identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
CC2FF234FFD380542F4DD13E87E9FF47.text	CC2FF234FFD380542F4DD13E87E9FF47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helodrilus Hoffmeister 1845	<div><p>GENUS HELODRILUS HOFFMEISTER, 1845 EMENDED</p> <p>Type species: Helodrilus oculatus Hoffmeister, 1845.</p> <p>Species included: Helodrilus cernosƲitoƲianus (Zicsi, 1967).</p> <p>Species putatiƲely included: Helodrilus samniticus Cognetti, 1914; Helodrilus phillipei Qiu &amp; Bouché, 1998; Helodrilus turquini Qiu &amp; Bouché, 1998.</p> <p>Diagnosis: Small worms (20–80 mm long, 1–2 mm wide) without pigmentation, setae lumbricine, closely paired. The clitellum begins around segment 20 and ends before 33. The tubercles cover 2–3 segments near the end of the clitellum. Spermathecae two or three pairs in 9/10, 10/11 or 9/10–11/12. Vesicles of two pairs in 11, 12. Calciferous glands adverticulate, weakly developed. Excretory system exoic, avesiculate.</p> <p>Remarks: We have putatively included the species Helodrilus samniticus described from Italy, Helodrilus phillipei and Helodrilus turquini described from France in the emended Helodrilus based on their morphological similarity to the type species and also their west European distribution. Further molecular studies are needed to confirm these placements and, as well, the identity of the latter two species considered synonyms of Helodrilus oculatus by Szederjesi et al. (2014).</p> <p>It is difficult to distinguish Helodrilus from the genera Gatesona Qiu &amp; Bouché, 1998 sensu Marchán et al., 2021, and the new CoƲentina and Imetescolex by considering only morphological characters. However, Helodrilus differs from all of these genera in its smaller size and weakly developed or vestigial calciferous glands.</p> <p>COVENTINA MARCHÁN, SZEDERJESI &amp; CSUZDI</p></div> 	https://treatment.plazi.org/id/CC2FF234FFD380542F4DD13E87E9FF47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Szederjesi, Tímea;Marchán, Daniel F.;Csuzdi, Csaba;Sarbu, Serban M.;Pavlíček, Tomáš;Krízsik, Virág;Martin, Patrick;Domínguez, Jorge	Szederjesi, Tímea, Marchán, Daniel F., Csuzdi, Csaba, Sarbu, Serban M., Pavlíček, Tomáš, Krízsik, Virág, Martin, Patrick, Domínguez, Jorge (2023): Three in one: molecular phylogeny of the genus Helodrilus (Crassiclitellata: Lumbricidae) with a description of two new genera and two new species. Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 197 (4): 899-908, DOI: 10.1093/zoolinnean/zlac069, URL: http://dx.doi.org/10.1093/zoolinnean/zlac069
