taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03B687B92D7FFFDA76BFFF09FE14A4E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901045/files/figure.png	http://doi.org/10.5281/zenodo.5901045	FIGURE 1. Morphological characters of the Bothynus villiersi species group: A, head in lateral view (mouthparts removed); B, head in frontal view; C, right maxilla in ventral view; D, head, prothoracic legs and pronotum in lateral view. Arrows indicate oblique downward clypeus in A, complete frontoclypeal ridge in B, palpomere 1 short in C, and conical process of mentum in D.	FIGURE 1. Morphological characters of the Bothynus villiersi species group: A, head in lateral view (mouthparts removed); B, head in frontal view; C, right maxilla in ventral view; D, head, prothoracic legs and pronotum in lateral view. Arrows indicate oblique downward clypeus in A, complete frontoclypeal ridge in B, palpomere 1 short in C, and conical process of mentum in D.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7FFFDA76BFFF09FE14A4E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901049/files/figure.png	http://doi.org/10.5281/zenodo.5901049	FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021).	FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021).	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7FFFDA76BFFF09FE14A4E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901053/files/figure.png	http://doi.org/10.5281/zenodo.5901053	FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype).	FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype).	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7FFFDA76BFFF09FE14A4E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901055/files/figure.png	http://doi.org/10.5281/zenodo.5901055	FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D.	FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7FFFDA76BFFF09FE14A4E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901059/files/figure.png	http://doi.org/10.5281/zenodo.5901059	FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite.	FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7FFFDA76BFFF09FE14A4E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901063/files/figure.png	http://doi.org/10.5281/zenodo.5901063	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7FFFDA76BFFF09FE14A4E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901065/files/figure.png	http://doi.org/10.5281/zenodo.5901065	FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species.	FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7FFFDA76BFFF09FE14A4E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901069/files/figure.png	http://doi.org/10.5281/zenodo.5901069	FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate.	FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7FFFDA76BFFF09FE14A4E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901071/files/figure.png	http://doi.org/10.5281/zenodo.5901071	FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record.	FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7EFFDC76BFFB6FFEA5A539.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901049/files/figure.png	http://doi.org/10.5281/zenodo.5901049	FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021).	FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021).	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7EFFDC76BFFB6FFEA5A539.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901053/files/figure.png	http://doi.org/10.5281/zenodo.5901053	FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype).	FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype).	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7EFFDC76BFFB6FFEA5A539.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901055/files/figure.png	http://doi.org/10.5281/zenodo.5901055	FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D.	FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7EFFDC76BFFB6FFEA5A539.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901059/files/figure.png	http://doi.org/10.5281/zenodo.5901059	FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite.	FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7EFFDC76BFFB6FFEA5A539.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901063/files/figure.png	http://doi.org/10.5281/zenodo.5901063	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7EFFDC76BFFB6FFEA5A539.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901065/files/figure.png	http://doi.org/10.5281/zenodo.5901065	FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species.	FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7EFFDC76BFFB6FFEA5A539.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901069/files/figure.png	http://doi.org/10.5281/zenodo.5901069	FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate.	FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D7EFFDC76BFFB6FFEA5A539.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901071/files/figure.png	http://doi.org/10.5281/zenodo.5901071	FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record.	FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D78FFDE76BFFA54FE57A7F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901049/files/figure.png	http://doi.org/10.5281/zenodo.5901049	FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021).	FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021).	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D78FFDE76BFFA54FE57A7F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901053/files/figure.png	http://doi.org/10.5281/zenodo.5901053	FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype).	FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype).	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D78FFDE76BFFA54FE57A7F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901055/files/figure.png	http://doi.org/10.5281/zenodo.5901055	FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D.	FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D78FFDE76BFFA54FE57A7F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901059/files/figure.png	http://doi.org/10.5281/zenodo.5901059	FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite.	FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D78FFDE76BFFA54FE57A7F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901063/files/figure.png	http://doi.org/10.5281/zenodo.5901063	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D78FFDE76BFFA54FE57A7F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901065/files/figure.png	http://doi.org/10.5281/zenodo.5901065	FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species.	FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D78FFDE76BFFA54FE57A7F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901069/files/figure.png	http://doi.org/10.5281/zenodo.5901069	FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate.	FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D78FFDE76BFFA54FE57A7F8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901071/files/figure.png	http://doi.org/10.5281/zenodo.5901071	FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record.	FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D75FFD076BFFB58FDB3A4AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901049/files/figure.png	http://doi.org/10.5281/zenodo.5901049	FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021).	FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021).	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D75FFD076BFFB58FDB3A4AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901053/files/figure.png	http://doi.org/10.5281/zenodo.5901053	FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype).	FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype).	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D75FFD076BFFB58FDB3A4AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901055/files/figure.png	http://doi.org/10.5281/zenodo.5901055	FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D.	FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D75FFD076BFFB58FDB3A4AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901059/files/figure.png	http://doi.org/10.5281/zenodo.5901059	FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite.	FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D75FFD076BFFB58FDB3A4AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901063/files/figure.png	http://doi.org/10.5281/zenodo.5901063	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D75FFD076BFFB58FDB3A4AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901065/files/figure.png	http://doi.org/10.5281/zenodo.5901065	FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species.	FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D75FFD076BFFB58FDB3A4AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901069/files/figure.png	http://doi.org/10.5281/zenodo.5901069	FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate.	FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D75FFD076BFFB58FDB3A4AB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901071/files/figure.png	http://doi.org/10.5281/zenodo.5901071	FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record.	FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D74FFD476BFFAA4FAAFA27C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901049/files/figure.png	http://doi.org/10.5281/zenodo.5901049	FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021).	FIGURE 2. Holotype dorsal habitus with their respective labels on the right side: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021).	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D74FFD476BFFAA4FAAFA27C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901053/files/figure.png	http://doi.org/10.5281/zenodo.5901053	FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype).	FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype).	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D74FFD476BFFAA4FAAFA27C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901063/files/figure.png	http://doi.org/10.5281/zenodo.5901063	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D74FFD476BFFAA4FAAFA27C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901071/files/figure.png	http://doi.org/10.5281/zenodo.5901071	FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record.	FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D70FFD676BFFD1DFAD3A4E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901053/files/figure.png	http://doi.org/10.5281/zenodo.5901053	FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype).	FIGURE 3. Male dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus spinophallicus (Ratcliffe & Cave, 2021); E, Bothynus belemensis new species (holotype).	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D70FFD676BFFD1DFAD3A4E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901055/files/figure.png	http://doi.org/10.5281/zenodo.5901055	FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D.	FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D70FFD676BFFD1DFAD3A4E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901059/files/figure.png	http://doi.org/10.5281/zenodo.5901059	FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite.	FIGURE 5. Spiculum gastrale: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species. hs = hemisternite.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D70FFD676BFFD1DFAD3A4E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901063/files/figure.png	http://doi.org/10.5281/zenodo.5901063	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D70FFD676BFFD1DFAD3A4E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901065/files/figure.png	http://doi.org/10.5281/zenodo.5901065	FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species.	FIGURE 7. Female dorsal habitus: A, Bothynus villiersi Endrödi, 1968; B, Bothynus horridus Endrödi, 1968; C, Bothynus alvarengai Martínez, 1983; D, Bothynus belemensis new species.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D70FFD676BFFD1DFAD3A4E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901069/files/figure.png	http://doi.org/10.5281/zenodo.5901069	FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate.	FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D70FFD676BFFD1DFAD3A4E0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901071/files/figure.png	http://doi.org/10.5281/zenodo.5901071	FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record.	FIGURE 9. Geographic distribution of the Bothynus villiersi species group. The star inscribed by a red circle indicates a possible mislabeled record.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D72FFD676BFFB6CFA3CA68A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901055/files/figure.png	http://doi.org/10.5281/zenodo.5901055	FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D.	FIGURE 4. Pronotal tubercle: A, Bothynus alvarengai Martínez, 1983; B, Bothynus horridus Endrödi, 1968. Prosternal process: C, Bothynus villiersi Endrödi, 1968; D, Bothynus belemensis new species. Arrows indicate tubercle bilobed in A, tubercle conical in B, prosternal process apex conical in C, and prosternal process apex rounded in D.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D72FFD676BFFB6CFA3CA68A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901069/files/figure.png	http://doi.org/10.5281/zenodo.5901069	FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate.	FIGURE 8. Female, detail of the anterior pronotal margin: A, Bothynus villiersi Endrödi, 1968; B, Bothynus belemensis new species. Female, internal view of the tergite 8: C, Bothynus villiersi Endrödi, 1968; D, Bothynus horridus Endrödi, 1968; E, Bothynus alvarengai Martínez, 1983; F, Bothynus belemensis. Arrows indicate arched margin in A, margin sinuous in B, and furrow in C. ip = internal plate.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
03B687B92D72FFD676BFFB6CFA3CA68A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/5901063/files/figure.png	http://doi.org/10.5281/zenodo.5901063	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	FIGURE 6. Aedeagus in lateral view and parameres in dorsal view, respectively: A–B, Bothynus villiersi Endrödi, 1968; C–D, Bothynus horridus Endrödi, 1968; E–F, Bothynus alvarengai Martínez, 1983; G–H, Bothynus belemensis new species. lmp = left medial process, rbp = right basal process, rmp = right medial process. Arrow indicating elongated parameres apex in B.	2022-01-25	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien		Zenodo	biologists	Duarte, Paulo R. M.;Grossi, Paschoal C.;Dupuis, Fabien			
