setae posteroventrally on hind basitarsus; and basisternum very small and shaped like a narrow triangle. Chaetotaxy. The chaetotaxy described is that of the Nearctic fauna; the chaetotaxy of the Paramyia from other regions has not been studied. Head: 2 pairs of frontal setae medioclinate; 3 pairs of orbital setae with the 2 lower pairs long and lateroclinate and upper pair shorter and medio-reclinate (less than half the length of lower orbital seta); 1 paired row of interfrontal setulae (stronger than upper pair of orbital seta), 1 pair of supra-antennal setae (anteriormost interfrontal setulae), 1 pair of setulae between supra-antennal seta and eye margin; ocellar setae as long as orbital setae and almost lateroclinate; postocellar setae cruciate; 1 pair of long lateral vertical setae and medial vertical setae. Thorax: 1 postpronotal seta, 2 notopleural setae, 1 presutural seta (close to notopleuron); 1 supraalar seta; 2 postalar setae; 2 dorsocentral setae (anterior more than half the length of posterior); 1 intrapostalar seta; 1 prescutellar seta (acrostichal); 2 scutellar setae (anterior more than half length of apical; apical pair divergent); 1 minute proepisternal seta and 1 minute proepimeral seta, anepisternum and anepimeron bare, 1 very strong katepisternal seta (upper posterior corner, some thinner seta along posterior side). Female postabdomen based on P. nitens (Fig. 2). For each species, the abdomen usually closely matches the color of the thorax. The sternites size are fairly standard across species, with sternite 1, 2 and 8 being quite short and sternite 5 being the longest. The amount of setae and their distribution is variable within species, making it unreliable as a character. However, there is a common basal configuration. Sternite 7 usually as a minimum of 6 setae arranged in two rows. First row, with 2 setae, with one at each lateral side of the sternite. Second row, with 4 setae, right at the apical end of the sternite, distributed in group of 2 with a bigger gap in the middle. There are often a few extra setae distributed in the bottom of the sternite, sometimes creating a 3 rd row located just above the 1 st one, but still with a variable configuration within species. Tergite 7 usually has a minimum of 8 setae arranged in two rows. First row, with a minimum of 2 setae, with one at each lateral side of the tergite, but often with a few randomly placed in between, often closer to center. Second row, with a minimum of 6 setae, right at the apical end of the tergite, separated in two groupings of 3 with a bigger gap in the middle. There are often extra setae located in the 2 nd row, while some create a 3 rd row just above the 1 st one. Still a lot of variation within species. The cerci usually have a pair of long setae apically, which are almost twice as long as the other surrounding setae.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFBDFFF8FF3AFC46C439A1E8.taxon	discussion	Remarks. Females are highly variable within species making it difficult to identify most species based on morphology; using sequencing is recommended to confirm female identity. The color pattern on the legs were consistent for the males (see key and species description), which is not the cases for females. Additionally, there was no obvious variation in the female post-abdomen.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFBBFFFFFF3AFD86C768A230.taxon	description	(Figs. 2 – 4, 25)	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFBBFFF3FF3AFDEFC7F8A1E8.taxon	description	Description. (Fig. 3) Total length ♂ 1.3 - 1.6 mm, ♀ 1.8 - 1.9 mm. Overall colour brown to dark brown, frontal trian- gle dark brown to black, shiny, reaching lunule; frons 1.25 times longer than wide and 1.44 times wider than flagellum; width at widest point 2.3 - 2.4 times the bottom eye width; gena brown with prunescence, 0.16 - 0.18 times eye height; postgena dark brown, with light prunescence and 0.30 - 0.33 width of head; scape and pedicel dark brown, first flagellomere trapezoidal, 1.2 times wider than long, dark brown with pale pubescence, arista dark brown and arista length 1.45 - 1.8 times width of first flagellomere; medium facial carina, visible laterally; palpus dark brown with 4 - 5 enlarged setae apically and 1 ventral seta pre-apically, labium brown 0.40 - 0.43 mm and labella brown 0.45 - 0.48 mm long. Scutum shiny dark brown; scutellum dark brown, microsetose; fore femur all brown; fore tibia all brown often slightly paler on anterior side; femur and tibia of mid and hind leg brown; all junctions of femur and tibia (“ knee ”) brown; all tarsi yellowish; hind basitarsus shorter than length of tarsi 2 - 4 combined; haltere brown basally and dark brown apically; wing 1.22 - 1.32 mm long; abdomen shiny dark. Male postabdomen (Fig. 4). Epandrium in lateral view 1.1 times wider than high, in posterior view 2.5 times wider than high (before cerci), small setae densely covering posterior portion of epandrium mixed with a low density of large setae; surstylus 1.3 times as high as epandrium, paddle shape with a large half-circular structure situated basally, on inner surface, with only a few short markings in the basal portion, setae medium length, covering anterior side and distal half of surstylus; cerci large and pointed apically, 0.70 times length of surstylus in posterior view, with a pair of very long preapical setae and other shorter setae, less than 1 / 3 its length, apically swollen in lateral view. Molecular barcode sequence can be accessed through GenBank Accession: KR 643351 and BOLD BIN: BOLD: AAG 0168 (average p-distance within BIN: 0.5 %).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFBBFFF3FF3AFDEFC7F8A1E8.taxon	distribution	Distribution (Fig. 25). Widespread throughout North America.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFBBFFF3FF3AFDEFC7F8A1E8.taxon	materials_examined	Material examined. UNITED STATES: Pennsylvania: 1869, ♂. (Holotype, MCZ) [MCZ-ENT 00013447]. CANADA: Alberta: Wood Buffalo National Park, Benchmark weather station, 59.5607 ° N, 112.261 ° W, 219 m, 2 ♀, 13 - 27. vii. 2012, N. Labine (BIOUG), 2 ♀, 13 - 20. vii. 2013, N. Labine (BIOUG); British Columbia: 10 km W Kamloops, New Afton Mine, 50.6719 ° N, 120.533 ° W, 1 ♀, 24. vii- 1. viii. 2013, C. Simon, Malaise trap (BIOUG), 10 mi. E. Creston, 49.1 ° N, 116.51 ° W, 1 ♂, 31. vii. 1980, G. Gibson, sweeping (DEBU); Manitoba: 2 km W Gardenton, Tall Grass Prairie Field Station, 49.0769 ° N, 96.7139 ° W, 1 ♂, 24. vi- 1. vii. 2008, H. D. White, Malaise trap (LEMQ); New Brunswick: Fundy, Maple Grove, 45.5822 ° N, 64.9859 ° W, 143 m, 1 ♀, 08. viii. 2009, BIObus 2009 (BIOUG), Kouchibouguac National Park, 46.8 ° N, 64.97 ° W, 1 ♂, 30. vii- 2. viii. 2013, O. Lonsdale (CNC); Newfoundland and Labrador: Terra Nova National Park, 48.598 ° N, 53.9702 ° W, 127 m, 1 ♀, 25. vi- 2. vii. 2013, E. Perry, Malaise trap (BIOUG), 1 ♀, 9 - 16. vii. 2013, E. Perry, Malaise trap (BIOUG); Nova Scotia: Cape Breton Highlands, Warren Lake, 46.7131 ° N, 60.3835 ° W, 59 m, 1 ♀, 24. vii. 2009, BIObus 2009, freehand (BIOUG), Halifax, Point Pleasant Park, 44.623 ° N, 63.5686 ° W, 30 m, 1 ♀, 15 - 22. vi. 2013, T. Zemlak, Malaise trap (BIOUG), 6 ♂, 4 ♀, 13 - 20. vii. 2013, T. Zemlak, Malaise trap (BIOUG), 1 ♂, 5 ♀, 6 - 13. vii. 2013, T. Zemlak, Malaise trap (BIOUG), 1 ♂, 29. vii- 6. viii. 1987, T. Zemlak, Malaise trap (BIOUG), 1 ♂, 22 - 26. vii. 2005, T. Zemlak, Malaise trap (BIOUG), 2 ♀, 27. vii- 3. viii. 2013, T. Zemlak, Malaise trap (BIOUG), 1 ♂, 2 ♀, 29. vi- 6. vii. 2013, T. Zemlak, Malaise trap (BIOUG), 1 ♂, 22 - 29. vi. 2013, T. Zemlak, Malaise trap (BIOUG), 2 ♂, 1 ♀, 20 - 27. vii. 2013, T. Zemlak, Malaise trap (BIOUG), Kejimkujik, Graf- ton Lake, 44.383 ° N, 65.2023 ° W, 103 m, 1 ♂, 01. viii. 2009, BIObus 2009 (BIOUG), 1 ♀, 03. viii. 2009, BIObus 2009 (BIOUG); Ontario: 4452 Rowsome Rd., Elizabethtown, 44.6213 ° N, 76.273 ° W, 120 m, 1 ♀, 19. vi. 2010, J. Sones (BIOUG), Algoma Distr., Witchdoctor Lake, 47.09 ° N, 83.76 ° W, 1 ♂, 29. viii. 1992, Cannings, Swann, Malaise trap (DEBU), Bruce Co., Bruce Peninsula National Park, Lake Scugog, 45.1167 ° N, 81.5333 ° W, 1 ♂, 1 - 17. viii. 2002, S. A. Marshall, Malaise trap (DEBU), Bruce Co., Cameron Lake, Fen, 45.219 ° N, 81.5465 ° W, 2 ♂, 28. vi- 16. vii. 1998, S. A. Marshall, Malaise trap (DEBU), Bruce Co., Dorcas Bay, 45.19 ° N, 81.58 ° W, 2 ♂, 21. vii- 1. viii. 2003, S. A. Marshall, Malaise trap (DEBU), 1 ♂, 1 - 22. viii. 2003, S. A. Marshall, Malaise trap (DEBU), Bruce Co., Dorcas Bay Dunes, 45.19 ° N, 81.58 ° W, 3 ♂, 2 - 25. viii. 1999, S. A. Marshall, Malaise trap (DEBU), 1 ♂, 20. vii- 2. viii. 1993, S. A. Mar- shall, Malaise trap (DEBU), 1 ♂, 20 - 27. vi. 2006, S. A. Marshall, Malaise trap (DEBU), 2 ♂, 11 - 20. vii. 1999, S. A. Marshall, Malaise trap (DEBU), Bruce Co., Dorcas Bay, dune, 45.19 ° N, 81.58 ° W, 1 ♂, 16 - 21. vii. 2012, S. A. Mar- shall, Malaise trap (DEBU), Bruce Co., Dorcas Bay, pond nr dune, 45.19 ° N, 81.58 ° W, 1 ♂, 11 - 20. vii. 1999, S. A. Marshall, Malaise trap (DEBU), 1 ♂, 11 - 18. vii. 2006, S. A. Marshall, Malaise trap (DEBU), Bruce Co., Fanthom Five National Park, Middle Island, 45.26 ° N, 81.66 ° W, 1 ♂, 23 - 30. vi. 2008, S. A. Marshall, pan trap (DEBU), Dorcas Bay, 45.19 ° N, 81.58 ° W, 2 ♂, 21 - 28. vii. 2008, S. A. Marshall, Malaise trap (DEBU), Fathom 5 National Park, Bears Rumpls., 45.26 ° N, 81.66 ° W, 1 ♂, 23 - 30. vi. 2008, T. Woodcock, S. A. Marshall, pan trap (DEBU), Gaudry Township, Witchdoctor Lake, 47.08 ° N, 83.76 ° W, 1 ♂, 29. viii. 1992, R. A. Cannings, J. E. Swann, Malaise trap (ROM), Halibur- ton District, Dorset, 1.6 km S. Leslie M. Frost Nat. Res. Centre, 45.16 ° N, 78.85 ° W, 1 ♂, 25. vi- 2. vii. 2013, B. Hub- ley, Malaise trap (ROM), Hilton Township, Tendy Bay, 46.09 ° N, 83.92 ° W, 1 ♂, 9. viii. 1992, J. E. Swann, Malaise trap (ROM), L. Cognashene Muskoka Dist., 44.96 ° N, 79.92 ° W, 1 ♂, 28. vii. 1981, J. T. Huber (CNC), Lambton Co., Pinery Provincial Park, power line, right-of-way, 44.55 ° N, 76.3667 ° W, 3 ♂, 7 - 16. vii. 2001, L. Packer, Malaise trap (ROM), 1 ♂, 24. vi- 9. vii. 1996, L. Packer, Malaise trap (ROM), 1 ♂, 28. vi- 8. vii. 1986, L. Packer, Malaise trap (ROM), 1 ♂, 28. vi- 12. vii. 1986, L. Packer, Malaise trap (ROM), Lambton Co., Pinery Provincial Park, Sand dunes, 44.55 ° N, 76.3667 ° W, 1 ♂, 28. vi- 16. vii. 1998, L. Packer, Malaise trap (ROM), 1 ♂, 19 - 26. vii. 2006, L. Packer, Malaise trap (ROM), Makonie Lake, 48.35 ° N, 83.24 ° W, 1 ♂, 11. vii. 2016, S. A. Marshall, hand collecting (DEBU), Manitoulin Distr., Manitoulin Is., Carter Bay, 45.6064 ° N, 82.1408 ° W, 1 ♂, 1 - 8. viii. 2003, Buck, Marshall, Malaise trap (DEBU), Manitoulin Island, Kip Fleming Tract, 8 km SW Gore Bay, 45.8703 ° N, 82.5419 ° W, 1 ♂, 27. vii- 3. viii. 2013, Mar- shall et al., Malaise trap (DEBU), Mer Bleu, 5 mi. E. Ottawa, 45.39 ° N, 75.51 ° W, 1 ♂, 17. vii. 1966, D. D. Munroe, Malaise trap (CNC), Northumberland Co., Cobourg, 3 km N Baltimore, 43.96 ° N, 78.17 ° W, 1 ♂, 17. vii. 1986, P. Danielsson (USNM), Parry Sound, Waubamik, 45.46 ° N, 80 ° W, 1 ♂, 7. viii. 1915, J. M. Aldrich (USNM), Pukaskwa National Park, 48.601 ° N, 86.2893 ° W, 631 m, 1 ♀, 6 - 12. viii. 2013, C. Harpur, Malaise trap (BIOUG), 1 ♂, 27. vi- 4. vii. 2006, C. Harpur, Malaise trap (BIOUG), Stonecliffe, Driftwood Provincial Park, 46.19 ° N, 77.85 ° W, 12 ♂, 2 - 25. viii. 1999, S. Peck, Malaise trap (DEBU), Thousand Islands National Park, 44.4534 ° N, 75.865 ° W, 1 ♂, 18 - 24. vi. 2015, M. Brown, Malaise trap (BIOUG), 1 ♂, 2 ♀, 2 - 9. vii. 2014, M. Brown, Malaise trap (BIOUG), 5 ♂, 5 ♀, 16 - 23. vii. 2014, M. Brown, Malaise trap (BIOUG), Thunder Bay Distr., Prairie River at Hwy 17, 38 km E Terrace Bay, 48.8011 ° N, 86.7844 ° W, 1 ♀, 15 - 19. vii. 2002, M. Buck, white pans (DEBU), 2 ♂, 15 - 19. vii. 2002, M. Buck, Mal- aise trap (DEBU), 1 ♂, 15 - 22. vii. 2005, M. Buck, Malaise trap (DEBU); Quebec: Beechgrove, 45.65 ° N, 76.1333 ° W, 1 ♂, 29. vi. 1962, J. R. Vockeroth (CNC), Bois-des-Bel Bog, 47.9683 ° N, 69.4274 ° W, 1 ♂, 21. vi. 2006, A. G. Taillefer, sweep (LEMQ), Gatineau Co., Masham Twp., 45.64 ° N, 76.01 ° W, 1 ♂, 10 - 13. vi. 1974, D. M. Wood (CNC), 1 ♂, 6 - 8. vii. 1974, D. M. Wood (CNC), James Bay Rte km 204.5, 50.9831 ° N, 77.6339 ° W, 1 ♂, 7 - 16. vii. 2001, M. Buck, B. Buck, Malaise trap (DEBU), James Bay Rte km 398.8, 52.3569 ° N, 77.1069 ° W, 1 ♂, 8 - 16. vii. 2001, M. Buck, B. Buck, white pans (DEBU), 1 ♂, 8 - 18. viii. 1971, M. Buck, B. Buck, white pans (DEBU), James Bay Rte km 570.3, 53.4942 ° N, 77.6208 ° W, 1 ♂, 9 - 18. xii. 1985, M. Buck, B. Buck, Malaise trap (DEBU), James Bay Rte., km 204.5, 50.9831 ° N, 77.6339 ° W, 4 ♂, 7 - 16. vii. 2001, M. Buck, B. Buck, Malaise trap (DEBU), Johnville Bog and Forest Park, 45.345 ° N, 71.7417 ° W, 1 ♀, 10 - 12. vii. 2005, J. Savage, J. Kuchta, Malaise trap (LEMQ), Laniel, 47.05 ° N, 79.28 ° W, 1 ♂, 1. vii. 1944, A. R. Brooks (CNC), Mt. St. Marie Low, 45.94 ° N, 75.87 ° W, 549 m, 1 ♂, 22. vi. 1965, J. R. Vock- eroth (CNC), Ogden, Marlington Bog, 45.04 ° N, 72.1733 ° W, 1 ♀, 4 - 11. vii. 2006, A. Moores, Malaise trap (LEMQ), Old Chelsea, Summit King Mt., 45.49 ° N, 75.86 ° W, 351 m, 1 ♂, 25. vi. 1962, J. R. Vockeroth (CNC), Rougemont, 45.4962 ° N, 73.0685 ° W, 1 ♀, 21 - 28. vii. 2008, V. Levesque, trunk trap (LEMQ), St-Daniel, Parc National de Fronte- nac, 45.9683 ° N, 71.1517 ° W, 13 ♂, 12 - 19. vii. 2006, A. Moores, pan trap (LEMQ), 1 ♂, 19 - 26. vii. 2013, A. Moores, pan trap (LEMQ), 1 ♂, 19 - 26. vii. 2006, A. Moores, pan trap (LEMQ), 1 ♂, 12 - 15. viii. 2005, A. Moores, pan trap (LEMQ), 1 ♂, 19 - 21. vii. 2016, A. Moores, pan trap (LEMQ). UNITED STATES: Arkansas: Montgomery Co., 20 mi W Glenwood, 34.33 ° N, 93.55 ° W, 3 ♂, 15. v- 28. viii. 1994, H. W. Robinson, sweep (LEMQ), 2 ♂, 15 - 21. vi. 2006, H. W. Robinson, sweep (LEMQ); Maine: Lincoln Co., East Boothbay, 43.85 ° N, 69.5833 ° W, 1 ♀, 9. viii. 2000, J. Sav- age, sweep (LEMQ); Maryland: Glen Echo, 38.97 ° N, 77.14 ° W, 1 ♂, 22. viii. 1922, J. R. Malloch (USNM); Mississippi: Scott Co., Forest, 32.36 ° N, 89.48 ° W, 1 ♂, 5 - 12. viii. 1971, S. A. Marshall, pan trap (DEBU); New Hampshire: Strafford Co., Spruce Hole, 3 mi. SW Durham, 43.14 ° N, 70.92 ° W, 2 ♂, 3 - 9. vii. 1987, D. S. Chandler, flight intercep- tion trap (DEBU); Tennessee: Rutledge, 36.28 ° N, 83.5 ° W, 1 ♂, 1 - 31. vii. 1954, M. R. Wheeler (AMNH); Vermont: East Franklin, Lake Carmi State Park, 44.9533 ° N, 72.8817 ° W, 1 ♀, 4 - 11. vii. 2006, A. Moores, pan trap (LEMQ).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFBBFFF3FF3AFDEFC7F8A1E8.taxon	discussion	Remarks. The holotype of P. nitens was examined and dissected for the first time. The holotype was not verified by Papp (2001). Based on Papp’s identification labels at the ROM and his re-description of P. nitens (e. g. en- larged male flagellum and genitalia drawings), it appears to correspond better to the new species (P. pseudonitens) described below, which is the most abundant and widely distributed species in North America. The status of P. nigra Williston is mentioned as a likely junior synonym of P. nitens by Sabrosky (1965), which was echoed by Papp (2001). As we could not locate the holotype of P. nigra, we do not feel we can realistically assess the status of P. nigra and we leave it as a valid species but this species will not be re-evaluated here. Females of P. anguliloba, P. lustrum, P. nitens, P. rectiloba, P. silvestris, P. wheeleri can only be reliably identified through sequencing.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFB7FFF0FF3AF8CAC56DA290.taxon	description	(Fig. 5 - 6; 26) Description. (Fig 5) Total length ♂ 1.3 mm, ♀ 1.6 - 2.2 mm. Overall colour yellowish, frontal triangle light brown to yellowish, shiny, almost reaching lunule; frons 1.1 times longer than wide and 2.2 times wider than flagellum; width at widest point 2.5 times the bottom eye width; gena yellowish and shiny, 0.29 times eye height; postgena yellowish and shiny and 0.32 width of head; scape and pedicel light brown, first flagellomere subquadrate, 1.3 times wider than long, dark grey around and yellowish at the base with pale pubescence, arista dark grey and 3.0 times width of first flagellomere; mild facial carina; palpus yellow with 4 enlarge setae apically, 1 longer ventral seta pre-apical and 4 short ventral setae, labium yellow 0.40 mm and labella yellow 0.43 mm long. Scutum shiny yellowish to light brown; scutellum yellowish; all legs yellowish; hind basitarsus equal length of tarsi 2 - 4 combined; haltere yellowish; wing 1.8 mm long; abdomen shiny yellowish to light brown. Male postabdomen (Fig. 6). Epandrium in lateral view 1.0 times wider than high, in posterior view 2.4 times wider than high (before cerci), small setae covering posterior portion of epandrium mixed with a low density of large setae; surstylus 0.4 times as high (at longest point) as epandrium, double-lobed tip, long setae apically; cerci large and pointed apically, 2.0 times length of surstylus in posterior view, with two pair of very long preapical setae. Molecular barcode sequence can be accessed through GenBank Accession: [pending] and BOLD BIN: BOLD: ADP 0751 (only 1 sequence).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFB7FFF0FF3AF8CAC56DA290.taxon	distribution	Distribution (Fig. 26). Southwestern United-States: Utah, New Mexico	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFB7FFF0FF3AF8CAC56DA290.taxon	materials_examined	Type material. Holotype ♂. UNITED STATES: New Mexico: Polio Carpio, Taos, 13. ix. 1965, R. G. Jones (USNM) [USNMENT 01204741]. Paratypes. UNITED STATES: Utah: Cache Co., USU School Forest, 41.77 ° N, 111.79 ° W, 1 ♀, 28. vii. 1977 (NHMLAC), 1 ♀, 1. i- 31. xii. 1977 (NHMLAC), Cache Co., W. Hodges Canyon, 41.92 ° N, 111.4 ° W, 1 ♀, 21. vii- 1. viii. 2003, Hanson, Knowlton (USNM), 2 ♀, 28. vii- 1. viii. 2005, Hanson, Knowlton (USNM), 1 ♀, 11 - 19. vii. 1983, Knowlton, Hanson (USNM), 1 ♀, 5 - 9. v. 1994, Knowlton, Hanson (USNM), 1 ♀, 12 - 16. vii. 1993, Hanson, Knowl- ton (USNM), 1 ♀, 4 - 8. viii. 1978, Knowlton, Hanson (USNM), 1 ♀, 18 - 21. vii. 1983, Hanson, Knowlton (USNM), 1 ♀, 5 - 12. viii. 1971, Knowlton, Hanson (USNM), Summit Co., Bear riv. R. S. 3 mi. SE, 40.92 ° N, 110.83 ° W, 4 ♀, 5 - 12. viii. 1971, Hanson, Knowlton, Malaise trap (USNM), 1 ♀, 6. vi- 1. vii. 1992, Hanson, Knowlton, Malaise trap (USNM), 1 ♀, 5 - 8. viii. 2005, Hanson, Knowlton, Malaise trap (USNM).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFB7FFF0FF3AF8CAC56DA290.taxon	etymology	Etymology. The epithet lutea comes from the Latin luteus meaning yellow, which refers to the color of the body.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFB7FFF0FF3AF8CAC56DA290.taxon	discussion	Remarks. Only two other described species of Paramyia are mostly yellow, Paramyia flava Papp, 2001 from Indonesia and Paramyia latigena Papp, 2001 from Vietnam.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFB4FFE8FF3AFCD6C6A9A534.taxon	description	(Fig. 7 - 8; 27) Description. (Fig. 7) Total length ♂ 1.5 - 1.6 mm, ♀ 1.8 - 2.0 mm. Overall colour brown to dark brown, frontal trian- gle dark brown to black, shiny, reaching lunule; frons 1.38 times longer than wide and 1.44 times wider than flagellum; width at widest point 2.0 - 2.4 times the bottom eye width; gena brown with prunescence, 0.18 - 0.21 times eye height; postgena dark brown, with light prunescence and 0.20 - 0.21 width of head; scape and pedicel dark brown, first flagellomere rectangular, 1.5 times longer than wide, dark brown with pale pubescence, arista dark brown and 1.9 - 2.1 times width of first flagellomere; strong facial carina, easily visible laterally; palpus dark brown with 4 - 5 enlarge setae apically and 1 ventral seta pre-apical, labium dark brown 0.40 - 0.53 mm and labella yellowish 0.48 - 0.53 mm long. Scutum shiny dark brown; scutellum dark brown, microsetose; fore femur brown with apical antero-dorsal part with a strong circle shape patch yellowish to light brown; fore tibia brown on dorsal side with base and apex yellowish and lateral side light brown to yellowish; femur and tibia on mid and hind leg brown; all junction of femur and tibia (“ knee ”) yellowish; all tarsi yellowish; hind basitarsus equal length of tarsi 2 – 4 combined; haltere light brown basally and dark brown apically; wing 1.25 – 1.5 mm long; abdomen shiny dark brown with first tergite often slightly paler. Male postabdomen (Fig. 8). Epandrium in lateral view 1.2 times wider than high, in posterior view 2.6 times wider than high (before cerci), small setae densely covering posterior portion of epandrium mixed with a low density of large setae; surstylus 1.6 times as high as epandrium, paddle shape with a large circular structure situated basally with many short markings within, setae medium length, covering anterior side and distal half of surstylus; cerci large and pointed apically, 0.85 times length of surstylus in posterior view, with a pair of very long preapical setae and other shorter setae, less than 1 / 3 its length, apically swollen in lateral view. Molecular barcode sequence can be accessed through GenBank Accession: KR 755530 and BOLD BIN: BOLD: AAG 0169 (average p-distance within BIN: 0.19 %).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFB4FFE8FF3AFCD6C6A9A534.taxon	distribution	Distribution (Fig. 27). Widespread throughout North America	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFB4FFE8FF3AFCD6C6A9A534.taxon	materials_examined	Type material. Holotype ♂. CANADA: Quebec: Gatineau Co., Masham Township, 27. vii. 1995, E. Ikeda (formerly in LEMQ, now deposited in CNC) [LEM- 0044908]. Paratypes. CANADA: Manitoba: Riding Mountain National Park, 50.6764 ° N, 99.8979 ° W, 663 m, 1 ♂, 2 – 9. vii. 2012, M. Tesar, R. Tesar (BIOUG); New Brunswick: Kouchibouguac National Park, 46.8 ° N, 64.97 ° W, 1 ♂, 30. vii – 2. viii. 2013, O. Lonsdale (CNC), 1 ♂, 30. vii – 8. viii. 2014, O. Lonsdale (CNC); Newfoundland and Labrador: Terra Nova National Park, 48.598 ° N, 53.9702 ° W, 127 m, 2 ♀, 9 – 16. vii. 2013, E. Perry, Malaise trap (BIOUG); Nova Scotia: Cape Breton Highland National Park, Lone Shielding, 46.81 ° N, 60.71 ° W, 1 ♂, 18. vii – 2. viii. 1987, D. Bright, J. Bright, flight interception trap (DEBU), Cape Breton Highland National Park, Mackenzie Mt., 46.82 ° N, 60.82 ° W, 300 m, 1 ♂, 7. vii. 1983, J. R. Vockeroth (CNC), Cape Breton Highlands National Park, 46.6553 ° N, 60.4285 ° W, 21 m, 1 ♂, 15. v – 28. viii. 1994, CBH staff, Malaise trap (BIOUG), Cape Breton Island, South Harbour, 46.87 ° N, 60.47 ° W, 1 ♂, 12. vii. 1983, J. R. Vockeroth, Malaise trap (DEBU), Halifax, Point Pleasant Park, 44.623 ° N, 63.5686 ° W, 30 m, 1 ♂, 27. vii – 3. viii. 2013, T. Zemlak, Malaise trap (BIOUG); Ontario: Algonquin Park, Swan Lake Stn, Scott Lake Survey, 45.4875 ° N, 78.7222 ° W, 1 ♂, 12 – 16. vii. 1993, Larson, Marshall, Barr, Malaise trap (DEBU), Algonquin Provincial Park, Swan Lake Stn, Scott Lake Survey, 45.49 ° N, 78.72 ° W, 1 ♂, 2. viii. 1994, Malaise trap (DEBU), Baptiste Lake, 45.1667 ° N, 78 ° W, 1 ♂, 23. vii. 2000, J. Forrest, sweep (LEMQ), Bruce Co., Bruce Peninsula National Park, High Dump Trail, 45.24 ° N, 81.4 ° W, 1 ♂, 26. vii. 2000, C. S. Onodera (DEBU), Bruce Co., Cameron Lake Rd., 45.2117 ° N, 81.55 ° W, 2 ♂, 5. vii. 1998, K. N. Barber (DEBU), Cambridge, rare Charitable Research Reserve, 43.3736 ° N, 80.3652 ° W, 304 m, 1 ♂, 18 – 24. vi. 2015, BIO Collections, Malaise trap (BIOUG), Hamilton, 43.25 ° N, 79.86 ° W, 1 ♂, 31. vii. 1980, M. Sanborne, Malaise trap (DEBU), Hilton Township, 46.09 ° N, 83.92 ° W, 2 ♂, 13. ix. 1992, J. E. Swann, Malaise trap (ROM), 1 ♂, 26. vii. 1992, J. E. Swann, Malaise trap (ROM), Hilton Township, Tendy Bay, 46.09 ° N, 83.92 ° W, 1 ♂, 27. viii. 1994, J. E. Swann, Malaise trap (ROM), 1 ♂, 22. viii. 1992, J. E. Swann, Malaise trap (ROM), Kaladar, 44.65 ° N, 77.12 ° W, 1 ♂, 9. vii. 1969, G. Steyskal (USNM), Kent Co., Rondeau Provincial Park, 42.32 ° N, 81.85 ° W, 1 ♂, 16. vii. 2003, O. Lonsdale (DEBU), Manitoulin Island, Misery Bay Provincial Nat. Res., 45.7911 ° N, 82.7494 ° W, 1 ♂, 4 – 8. viii. 1978, Marshall et al., Malaise pan traps (DEBU), Thousand Islands National Park, 44.4534 ° N, 75.865 ° W, 1 ♂, 30. vii – 8. viii. 2014, M. Brown, Malaise trap (BIOUG); Quebec: Bois-des-Bel Bog, 47.9683 ° N, 69.4274 ° W, 2 ♂, 30. vi – 6. vii. 2006, A. G. Taillefer, pan trap (LEMQ), Cascapedia, 48.21 ° N, 65.91 ° W, 2 ♂, 2. viii. 1954, J. E. H. Martin (USNM), Forillon National Park, 48.857 ° N, 64.376 ° W, 135 m, 1 ♂, 30. vii – 2. viii. 2013, D. Brodeur, Malaise trap (BIOUG), Gatineau Co., Masham Twp., 45.64 ° N, 76.01 ° W, 1 ♂, 14. vii. 1974 (CNC), Gatineau Park, 45.5667 ° N, 75.95 ° W, 1 ♂, 28. vi. 1995, E. Ikeda (LEMQ), Johnville Bog and Forest Park, 45.345 ° N, 71.7417 ° W, 1 ♂, 22 – 30. vii. 2013, J. Savage, J. Kuchta, Malaise trap (LEMQ), 1 ♀, 16 – 28. vii. 1993, J. Savage, J. Kuchta, Malaise trap (LEMQ), La Mauricie National Park, 46.6507 ° N, 72.9698 ° W, 219 m, 1 ♂, 1 ♀, 24. vii – 1. viii. 2013, M. Toure, Malaise trap (BIOUG), Lac Phillipe, 45.6167 ° N, 76 ° W, 1 ♂, 7. vii. 1968, J. R. Vockeroth (CNC), Mont St. Hilaire, 45.5458 ° N, 73.1604 ° W, 1 ♂, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), Ogden, Marlington Bog, 45.04 ° N, 72.1733 ° W, 1 ♂, 1 ♀, 4 – 11. vii. 2006, A. Moores, Malaise trap (LEMQ), Old Chelsea, Summit King Mt., 45.49 ° N, 75.86 ° W, 351 m, 1 ♂, 25. vi. 1962, J. R. Vockeroth (CNC), Rougemont, 45.4604 ° N, 73.0685 ° W, 1 ♂, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ); Saskatchewan: Prince Albert National Park, Narrows Peninsula Trail, 53.9872 ° N, 106.282 ° W, 530 m, 1 ♂, 11. vii. 2012, BIOBus 2012, sweep (BIOUG). UNITED STATES: Connecticut: Fairfield Co., New Canaan, 41.16 ° N, 73.54 ° W, 1 ♂, 28. vii. 1962, M. Statnam (AMNH); New Hampshire: Coos Co., White Mt. Nat. For., Dolly Copp, 44.3167 ° N, 71.2167 ° W, 1 ♂, 8. viii. 2000, J. Sav- age, sweep (LEMQ), Gorham, 44.39 ° N, 71.17 ° W, 1 ♂, 14. viii. 1958, J. R. Vockeroth (CNC), Mt. Washington, Dolly Copp, 44.33 ° N, 71.22 ° W, 2 ♂, 4. vii. 1936, A. L. Melander (USNM); New Jersey: Morris Co., Pompton Plains, 41 ° N, 74.29 ° W, 3 ♀, 10 – 13. viii. 1987, D. Grimaldi (AMNH); New York: Lake Placid, 44.28 ° N, 73.98 ° W, 610 m, 1 ♂, 19. vii. 1962, J. R. Vockeroth (CNC); North Carolina: Transylvania, Sapphire, 35.11 ° N, 83 ° W, 2 ♂, 20. vii. 1990, W. W. Wirth, UV light trap (USNM); Tennessee: Cades Cave, GSMNP, 35.59 ° N, 83.84 ° W, 1 ♂, 1. vii – 31. viii. 1953, J. M. Carpenter (AMNH); Vermont: East Franklin, Lake Carmi State Park, 44.9533 ° N, 72.8817 ° W, 1 ♂, 4 – 11. vii. 2006, A. Moores, pan trap (LEMQ), Orleans Co. nr Bald Mtn., 44.7939 ° N, 71.988 ° W, 503 m, 1 ♂, 2 – 25. viii. 1999, D. Grimaldi, sweep netted (AMNH); Virginia: Patrick Co., Vesta, 36.72 ° N, 80.36 ° W, 853 m, 1 ♂, 30. v. 1962, J. R. Vock- eroth (CNC), Washington Co., Mt. Rogers N. R. A., Beartree, 36.656 ° N, 81.69 ° W, 1 ♀, 16. viii. 2007, A. MacLeod, sweep (LEMQ); West Virginia: Ritchie Co., North Bend State Park, 39.23 ° N, 81.1 ° W, 1 ♂, 23. vi. 1970, G. Steyskal (USNM); Wisconsin: St. Germain, 45.94 ° N, 89.5 ° W, 1 ♂, 20. vii. 1962, S. Camras (FMNH). Other material examined. CANADA: Manitoba: Riding Mountain National Park, 50.6764 ° N, 99.8979 ° W, 663 m, 1 ♂, 2 – 9. vii. 2012, M. Tesar, R. Tesar (BIOUG); New Brunswick: Fundy National Park, 45.5894 ° N, 64.9556 ° W, 61 m, 1 ♀, 16 – 23. vii. 2014, C. King, Malaise trap (BIOUG), Kouchibouguac National Park, 46.7707 ° N, 65.0064 ° W, 61 m, 1 ♀, 16 – 23. vii. 2013, B. Martin, Malaise trap (BIOUG), 3 ♂, 11 ♀, 30. vii – 2. viii. 2013, O. Lonsdale (CNC), 5 ♂, 2. vii. 1977, J. R. Vockeroth (CNC), 1 ♂, 16 – 23. vii. 2014, B. Martin, Malaise trap (BIOUG), 1 ♀, 30. vii – 8. viii. 2014, O. Lonsdale (CNC), Kouchibouguac National Park, 46.8 ° N, 64.97 ° W, 1 ♀, 16 – 23. vii. 2013, B. Martin, Malaise trap (BIOUG), 3 ♂, 11 ♀, 30. vii – 2. viii. 2013, O. Lonsdale (CNC), 5 ♂, 2. vii. 1977, J. R. Vockeroth (CNC), 1 ♂, 16 – 23. vii. 2014, B. Martin, Malaise trap (BIOUG), 1 ♀, 30. vii – 8. viii. 2014, O. Lonsdale (CNC), Madawaska Co., Saint-Jacques, Baisley Rd, 47.46 ° N, 68.43 ° W, 1 ♂, 27. vii. 2013, O. Lonsdale (CNC); Newfoundland and Labrador: Terra Nova, Newman Sound, 48.5429 ° N, 53.9776 ° W, 41 m, 1 ♂, 11. vii. 2009, BIObus 2009 (BIOUG); Terra Nova National Park, 48.598 ° N, 53.9702 ° W, 127 m, 2 ♀, 9 – 16. vii. 2013, E. Perry, Malaise trap (BIOUG), 1 ♀, 25. vi – 2. vii. 2013, E. Perry, Malaise trap (BIOUG); Nova Scotia: Cape Breton Highland National Park, Lone Shielding, 46.81 ° N, 60.71 ° W, 1 ♀, 18 – 25. vii. 2013, D. Bright, J. Bright, flight interception trap (DEBU), Cape Breton Highland National Park, Mackenzie Mt., 46.82 ° N, 60.82 ° W, 300 m, 1 ♀, 7. vii. 1983, J. R. Vockeroth (CNC), 1 ♀, 11. vii. 1983, J. R. Vockeroth, Malaise trap (CNC), Cape Breton Highland National Park, N. Mtn., 46.83 ° N, 60.6 ° W, 1 ♀, 11. vii. 1983, D. M. Wood, flight trap (CNC), Cape Breton Highland National Park, North Mt., 46.83 ° N, 60.6 ° W, 1 ♂, 19. vii. 1983 (CNC), 1 ♂, 11 – 20. vii. 1999, Masner, intercept and pan trap (DEBU), Cape Breton Highland National Park, North Mt., 46.83 ° N, 60.6 ° W, 400 m, 1 ♂, 19. vii. 1983 (CNC), 1 ♂, 11 – 20. vii. 1999, Masner, intercept and pan trap (DEBU), Cape Breton Highlands National Park, 46.6553 ° N, 60.4285 ° W, 21 m, 1 ♀, 2. ix – 5. x. 1979, CBH staff, Malaise trap (BIOUG), Cape Breton Island, South Harbour, 46.87 ° N, 60.47 ° W, 1 ♂, 12. vii. 1983, J. R. Vockeroth, Malaise trap (DEBU), Halifax, Point Pleasant Park, 44.623 ° N, 63.5686 ° W, 30 m, 1 ♀, 15 – 21. vii. 1986, T. Zemlak, Malaise trap (BIOUG), 1 ♀, 6 – 13. vii. 2013, T. Zemlak, Malaise trap (BIOUG), 1 ♂, 13 – 20. vii. 2013, T. Zemlak, Mal- aise trap (BIOUG), 1 ♀, 29. vii – 6. viii. 1987, T. Zemlak, Malaise trap (BIOUG), 1 ♂, 28. vi – 12. vii. 1986, T. Zemlak, Malaise trap (BIOUG), S. Harbour, 46.87 ° N, 60.47 ° W, 1 ♂, 12. vii. 1983, J. R. Vockeroth (CNC), Shelburne, 43.76 ° N, 65.32 ° W, 1 ♂, 10. viii. 1958, J. R. Vockeroth (CNC); Ontario: 7 mi. E Griffith, 45.24 ° N, 77.17 ° W, 1 ♂, 8. vii. 1989, B. E. Cooper (DEBU), 7 mi. E. Griffith, 45.24 ° N, 77.17 ° W, 2 ♂, 5. vii. 1982, B. E. Cooper (CNC), 1 ♂, 1 ♀, 14. vii. 1990, B. E. Cooper (CNC), 2 ♂, 16. vii. 1984, B. E. Cooper (CNC), Algonquin Park, Swan Lake Stn, Scott Lake Survey, 45.4875 ° N, 78.7222 ° W, 1 ♂, 12 – 15. vii. 2005, Larson, Marshall, Barr, Malaise trap (DEBU), 1 ♀, 16. vii – 2. viii. 1999, Larson, Marshall, Barr, Malaise trap (DEBU), 1 ♂, 28. vi – 16. vii. 1998, Larson, Marshall, Barr, pan trap (DEBU), 2 ♂, 16 – 28. vii. 1993, B. Larson, Malaise trap (DEBU), 1 ♀, 20 – 27. vii. 2013, Larson, Marshall, Barr, Malaise trap (DEBU), 3 ♂, 16. vii. 1993, B. Larson, Malaise trap (DEBU), 3 ♀, 12 – 19. vii. 2006, Larson, Marshall, Barr, Malaise trap (DEBU), 1 ♂, 16 – 21. vii. 2012, Larson, Marshall, Barr, intercept trap (DEBU), 11 ♂, 2 ♀, 15. vii. 1993, B. Larson (DEBU), Al- gonquin Provincial Park, Scott Lake, Outlet, 45.49 ° N, 78.72 ° W, 1 ♂, 6. viii. 1994, S. A. Marshall, sweep (DEBU), Algonquin Provincial Park, Swan Lake Stn, Scott Lake, 45.49 ° N, 78.72 ° W, 1 ♂, 1 ♀, 6. viii. 1994, S. A. Marshall, sweep (DEBU), Algonquin Provincial Park, Swan Lake Stn, Scott Lake Survey, 45.49 ° N, 78.72 ° W, 2 ♂, 1 ♀, 1 – 10. vii. 1994, Malaise trap (DEBU), 7 ♂, 2. viii. 1994, Malaise trap (DEBU), 3 ♂, 1 ♀, 15. viii. 1994, Malaise trap (DEBU), Algonquin Provincial Park, Swan Lake Stn, Scott Lake, Swan Lake margin, 45.49 ° N, 78.72 ° W, 3 ♂, 6. viii. 1994, S. A. Marshall (DEBU), Algonquin Provincial Park, Swan Stn., Scott Lake Survey, 45.49 ° N, 78.72 ° W, 1 ♂, 9. vii. 1995, D. C. Caloren, J. M. Dow, horizontal pan trap (DEBU), Algonquin Provincial Park, Wildlife Research Station, 45.46 ° N, 78.8 ° W, 1 ♀, 7 – 8. vi. 1986, A. B. T. Smith (LEMQ), Bruce Co., Bruce Peninsula National Park, High Dump Trail, 45.24 ° N, 81.4 ° W, 1 ♀, 26. vii. 2000, C. S. Onodera (DEBU), Bruce Co., Cameron Lake Rd., 45.2117 ° N, 81.55 ° W, 8 ♂, 3 ♀, 5. vii. 1998, K. N. Barber (DEBU), Bruce Co., Cameron Lake, Fen, 45.219 ° N, 81.5465 ° W, 3 ♂, 28. vi – 16. vii. 1998, S. A. Marshall, Malaise trap (DEBU), 2 ♂, 28 – 31. vii. 1978, S. A. Marshall, Mal- aise trap (DEBU), Bruce Co., Covels, 45.25 ° N, 81.7667 ° W, 1 ♀, 26 – 28. vii. 2005, S. A. Marshall, Malaise trap (DEBU), Bruce Co., Dunks Bay, 45.2497 ° N, 81.6408 ° W, 1 ♀, 16. viii. 2005, S. A. Marshall (DEBU), Bruce Co., S. Cameron Lake Rd., 45.22 ° N, 81.57 ° W, 1 ♂, 31. vii. 1997, S. A. Marshall (DEBU), Camelot Island, St. Lawrence Is- lands National Park, 44.3 ° N, 76.11 ° W, 1 ♀, 3. viii. 1976, W. Reid, Malaise trap (CNC), Carleton Co., 8 km S. Rich- mond, 45.19 ° N, 75.84 ° W, 1 ♂, 17. vi. 1987, J. M. Cumming (CNC), Dornoch, Anderson Lake, 44.31 ° N, 80.86 ° W, 8 ♀, 12. vii. 1992, S. A. Marshall (DEBU), Dornoch, Anderson Lake Fen, 44.31 ° N, 80.86 ° W, 2 ♂, 4 ♀, 6 – 8. vii. 1974, S. A. Marshall, pan trap (DEBU), 3 ♂, 12. vii. 1992, S. A. Marshall (DEBU), 1 ♂, 12. vii. 1992, S. A. Marshall, pan trap (DEBU), Fathom 5 National Park, Covels Trail to Bass Bay, 45.26 ° N, 81.66 ° W, 1 ♂, 27. vii. 1995, D. C. Caloren (DEBU), Frontenac Co., Skycroft Campground, 5 km W Chaffeys Locks, 44.55 ° N, 76.3667 ° W, 2 ♀, 9 – 15. vii. 2001, B. Hubley, Malaise trap (ROM), 1 ♂, 14 – 19. vii. 1987, B. Hubley, Malaise trap (ROM), Hilton Beach, 46.25 ° N, 83.88 ° W, 1 ♀, 18 – 25. vii. 2013, J. E. Swann, pan trap (ROM), Hilton Township, 46.09 ° N, 83.92 ° W, 1 ♂, 2 ♀, 13. ix. 1992, J. E. Swann, Malaise trap (ROM), 1 ♀, 9. viii. 1992, J. E. Swann, Malaise trap (ROM), 1 ♀, 24. viii. 1992, J. E. Swann, Malaise trap (ROM), Hilton Township, Tendy Bay, 46.09 ° N, 83.92 ° W, 3 ♀, 13. ix. 1992, J. E. Swann, Malaise trap (ROM), 1 ♀, 6. viii. 1992, J. E. Swann, Malaise trap (ROM), 1 ♀, 4. ix. 1992, J. E. Swann, Malaise trap (ROM), 3 ♀, 27. ix. 1992, J. E. Swann, Malaise trap (ROM), 2 ♀, 8. x. 1992, J. E. Swann, Malaise trap (ROM), Icewater Creek WS 12.7 km NNE Searchmont, mi. 10.5 Whitman Dam Rd, 46.79 ° N, 84.03 ° W, 1 ♂, 15 – 19. vii. 2002, Malaise trap (DEBU), 1 ♂, 29. vi – 6. vii. 2013, Malaise trap (DEBU), Killbear Park, 45.36 ° N, 80.21 ° W, 2 ♂, 30. vi. 1979, W. A. Attwater (DEBU), L. Cognashene Muskoka Dist., 44.96 ° N, 79.92 ° W, 3 ♂, 28. vii. 1981, J. T. Huber (CNC), Lambton Co., Pinery Provincial Park, Sand dunes, 44.55 ° N, 76.3667 ° W, 1 ♀, 28. vi – 2. vii. 1993, L. Packer, Malaise trap (ROM), Makonie Lake, 48.35 ° N, 83.24 ° W, 4 ♀, 11. vii. 2016, S. A. Marshall (DEBU), 2 ♂, 11. vii. 2016, S. A. Mar- shall, hand collecting (DEBU), Manitoulin Island, Kip Fleming Tract, 8 km SW Gore Bay, 45.8703 ° N, 82.5419 ° W, 1 ♂, 15 – 22. vi. 2013, Marshall et al., Malaise trap (DEBU), Manitoulin Island, Misery Bay Provincial Nat. Res., 45.7911 ° N, 82.7494 ° W, 1 ♂, 4 – 8. viii. 1978, Paiero, Pivar, white pans (DEBU), Marmora, 44.49 ° N, 77.68 ° W, 3 ♂, 7. vii. 1957, K. D. Southern (DEBU), McDonald Island, St. Lawrence Islands National Park, 44.31 ° N, 76.17 ° W, 1 ♀, 5. viii. 1976, A. Carter, Malaise trap (CNC), McDonald Island, St. Lawrence Islands National Park, 44.31 ° N, 76.17 ° W, 1 ♂, 5. viii. 1976, W. Reid, Malaise trap (CNC), Mt. Picton, 44.04 ° N, 77.06 ° W, 1 ♂, 7. vii. 1970, J. F. McAl- pine (CNC), Muskoka Dist., L. Cognashene, 44.96 ° N, 79.92 ° W, 1 ♂, 28. vii. 1981, J. T. Huber (CNC), Ottawa, 45.43 ° N, 75.68 ° W, 1 ♂, 21. vii. 1989, J. R. Vockeroth (CNC), 1 ♂, 24. vi. 1991, J. R. Vockeroth (CNC), 1 ♀, 10. vii. 1989, J. R. Vockeroth (CNC), Thousand Islands National Park, 44.4534 ° N, 75.865 ° W, 1 ♂, 18 – 24. vi. 2015, M. Brown, Malaise trap (BIOUG), Thwartway Island, St. Lawrence Islands National Park, 44.29 ° N, 76.15 ° W, 1 ♀, 4. viii. 1976, A. Carter, Malaise trap (CNC), 1 ♀, 10. viii. 1976, G. Calderwood, Malaise trap (CNC), 1 ♂, 1 ♀, [no date] (CNC); Quebec: Baie Comeau, Ethier Creek, 49.22 ° N, 68.14 ° W, 1 ♂, 20. viii. 1954, C. C. Steward (CNC), Beechgrove, 45.65 ° N, 76.1333 ° W, 5 ♂, 29. vi. 1962, J. R. Vockeroth (CNC), Bois-des-Bel Bog, 47.9683 ° N, 69.4274 ° W, 27 ♂, 33 ♀, 30. vi – 6. vii. 2006, A. G. Taillefer, pan trap (LEMQ), 1 ♂, 21 – 25. vii. 1978, A. G. Taillefer, pan trap (LEMQ), 1 ♀, 21 – 28. vi. 2008, A. G. Taillefer, pan trap (LEMQ), 1 ♂, 4 ♀, 21 – 30. vi. 2006, A. G. Taillefer, pan trap (LEMQ), 2 ♀, 30. vi – 7. vii. 2008, A. G. Taillefer, pan trap (LEMQ), Cascapedia, 48.21 ° N, 65.91 ° W, 4 ♂, 1 ♀, 2. viii. 1954, J. E. H. Martin (CNC), Chemin-du-Lac Bog, 47.7602 ° N, 69.5265 ° W, 2 ♂, 3 ♀, 21 – 30. vi. 2006, A. G. Taillefer, pan trap (LEMQ), 1 ♀, 19. vii – 5. viii. 1986, A. G. Taillefer, Malaise trap (LEMQ), 1 ♀, 21. vi. 2006, A. G. Taillefer, sweep (LEMQ), Duncan Lake nr. Rupert, 53.48 ° N, 77.95 ° W, 1 ♂, 4. viii. 1970, J. F. McAlpine (CNC), 1 ♂, 6. viii. 1970, J. F. McAlpine (CNC), Forillon National Park, 48.857 ° N, 64.376 ° W, 135 m, 2 ♂, 4 ♀, 15 – 22. vii. 2013, F. Tremblay, Mal- aise trap (BIOUG), 1 ♀, 30. vii – 2. viii. 2013, D. Brodeur, Malaise trap (BIOUG), 1 ♀, 23. v – 8. vi. 1991, F. Tremblay, Malaise trap (BIOUG), 1 ♂, 2 ♀, 30. vii – 5. viii. 2013, D. Brodeur, Malaise trap (BIOUG), 3 ♂, 6 ♀, 22 – 30. vii. 2013, F. Tremblay, Malaise trap (BIOUG), 1 ♂, 1 ♀, 15 – 17. viii. 2005, F. Tremblay, Malaise trap (BIOUG), 1 ♂, 13 – 18. v. 2008, F. Tremblay, Malaise trap (BIOUG), 1 ♂, 5 – 12. viii. 2013, F. Tremblay, Malaise trap (BIOUG), Gaspe, Petit-Cap, 49.04 ° N, 64.45 ° W, 1 ♀, 8. viii. 2001, H. Varady-Szabo, sweep (LEMQ), Gaspe, ZEC York-Baillargeon, shore of Lac Lindsay, 48.8167 ° N, 64.8667 ° W, 1 ♂, 26. vii. 2000, H. Varady-Szabo, sweep (LEMQ), Gaspesie, Mont Albert, camp- ground, 48.9333 ° N, 66.1 ° W, 1 ♂, 31. vii. 2001, S. Boucher, V. Dion, sweep (LEMQ), Gatineau Co., Masham Town- ship, 45.64 ° N, 76.01 ° W, 4 ♂, 27. vii. 1995, E. Ikeda (LEMQ), Gatineau Co., Masham Twp., 45.64 ° N, 76.01 ° W, 1 ♂, 22 – 29. vi. 2008, D. M. Wood (CNC), 1 ♂, 7. ix – 7. x. 2003, D. M. Wood (CNC), 1 ♂, 6 – 8. vii. 1998, D. M. Wood (CNC), 1 ♂, 10 – 13. vi. 1974, D. M. Wood (CNC), 5 ♂, 1 ♀, 14. vii. 1974 (CNC), Gatineau Park, 45.5667 ° N, 75.95 ° W, 8 ♂, 1 ♀, 28. vi. 1995, E. Ikeda (LEMQ), Hull, 45.43 ° N, 75.71 ° W, 1 ♀, 6. viii. 1965, Malaise trap (USNM), James Bay Rte km 204.5, 50.9831 ° N, 77.6339 ° W, 1 ♂, 7 – 16. vii. 2001, M. Buck, B. Buck, Malaise trap (DEBU), Johnville Bog and Forest Park, 45.345 ° N, 71.7417 ° W, 2 ♂, 15 – 22. vii. 2013, J. Savage, J. Kuchta, Malaise trap (LEMQ), 1 ♂, 29. vi – 6. vii. 2013, J. Savage, J. Kuchta, Malaise trap (LEMQ), 1 ♂, 27. v – 3. vi. 1987, J. Savage, J. Kuchta, Malaise trap (LEMQ), 1 ♀, 16 – 21. vii. 1992, J. Savage, J. Kuchta, Malaise trap (LEMQ), 1 ♂, 1 – 5. viii. 2005, J. Savage, J. Kuchta, Malaise trap (LEMQ), King Mt. footpath, Gatineau Hills, 45.49 ° N, 75.86 ° W, 2 ♂, 30. vii. 1959, L. K. Smith (CNC), La Mauricie National Park, 46.6507 ° N, 72.9698 ° W, 219 m, 1 ♀, 11 – 18. vii. 2013, M. Toure, Malaise trap (BIOUG), 2 ♀, 3 – 11. vii. 2013, M. Toure, Malaise trap (BIOUG), 10 ♀, 24. vii – 1. viii. 2013, M. Toure, Malaise trap (BIOUG), Lac Brule, 46.1 ° N, 74.28 ° W, 1 ♂, 21. vii. 1947, O. Peck (CNC), Lac Phillipe, 45.6167 ° N, 76 ° W, 3 ♂, 2 ♀, 7. vii. 1968, J. R. Vockeroth (CNC), 1 ♂, 2 ♀, 4. vii. 1966, D. D. Munroe (CNC), 1 ♂, 5. vii. 1966, D. D. Munroe (CNC), Laniel, 47.05 ° N, 79.28 ° W, 1 ♂, 3. vii. 1944, A. R. Brooks (CNC), 1 ♂, 3. vii. 1941, A. R. Brooks (CNC), Messines, 46.27 ° N, 76.05 ° W, 1 ♂, 10. vii. 1947, W. R. M. Mason (CNC), Mont St. Bruno, 45.5479 ° N, 73.3191 ° W, 1 ♀, 24. vii – 1. viii. 2013, V. Levesque, flight interception trap (LEMQ), 2 ♀, 22 – 29. vi. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♀, 22 – 29. vi. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♀, 15 – 22. vi. 2013, V. Levesque, trunk trap (LEMQ), Mont St. Bruno, 45.548 ° N, 73.3104 ° W, 1 ♀, 24. vii – 1. viii. 2013, V. Levesque, flight interception trap (LEMQ), 2 ♀, 22 – 29. vi. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♀, 22 – 29. vi. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♀, 15 – 22. vi. 2013, V. Levesque, trunk trap (LEMQ), Mont St. Bruno, 45.548 ° N, 73.3103 ° W, 1 ♀, 24. vii – 1. viii. 2013, V. Levesque, flight interception trap (LEMQ), 2 ♀, 22 – 29. vi. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♀, 22 – 29. vi. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♀, 15 – 22. vi. 2013, V. Levesque, trunk trap (LEMQ), Mont St. Bruno, 45.5521 ° N, 73.3339 ° W, 1 ♀, 24. vii – 1. viii. 2013, V. Levesque, flight interception trap (LEMQ), 2 ♀, 22 – 29. vi. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♀, 22 – 29. vi. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♀, 15 – 22. vi. 2013, V. Levesque, trunk trap (LEMQ), Mont St. Hilaire, 45.5436 ° N, 73.1448 ° W, 11 ♂, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 15 ♂, 11 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), 4 ♂, 30. vi – 7. vii. 2008, V. Levesque, trunk trap (LEMQ), 1 ♂, 1 ♀, 22 – 23. ix. 1996, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 6. viii. 2014, V. Levesque, flight interception trap (LEMQ), 2 ♀, 30. vii – 5. viii. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 13. viii. 1986, V. Levesque, flight interception trap (LEMQ), 1 ♀, 1 – 8. vii. 2008, V. Levesque, flight inter- ception trap (LEMQ), Mont St. Hilaire, 45.5457 ° N, 73.1608 ° W, 11 ♂, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight inter- ception trap (LEMQ), 15 ♂, 11 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), 4 ♂, 30. vi – 7. vii. 2008, V. Levesque, trunk trap (LEMQ), 1 ♂, 1 ♀, 22 – 23. ix. 1996, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 6. viii. 2014, V. Levesque, flight interception trap (LEMQ), 2 ♀, 30. vii – 5. viii. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 13. viii. 1986, V. Levesque, flight interception trap (LEMQ), 1 ♀, 1 – 8. vii. 2008, V. Levesque, flight interception trap (LEMQ), Mont St. Hilaire, 45.5458 ° N, 73.161 ° W, 11 ♂, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 15 ♂, 11 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), 4 ♂, 30. vi – 7. vii. 2008, V. Levesque, trunk trap (LEMQ), 1 ♂, 1 ♀, 22 – 23. ix. 1996, V. Levesque, flight inter- ception trap (LEMQ), 1 ♂, 23. vii – 6. viii. 2014, V. Levesque, flight interception trap (LEMQ), 2 ♀, 30. vii – 5. viii. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 13. viii. 1986, V. Levesque, flight interception trap (LEMQ), 1 ♀, 1 – 8. vii. 2008, V. Levesque, flight interception trap (LEMQ), Mont St. Hilaire, 45.5458 ° N, 73.1604 ° W, 11 ♂, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 15 ♂, 11 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight in- terception trap (LEMQ), 4 ♂, 30. vi – 7. vii. 2008, V. Levesque, trunk trap (LEMQ), 1 ♂, 1 ♀, 22 – 23. ix. 1996, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 6. viii. 2014, V. Levesque, flight interception trap (LEMQ), 2 ♀, 30. vii – 5. viii. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 13. viii. 1986, V. Levesque, flight interception trap (LEMQ), 1 ♀, 1 – 8. vii. 2008, V. Levesque, flight interception trap (LEMQ), Mont St. Hilaire, 45.5459 ° N, 73.1605 ° W, 11 ♂, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 15 ♂, 11 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), 4 ♂, 30. vi – 7. vii. 2008, V. Levesque, trunk trap (LEMQ), 1 ♂, 1 ♀, 22 – 23. ix. 1996, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 6. viii. 2014, V. Levesque, flight interception trap (LEMQ), 2 ♀, 30. vii – 5. viii. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 13. viii. 1986, V. Levesque, flight interception trap (LEMQ), 1 ♀, 1 – 8. vii. 2008, V. Levesque, flight interception trap (LEMQ), Mont St. Hilaire, 45.546 ° N, 73.161 ° W, 11 ♂, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 15 ♂, 11 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), 4 ♂, 30. vi – 7. vii. 2008, V. Levesque, trunk trap (LEMQ), 1 ♂, 1 ♀, 22 – 23. ix. 1996, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 6. viii. 2014, V. Levesque, flight interception trap (LEMQ), 2 ♀, 30. vii – 5. viii. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 13. viii. 1986, V. Levesque, flight interception trap (LEMQ), 1 ♀, 1 – 8. vii. 2008, V. Levesque, flight interception trap (LEMQ), Mont St. Hilaire, 45.551 ° N, 73.1537 ° W, 11 ♂, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 15 ♂, 11 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), 4 ♂, 30. vi – 7. vii. 2008, V. Levesque, trunk trap (LEMQ), 1 ♂, 1 ♀, 22 – 23. ix. 1996, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 6. viii. 2014, V. Levesque, flight interception trap (LEMQ), 2 ♀, 30. vii – 5. viii. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 13. viii. 1986, V. Levesque, flight interception trap (LEMQ), 1 ♀, 1 – 8. vii. 2008, V. Levesque, flight interception trap (LEMQ), Mont St. Hilaire, 45.551 ° N, 73.1532 ° W, 11 ♂, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 15 ♂, 11 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight intercep- tion trap (LEMQ), 4 ♂, 30. vi – 7. vii. 2008, V. Levesque, trunk trap (LEMQ), 1 ♂, 1 ♀, 22 – 23. ix. 1996, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 6. viii. 2014, V. Levesque, flight interception trap (LEMQ), 2 ♀, 30. vii – 5. viii. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 13. viii. 1986, V. Levesque, flight interception trap (LEMQ), 1 ♀, 1 – 8. vii. 2008, V. Levesque, flight interception trap (LEMQ), Mont St. Hilaire, 45.5513 ° N, 73.1537 ° W, 11 ♂, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 15 ♂, 11 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), 4 ♂, 30. vi – 7. vii. 2008, V. Levesque, trunk trap (LEMQ), 1 ♂, 1 ♀, 22 – 23. ix. 1996, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 6. viii. 2014, V. Levesque, flight interception trap (LEMQ), 2 ♀, 30. vii – 5. viii. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 13. viii. 1986, V. Levesque, flight interception trap (LEMQ), 1 ♀, 1 – 8. vii. 2008, V. Levesque, flight interception trap (LEMQ), Mont St. Hilaire, 45.5514 ° N, 73.1535 ° W, 11 ♂, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 15 ♂, 11 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), 4 ♂, 30. vi – 7. vii. 2008, V. Levesque, trunk trap (LEMQ), 1 ♂, 1 ♀, 22 – 23. ix. 1996, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 6. viii. 2014, V. Levesque, flight interception trap (LEMQ), 2 ♀, 30. vii – 5. viii. 2013, V. Levesque, flight interception trap (LEMQ), 1 ♂, 23. vii – 13. viii. 1986, V. Levesque, flight interception trap (LEMQ), 1 ♀, 1 – 8. vii. 2008, V. Levesque, flight interception trap (LEMQ), Mont St. Hilaire Reserve, Pain de Sucre Trail, 45.55 ° N, 73.17 ° W, 1 ♂, 27. vi. 2001, S. E. Brooks, sweep (LEMQ), Notre-Dame-du-Lac, halte du Lac Temiscouata, 47.5833 ° N, 68.7333 ° W, 1 ♂, 27. vii. 2001, M. Giroux, sweep (LEMQ), Ogden, Marlington Bog, 45.04 ° N, 72.1733 ° W, 3 ♂, 12 ♀, 4 – 11. vii. 2006, A. Moores, Malaise trap (LEMQ), 15 ♂, 6 ♀, 4 – 11. vii. 2006, A. Moores, pan trap (LEMQ), 1 ♀, 27. vii – 5. viii. 2010, A. Moores, pan trap (LEMQ), 26 ♂, 11 ♀, 27. vi – 4. vii. 2006, A. Moores, pan trap (LEMQ), 1 ♀, 20 – 27. vii. 2013, A. Moores, Malaise trap (LEMQ), 1 ♀, 27. vii – 3. viii. 2013, A. Moores, pan trap (LEMQ), Old Chelsea, 45.5 ° N, 75.81 ° W, 2 ♂, 9. vii. 1959, J. R. Vockeroth (CNC), 1 ♂, 30. viii. 1985, J. R. Vockeroth (CNC), 1 ♂, 18. vii. 1961, J. R. Vockeroth (CNC), 1 ♂, 25. vi. 1959, J. G. Chillcott (CNC), Old Chelsea, King Mt., 45.49 ° N, 75.86 ° W, 2 ♂, 13. viii. 1969, B. V. Peterson (CNC), Old Chel- sea, Summit King Mt., 45.49 ° N, 75.86 ° W, 351 m, 1 ♂, 24. vi. 1964, J. R. Vockeroth (CNC), 1 ♂, 1 ♀, 9. viii. 1961, J. R. Vockeroth (CNC), 3 ♂, 25. vi. 1962, J. R. Vockeroth (CNC), 1 ♂, 18. vii. 1961, J. R. Vockeroth (CNC), Reserve ecologique des Tourbieres de Lanoraie, 45.9967 ° N, 73.3 ° W, 1 ♀, 3. vii. 2006, A. Moores, sweep (LEMQ), Rouge- mont, 45.4559 ° N, 73.0613 ° W, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♀, 21. vii – 1. viii. 2003, V. Levesque, flight interception trap (LEMQ), 1 ♂, 21 – 28. vii. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♂, 1 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), Rougemont, 45.4602 ° N, 73.0689 ° W, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♀, 21. vii – 1. viii. 2003, V. Levesque, flight interception trap (LEMQ), 1 ♂, 21 – 28. vii. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♂, 1 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), Rougemont, 45.4604 ° N, 73.0685 ° W, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♀, 21. vii – 1. viii. 2003, V. Levesque, flight interception trap (LEMQ), 1 ♂, 21 – 28. vii. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♂, 1 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), Rougemont, 45.4606 ° N, 73.0688 ° W, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight intercep- tion trap (LEMQ), 1 ♀, 21. vii – 1. viii. 2003, V. Levesque, flight interception trap (LEMQ), 1 ♂, 21 – 28. vii. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♂, 1 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), Rougemont, 45.4934 ° N, 73.0647 ° W, 2 ♀, 23 – 30. vi. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♀, 21. vii – 1. viii. 2003, V. Levesque, flight interception trap (LEMQ), 1 ♂, 21 – 28. vii. 2008, V. Levesque, flight interception trap (LEMQ), 1 ♂, 1 ♀, 30. vi – 7. vii. 2008, V. Levesque, flight interception trap (LEMQ), St-Charles Bog, 46.7478 ° N, 70.995 ° W, 1 ♀, 14 – 19. vii. 1987, A. G. Taillefer, pan trap (LEMQ), Ste-Barbe, Large Teafield, 45.125 ° N, 74.225 ° W, 1 ♂, 30. vi – 6. vii. 2006, A. Moores, Malaise trap (LEMQ); Saskatchewan: Prince Albert National Park, Narrows Peninsula Trail, 53.9872 ° N, 106.282 ° W, 530 m, 1 ♂, 9 – 14. vii. 2012, BIOBus 2012, Malaise trap (BIOUG), 3 ♀, 09. vii. 2012, BIOBus 2012, sweep (BIOUG), Prince Albert National Park, Spruce River Highlands Trail, 53.7125 ° N, 106.048 ° W, 541 m, 1 ♂, 10 – 14. vii. 2012, BIOBus 2012, Malaise trap (BIOUG). UNITED STATES: Connecticut: Redding, 41.3 ° N, 73.39 ° W, 1 ♂, 27. vii. 1933, A. L. Melander (USNM), Terryville, 41.68 ° N, 73.01 ° W, 1 ♂, 22. vi. 1957, R. I. Sailer (USNM); District of Columbia: [no locality given]: 1 ♂, [no date], Coquillett (USNM), Kenilworth Park, 38.9167 ° N, 76.95 ° W, 1 ♀, 20. ix. 1996, G. F. Hevel (USNM); Florida: Highlands Co., Archbold Biol. Sta, 27.18 ° N, 81.35 ° W, 1 ♂, 26. iv. 1967, B. V. Peterson (CNC), Jackson Co., Florida Caverns State Park, 30.82 ° N, 85.23 ° W, 1 ♂, 26. v. 1973, W. W. Wirth, Malaise trap (USNM), Sebring, Highlands Hamm State Park, 27.47 ° N, 81.53 ° W, 3 ♀, 15. iv. 1970, W. W. Wirth, Malaise trap (USNM); Georgia: Forsyth, 33.03 ° N, 83.94 ° W, 2 ♂, 2. vi. 1970 (CNC); Indiana: La Fayette, 40.42 ° N, 86.87 ° W, 1 ♀, 9. ix. 1916, J. M. Aldrich (USNM); Maine: Oldtown, 44.93 ° N, 68.64 ° W, 1 ♀, 3. viii. 1966, T. J. Spilman (USNM); Maryland: Glen Echo, 38.97 ° N, 77.14 ° W, 1 ♂, 9. vii. 1922, J. R. Malloch (USNM), Montgomery Co., Bethesda, 38.99 ° N, 77.1 ° W, 1 ♂, 26. vii. 1970, G. Steyskal (USNM), 1 ♂, 3. vii. 1970, G. Steyskal (USNM), Worcester Co., Snow Hill, 38.18 ° N, 75.39 ° W, 1 ♀, 16. vii. 1968, W. W. Wirth (USNM); Massachusetts: Waquoit, 41.58 ° N, 70.53 ° W, 1 ♂, [no date], A. H. Sturtevant (USNM), Woods Hole, 41.53 ° N, 70.67 ° W, 1 ♀, 23. vii. 1948, A. H. Sturtevant (USNM); Michigan: Cheboygan Co., 45.65 ° N, 84.47 ° W, 1 ♂, 17. vii. 1942, C. W. Sabrosky (USNM), Isle Royale, 48 ° N, 88.93 ° W, 1 ♂, 30. vi – 6. vii. 2006, C. Sabrosky (USNM), Roscommon Co., 44.5 ° N, 84.59 ° W, 2 ♂, 22. vii. 1951, R. R. Dreisbach (USNM); Minnesota: Eaglesnest, 47.84 ° N, 92.1 ° W, 1 ♀, 12. viii. 1958, W. V. Balduf (USNM), Ely, 47.9 ° N, 91.87 ° W, 1 ♀, 14. viii. 1942, W. V. Balduf (USNM); New Hampshire: Benton, 44.1 ° N, 71.9 ° W, 1 ♂, 2 ♀, 6. vii. 1931, A. L. Melander (USNM), Carroll, Co., White Moun- tain Nat. For., 3 km E Sabbaday Falls, 43.9833 ° N, 71.3833 ° W, 1 ♀, 10. viii. 2000, T. A. Wheeler, sweep (LEMQ), Cheshire Co., Alstead, 43.15 ° N, 72.36 ° W, 1 ♀, 25 – 26. vii. 1976, D. E. Boufford (USNM), 1 ♂, 26. vi – 3. vii. 2006, D. E. Boufford (USNM), 1 ♂, 25. vii – 1. viii. 2013, D. E. Boufford (USNM), Coos Co., 1 km E Stark, 44.6 ° N, 71.4 ° W, 2 ♂, 1 ♀, 8. viii. 2000, J. Savage, sweep (LEMQ), Coos Co., White Mt. Nat. For., Dolly Copp, 44.3167 ° N, 71.2167 ° W, 11 ♂, 4 ♀, 8. viii. 2000, J. Savage, sweep (LEMQ), Fabyan, 44.27 ° N, 71.46 ° W, 1 ♂, 30. vi. 1936, A. L. Melander (USNM), Franconia, 44.23 ° N, 71.75 ° W, 1 ♂, 6. vii. 1936, A. L. Melander (USNM), Gorham, 44.39 ° N, 71.17 ° W, 1 ♂, 16. vii. 1958, J. R. Vockeroth (CNC), Grafton Co., White Mountain N. F., Kancamagus Pass, 44.0167 ° N, 71.4833 ° W, 1 ♀, 10. viii. 2000, T. A. Wheeler (LEMQ), Jefferson Notch, 44.3 ° N, 71.35 ° W, 2 ♂, 30. vii. 1961, W. W. Wirth (USNM), Mt. Washington, Dolly Copp, 44.33 ° N, 71.22 ° W, 3 ♂, 2 ♀, 4. vii. 1936, A. L. Melander (USNM), White Mts., Dolly Copp, 44.33 ° N, 71.22 ° W, 1 ♂, 11. vii. 1931, A. L. Melander (USNM), White Mts., Stinson Lake, 43.86 ° N, 71.81 ° W, 5 ♂, 3 ♀, 23. vii. 1961, W. W. Wirth (USNM), White Mts., The Flume, 44.11 ° N, 71.64 ° W, 1 ♀, 7. vii. 1936, A. L. Me- lander (USNM); New Jersey: Morris Co., Pompton Plains, 41 ° N, 74.29 ° W, 1 ♀, 11 – 14. vii. 1978, D. Grimaldi (AMNH); New York: Adirondack Park, Blue Mountain, 43.875 ° N, 74.43 ° W, 1 ♀, 31. vii. 2002, J. Forrest, T. A. Wheeler, sweep (LEMQ), Adirondacks, Avalanche Trail, 44.14 ° N, 73.94 ° W, 3 ♂, 30. vii. 1929, A. L. Melander (USNM), Allegany St. Pk., 42.06 ° N, 78.76 ° W, 1 ♀, 13. viii. 1939 (AMNH), Franklin Co., Paul Smiths, 44.44 ° N, 74.25 ° W, 3 ♂, 20. vii. 1962, J. R. Vockeroth (CNC), Kalbfleisch R. S., Hunthington, L. I., 40.86 ° N, 73.35 ° W, 1 ♀, 16. viii. 1962, P. H. Arnaud, Malaise trap (AMNH), Keene Valley, 44.19 ° N, 73.79 ° W, 7 ♂, 20. vii. 1962, J. G. Chillcott (CNC), Lake Placid, 44.28 ° N, 73.98 ° W, 20 ♂, 3 ♀, 19. vii. 1962, J. G. Chillcott (CNC), 2 ♂, 1. vii. 1922, J. M. Aldrich (USNM), 5 ♂, 3 ♀, 2. vii. 1922, J. M. Aldrich (USNM), 1 ♂, 20. vii. 1962, J. G. Chillcott (CNC), 1 ♂, 26. vii. 1929, A. L. Melander (USNM), Lake Placid, 44.28 ° N, 73.98 ° W, 610 m, 20 ♂, 3 ♀, 19. vii. 1962, J. G. Chillcott (CNC), 2 ♂, 1. vii. 1922, J. M. Aldrich (USNM), 5 ♂, 3 ♀, 2. vii. 1922, J. M. Aldrich (USNM), 1 ♂, 20. vii. 1962, J. G. Chillcott (CNC), 1 ♂, 26. vii. 1929, A. L. Melander (USNM), nr Mt. Marcy, 44.11 ° N, 73.92 ° W, 1 ♂, 19. viii. 1972, L. Knutson (USNM), Ulster Co., Cherrytown, 4 mi. NNW Kerhonkson, 41.83 ° N, 74.33 ° W, 1 ♀, 9 – 13. vi. 2006, P. Wygodzinsky, B. Wy- godzinsky (AMNH); North Carolina: 9 mi. N. Linville, Hwy 221, 36.1 ° N, 81.83 ° W, 1 ♂, 13. vi. 1962, J. G. Chillcott (CNC), Alleghany Co., Cherry Lane, 36.44 ° N, 81.02 ° W, 884 m, 3 ♂, 6. vi. 1962 (CNC), Blue Ridge Pkwy., Mt. Rich- land-Balsam, 35.37 ° N, 82.99 ° W, 1951 m, 1 ♀, 30. v. 1965, J. G. Chillcott (CNC), Haywood Co., Pisgah National For- est, Chestnut Bald, 35.41 ° N, 82.63 ° W, 1798 m, 1 ♂, 2. viii. 1957, J. G. Chillcott (CNC), Highlands, 35.05 ° N, 83.2 ° W, 1158 m, 1 ♂, 26. vi. 1957, J. R. Vockeroth (CNC), 1 ♂, 29. vi. 1957, J. R. Vockeroth (CNC), 1 ♂, 24. vi. 1957, J. R. Vock- eroth (CNC), 11 ♂, 1 ♀, 25. vi. 1957, J. R. Vockeroth (CNC), Highlands, Blue Valley Lookout, 35.01 ° N, 83.22 ° W, 1067 m, 1 ♂, 29. vi. 1957, J. R. Vockeroth (CNC), Jackson Co., Highlands, 6 mi. E., 35.05 ° N, 83.2 ° W, 2 ♂, 5. vi. 1962, J. G. Chillcott (CNC), Lake Toxaway, 35.13 ° N, 82.93 ° W, 1 ♂, 12. vii. 1957, J. G. Chillcott (USNM), Macon Co., Highlands Wightman Cottage, 35.05 ° N, 83.19 ° W, 1 ♂, 1 ♀, 10. vii. 1987, W. W. Wirth, UV light trap (USNM), Macon Co., Wayah Gap, 35.15 ° N, 83.58 ° W, 945 m, 1 ♂, 31. v. 1965, J. G. Chillcott (CNC), Mitchell Co., Penland, 35.93 ° N, 82.11 ° W, 914 m, 1 ♂, 1 ♀, 20. vi. 1957, G. Steyskal (USNM), Transylvania, Sapphire, 35.11 ° N, 83 ° W, 6 ♂, 1 ♀, 20. vii. 1990, W. W. Wirth, UV light trap (USNM), Wayah Gap, 35.15 ° N, 83.58 ° W, 1372 m, 2 ♂, 5. vi. 1962, J. G. Chill- cott (CNC); Pennsylvania: Lycoming Co., Ralston, 41.51 ° N, 76.95 ° W, 1 ♂, 8. vi. 1962, J. G. Chillcott (CNC); South Dakota: Flynn Creek, 8 mi. N. Pringle, 43.66 ° N, 103.41 ° W, 1646 m, 1 ♂, 8 – 28. vii. 1980, H. Howden, A. Howden (CNC), 1 ♂, 8 – 15. vii. 2001, H. Howden, A. Howden (CNC); Vermont: East Franklin, Lake Carmi State Park, 44.9533 ° N, 72.8817 ° W, 1 ♂, 1 ♀, 11 – 18. vii. 2006, A. Moores, Malaise trap (LEMQ), 2 ♂, 11 – 18. vii. 2006, A. Moores, pan trap (LEMQ), 1 ♂, 4 – 11. vii. 2010, A. Moores, pan trap (LEMQ); Virginia: Alexandria, 38.81 ° N, 77.05 ° W, 1 ♂, 14. vi. 1951, W. W. Wirth (USNM), Bath Co., 9.6 km N Clifton Forge, 37.82 ° N, 79.82 ° W, 1 ♂, 13 – 20. vi. 1998, S. Peck, J. Peck, flight interception trap (DEBU), Brush Mt., Black shurg, 37.29 ° N, 80.44 ° W, 853 m, 1 ♂, 27. v. 1962, J. G. Chillcott (CNC), Falls Church, Holmes Run, 38.85 ° N, 77.19 ° W, 1 ♂, 22. vi. 1961, W. W. Wirth, light trap (USNM), 2 ♂, 6. vii. 1961, W. W. Wirth, light trap (USNM), Patrick Co., Vesta, 36.72 ° N, 80.36 ° W, 853 m, 10 ♂, 2 ♀, 30. v. 1962, J. R. Vockeroth (CNC), Rockbridge Co., Steeles Tavern, 37.93 ° N, 79.2 ° W, 701 m, 1 ♂, 1 ♀, 7. vi. 1962, J. R. Vockeroth (CNC); West Virginia: Franklin, 38.64 ° N, 79.33 ° W, 1 ♂, 26. vi. 1951, W. W. Wirth (USNM), Morgantown, 39.63 ° N, 79.95 ° W, 1 ♂, 27. vi. 1970, G. Steyskal (USNM), Pocahontas Co., Cranberry Glades, 38.2 ° N, 80.27 ° W, 1 ♀, 16. vii. 1955, C. W. Sabrosky (USNM), Ritchie Co., North Bend State Park, 39.23 ° N, 81.1 ° W, 1 ♂, 4 ♀, 23. vi. 1970, G. Steyskal (USNM); Wisconsin: St. Germain, 45.94 ° N, 89.5 ° W, 1 ♂, 2 ♀, 20. vii. 1962, S. Camras (FMNH).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFB4FFE8FF3AFCD6C6A9A534.taxon	etymology	Etymology. The species name reflects the historical misidentification of this species as Paramyia nitens.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFB4FFE8FF3AFCD6C6A9A534.taxon	discussion	Remarks. Paramyia pseudonitens is the most commonly encountered and most abundant species in North America.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFACFFECFF3AFA0DC7BBA5A0.taxon	description	(Fig. 9 – 10; 28) Description. (Fig. 9) Total length ♂ 1.4 – 1.7 mm, ♀ 1.6 – 2.3 mm. Overall colour brown to dark brown, frontal tri- angle dark brown, shiny, reaching lunule; frons 1.0 – 1.13 times longer than wide and 1.9 – 2.0 times wider than flagellum; width at widest point 2.0 – 2.1 times the bottom eye width; gena brown with prunescence, 0.12 – 0.16 times eye height; postgena dark brown, with light prunescence and 0.24 – 0.28 width of head; scape and pedicel dark brown, first flagellomere subquadrate, same length than width, dark brown with pale pubescence, arista dark brown and 2.0 – 2.2 times width of first flagellomere; medium facial carina, easily visible laterally; palpus dark brown with 4 – 5 enlarge setae apically and 1 ventral seta pre-apical, labium brown to dark brown 0.35 – 40 mm and labella brown 0.45 – 0.48 mm long. Scutum shiny brown to dark brown; scutellum dark brown, microsetose; fore femur brown with apical anterodorsal part with a strong circle shape patch yellowish to light brown; fore tibia brown on dorsal and posterior side with base and apex yellowish and anterior side light brown to yellowish; femur and tibia on mid and hind leg brown; all junction of femur and tibia (“ knee ”) yellowish; all tarsi yellowish; hind basitarsus shorter than length of tarsi 2 – 4 combined; haltere uniformly brown; wing 1.42 – 1.50 mm long; abdomen shiny dark brown with first tergite often slightly paler. Male postabdomen (Fig. 10). Epandrium in lateral view 1.06 times higher than wide, in posterior view 2.3 times wider than high (before cerci), small setae densely covering posterior portion of epandrium mixed with a low density of large setae; surstylus 1.2 times as high as epandrium, paddle shape with a large circular structure situated basally with lots of short markings within and a distinct dent on the anterior side of the apical portion, setae medium length, covering anterior side and distal half of surstylus; cerci small and slightly blunt apically, 0.6 times length of surstylus in posterior view, with a pair of very long preapical setae and other shorter setae, less than 1 / 3 its length, apically swollen in lateral view. Molecular barcode sequence can be accessed through GenBank Accession: KR 391632 and BOLD BIN: BOLD: AAG 0176 (average p-distance within BIN: 0.09 %). Distribution (Fig. 28). Widespread throughout North America Type material. Holotype ♂. CANADA: Quebec: St-Charles Bog, 22. vi. 2006, A. G. Taillefer, sweep, (formerly in LEMQ, now deposited in CNC) [LEM- 0043800]. Paratypes. CANADA: Nova Scotia: Kejimkujik, Grafton Lake, 44.383 ° N, 65.2023 ° W, 103 m, 1 ♀, 03. viii. 2009, BIObus 2009 (BIOUG); Ontario: Frontenac Co., Skycroft Campground, 5 km W Chaffeys Locks, 44.55 ° N, 76.3667 ° W, 2 ♂, 14 – 19. vii. 1987, B. Hubley, Malaise trap (ROM), Guelph, U of G Arboretum, 43.54 ° N, 80.21 ° W, 1 ♂, 22 – 29. vi. 2008, B. V. Brown, unbaited pitfalls (DEBU), Hilton Township, Tendy Bay, 46.09 ° N, 83.92 ° W, 1 ♂, 27. viii. 1994, J. E. Swann, Malaise trap (ROM), Mer Bleu, 5 mi. E. Ottawa, 45.39 ° N, 75.51 ° W, 1 ♂, 6. viii. 1966, D. D. Munroe, Malaise trap (CNC), 1 ♂, 11. vii. 1966, D. D. Munroe, Malaise trap (CNC), 1 ♂, 25. vi. 1964 (CNC), 1 ♂, 29. vii. 1966, D. D. Munroe, Malaise trap (CNC), 2 ♂, 17. vii. 1966, D. D. Munroe, Malaise trap (CNC), 1 ♂, 25. vii. 1966, D. D. Munroe, Malaise trap (CNC), Oliver bog, 3 km S. Galt, 43.34 ° N, 80.3 ° W, 1 ♂, 27. vi – 4. vii. 2006, D. Blades (DEBU), 1 ♂, 3 – 10. vii. 2006, D. Blades (DEBU), Ottawa, 45.43 ° N, 75.68 ° W, 1 ♂, 30. viii. 1993, J. R. Vockeroth (CNC), 1 ♂, 8. viii. 1993, J. R. Vockeroth (CNC); Quebec: Johnville Bog and Forest Park, 45.345 ° N, 71.7417 ° W, 2 ♂, 15 – 22. vii. 2005, J. Savage, J. Kuchta, Malaise trap (LEMQ), Reserve ecologique des Tourbieres de Lanoraie, 45.9967 ° N, 73.3 ° W, 3 ♂, 10 – 17. vii. 2006, A. Moores, Malaise trap (LEMQ), St-Charles Bog, 46.7478 ° N, 70.995 ° W, 2 ♂, 14. vi. 2006, A. G. Taillefer, sweep (LEMQ), 2 ♂, 22. vi. 2006, A. G. Taillefer, sweep (LEMQ), Ste-Barbe, Large Teafield, 45.125 ° N, 74.225 ° W, 2 ♂, 17 – 24. vii. 2006, A. Moores, Malaise trap (LEMQ). UNITED STATES: Florida: Alachua Co., Chantilly Acres, 29.71 ° N, 82.42 ° W, 1 ♂, 25. iv. 1970, W. W. Wirth, Malaise trap (USNM), Gaines- ville, 29.65 ° N, 82.32 ° W, 1 ♂, 21 – 30. vi. 2006, W. Mason, Malaise trap (DEBU), Highlands Co., Archbold Biol. Sta, 27.18 ° N, 81.35 ° W, 1 ♂, 13 – 20. vii. 2012, S. M. Paiero, Malaise trap (DEBU), Ocean Pond, Olustee, 30.21 ° N, 82.45 ° W, 1 ♂, 29. vi. 1953, M. R. Wheeler (AMNH), Orange Co., Rock Springs, 28.76 ° N, 81.5 ° W, 1 ♂, 21. iv. 1970, W. W. Wirth (USNM), Sebring, Highlands Hamm State Park, 27.47 ° N, 81.53 ° W, 2 ♂, 15. iv. 1970, W. W. Wirth, Mal- aise trap (USNM); Illinois: Macomb, 40.46 ° N, 90.68 ° W, 1 ♂, 29. v. 1962, W. W. Wirth (USNM); Indiana: La Fayette, 40.42 ° N, 86.87 ° W, 1 ♂, 4. ix. 1916, J. M. Aldrich (USNM); Maryland: Montgomery Co., Bethesda, 38.99 ° N, 77.1 ° W, 1 ♂, 20. vii. 1968, G. Steyskal (USNM); Michigan: Hunt Cr. Exp. Sta. nr. Lewiston, 44.88 ° N, 84.31 ° W, 1 ♂, 20. vii. 1942, C. W. Sabrosky (USNM); North Carolina: Bladen Co., Singletary Lake State Park, 34.5833 ° N, 78.4583 ° W, 1 ♂, 19 – 21. vi. 2006, Marshall, Paiero, pan trap (DEBU), Highlands, 35.05 ° N, 83.2 ° W, 1158 m, 1 ♂, 29. vi. 1957, J. R. Vockeroth (CNC); Tennessee: Cades Cave, GSMNP, 35.59 ° N, 83.84 ° W, 1 ♂, 1. vii – 31. viii. 1953, J. M. Carpenter (AMNH); Texas: Woodville, 30.76 ° N, 94.43 ° W, 1 ♂, 1 – 30. ix. 1954, M. R. Wheeler (AMNH); Vermont: East Franklin, Lake Carmi State Park, 44.9533 ° N, 72.8817 ° W, 1 ♂, 11 – 18. vii. 2006, A. Moores, Malaise trap (LEMQ); Virginia: Cape Henry, 36.93 ° N, 76.02 ° W, 2 ♂, 28. vi. 1939, A. L. Melander (USNM). Other material examined. CANADA: British Columbia: Lakelse L. bog nr. Terrace, 54.37 ° N, 128.53 ° W, 1 ♂, 14. vi. 1960, C. H. Mann (CNC), Skagit Val., 8 mi. W Hope, 49.38 ° N, 121.44 ° W, 1 ♂, 8 – 15. vii. 2001, intercept trap (DEBU); Ontario: Mer Bleu, 5 mi. E. Ottawa, 45.39 ° N, 75.51 ° W, 1 ♂, 6. viii. 1966, D. D. Munroe, Malaise trap (CNC), 2 ♂, 19. vii. 1966, D. D. Munroe, Malaise trap (CNC), 1 ♂, 25. vii. 1966, D. D. Munroe, Malaise trap (CNC), 17 ♂, 17. vii. 1966, D. D. Munroe, Malaise trap (CNC), 1 ♂, 15. vii. 1966, D. D. Munroe, Malaise trap (CNC), 8 ♂, 29. vii. 1966, D. D. Munroe, Malaise trap (CNC), 3 ♂, 25. vi. 1964, D. D. Munroe, Malaise trap (CNC), 3 ♂, 25. vi. 1964 (CNC), 2 ♂, 18. viii. 1966, D. D. Munroe, Malaise trap (CNC), Mer Bleu, Ottawa, 45.39 ° N, 75.51 ° W, 1 ♂, 26. v. 1904, W. Metcalfe (CNC), Mer Bleue, Ottawa, 45.39 ° N, 75.51 ° W, 7 ♂, 25. vii. 1963, J. G. Chillcott (CNC), Ottawa, 45.43 ° N, 75.68 ° W, 1 ♂, 31. vii. 1989, J. R. Vockeroth (CNC); Quebec: Gatineau Co., Masham Twp., 45.64 ° N, 76.01 ° W, 1 ♂, 6 – 8. vii. 1974, D. M. Wood (CNC), Johnville Bog and Forest Park, 45.345 ° N, 71.7417 ° W, 11 ♂, 1 ♀, 15 – 22. vii. 2005, J. Savage, J. Kuchta, Malaise trap (LEMQ), 1 ♂, 28. vii – 1. viii. 2005, J. Savage, J. Kuchta, Malaise trap (LEMQ), 1 ♀, 6. vi – 1. vii. 1992, J. Savage, J. Kuchta, Malaise trap (LEMQ), 1 ♂, 9 – 13. vi. 2006, J. Savage, J. Kuchta, Malaise trap (LEMQ), 1 ♂, 26. viii – 2. ix. 2013, J. Savage, J. Kuchta, Malaise trap (LEMQ), 1 ♂, 29. vi – 3. vii. 1986, J. Savage, J. Kuchta, Malaise trap (LEMQ), 6 ♂, 1 ♀, 22 – 26. vii. 2005, J. Savage, J. Kuchta, Malaise trap (LEMQ), 1 ♂, 22 – 30. vii. 2013, J. Savage, J. Kuchta, Malaise trap (LEMQ), 1 ♂, 13. v – 21. vii. 1983, J. Savage, J. Kuchta, Malaise trap (LEMQ), Ogden, Marlington Bog, 45.04 ° N, 72.1733 ° W, 1 ♂, 27. vi – 4. vii. 2006, A. Moores, pan trap (LEMQ), Old Chelsea, Summit King Mt., 45.49 ° N, 75.86 ° W, 351 m, 1 ♂, 25. vi. 1962, J. R. Vockeroth (CNC), Reserve ecologique des Tourbieres de Lanoraie, 45.9967 ° N, 73.3 ° W, 1 ♀, 26 – 28. vii. 2005, A. Moores, Malaise trap (LEMQ), 1 ♂, 10 – 17. vii. 2006, A. Moores, pan trap (LEMQ), 41 ♂, 10 – 17. vii. 2006, A. Moores, Malaise trap (LEMQ), St-Charles Bog, 46.7478 ° N, 70.995 ° W, 35 ♂, 2 ♀, 22. vi. 2006, A. G. Taillefer, sweep (LEMQ), 4 ♂, 14. vi. 2006, A. G. Taillefer, sweep (LEMQ), Ste-Barbe, Large Teafield, 45.125 ° N, 74.225 ° W, 30 ♂, 17 – 24. vii. 2006, A. Moores, Malaise trap (LEMQ), 2 ♂, 18 – 25. vi. 2014, A. Moores, Malaise trap (LEMQ), 1 ♂, 3 – 7. viii. 1936, A. Moores, Malaise trap (LEMQ), 6 ♂, 2 ♀, 3 – 10. vii. 2006, A. Moores, Malaise trap (LEMQ). UNITED STATES: Arizona: 10 mi. NW Flagstaff, San Francisco Mts, 35.34 ° N, 111.68 ° W, 2896 m, 1 ♂, 18 – 21. vii. 1983, S. Peck, J. Peck, Malaise trap (DEBU); Arkansas: Lo- gan Co., Ozark National Forest, Magazine Mountain, 35.17 ° N, 93.64 ° W, 1 ♂, 23. vi – 31. viii. 1995, J. E. Swann, flight interception trap (ROM); District of Columbia: Washington, Rock Creek Park, 38.94 ° N, 77.05 ° W, 1 ♂, 4. viii. 1956, P. H. Arnaud, Jr. (USNM); Florida: Archbold Biol. Sta., 27.18 ° N, 81.35 ° W, 1 ♂, 28. iv. 1967, B. V. Peterson (CNC), 1 ♂, 29. iv. 1967, B. V. Peterson (CNC), Sebring, Highlands Hamm State Park, 27.47 ° N, 81.53 ° W, 4 ♂, 15. iv. 1970, W. W. Wirth, Malaise trap (USNM); Georgia: Forsyth, 33.03 ° N, 83.94 ° W, 2 ♂, 2. vi. 1970 (CNC), Warwoman Creek, 34.89 ° N, 83.29 ° W, 1 ♂, 4. viii. 1957, W. R. Richards (CNC), Waycross, 31.2 ° N, 82.4 ° W, 1 ♂, 30. vi. 1953, M. R. Wheeler (AMNH); Illinois: Carlinville, 39.27 ° N, 89.86 ° W, 1 ♂, 1 – 31. vii. 1956, M. R. Wheeler (AMNH), Macomb, 40.46 ° N, 90.68 ° W, 1 ♂, 29. v. 1962, W. W. Wirth (USNM); Indiana: La Fayette, 40.42 ° N, 86.87 ° W, 1 ♂, 8. vii. 1916, J. M. Al- drich (USNM), 1 ♂, 24. viii. 1916, J. M. Aldrich (USNM); Maryland: Colesville, 39.08 ° N, 77 ° W, 1 ♂, 14. vi. 1975, W. W. Wirth, Malaise trap (USNM), Montgomery Co., Bethesda, 38.99 ° N, 77.1 ° W, 1 ♂, 20. vii. 1968, G. Steyskal (USNM), 1 ♂, 24. viii. 1970, G. Steyskal (USNM), Montgomery Co., Colesville, 39.08 ° N, 77 ° W, 1 ♂, 13. viii. 1977, W. W. Wirth, Malaise trap (USNM), Prince Georges Co., Patuxent Wildlife Research Center, 39.03 ° N, 76.8 ° W, 1 ♂, 8. vii. 1978, W. W. Wirth (USNM); Michigan: Gd. Traverse Co., 44.76 ° N, 85.62 ° W, 1 ♂, 19. vii. 1953, G. Steyskal (USNM); New Hampshire: Strafford Co., Spruce Hole, 3 mi. SW Durham, 43.14 ° N, 70.92 ° W, 1 ♂, 3 – 7. vi. 1974, D. S. Chandler, flight interception trap (DEBU); New York: Lake Placid, 44.28 ° N, 73.98 ° W, 1 ♂, 19. vii. 1962, J. G. Chillcott (CNC), 1 ♂, 28. vii. 1929, A. L. Melander (USNM); North Carolina: Bladen Co., Singletary Lake State Park, 34.5833 ° N, 78.4583 ° W, 1 ♂, 19. vii – 5. viii. 1986, Marshall, Paiero, pan trap (DEBU), Macon Co., Highlands Lake Ravenel, 35.06 ° N, 83.19 ° W, 1 ♂, 14. vi. 1986, W. W. Wirth, Malaise trap (USNM), Macon Co., Highlands Wightman Cottage, 35.05 ° N, 83.19 ° W, 2 ♂, 10. vii. 1987, W. W. Wirth, UV light trap (USNM); Texas: San Jacinto Co., 6 km S Coldspring, Double Lake Camp, 30.55 ° N, 95.13 ° W, 2 ♂, 22. v – 8. vii. 1983, S. Peck, J. Peck, flight interception trap (DEBU), San Jacinto Co., Coldspring, Double Lake Camp, 30.55 ° N, 95.13 ° W, 2 ♂, 22 – 26. vi. 1974, S. Peck, J. Peck, flight interception trap (DEBU), Woodville, 30.76 ° N, 94.43 ° W, 1 ♂, 1 – 30. ix. 1954, M. R. Wheeler (AMNH); Vermont: East Franklin, Lake Carmi State Park, 44.9533 ° N, 72.8817 ° W, 6 ♂, 2 ♀, 11 – 18. vii. 2006, A. Moores, Malaise trap (LEMQ); Virginia: Cape Henry, 36.93 ° N, 76.02 ° W, 6 ♂, 28. vi. 1939, A. L. Melander (USNM), Chain Bridge, 38.93 ° N, 77.11 ° W, 1 ♂, 20. viii. 1922, J. R. Malloch (USNM), Falls Church, Holmes Run, 38.85 ° N, 77.19 ° W, 1 ♂, 2. vi. 1962, W. W. Wirth, light trap (USNM), 1 ♂, 6. viii. 1961, W. W. Wirth, light trap (USNM), 1 ♂, 23. viii. 1961, W. W. Wirth, light trap (USNM), 1 ♂, 17. ix. 1961, W. W. Wirth, light trap (USNM), Suitland bog, 38.84 ° N, 76.9 ° W, 4 ♂, 14. vi. 1951, W. W. Wirth (USNM); West Virginia: Franklin, 38.64 ° N, 79.33 ° W, 1 ♂, 26. vi. 1951, W. W. Wirth (USNM).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFACFFECFF3AFA0DC7BBA5A0.taxon	etymology	Etymology. The species name lustrum, a Latin noun, refers to bogs from which most of the species have been collected.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFACFFECFF3AFA0DC7BBA5A0.taxon	discussion	Remarks. The male genitalia are similar to those of Paramyia silvestris. Paramyia lustrum has a lot more marking within the overall surface of circular structure (Fig. 10 D). The surstylus is often twisting and not flat in appearance; take care to adjust the position to see the anterior edge properly. Genitalia often tucked within the abdomen, making it harder to see all characters. Females of Paramyia anguliloba, P. lustrum, P. nitens, P. rectiloba, P. silvestris, P. wheeleri can only be reliably identified through sequencing.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA8FFE1FF3AF999C408A1E8.taxon	description	(Fig. 11 – 12; 26) Description. (Fig. 11) Total length ♂ 1.3 – 1.4 mm, ♀ 1.7 – 1.8 mm. Overall colour dark brown to black, frontal tri- angle dark brown to black, shiny, reaching lunule; frons 1.05 – 1.25 times longer than wide and 1.7 – 1.8 times wider than flagellum; width at widest point 2.0 – 2.5 times the bottom eye width; gena dark brown with prunescence, 0.18 times eye height; postgena dark brown, with light prunescence and 0.25 – 0.27 width of head; scape and pedicel dark brown, first flagellomere trapezoidal, 1.1 times wider than long, dark brown with pale pubescence, arista dark brown and 1.63 – 1.78 times width of first flagellomere; strong facial carina, easily visible laterally; palpus dark brown with 4 enlarge setae apically, 1 longer ventral seta pre-apical and a row of short ventral setae, labium dark brown 0.36 – 0.39 mm and labella dark brown 0.47 – 0.51 mm long. Scutum shiny dark brown to black; scutellum dark brown to black, microsetose; all legs: femur, tibia and tarsi dark brown; hind basitarsus equal length of tarsi 2 – 4 combined; haltere uniformly dark brown; wing 1.33 – 1.42 mm long; abdomen shiny dark brown. Male postabdomen (Fig. 12). Epandrium in lateral view 1.04 times wider than high, in posterior view 2.2 times wider than high (before cerci), small setae densely covering posterior portion of epandrium mixed with a low density of large setae; surstylus 1.5 times as high as epandrium, paddle shape with a large wavy structure situated basally with many short markings within and a large dent on the apical anterior side starting near mid-length, setae medium length, covering anterior side and distal half of surstylus; cerci short and blunt apically, 0.57 times length of surstylus in posterior view, with a pair of very long preapical setae and other shorter setae, less than 1 / 3 its length, apically swollen in lateral view. Molecular barcode sequence can be accessed through GenBank Accession: [pending] and BOLD BIN: N / A (sequence too short for the BIN algorithm). Distribution (Fig. 26). Southwestern United-States: New Mexico, Arizona Type material. Holotype ♂. UNITED STATES: New Mexico: Catron Co., 8 mi. SE. Luna, 33.82 ° N, 108.95 ° W, 2286 m, 9 – 14. vii. 1979, S. Peck, J. Peck (DEBU). Paratypes. UNITED STATES: Arizona: Catalina Mts., Htchk. Hwy. mi. 27, 32.44 ° N, 110.79 ° W, 1 ♀, 12. viii. 1958, M. S. Adachi (USNM), Cochise Co., Rustler Park, Chiricahua Mts., 31.84 ° N, 109.29 ° W, 2560 m, 1 ♀, 21. viii. 1964 (AMNH); New Mexico: Catron Co., 5 mi. W Luna, San Francisco river, 33.82 ° N, 108.95 ° W, 2256 m, 1 ♂, 9 – 14. vii. 1987, S. Peck, J. Peck (DEBU), Catron Co., 8 mi. SE. Luna, 33.82 ° N, 108.95 ° W, 2286 m, 2 ♂, 9 – 14. vii. 1979, S. Peck, J. Peck (DEBU).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA8FFE1FF3AF999C408A1E8.taxon	etymology	Etymology. The species name nigritarsi comes from the Latin nigra, meaning black, combined with tarsi, which refers to the dark tarsi for all legs.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA4FFE7FF3AFF0AC481A454.taxon	description	(Fig. 13 – 14; 29) Description. (Fig. 13) Total length ♂ 1.8 mm, ♀ 2.0 – 2.3 mm. Overall colour dark brown to black, frontal triangle dark brown to black, shiny, reaching lunule; frons 1.0 – 1.1 times longer than wide and 2.0 – 2.2 times wider than flagellum; width at widest point 2.3 – 2.5 times the bottom eye width; gena shiny brown, 0.21 – 0.22 times eye height; postgena shiny brown, with light prunescence and 0.28 – 0.32 width of head; scape and pedicel brown to dark brown, first flagellomere oblong, 1.2 times wider than long, dark brown with pale pubescence, arista dark brown and 1.7 – 2.1 times width of first flagellomere; moderate facial carina, visible laterally; palpus dark brown with 4 enlarge setae apically, 1 ventral seta pre-apical and a ventral row of slender setae, labium brown 0.43 – 0.45 mm and labella brown 0.45 – 0.49 0.48 – 0.53 mm long. Scutum shiny dark brown to black; scutellum dark brown to black, microsetose; fore femur dark brown; fore tibia brown on dorsal and posterior side with apex lighter and anterior side light brown; femur and tibia on mid and hind leg dark brown; all tarsi light brown; hind basitarsus shorter than length of tarsi 2 – 4 combined; haltere uniformly dark brown; wing 1.6 – 1.7 mm long; abdomen shiny brown to dark brown. Male postabdomen (Fig. 14). Epandrium in lateral view 1.1 times longer than wide, in posterior view 2.3 times wider than high (before cerci), small setae densely covering posterior portion of epandrium mixed with a low density of large setae; surstylus 1.1 times as high as epandrium, paddle shape with anterior edge straight and posterior edge curved (opposite from the other species), and with a large egg shape structure situated basally and covering about half the surstylus length, short setae covering anterior side and distal half of surstylus and with 3 very long apical setae; cerci small, narrow and pointed apically, 0.53 times length of surstylus in posterior view, with short strong setae on bottom half, apically swollen in lateral view. Molecular barcode sequence can be accessed through GenBank Accession: HQ 981516 and BOLD BIN: BOLD: AAN 8613 (average p-distance within BIN: 0.3 %).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA4FFE7FF3AFF0AC481A454.taxon	distribution	Distribution (Fig. 29). Eastern North America: Canada: Ontario; United States: Georgia, Florida	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA4FFE7FF3AFF0AC481A454.taxon	materials_examined	Type material. Holotype ♂. CANADA: Ontario: Kent Co., Rondeau Provincial Park, South Point trail nr east parking lot, 15. v – 28. viii. 1994, Buck, Marshall, Malaise trap, (DEBU) [debu 01140022]. Paratypes. CANADA: Ontario: 4452 Rowsome Rd., Elizabethtown, 44.6213 ° N, 76.273 ° W, 120 m, 1 ♂, 12. vii. 2010, J. Sones (BIOUG), Kent Co., Rondeau Provincial Park, South Point trail nr east parking lot, 42.2617 ° N, 81.8469 ° W, 1 ♀, 17 – 24. vii. 2006, S. A. Marshall, Malaise trap (DEBU), 1 ♀, 7 – 16. vii. 2001, Marshall et al., Malaise trap (DEBU), Puslinch, Property of Bob Hanner, 43.4464 ° N, 80.2512 ° W, 335 m, 1 ♂, 1 ♀, 04. ix. 2008, T. Terzin (BIOUG), 1 ♂, 18. ix. 2008, T. Terzin (BIOUG), Thousand Islands National Park, 44.4534 ° N, 75.865 ° W, 1 ♂, 2 – 9. vii. 2014, M. Brown, Malaise trap (BIOUG), Wellington County, Puslinch Township, 43.54 ° N, 80.14 ° W, 320 m, 2 ♀, 10. vii. 2010, P. Hebert (BIOUG). UNITED STATES: Florida: Archbold Biol. Sta., 27.18 ° N, 81.35 ° W, 1 ♀, 9 – 18. xii. 1985, S. A. Marshall, flight interception trap (DEBU); Georgia: Forsyth, 33.03 ° N, 83.94 ° W, 2 ♀, 2. vi. 1970 (CNC).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA4FFE7FF3AFF0AC481A454.taxon	etymology	Etymology. The specific name refers to the very short flagellum, from the Latin brevis meaning short and Greek keraia meaning antenna, or any horn-like structure.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA3FFE4FF3AFB1AC473A1E8.taxon	description	(Fig. 15 – 16; 30) Description. (Fig. 15) Total length ♂ 1.4 – 1.7 mm, ♀ 1.6 – 1.8 mm. Overall colour brown to dark brown, frontal triangle brown to dark brown, shiny, reaching lunule; frons 1.14 – 1.25 times longer than wide and 1.6 – 1.74 times wider than flagellum; width at widest point 2.3 – 3.0 times the bottom eye width; gena brown with prunescence, 0.21 – 0.22 times eye height; postgena dark brown, with light prunescence and 0.25 width of head; scape and pedicel dark brown, first flagellomere trapezoidal, 1.2 times wider than long, dark brown with pale pubescence, arista dark brown and 1.6 – 1.7 times width of first flagellomere; mild facial carina, visible laterally on the upper part only; palpus dark brown with 4 enlarge setae apically and 1 ventral seta pre-apical, labium dark brown 0.43 – 0.45 mm and labella dark brown 0.49 – 0.53 mm long. Scutum shiny dark brown; scutellum dark brown, microsetose; all femur and tibia dark brown; all tarsi brown to light and paler than tibia; hind basitarsus shorter than length of tarsi 2 – 4 combined; haltere uniformly dark brown; wing 1.42 – 1.50 mm long; abdomen shiny dark brown. Male postabdomen (Fig. 16). Epandrium in lateral view 1.1 times wider than high, in posterior view 2.4 times wider than high (before cerci), small setae densely covering posterior portion of epandrium mixed with a low density of large setae; surstylus 1.3 times as high as epandrium, paddle shape with a strong elbow shape on the anterior edge and with a large circular structure situated basally with many short markings within, setae medium length, covering anterior side and distal half of surstylus; cerci medium size and pointed apically, 0.74 times length of surstylus in posterior view, with a pair of very long preapical setae and other shorter setae, less than 1 / 3 its length, apically swollen in lateral view. Molecular barcode sequence can be accessed through GenBank Accession: KP 049708 and BOLD BIN: BOLD: ACI 8776 (average p-distance within BIN: 0.23 %).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA3FFE4FF3AFB1AC473A1E8.taxon	distribution	Distribution (Fig. 30). East Coast: Canada: Ontario; United States: Florida, Maryland, Virginia	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA3FFE4FF3AFB1AC473A1E8.taxon	materials_examined	Type material. Holotype ♂. CANADA: Ontario: L. Cognashene Muskoka Dist., 28. vii. 1981, J. T. Huber (CNC) [CNC 263762]. Paratypes. CANADA: Ontario: Frontenac Co., Skycroft Campground, 5 km W Chaffeys Locks, 44.55 ° N, 76.3667 ° W, 2 ♂, 14 - 19. vii. 1987, B. Hubley, Malaise trap (ROM), Hamilton, 43.3474 ° N, 80.1801 ° W, 261 m, 1 ♂, 10 - 11. vii. 2017, African Lion Safari Staff, Malaise trap (BIOUG), L. Cognashene Muskoka Dist., 44.96 ° N, 79.92 ° W, 12 ♂, 28. vii. 1981, J. T. Huber (CNC), Ottawa, Montford Hospital, 45.4478 ° N, 75.6408 ° W, 1 ♂, 15. vii. 2002, J. R. Vockeroth (CNC), Point Pelee National Park, Woodland Trail, 41.9641 ° N, 82.5287 ° W, 170 m, 1 ♂, 19. vi. 2014, BIObus 2014, sweep (BIOUG), Thousand Islands National Park, Jones Creek by Mallorytown, County Rd. 5, 44.4747 ° N, 75.8652 ° W, 117 m, 1 ♀, 13 - 27. vii. 2012, BIOBus 2012 (BIOUG), Toronto, 43.703 ° N, 79.402 ° W, 176 m, 1 ♂, 3 - 10. vii. 2017, P. Hebert, Malaise trap (BIOUG), Warsaw, Ferris Provincial Park, 44.2829 ° N, 77.7963 ° W, 131 m, 2 ♀, 1 - 15. viii. 2014, CBG Collections Staff, Malaise trap (BIOUG), Windsor, Ojibway Prairie Provincial Park, 42.2628 ° N, 83.0725 ° W, 175 m, 1 ♀, 21 - 30. vi. 2006, CBG Collections Staff, Malaise trap (BIOUG); Quebec: Mont St. Bruno, 45.548 ° N, 73.3101 ° W, 1 ♀, 1 - 8. vii. 2008, V. Levesque, trunk trap (LEMQ). UNITED STATES: Florida: Ocean Pond, Olustee, 30.21 ° N, 82.45 ° W, 1 ♂, 29. vi. 1953, M. R. Wheeler (AMNH); Maryland: Calvert Co., Scientists Cliffs, 8 km S. Prince Frederick, 38.5167 ° N, 76.5133 ° W, 2 ♂, 22. v- 16. viii. 1983, J. M. Cumming (CNC); Ohio: Washington Co., S of New Metamoras on Hwy 7, Wayne Nat. For., 39.444 ° N, 81.162 ° W, 1 ♀, 14. viii. 2007, T. A. Wheeler, sweep (LEMQ); Virginia: Falls Church, 38.88 ° N, 77.17 ° W, 1 ♂, 26. vii. 1951, W. W. Wirth (USNM). Other material examined. CANADA: Ontario: Warsaw, Ferris Provincial Park, 44.2829 ° N, 77.7963 ° W, 131 m, 1 ♀, 4 - 18. vii. 2014, CBG Collections Staff, Malaise trap (BIOUG), Windsor, Ojibway Prairie Provincial Park, 42.2628 ° N, 83.0725 ° W, 175 m, 1 ♀, 24. vi- 1. vii. 2008, CBG Collections Staff, Malaise trap (BIOUG).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA3FFE4FF3AFB1AC473A1E8.taxon	etymology	Etymology. The specific name anguliloba is derived from the Latin nouns angulus meaning angle or bend, and lobus meaning lobe or rounded projection, reflecting the strong angle / elbow on the anterior side of the surstylus.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA3FFE4FF3AFB1AC473A1E8.taxon	discussion	Remarks. Females of P. anguliloba, P. lustrum, P. nitens, P. rectiloba, P. silvestris, P. wheeleri can only be reliably identified through sequencing.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA0FFD9FF3AFE4EC7F8A33C.taxon	description	(Fig. 17 - 18; 29) Description. (Fig. 17) Total length ♂ 1.3 - 1.4 mm, ♀ 1.6 - 1.7 mm. Overall colour brown to dark brown, frontal tri- angle dark brown to black, shiny, reaching lunule; frons 1.05 - 1.13 times longer than wide and 1.67 - 1.75 times wider than flagellum; width at widest point 2.0 - 3.33 times the bottom eye width; gena brown with prunescence, 0.13 - 0.17 times eye height; postgena dark brown, with light prunescence and 0.25 - 0.36 width of head; scape and pedicel dark brown, first flagellomere subquadrate, 1.2 times longer than wide, brown to dark brown with pale pubescence, arista dark brown and 1.67 - 1.8 times width of first flagellomere; moderate facial carina, easily visible laterally; palpus dark brown with 4 enlarge setae apically and 1 ventral seta pre-apical, labium dark brown 0.35 - 0.36 mm and labella dark brown 0.37 - 0.40 mm long. Scutum shiny dark brown; scutellum dark brown, microsetose; all femora and tibiae fully dark brown; all tarsi brown to light brown, fore tarsi color identical to fore tibia, more clearly visible on dorsal side; hind basitarsus shorter than length of tarsi 2 - 4 combined; haltere uniformly dark brown; wing 1.27 - 1.3 mm long; abdomen shiny dark. Male postabdomen (Fig. 18). Epandrium in lateral view 1.1 times wider than high, in posterior view 2.8 times wider than high (before cerci), small setae densely covering posterior portion of epandrium mixed with a low density of large setae; surstylus 1.3 times as high as epandrium, paddle shape with a small dent on the apical third of anterior side and with a large circular structure situated basally with short markings within, more limited to the distal half, setae medium length, covering anterior side and distal half of surstylus; cerci medium size and pointed apically, 0.7 times length of surstylus in posterior view, with a pair of very long preapical setae and other shorter setae, less than 1 / 3 its length, apically swollen in lateral view. Molecular barcode sequence can be accessed through GenBank Accession: HM 860793 and BOLD BIN: BOLD: ABX 4879 (average p-distance within BIN: 0.38 %).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA0FFD9FF3AFE4EC7F8A33C.taxon	distribution	Distribution (Fig. 29). Widespread throughout North America	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA0FFD9FF3AFE4EC7F8A33C.taxon	materials_examined	Type material. Holotype ♂. CANADA: Quebec: Bois-des-Bel Bog, 15. vi. 2006, A. G. Taillefer, sweep, (formerly in LEMQ, now deposited in CNC) [LEM- 0043712]. Paratypes. CANADA: Alberta: Jasper National Park, Pocahontas Campground, 53.1954 ° N, 117.914 ° W, 1131 m, 1 ♀, 16 - 22. vii. 2013, BIOBus 2012, Malaise trap (BIOUG); British Columbia: Silver Lake Hope, 49.31 ° N, 121.41 ° W, 1 ♂, 2. vii. 1968, W. W. Wirth (USNM); Newfoundland and Labrador: Gros Morne National Park, Lomond River Trail, 49.429 ° N, 57.7405 ° W, 35 m, 1 ♀, 10 - 20. vii. 2013, BIObus 2013, intercept trap (BIOUG), Terra Nova National Park, 48.598 ° N, 53.9702 ° W, 127 m, 1 ♂, 1 ♀, 25. vi- 2. vii. 2013, E. Perry, Malaise trap (BIOUG), 2 ♂, 1 ♀, 9 - 16. vii. 2013, E. Perry, Malaise trap (BIOUG), 1 ♂, 25 - 26. vii. 1976, E. Perry, Malaise trap (BIOUG); Nova Scotia: Cape Breton Highland National Park, Mackenzie Mt., 46.82 ° N, 60.82 ° W, 1 ♂, 4. vii. 1983, Vockervoth, Mal- aise trap (DEBU), Halifax, Point Pleasant Park, 44.623 ° N, 63.5686 ° W, 30 m, 1 ♂, 1 - 8. vi. 2013, T. Zemlak, Malaise trap (BIOUG); Ontario: Bruce Co., Cameron Lake Rd., fen, 45.22 ° N, 81.57 ° W, 1 ♂, 13 - 27. vii. 2012, S. A. Marshall, Malaise trap (DEBU), Thunder Bay Distr., Prairie River at Hwy 17, 38 km E Terrace Bay, 48.8011 ° N, 86.7844 ° W, 1 ♂, 15 - 22. vii. 2005, M. Buck, Malaise trap (DEBU); Quebec: Beechgrove, 45.65 ° N, 76.1333 ° W, 1 ♂, 29. vi. 1962, J. R. Vockeroth (CNC), Bois-des-Bel Bog, 47.9683 ° N, 69.4274 ° W, 7 ♂, 15. vi. 2006, A. G. Taillefer, sweep (LEMQ), 1 ♂, 9 - 14. vii. 1979, A. G. Taillefer, Malaise trap (LEMQ), 1 ♂, 21. vi. 2006, A. G. Taillefer, sweep (LEMQ), 1 ♂, 9 - 13. vi. 2006, A. G. Taillefer, Malaise trap (LEMQ), Bolton Pass, Knowlton, 45.22 ° N, 72.51 ° W, 244 m, 1 ♂, 5. vi. 1963, J. R. Vockeroth (CNC), Forillon National Park, 48.857 ° N, 64.376 ° W, 135 m, 1 ♀, 5 - 15. vii. 2013, F. Tremblay, Mal- aise trap (BIOUG), Gatineau Co., Masham Twp., 45.64 ° N, 76.01 ° W, 1 ♂, 3 - 10. vii. 2006, D. M. Wood (CNC), James Bat Rte., km 307, Pontax II River, 51.7753 ° N, 77.4242 ° W, 1 ♂, 8 - 10. viii. 2005, M. Buck, B. Buck, yellow pans (DEBU), James Bay Rte., km 204.5, 50.9831 ° N, 77.6339 ° W, 2 ♂, 7 - 16. vii. 2001, M. Buck, B. Buck, Malaise trap (DEBU), 2 ♂, 8 - 9. vii. 1961, M. Buck, B. Buck, Malaise trap (DEBU). UNITED STATES: Colorado: Rio Grande Co., Beaver Creek, 37.58 ° N, 106.65 ° W, 3048 m, 4 ♂, 21. vi. 1972, W. W. Wirth, Malaise trap (USNM). Other material examined. CANADA: Alberta: Jasper National Park, Pocahontas Campground, 53.1954 ° N, 117.914 ° W, 1131 m, 1 ♀, 16 - 22. vii. 2013, BIOBus 2012, Malaise trap (BIOUG); British Columbia: Yoho National Park, 51.3653 ° N, 116.528 ° W, 1268 m, 1 ♀, 7 - 16. vii. 2014, Jamie, Malaise trap (BIOUG); Newfoundland and Labrador: Terra Nova National Park, 48.598 ° N, 53.9702 ° W, 127 m, 1 ♀, 25 - 26. vii. 1976, F. Tremblay, Malaise trap (BIOUG), 2 ♀, 9 - 16. vii. 2013, E. Perry, Malaise trap (BIOUG); Nova Scotia: Halifax, Point Pleasant Park, 44.623 ° N, 63.5686 ° W, 30 m, 1 ♀, 15 - 19. vii. 2002, T. Zemlak, Malaise trap (BIOUG); Ontario: Griffith 7 mi. E, 45.24 ° N, 77.17 ° W, 1 ♂, 19. vii. 1990, B. E. Cooper (DEBU); Quebec: Bois-des-Bel Bog, 47.9664 ° N, 69.4305 ° W, 16 ♂, 9 - 13. vi. 2006, A. G. Taillefer, Malaise trap (LEMQ), 1 ♂, 9 - 14. vii. 1979, A. G. Taillefer, Malaise trap (LEMQ), 1 ♂, 15. vi- 2. vii. 2010, A. G. Taillefer, pan trap (LEMQ), 1 ♂, 21. vi. 2006, A. G. Taillefer, sweep (LEMQ), 1 ♂, 15 - 22. vi. 2008, A. G. Taillefer, pan trap (LEMQ), 5 ♂, 15. vi. 2006, A. G. Taillefer, sweep (LEMQ), 1 ♂, 15 - 21. vi. 2006, A. G. Taillefer, pan trap (LEMQ), Bois-des-Bel Bog, 47.9683 ° N, 69.4274 ° W, 16 ♂, 9 - 13. vi. 2006, A. G. Taillefer, Malaise trap (LEMQ), 1 ♂, 9 - 14. vii. 1979, A. G. Taillefer, Malaise trap (LEMQ), 1 ♂, 15. vi- 2. vii. 2010, A. G. Taille- fer, pan trap (LEMQ), 1 ♂, 21. vi. 2006, A. G. Taillefer, sweep (LEMQ), 1 ♂, 15 - 22. vi. 2008, A. G. Taillefer, pan trap (LEMQ), 5 ♂, 15. vi. 2006, A. G. Taillefer, sweep (LEMQ), 1 ♂, 15 - 21. vi. 2006, A. G. Taillefer, pan trap (LEMQ), Forillon National Park, 48.857 ° N, 64.376 ° W, 135 m, 1 ♂, 23. vi- 26. vii. 1995, F. Tremblay, Malaise trap (BIOUG), Gaspe, ZEC York-Baillargeon, forest edge near Lac Pauline, 48.8167 ° N, 64.8667 ° W, 1 ♂, 26. vii. 2000, H. Varady- Szabo (LEMQ), James Bay Rte. km 570.3, 53.4942 ° N, 77.6208 ° W, 1 ♂ ,, M. Buck, B. Buck, Malaise trap (DEBU), James Bay Rte., km 204.5, 50.9831 ° N, 77.6339 ° W, 1 ♂, 7 - 16. vii. 2001, M. Buck, B. Buck, Malaise trap (DEBU), Mont St. Hilaire, 45.546 ° N, 73.161 ° W, 1 ♀, 10 - 17. vi. 2008, V. Levesque, trunk trap (LEMQ); Saskatchewan: Prince Albert National Park, 53.8497 ° N, 106.078 ° W, 627 m, 1 ♀, 10 - 12. vii. 2012, R. Thompson, E. Knight (BIOUG).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA0FFD9FF3AFE4EC7F8A33C.taxon	etymology	Etymology. The Latin name silvestris referring to forests where this species is collected most often.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FFA0FFD9FF3AFE4EC7F8A33C.taxon	discussion	Remarks. The male genitalia can appear similar to species P. lustrum; however, P. silvestris has less marking within the overall surface of circular structure and they are more situated in the bottom half (Fig. 18 C). The epandrium also has a curve before the cerci on lateral view. Make sure to adjust the position on the anterior edge properly to see the dent. The surstylus apex is also flatter like P. rectiloba while P. lustrum is more pointed. Genitalia often tucked within the abdomen, making it harder to see all characters. Females of P. anguliloba, P. lustrum, P. nitens, P. rectiloba, P. silvestris, P. wheeleri can only be reliably identified through sequencing.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF9DFFDFFF3AFC22C473A76B.taxon	description	(Fig. 19 - 20; 26) Description. (Fig. 19) Total length ♂ 1.2 mm, ♀ 1.5 - 1.8 mm. Overall colour brown to dark brown, frontal triangle brown to dark brown, shiny, reaching lunule; frons 1.31 times longer than wide and 1.6 times wider than flagellum; width at widest point 2.0 times the bottom eye width; gena brown with prunescence, 0.09 times eye height; postgena brown to dark brown, with light prunescence and 0.24 width of head; scape and pedicel brown to dark brown, first flagellomere subquadrate, 1.1 times wider than long, brown with pale pubescence, arista dark brown and 1.5 times width of first flagellomere; moderate facial carina, easily visible laterally; palpus dark brown with 4 enlarge setae apically, 1 ventral seta pre-apical and a ventral row of slender setae, labium brown to dark brown 0.33 - 0.40 mm and labella brown to dark brown 0.36 mm long. Scutum shiny brown; scutellum brown, microsetose; all femur brown; fore tibia brown with apex paler and anterior side mostly light brown to yellowish; mid and hind tibia all brown; all tarsi light brown to yellowish; hind basitarsus shorter than length of tarsi 2 - 4 combined; haltere uniformly brown; wing 1.14 mm long; abdomen shiny brown to dark brown. Male postabdomen (Fig. 20). Epandrium in lateral view 1.25 times longer than wide, in posterior view 2.3 times wider than high (before cerci), small setae densely covering posterior portion of epandrium mixed with a low density of large setae; surstylus 1.1 times as high as epandrium, elephant trunk shape tapering in the first 1 / 3 and remaining thin until tip, with a large thick half circular structure situated basally with few short markings within, with only a few apical setae; cerci small and slightly rounded apically, 0.46 times length of surstylus in posterior view, medium length setae along the edge, uniform thickness in lateral view. Molecular barcode sequence can be accessed through GenBank Accession: HM 417566.1 and BOLD BIN: BOLD: AAG 0166 (average p-distance within BIN: 0.17 %).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF9DFFDFFF3AFC22C473A76B.taxon	distribution	Distribution (Fig. 26). Canada: Ontario; United States: Arizona	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF9DFFDFFF3AFC22C473A76B.taxon	materials_examined	Type material. Holotype ♂. CANADA: Ontario: Cambridge, rare Charitable Research Reserve, 43.3736 ° N, 80.3652 ° W, 304 m, 4 - 11. vi. 2015, BIO Collections, Malaise trap, (formerly in BIOUG, now deposited in CNC) [BIOUG 22726 - B 04]. Paratypes. CANADA: Ontario: Cambridge, rare Charitable Research Reserve, 43.3736 ° N, 80.3652 ° W, 304 m, 2 ♀, 18 - 24. vi. 2015, BIO Collections, Malaise trap (BIOUG), 1 ♀, 18 - 24. vii. 1979, BIO Collec- tions, Malaise trap (BIOUG), 1 ♀, 18 - 21. vii. 1978, BIO Collections, Malaise trap (BIOUG), Thousand Islands Na- tional Park, 44.4534 ° N, 75.865 ° W, 1 ♀, 2 - 9. vii. 2014, M. Brown, Malaise trap (BIOUG). UNITED STATES: Ari- zona: Santa Cruz Co., Patagonia, 31.53 ° N, 110.77 ° W, 1 ♂, 14. x. 1995, B. V. Brown, E. Wilk, Malaise trap (LEMQ).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF9DFFDFFF3AFC22C473A76B.taxon	etymology	Etymology. The species name is in honour of Dr. Terry A. Wheeler who was involved in the early stages of this study but could not see the final product because of a brain tumor. He was quite interested in this species because the genitalia are so drastically different from the other species.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF9DFFDFFF3AFC22C473A76B.taxon	discussion	Remarks. Females of P. anguliloba, P. lustrum, P. nitens, P. rectiloba, P. silvestris, P. wheeleri can only be reliably identified through sequencing.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF9BFFDCFF3AF8C9C29EA0B4.taxon	description	(Fig. 21 - 22; 26) Description. (Fig. 21) Total length ♂ 1.8 mm. Female unknown. Overall colour dark brown, frontal triangle dark brown to black, shiny, reaching lunule; frons 1.33 times longer than wide and 1.5 times wider than flagellum; width at widest point 1.81 times the bottom eye width; gena dark brown with prunescence, 0.12 times eye height; postgena dark brown, with light prunescence and 0.15 width of head; scape and pedicel dark brown, first flagellomere trapezoidal, 1.2 times wider than long, dark brown with pale pubescence, arista dark brown and 1.67 times width of first flagellomere; moderate facial carina, easily visible laterally; palpus dark brown with 4 enlarge setae apically and 1 ventral seta close to mid-point, labium dark brown 0.43 mm and labella dark brown 0.45 mm long. Scutum shiny dark brown; scutellum dark brown, microsetose; fore femur dark brown with apical antero-dorsal part with a slightly paler circle shape; fore tibia dark brown and slightly paler on the anterior apical section; femur and tibia on mid and hind leg dark brown; all tarsi light brown to yellowish; hind basitarsus shorter than length of tarsi 2 - 4 combined; haltere uniformly dark brown; wing 1.52 mm long; abdomen shiny dark brown. Male postabdomen (Fig. 22). Epandrium in lateral view 1.02 times wider than high, in posterior view 2.6 times wider than high (before cerci), small setae densely covering posterior portion of epandrium mixed with a low density of large setae; surstylus 1.4 times as high as epandrium, paddle shape increasing in width from base to apex, with a large wavy circular structure situated basally with many wavy short markings within, setae medium length, covering anterior side and distal half of surstylus; cerci large and pointed apically, 0.86 times length of surstylus in posterior view, with two pair of medium length setae along the edge, apically swollen in lateral view. Molecular barcode sequence can be accessed through GenBank Accession: [pending] and BOLD BIN: origi- nally BOLD: ADO 8669, now merged with BOLD: AAG 0168 (P. nitens).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF9BFFDCFF3AF8C9C29EA0B4.taxon	distribution	Distribution (Fig. 26). Southern United-States: Mississippi	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF9BFFDCFF3AF8C9C29EA0B4.taxon	materials_examined	Type material. Holotype ♂. UNITED STATES: Mississippi: Winston Co., Noxubee National Wildlife E., Tripletts, 19. v. 2013, J. M. Cumming (CNC).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF9BFFDCFF3AF8C9C29EA0B4.taxon	etymology	Etymology. The species name comes from incrasso (from Latin) meaning fatten, thicken and lobos (from Greek) meaning lobe, which refers to the surstylus shape increasing in width from base to apex.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF98FFD1FF3AFEBAC473A25C.taxon	description	(Fig. 23 – 24; 30) Description. (Fig. 23) Total length ♂ 1.4 – 1.5 mm, ♀ 1.7 – 1.8 mm. Overall colour brown to dark brown, frontal tri- angle dark brown to black, shiny, reaching lunule; frons 1.1 – 1.33 times longer than wide and 1.75 – 1.8 times wider than flagellum; width at widest point 2.13 – 2.33 times the bottom eye width; gena brown to dark brown with prunescence, 0.14 – 0.19 times eye height; postgena dark brown, with light prunescence and 0.23 – 0.30 width of head; scape and pedicel dark brown, first flagellomere subquadrate to slightly trapezoidal, same length than width, dark brown with pale pubescence, arista dark brown and 1.64 – 1.82 times width of first flagellomere; moderate facial carina, easily visible laterally; palpus brown to dark brown with 5 enlarge setae apically and 1 ventral seta pre-apical, labium dark brown 0.37 – 0.43 mm and labella brown and apically paler 0.46 – 0.48 mm long. Scutum shiny dark brown; scutellum dark brown, microsetose; fore femur brown with apical antero-dorsal part with a slightly paler circle shape, mid and hind femur brown and all tibia brown; all tarsi light brow to yellowish; hind basitarsus shorter than length of tarsi 2 – 4 combined; haltere brown and slightly darker apically; wing 1.21 – 1.33 mm long; abdomen shiny dark brown. Male postabdomen (Fig. 24). Epandrium in lateral view 1.0 times wider than high, in posterior view 2.4 times wider than high (before cerci), small setae densely covering posterior portion of epandrium mixed with a low density of large setae; surstylus 1.3 times as high as epandrium, rounded rectangular shape with a large circular structure, thicker on the posterior side, situated basally with just a few short markings in the middle, setae medium length, covering anterior side and distal half of surstylus; cerci large and pointed apically, 0.89 times length of surstylus in posterior view, with a pair of very long preapical setae and other shorter setae, less than 1 / 3 its length, apically swollen in lateral view. Molecular barcode sequence can be accessed through GenBank Accession: HQ 981642 and BOLD BIN: BOLD: AAG 0170 (average p-distance within BIN: 0.36 %).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF98FFD1FF3AFEBAC473A25C.taxon	distribution	Distribution (Fig. 30). East Coast: Canada: Nova Scotia, Quebec, Ontario; United States: Vermont, New Hampshire, Michigan, Florida / West Coast: Canada: British Columbia; United States: New Mexico	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF98FFD1FF3AFEBAC473A25C.taxon	materials_examined	Type material. Holotype ♂. CANADA: British Columbia: 10 mi. E. Creston, 31. vii. 1980, G. Gibson, sweeping, (DEBU). Paratypes. CANADA: British Columbia: 10 km W Kamloops, New Afton Mine, 50.6719 ° N, 120.533 ° W, 1 ♂, 24. vii – 1. viii. 2013, C. Simon, Malaise trap (BIOUG), 10 mi. E. Creston, 49.1 ° N, 116.51 ° W, 4 ♂, 31. vii. 1980, G. Gibson, sweeping (DEBU), 10 mi. E. Osoyoos, 49.03 ° N, 119.44 ° W, 1 ♂, 30. vii. 1980, G. Gibson, sweeping (DEBU); Nova Scotia: Halifax, Point Pleasant Park, 44.623 ° N, 63.5686 ° W, 30 m, 1 ♀, 14 – 22. vi. 2006, T. Zemlak, Malaise trap (BIOUG), 1 ♂, 20 – 27. vii. 2013, T. Zemlak, Malaise trap (BIOUG), 1 ♀, 22 – 26. vii. 2005, T. Zemlak, Malaise trap (BIOUG), 1 ♂, 20 – 30. xi. 1986, T. Zemlak, Malaise trap (BIOUG), 1 ♂, 14 – 19. vii. 1987, T. Zemlak, Malaise trap (BIOUG), 1 ♀, 20. viii – 3. ix. 2014, T. Zemlak, Malaise trap (BIOUG); Ontario: Frontenac Co., Skycroft Campground, 5 km W Chaffeys Locks, 44.55 ° N, 76.3667 ° W, 1 ♂, 14 – 19. vii. 1987, B. Hubley, Malaise trap (ROM), 1 ♂, 14. viii – 7. ix. 2003, B. Hubley, Malaise trap (ROM), 1 ♂, 14 – 19. vii. 2013, B. Hubley, Malaise trap (ROM), Killar- ney Provincial Park, nr hwy 637, 46.01 ° N, 81.1 ° W, 1 ♂, 11. vii. 2004, J. Forrest, sweep (LEMQ), Lambton Co., Pinery Provincial Park, Sand dunes, 44.55 ° N, 76.3667 ° W, 1 ♂, 19 – 26. vii. 2006, L. Packer, Malaise trap (ROM), Thousand Islands National Park, 44.4534 ° N, 75.865 ° W, 3 ♂, 1 ♀, 16 – 23. vii. 2014, M. Brown, Malaise trap (BI- OUG), 3 ♂, 30. vii – 8. viii. 2014, M. Brown, Malaise trap (BIOUG), 2 ♂, 2 – 9. vii. 2014, M. Brown, Malaise trap (BI- OUG), 1 ♂, 17. vi – 3. vii. 2003, M. Brown, Malaise trap (BIOUG), 1 ♂, 31. vii – 18. viii. 2014, M. Brown, Malaise trap (BIOUG); Quebec: Gatineau Co., Masham Township, 45.64 ° N, 76.01 ° W, 1 ♂, 27. vii. 1995, E. Ikeda (LEMQ), Mont St. Bruno, 45.5524 ° N, 73.3339 ° W, 1 ♀, 24. vii – 1. viii. 2013, V. Levesque, flight interception trap (LEMQ), Ogden, Marlington Bog, 45.04 ° N, 72.1733 ° W, 1 ♀, 4 – 5. vii. 2010, A. Moores, Malaise trap (LEMQ), Old Chelsea, King Mt., 45.49 ° N, 75.86 ° W, 1 ♂, 13. viii. 1969, B. V. Peterson (CNC), St-Daniel, Parc National de Frontenac, 45.9683 ° N, 71.1517 ° W, 1 ♂, 12 – 19. viii. 2013, A. Moores, pan trap (LEMQ). UNITED STATES: Florida: Miami, 25.76 ° N, 80.19 ° W, 1 ♂, 20. ii. 1912, F. Knal (USNM); Michigan: Hunt Cr. Exp. Sta. nr. Lewiston, 44.88 ° N, 84.31 ° W, 1 ♂, 20. vii. 1942, C. W. Sabrosky (USNM); New Mexico: Catron Co., 8 mi. SE. Luna, 33.82 ° N, 108.95 ° W, 2286 m, 4 ♂, 9 – 14. vii. 1987, S. Peck, J. Peck (DEBU); Vermont: East Franklin, Lake Carmi State Park, 44.9533 ° N, 72.8817 ° W, 1 ♂, 11 – 18. vii. 2006, A. Moores, pan trap (LEMQ), 1 ♂, 11 – 17. vii. 2013, A. Moores, Malaise trap (LEMQ). Other material examined. CANADA: British Columbia: 10 km W Kamloops, New Afton Mine, 50.6719 ° N, 120.533 ° W, 1 ♂, 24 – 30. vii. 2013, C. Simon, Malaise trap (BIOUG), 1 ♂, 24 – 28. viii. 1987, C. Simon, Malaise trap (BIOUG), Mount Revelstoke National Park, Off trail behind old staff housing, 51.0181 ° N, 118.205 ° W, 608 m, 1 ♂, 4 – 11. vii. 2006, S. Devita, Malaise trap (BIOUG); Nova Scotia: Halifax, Point Pleasant Park, 44.623 ° N, 63.5686 ° W, 30 m, 3 ♂, 1 ♀, 6 – 13. vii. 2013, T. Zemlak, Malaise trap (BIOUG), 1 ♀, 3 – 10. viii. 2013, T. Zemlak, Malaise trap (BIOUG), 1 ♀, 28. vi – 12. vii. 1986, T. Zemlak, Malaise trap (BIOUG), Kejimkujik, Hemocks and Hardwoods, 44.4425 ° N, 65.2535 ° W, 97 m, 1 ♂, 31. vii. 2009, BIObus 2009 (BIOUG); Ontario: 4452 Rowsome Rd., Elizabethtown, 44.6213 ° N, 76.273 ° W, 120 m, 1 ♀, 19. vi. 2010, J. Sones (BIOUG), Bruce Co., Bruce Peninsula National Park, Lake Scugog, 45.1167 ° N, 81.5333 ° W, 1 ♂, 1 – 17. viii. 2002, S. A. Marshall, Malaise trap (DEBU), Puslinch, Property of Bob Hanner, 43.4464 ° N, 80.2512 ° W, 335 m, 2 ♀, 03. viii. 2008, T. Terzin (BIOUG), 1 ♂, 21. viii. 2008, T. Terzin (BIOUG), Thousand Islands National Park, 44.4534 ° N, 75.865 ° W, 6 ♂, 5 ♀, 16 – 23. vii. 2014, M. Brown, Malaise trap (BIOUG), 1 ♂, 2 ♀, 30. vii – 8. viii. 2014, M. Brown, Malaise trap (BIOUG), 1 ♂, 1 ♀, 2 – 9. vii. 2014, M. Brown, Malaise trap (BIOUG), 1 ♂, 17 – 24. vii. 2006, M. Brown, Malaise trap (BIOUG), 1 ♂, 4 – 11. vii. 2006, M. Brown, Mal- aise trap (BIOUG); Quebec: Johnville Bog and Forest Park, 45.345 ° N, 71.7417 ° W, 1 ♂, 12 – 19. vii. 2006, J. Savage, J. Kuchta, Malaise trap (LEMQ). UNITED STATES: New Hampshire: Coos Co., 1 km E Stark, 44.6 ° N, 71.4 ° W, 1 ♂, 8. viii. 2000, J. Savage, sweep (LEMQ).	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF98FFD1FF3AFEBAC473A25C.taxon	etymology	Etymology. The species name come from the Latin rectus, meaning straight and lobos (from Greek) meaning lobe, which refers to the fairly straight / parallel side of the surstylus.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
03B187B7FF98FFD1FF3AFEBAC473A25C.taxon	discussion	Remarks. Females of P. anguliloba, P. lustrum, P. nitens, P. rectiloba, P. silvestris, P. wheeleri can only be reliably identified through sequencing.	en	Levesque-Beaudin, Valerie, Mlynarek, Julia J. (2020): Revision of Nearctic Paramyia Williston (Diptera: Milichiidae). Zootaxa 4732 (1): 1-56, DOI: https://doi.org/10.11646/zootaxa.4732.1.1
