taxonID	type	description	language	source
03949824997BFF83FE2D2620FE2DF955.taxon	type_taxon	Type species Devolana hemicycla DeLongı 1967 ı p. 23 ı original description. Distribution	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
03949824997BFF83FE2D2620FE2DF955.taxon	distribution	Endemic to Mexicoı reported from the following states: Guerrero (Iguala de la Independencia and Zincauro) and Jalisco (Sierra de Tuxcacuesco) ı Figure 4.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
03949824997BFF83FE2D2620FE2DF955.taxon	diagnosis	Diagnosis Devolana is easily distinguished from other similar genera of Athysanini by the following combination of characteristics: (1) head very short with anterior and posterior margins of crown parallel; (2) face with arcuate black line extended from midline laterad through ocelli and along border of frontoclypeus; (3) pronotum with pair of large black spots; (4) pygofer without processes; and (5) aedeagus with a set of apical processes.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
03949824997BFF83FE2D2620FE2DF955.taxon	description	Morphology Moderately robust leafhoppers with overall appearance yellowish with black spots on crown and pronotum. Crown roundedı paralleled to posterior margin. Transition of crown to face angledı shagreen. Ocellocular area parallel and wide. Ocelli slightly below margin of headı distance to adjacent eye approximately 2 x ocellar diameter. Frontoclypeus wideı lateral frontal sutures reaching ocellusı fine setae laterad frontal suture. Anteclypeus widening apicallyı apex curved slightly surpassing natural curve of genae. Lorum subequal than anteclypeus near base. Genae incised laterad. Pronotum weakly produced mediallyı laterad margins carinateı yellowish with a symmetrical pair of large black spots and a paired diffuse palebrown line between them. Scutellum broad. Forewing macropterousı translucent with black-brown colour on vein-scheme or yellowish pigmentation near the base of clavusı appendix restricted to anal marginı apex roundedı apical and anteapical cells largeı Cs – Pcu crossvein absentı Pcu – A 1 crossvein only presented in D. tuxcacuensis. Front femur AM 1 near mid-height of apexı long; IC longı thin; row AV small. Front tibia dorsal macrosetal formula (AD + PD) 1 + 4 ı PV small stout setae. Hind femur macrosetal formula 2 + 2 + 1. Pygofer broadı shortı square or pointedı without processesı long and small stout macrosetae near apexı basolateral cleft present. Segment tenth smallı broadı wellsclerotised with membranous tip. Valve and subgenital plates free articulated to pygofer. Valve rounded and slightly produced. Subgenital plate trianguları uniseriate laterad with robust and fine setae. Connective Y-shapedı stem short. Style slenderı basal and preapical lobe weakly developed with setae on posterad marginı apophysis short and curved mesadı apex truncate. Aedeagus curved dorsadı aedeagal shaft with lateral flanges devolved or notı two or three pairs of apical processes. Aedeagal base without processes. Gonoduct sclerotised. Gonophore subapical on ventral surface.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
03949824997BFF83FE2D2620FE2DF955.taxon	discussion	Remarks DeLong (1967) compared this taxon to Exitianus and Spinulana ı which differ by combinations of genital structures and external features. Exitianus was included in tribe Chiasmini by Zahniser and Dietrich (2013); members of this tribe have a tapering or parallel-sided anteclypeus and an aedeagus hinged at the base. Spinulana and Devolana ı both of which are currently placed in Athysanini differ in general body colour and in the structure of the male pygofer and aedeagus.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249976FF8FFE6021BDFCD2FAF7.taxon	description	(Figure 10)	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249976FF8FFE6021BDFCD2FAF7.taxon	distribution	Distribution known Iguala de la Independenciaı Guerrero (Mexico) ı Figure 4.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249976FF8FFE6021BDFCD2FAF7.taxon	description	Measurements Length male 6 mmı female unknown.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249976FF8FFE6021BDFCD2FAF7.taxon	materials_examined	Material examined Holotype male (OSUC) – MEXICO: Guerreroı Igualaı X – 25 – 1941 ı DeLongı Goodı Caldwell and Plummer Colls. E- 103. D. M. DeLong Collection. # OSUC 209432.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249976FF8FFE6021BDFCD2FAF7.taxon	discussion	Remarks Label data from the only previously dissected specimen in the DeLong collection at Ohio State Universityı labelled as the holotype of this speciesı was misquoted in DeLong’ s (1967) publication. This specimen is unquestionably the holotype because DeLong (1967) illustrated the male genitalia with his original description and none of the paratypes were dissected. The label data published with the original description instead apply to three of the four specimens labelled as paratypes. DeLong’ s type series of D. hemicycla includes two additional undescribed species. Study of the entire type series of five specimens indicates that the specimen originally labelled as the holotype is the only representative of D. hemicycla speciesı whereas other specimens in the type series belong to the species D. youajla and D. xajxayakamej ı described as new above.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
03949824997DFF88FEDF2323FC6AFB6A.taxon	description	Description Measurements. Length male 5 mmı and female unknown. Male morphology. Overall colour light-yellowish dorsally with white venter (Figures 1 – 2). Crown with black transverse band at apex extended laterally onto faceı and pair of tiny black spots next to eyes (Figures 1 (a) and 2). Face with upper-half yellowish and lower half white. Frontoclypeal and anteclypeal margins marked with black band extended dorsad over ocelli to midlineı multiple black transverse bands over frontoclypeal surface and anteclypeus with one dark-brown medial vertical line. Gena and lorum white with diffuse blackı yellow and brown minute spots (Figure 1 (b )). Forewing translucent over almost entire surface except with patches of yellowish opaque pigment near base of clavus and veins Cı Mı CuP and A dark brown (Figure 1 (c )). Legs white with black stripes along tibiae and across front femur. Female. Unknown. Male genitalia. Pygofer broad with macrosetae mixed with small setae beyond midlength (Figure 3 (a – b )). Valve slightly projected and roundedı articulated with pygofer and subgenital plates (Figure 3 (c )). Plate extended to apex of pygoferı with mix of fine and stout long setae along lateral margin (Figure 3 (d )). Style slender with basal and preapical lobe weakly developedı apophysis slender and slightly curved mesad (Figure 3 (h )). Connective Y-form (Figure 3 (i )). Aedeagus robustı short and curved with two long pairs of apical processes directed anterodorsad (Figure 3 (e – g )). Immature stages. Unknown.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
03949824997DFF88FEDF2323FC6AFB6A.taxon	distribution	Type locality. Sierra de Tuxcacuescoı Jalisco (Mexico) ı Figure 4. Seasonality. This species occurs between September and October.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
03949824997DFF88FEDF2323FC6AFB6A.taxon	etymology	Etymology. The species epithet refers to the type locality.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
03949824997DFF88FEDF2323FC6AFB6A.taxon	materials_examined	Type material. Holotype male (INHS) – MEXICO: Jaliscoı 13 km ESE Tuxcacuesco 900 mı 19 ° 40 ʹ 6 ” N 104 ° 1 ʹ 53 ” Wı 13 October 2001 ı S. H. McKamey et al. Colls. ı fogging 1 E-T. Paratype male (CAJAPE) – MEXICO: Jaliscoı Región Sierra de Amulaı Municipio Tuxcacuescoı Sierra de Tuxcacuesco a 5.5 km del pueblo Tuxcacuescoı 19 ° 42 ʹ 02.1 ” N 104 ° 01 ʹ 41.6 ” Wı 864 m. 08 de septiembre de 2018. Vegetación BTC. J. A. Pinedo-Escatel Col. Trampa de Luz [MEXJAL 88].	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
03949824997DFF88FEDF2323FC6AFB6A.taxon	discussion	Remarks. This species is similar in dorsal appearance to D. youajla ı but D. tuxcacuensis differs in lacking lateral flanges on the aedeagal shaft (Figure 3 (f – g )).	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249975FF8FFEBB21D1FBA1FEBD.taxon	description	(Figures 8 and 9) Description Measurements. Length male 5.2 mmı female unknown. Male morphology. Overall colour light-yellowish (Figure 8 (a – b )). Crown yellowish with black transverse band at apex and pair of black spots next to eyes (Figure 8 (a )). Face entire yellowish. Frontoclypeal and anteclypeal sides unmarkedı several black transverse bands over frontoclypeal surface and anteclypeus yellowish. Gena and lorum yellowishı unmarked (Figure 8 (b )). Pronotum brown-yellowish with a symmetrical pair of black spots with a paired diffuse pale-brown line between them. Forewing translucent with dark-brown pigment on Cı Mı CuAı Csı Pcu and A veins (Figure 8 (c )). Legs brown with black stripe along front tibia. Female. Unknown. Male genitalia. Pygofer somewhat conical with macrosetae on posterior half (Figure 7 (a – b )). Valve projected articulated with pygofer and plates (Figure 9 (c )). Plate extended near to apex of pygofer with row of long and stout setae beside fine long setae on lateral margin (Figure 9 (d )). Style slender with lobes weakly developedı apophysis short and curved mesad (Figure 9 (h )). Connective Y-form (Figure 9 (i )). Aedeagus robustı shortı apex pointed and flanges developed with three pairs of apical processes; two processes emerging from flanges on dorsal surface directed dorsad and anteradı respectively; third process arising close from apex on ventral surfaceı long surpassing midlength of aedeagal shaft and subapically expanded (Figure 9 (e – g )). Immature stages. Unknown.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249975FF8FFEBB21D1FBA1FEBD.taxon	distribution	Type locality. Zincauroı Guerrero (Mexico) ı Figure 4. Seasonality. This species occurs in October.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249975FF8FFEBB21D1FBA1FEBD.taxon	etymology	Etymology. The species epithet derives from the Náhuatl word maskı in allusion to its black markings of the crown	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249975FF8FFEBB21D1FBA1FEBD.taxon	materials_examined	Type material. Holotype male (OSUC) – MEXICO: Guerreroı Zincauroı IX – 2 – 1930 ı J. Parra Coll. ı D. M. DeLong Collection. # OSUC 356604. Note: Paratype previously designated to Devolana hemycicla	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249975FF8FFEBB21D1FBA1FEBD.taxon	discussion	Remarks. This species is very similar in external appearance to D. hemycicla species but differs in lacking lobes on the posterior margin of the aedeagal flanges (Figure 9 (f – g )).	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249971FF8CFE4324ECFC8CFE56.taxon	description	(Figures 6 and 7) Description Measurements. Length male 5.9 mmı female unknown. Male morphology. Overall colour brown-yellowish. Crown shortı paralleled to anterior margin with two pairs of black spotsı one next to midline and second next to eyes (Figure 6 (a )). Face yellowish. Frontoclypeal and anteclypeal margins marked with black band extended dorsad over ocelli and almost connecting to midlineı multiple black transverse bands over frontoclypeal surface presented and anteclypeus longitudinally brown marked. Lorum yellowish. Gena mostly yellowish with a brown line running from eye corner towards midlength of lorum and diffuse black spot around maxillary sensillum (Figure 6 (b )). Pronotumı anterior half gold-yellowish with posterior area white. Forewing translucent with dark-brown pigment on Cı Mı CuAı Csı Pcu and A veins (Figure 6 (c )). Legs yellowish with black stripes along tibiae and front femur. Female. Unknown. Male genitalia. Pygofer broad with stout macrosetae distributed near midlength towards posterad margin (Figure 7 (a – b )). Valve medially producedı articulated with pygofer and subgenital plates (Figure 7 (c )). Plate thinı length similar to posterad margin of pygoferı few small fine and stout setae along lateral margin (Figure 7 (d )). Style slender with posterad lobes weakly developedı apophysis short and curved (Figure 7 (h )). Connective Y-form (Figure 3 (i )). Aedeagus robustı short and apex pointed with broad lateral flanges developed with two apical processes; one emerging of distant flange corner directed dorsad and second arising near apex on ventral surface running ventromesad until midlength of aedeagal shaftı cylindrical (Figure 7 (e – g )). Immature stages. Unknown.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249971FF8CFE4324ECFC8CFE56.taxon	etymology	Etymology. The species epithet derives from the Náhuatl word Igualaı meaning to the type locality.	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249971FF8CFE4324ECFC8CFE56.taxon	materials_examined	Type material. Holotype male (OSUC) – MEXICO: Guerreroı Igualaı IX – 11 – 1939 ı D. M. DeLong Collectionı D. M. DeLong Coll. # OSUC 356605. Paratype 2 males (OSUC) – MEXICO: Guerreroı Igualaı IX – 11 – 1939 ı D. M. DeLong Collectionı D. M. DeLong Coll. # OSUC 356606 and 356607 ı respectively. Note: all specimens examined were previously designated as paratypes of Devolana hemycicla DeLong by DeLong (1967).	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
039498249971FF8CFE4324ECFC8CFE56.taxon	discussion	Remarks. Devolana youajla differs from other species in having an aedeagus with flanges developed and two pairs of apical processes (Figure 7 (f – g )).	en	Pinedo-Escatel, José Guadalupe Aguilar-Pérez ı Jorge Adilson, Valdez-Quezada, Brianda Catalina (2019): Three new Mexican species of the endemic Athysanini leafhopper genus Devolana DeLong (Hemiptera: Cicadellidae) from the tropical dry forest. Journal of Natural History 53 (33): 2039-2056, DOI: 10.1080/00222933.2019.1683244
