identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C2CE00FFFEFF9C3EFA0E57FBA64363.text	03C2CE00FFFEFF9C3EFA0E57FBA64363.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cerasommatidiidae STATUS RESTORED	<div><p>CERASOMMATIDIIDAE STATUS RESTORED</p> <p>The BI, ML and MP analyses provide unequivocal support for the monophyly of Cerasommatidia clade and, therefore, we propose the restoration of family status for Cerasommatidiidae.</p> <p>Cerasommatidiidae are divided into four clades corresponding to the four genera proposed here (Fig. 2). The new monotypic genus from Madagascar, MahaƲelo, is recovered as sister-taxon to the Neotropical, type genus of the family, Cerasommatidia (= Ibicarella, synon. nov.), albeit with weak support (PP = 0.83). Morphologically, both genera share the labial palp with oval, inflated second palpomere (12:1). MahaƲelo is characterized by one uncontroverted synapomorphy, scutellar shield invisible (25:1) and one homoplastic feature, abdominal ventrite 1 without postcoxal lines (34:0). Cerasommatidia is supported by one homoplastic character: mesotrochantin at least partially exposed (28:0). Support for the clade comprising the new genera Yamuy and Karumbe is low (PP = 0.47) and lacks supporting morphological features. The new genus Karumbe is supported by the distinctly bordered pronotal base (19:1) and pronotal anterior margin with bordering line crenulate (21:1). Yamuy is characterized by a single uncontroverted synapomorphy, elytral vestiture comprising multiple setal forms (1:1) and one homoplastic character, the abdominal ventrite 1 with postcoxal lines (34:1).</p> </div>	https://treatment.plazi.org/id/03C2CE00FFFEFF9C3EFA0E57FBA64363	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFFEFF933D8D0991FE114730.text	03C2CE00FFFEFF933D8D0991FE114730.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cerasommatidiidae STAT. RESTORED WITHIN	<div><p>THE PHYLOGENETIC PLACEMENT OF CERASOMMATIDIIDAE STAT. RESTORED WITHIN THE COCCINELLID LINEAGE OF COCCINELLOIDEA</p> <p>The results from our molecular (Fig. 1), morphological (Supporting Information, File S12a–c) and combined data (molecules and morphology) (Fig. 2) analyses strongly and unambiguously support the status of Cerasommatidiidae as a distinct clade at the family level. Morphologically, the monophyly of Cerasommatidiidae is supported by several characters (Fig. 2) (listed in Results) that set cerasommatidiids apart from Eupsilobiidae and other coccinelloid groups. In particular, the ovipositor lacking coxites is unique to this group and has not been found in any other Coccinelloidea. The sister-relationship of Cerasommatidiidae with Mycetaeidae is strongly supported in both the molecular and combined analyses. Yet, only one morphological, homoplastic character was recovered in the combined analyses as supporting this relationship: the mandible with subapical teeth or serrations (9:0). At this point it is difficult to elucidate shared morphological characters between members of these two families.</p> <p>It is noteworthy that the position of Eupsilobiidae differs between the combined and molecular trees. In the molecular tree (Fig. 1) it is recovered as the sistergroup to Coccinellidae with strong support (PP = 1.0), whereas in the combined analysis it forms the sistergroup to the Mycetaeidae + Cerasommatidiidae, although with low nodal support (PP = 0.69) and no corresponding anatomical synapomorphies.</p> <p>The monophyly of Eupsilobiidae is supported by two uncontroverted synapomorphies: aedeagus with penis coiled (38:1) and ovipositor with infundibulumlike structure/sperm duct modified (41:1) and by one homoplastic character: labial mentum with large, triangular, raised area (11:2). The position of Eupsilobiidae within the coccinellid group, as well as the internal family relationships, remains uncertain and ought to be readdressed in future studies. The genus Chileolobius, the only member of Eupsilobiidae represented in the molecular analysis, was supported in our combined data analysis as sister-group to the rest of Eupsilobiidae (‘core Eupsilobiidae’ composed of Eidoreus, EƲolocera, Microxenus and Natalinus). This result could be an artefact of the remaining eupsilobiid exemplars being represented in the combined data analysis by morphological data only. However, Chileolobius is an enigmatic taxon, exhibiting several morphological features shared with all or some Coccinellidae but no other Eupsilobiidae, including frontoclypeal suture absent (6:0), mandibular mola absent or reduced (7:1) and mesotrochantin concealed (28:1). The core eupsilobiids are recovered with strong nodal support (PP = 0.99) and multiple anatomical apomorphies, including antennae composed of ten articles (4:2) and mesotrochantin at least partially exposed (28:0). A closer relationship between Eidoreus and Microxenus, with respect to Chileolobius and Ibicarella, was previously hypothesized by Pakaluk &amp; Ślipiński (1990). Including molecular data for exemplars of core Eupsilobiidae will be the next step to elucidate their relationships with Chileolobius and corroborate the position of Eupsilobiidae among the coccinellid group of families.</p> <p>The results of this study demonstrate the separation of Cerasommatidiidae from Eupsilobiidae, and justify the restoration of this lineage to family status. Cerasommatidiidae are corroborated by both high topological support as the sister-group to Mycetaeidae in the molecular and combined analysis and by their unique morphological features, as outlined above.</p> <p>The relationships between species of Cerasommatidiidae, as revealed by the morphological and combined analyses, justify the recognition of four genera despite an unclear relationship between them. Cerasommatidia in the new sense (including Ibicarella) is a homogeneous group, with species differing only in the structures of the male genitalia, the body proportions and the degree of development of the pronotal border. A sister-relationship recovered for Cerasommatidia + MahaƲelo is based on single homoplastic character: labial palp with palpomere 2 oval, inflated (12:1) and is not strongly supported (PP = 0.83). Similarly, the sisterrelationship of Yamuy with Karumbe has negligible nodal support (PP = 0.47) and no morphological characters to bolster the relationship.</p> </div>	https://treatment.plazi.org/id/03C2CE00FFFEFF933D8D0991FE114730	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFF1FF923D6B0F66FBFF4307.text	03C2CE00FFF1FF923D6B0F66FBFF4307.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cerasommatidiidae BRETHES 1925	<div><p>CERASOMMATIDIIDAE BRÈTHES, 1925 STAT. RESTORED</p> <p>(FIGS 1–20)</p> <p>Cerasommatidiidae Brèthes, 1925: 199.</p> <p>Type genus: Cerasommatidia Brèthes, 1925 by original designation.</p> <p>As synonym of Eupsilobiinae Casey, 1895: 452.– Pakaluk et al., 1994: 228; Shockley et al., 2009: 27; Bouchard et al., 2011: 372.</p> <p>Diagnostic combination: Body short oval (0.80– 1.45 mm) and convex (Figs 3, 6, 12, 17). Head with frontoclypeal suture present (Figs 9A, 15B, 18B). Eyes coarsely faceted (Figs 4A, 5A, 7A, 9A, 15B, 16A, 17D). Antennal sockets visible from above (Figs 4A, 5A, 9A, 16A). Ventral antennal grooves deep between eyes and mouthparts (Figs 3C, 18B). Gular sutures short, widely separated, weakly convergent anteriorly (Figs 7A, 15A). Tentorium with anterior arms fused medially and widely divergent anteriorly, posterior arms connected throughout their length to posterior weakly sinuate transverse bridge/corpotentorium (Figs 3C, 7A, 17D). Antenna composed of 11 antennomeres with distinct two-antennomere club (Figs 3C, 5E, 7A, 11A, 12C, 13A, 15A, 18A). Labrum large, sparsely setose, rounded anteriorly with or without small, median emargination (Figs 7B, 10A, 13B, 15B, 18B). Maxillary palpi (Figs 3C, 5B, 7C, 9B, 10A, 13C, 15A, 17E) with four palpomeres: palpomere 1 small; palpomere 2 weakly transverse and somewhat bulbous; palpomere 3 transverse; terminal palpomere distinctly elongate, swollen near base and then narrowing/flattened laterally toward apex; apex with short, sclerotized sensilla. Mandible stout (Figs 7E, 13E, 17F), sclerotized, with two apical teeth and two small subapical teeth; prostheca large, membranous with fringe of stout setae along its inner margin and with a series of long sclerotized, teeth-like projections apically; mola large, strongly sclerotized, with distinct transverse ridges. Labium with mentum irregularly hexagonal, with anterior margin straight, surface flat; prementum weakly elongate, sclerotized with ligula submembranous, lobed at sides (Figs 5B, 7D, 9B, 10A, 15A). Palpi with three palpomeres, widely separated by a distance more than width of palpiger: palpomere 1 smallest, palpomere 2 bulbous, palpomere 3 sometimes slightly bulbous.</p> <p>Pronotum strongly transverse, widest at or near base and strongly convergent anteriorly (Figs 3A, 4B, 5C, 10C, 11B, 18C). Front angles weakly produced anteriorly, obtuse; hind angles often with small, oblique indentation to receive humeral corner of elytron. Prosternum large, in form of chin piece covering mouthparts (Figs 5D, 7F, 9D, 10B, 11C, 15D, 16C, 18D); prosternal process narrowest in about its half-length, continuously widened apically, provided with raised, paired lateral carinae or single median carina. Hypomeron with deep and long antennal grooves (Figs 7F, 9D, 10B, 15D, 18D). Mesoventral intercoxal process widely separates mesocoxae. Mesocoxa circular in outline, its cavity outwardly open; trochantin concealed (Figs 3D, 4E, 5H, 7I, 9F, 10E, 11F, 13D, 15G, 16D, 17G, 18E). Meso-metaventral junction arcuate anteriorly. Metaventrite strongly transverse, weakly convex, without postcoxal lines. Metanepisternum with small, outer blunt projection near anterior margin (Figs 3D, 7I, 17G). Metacoxae transverse, widely separated (Figs 3D, 7I, 9F, 10F, 16D, 18E). Metendosternite with short stalk and widely separated anterior arms and anterior tendons (Fig. 7J, 13D). Epipleura not foveate. Legs with trochanterofemoral attachment subheteromeroid (Figs 3D, 5H, 7K, 9F, 10F, 11F, 13D, 15D, 16D, 18E). Tibiae slender, gradually weakly widening towards tarsus; surrounded by short, stout spines. Tarsal formula 4-4- 4 in both sexes with tarsomere 1 weakly and tarsomere 2 distinctly lobed ventrally, tarsomere 3 short and tarsomere 4 about as long as tarsomere 1 (Figs 4F, 5F, 7L, 10D, 11E, 15E, 18G). Claws toothed at base (Figs 5F, 13F, 15E, 16E).</p> <p>Abdomen with six ventrites in both sexes (Figs 4G, 8M, 15H, 18G). Ventrite 1 with (Figs 3E, 4G, 5H, 7M) or without distinctly developed, rounded postcoxal lines (Figs 15H, 16F, 18G); at midline about as long as three next ventrites together. Ventrite 5 with posterolateral angles strongly produced backwards and somewhat overlapping antero-lateral parts of ventrite 6 (anterior margin of ventrite 6 much narrower than posterior margin of ventrite 5); ventrite 6 arcuate apically, surrounded by anterior margin of tergite VIII bent downwards, appearing as narrow, false ‘ventrite 7’. Male genital segment (IX) reduced but with strongly developed, sclerotized, long, narrow, single apophysis (Figs 3E, 8A, D, 14A, D, 17I). Aedeagus (Figs 3F, 8B, E, 14B, E, 17H) with penis resting on its side when retracted.Ovipositor of female genitalia without coxites (Figs 8C, G, 14C, F, 17J).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFF1FF923D6B0F66FBFF4307	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFF0FF963D8509BBFE1E4633.text	03C2CE00FFF0FF963D8509BBFE1E4633.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cerasommatidia Brethes 1925	<div><p>CERASOMMATIDIA BRÈTHES, 1925</p> <p>(FIGS 3–5)</p> <p>Cerasommatidia Brèthes, 1925: 199.</p> <p>Type species, by monotypy, Cerasommatidia arroaei Brèthes, 1925. – Pakaluk et al., 1994: 228; Shockley et al., 2009: 27.</p> <p>Ibicarella Pakaluk &amp; Ślipiński, 1990: 717, synon. nov.</p> <p>Type species, by original designation, Ibicarella plaumanni Pakaluk and Ślipiński, 1990. – Pakaluk et al., 1994: 228; Shockley et al., 2009: 27.</p> <p>Diagnosis: Cerasommatidia can be separated from other genera of Cerasommatidiidae by the following combination of characters: dorsal surface of the body covered by inconspicuous tiny decumbent hairs (often broken or missing) (Figs 3A, B, 4D, 5C, G), pronotum with anterior and lateral sides narrowly marginate (Figs 4B, 5C); prosternal process moderately wide (Fig. 5D); trochanters flattened and expanded posteriorly to cover the tibio-tarsal joint in repose (Figs 4E, G, 5H); abdominal ventrite 1 with complete, semicircular postcoxal lines (Figs 3E, 4G, 5H); aedeagus with tegmen membranous, not fused with penis (Fig. 3E, F). Cerasommatidia resembles MahaƲelo in the body shape, coloration, short, inconspicuous dorsal vestiture and the pronotal lateral margins without internal carina. However, it can be distinguished from the later by having the prosternal process with a pair of lateral carinae extending anteriorly beyond level of anterior margin of procoxal cavities (a single, long, median, longitudinal carina present in MahaƲelo), hind angles of pronotum without indentation (in MahaƲelo hind angles of pronotum possess small indentation to receive humeral corner of elytron), scutellar shield visible (invisible/absent in MahaƲelo) and complete semicircular femoral lines in the abdominal ventrite 1.</p> <p>Description: Length 1.15–1.70 mm. Body moderately short oval, convex, 1.3–1.4 times as long as wide, 1.8–1.9 times as long as high. Coloration homogeneous: dark brown to black (Fig. 3A, B).</p> <p>Head with dorsal surface uniformly covered with short fine setae (Figs 4A, 5A). Clypeus large, rectangular. Frontoclypeal suture present but feebly marked, almost straight. Ventral antennal grooves short, reaching close to posterior eye level (Fig. 5B). Antenna approximately 0.35 of length of body (Figs 3A, C, 5E); antennal club one quarter of total antennal length; antennomeres 1–5 longer than wide, with antennomere 5 longer than antennomere 6; antennomeres 6–9 variable. Eyes comparatively small, moderately prominent (Figs 3C, 4A, 5A). Galea large, densely setose apically. Lacinia weakly narrower than galea with dense setae at apex and inner margin. Labium (Fig. 5B) with mentum widest near midlength or basal third, palpomere 2 and 3 large, bulbous, somewhat oval and subrectangular respectively; terminal palpomere short and subtruncate to weakly elongate, acuminate, 1.2–1.5 times as long as wide.</p> <p>Prothorax. Pronotum 2.2–2.5 times as wide as long, widest at base and strongly convergent anteriorly, 1.9–2.2 times wider at widest part than on front angles (Figs 4B, 5C). Anterior margin (at least partly) and lateral margins narrowly bordered; base of pronotum regularly rounded, without distinct bordering line or with faint bordering line present medially. Pronotal sides scarcely rounded; hind angles without indentation to receive humeral corner of elytron. Pronotal disc moderately convex, smooth, with sparse fine punctures provided with thin, tiny hairs. Prosternum (Fig. 5D) with anterior margin weakly arcuate posteriorly; prosternal process extending posteriorly to hind level of procoxae, comparatively wide, at apex about 0.7–0.8 of the width of procoxal cavity; with raised lateral carinae reaching anteriorly well beyond level of anterior margin of procoxal cavities; area between carinae weakly concave. Hypomeron with deep, long, straight antennal grooves (Fig. 5D).</p> <p>Pterothorax. Mesonotum with scutellar shield small, transverse, weakly rounded at apex (Fig. 5C). Mesoventrite (Figs 3D, 4E, 5H) with intercoxal process smooth, almost flat with anterior raised border incomplete medially; 1.2–1.4 times as wide as mesocoxal diameter. Metaventrite (Figs 3D, 4E, 5H) as long as abdominal ventrites 1–3 together; with remnants of discrimen posteriorly; anterior margin with bordering carina widening towards lateral corners; central area with small to large and deep setiferous punctures. Anterior part of metanepisternum with small outer blunt like projection (Fig. 3D). Elytra 0.85–1.00 mm long (Figs 3A, 4C, 5G), about as long a wide, 2.9–3.3 times as long and 1.2 times as wide as pronotum, with lateral margins narrowly explanate. Surface with small setiferous punctures bearing tiny, thin setae. Epipleura almost reaching elytral apex but incomplete, narrow, with internal bordering line narrow, present from the level of mid coxae to apex. Hindwings reduced or absent.</p> <p>Legs (Fig. 3D). Trochanters angulately produced posteriorly. Meso- and metatrochanters with posterior margin as a conspicuous laminar expansion that covers tibio-tarsal joint in repose. Femora flattened (mid and hind femora more distinctly than fore femora); with grooves for tibiae present throughout whole length. Claws with comparatively small blunt tooth at base (Figs 4F, 5F).</p> <p>Abdomen (Figs 3E, 4G) with ventrite 1 anteriorly with bordering carina forming distinct, rounded and complete laterally postcoxal lines, extending posteriorly beyond midlength of ventrite 1.</p> <p>Male genitalia (Fig. 3E, F). Aedeagus with penis long and narrow, sclerotized, curved or asymmetrically sinuate, not-ramificate apically. Tegmen: large, in form of submembranous ring; tegminal strut long, membranous, flattened.</p> <p>Female genitalia not studied.</p> <p>Species included: Cerasommatidia arroaei Brèthes, 1925, C. plaumanni (Pakaluk &amp; Ślipiński, 1990), C. rotundata (Pakaluk &amp; Ślipiński, 1990).</p> <p>Distribution: Brazil (Fig. 20B).</p> <p>Comment: When Pakaluk &amp; Ślipiński (1990) described Ibicarella in Eupsilobiinae they were not aware of its resemblance to Cerasommatidia Brèthes, 1925. Subsequently, when the holotype of Cerasommatidia arroaei was rediscovered its resemblance with Ibicarella was noticed and served as basis for the synonymy of Cerasommatidiidae under Eupsilobiinae (Pakaluk et al., 1994). Nevertheless, in part due to the state of preservation of the holotype of C. arroaei, its morphology was not thoroughly studied and both Cerasommatidia and Ibicarella were retained as valid genera. In the present study, we carefully examined the morphology of the type material of all Cerasommatidia and Ibicarella species and, following the results of our morphological (Supporting Information, File S12a–c) and combined (Fig. 2) analyses, we consider both represent the same genus and Ibicarella Pakaluk &amp; Ślipiński is considered a subjective junior synonym of Cerasommatidia Arrow.</p></div> 	https://treatment.plazi.org/id/03C2CE00FFF0FF963D8509BBFE1E4633	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFF4FF963D740BCEFB3A47F9.text	03C2CE00FFF4FF963D740BCEFB3A47F9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cerasommatidia arrowi Brèthes 1925	<div><p>CERASOMMATIDIA ARROWI BRÈTHES, 1925</p> <p>(FIGS 3A, F, 4A–G, 20B)</p> <p>Cerasommatidia arroaei Brèthes, 1925: 201. – Pakaluk et al., 1994: 228; Shockley et al., 2009: 27.</p> <p>Material examined: Holotype, male, BRAZIL: Rio de Janeiro (Corcovado), 11.V.1912 // Cerasommatidia arroaei nov. spec. Brèthes (BMNH).</p> <p>Diagnosis: Cerasommatidia arroaei is most similar to C. plaumanni in overall appearance and by sharing the anterior bordering line of the pronotum disappearing medially (Fig. 4B). However, it can be distinguished from C. plaumanni by having the posterior margin of the pronotum not bordered and by the aedeagus regularly curved and possessing a much longer tegmen (Fig. 3F) than in C. plaumanni.</p> <p>Description: Body: length 1.4 mm, 1.4 times as long as wide, 1.8 times as long as high, short oval and moderately convex, black with dark-brown legs, antennae and palpi (Fig. 3A).</p> <p>Antenna nearly 0.35 of length of body; antennomeres 1–5 longer than wide; antennomeres 6, 7, 9 subquadrate and antennomere 8 slightly transverse.Apical labial palpomere short and subtruncate, 1.3 times as long as wide.</p> <p>Pronotum 2.2 times as wide as long, 2.2 times as wide at widest part than at front angles (Figs 3A, 4B). Anterior margin with fine bordering line vanishing medially; lateral margins narrowly, gently bordered; base not bordered. Prosternal process with apex about 0.65 of width of procoxal cavity, narrowest near halflength, weakly widened apically, with raised lateral carinae, reaching almost apical quarter of prosternum, with central part between carinae slightly depressed.</p> <p>Elytra 1.0 mm long, about as long a wide, 2.9 times as long and 1.2 times as wide as pronotum; lateral margins visible from above at basal two-thirds (Fig. 4C). Metaventrite with fine setiferous punctures (Fig. 4E). Mesoventral process about 1.1 times as wide as mesocoxal diameter (Fig. 4E).</p> <p>Legs. Meso- and metatrochanters flattened, weakly angulately produced posteriorly.</p> <p>Male genitalia (Fig. 3F). Aedeagus with penis long and comparatively narrow, sclerotized, curved, with simple apex. Tegmen large, submembranous, long; tegminal strut large, flattened, membranous.</p> <p>Female unknown.</p> <p>Distribution: Brazil (Fig. 20B).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFF4FF963D740BCEFB3A47F9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFF4FF953D470DB6FDBA47D3.text	03C2CE00FFF4FF953D470DB6FDBA47D3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cerasommatidia plaumanni (Pakaluk & Slipinski 1990) COMB. NOV.	<div><p>CERASOMMATIDIA PLAUMANNI (PAKALUK &amp; ŚLIPIŃSKI, 1990) COMB. NOV.</p> <p>(FIGS 3B–E, 5A–H, 20B)</p> <p>Ibicarella plaumanni Pakaluk &amp; Ślipiński, 1990: 718. – Shockley et al., 2009: 28.</p> <p>Material examined: Paratypes: B RAZIL: Santa Catarina: Nova Teutonia, F. Plaumann, X.72/ Ibicarella plaumanni sp. n., det. S.A. Ślipiński 87/ Paratype (2 ex., MIZ); same locality data/ Cerylonid group? Endomychidae, Ibicarella sp. [R. A. Crowson adult Coleoptera slide] (1 male, BMNH).</p> <p>Diagnosis: Cerasommatidia plaumanni is most similar to C. arroaei in overall appearance and by sharing the anterior bordering of the pronotum almost completely absent or becoming feebler medially. However, it can be distinguished by having the posterior margin of the pronotum distinctly bordered (Fig. 5C) and the aedeagus sinuate with a much shorter tegmen (Fig. 3E) than in C. arroaei (Fig. 3F).</p> <p>Description: Body: length 1.15 mm, 1.25 times as long as wide, 1.8 times as long as high, short oval and moderately convex, black with dark-brown legs, antennae and palpi.</p> <p>Antenna nearly 0.35 of length of body (Fig. 3B); antennomeres 1–5 longer than wide; antennomeres 6, 7, 9 subquadrate and antennomere 8 slightly transverse (Fig. 5E). Apical labial palpomere short and briefly truncate, 1.2 as long as wide (Fig. 5B).</p> <p>Pronotum 2.5 times as wide as long, 1.9 times wider at widest part than on front angles (Fig. 5C). Anterior margin with fine bordering line vanishing medially; lateral margins narrowly, gently bordered. Base bordered with feeble carina. Prosternal process (Fig. 5D), with apex about 0.75 of width of procoxal cavity, narrowest near half-length, weakly widened apically, with lateral carinae raised, almost reaching apical quarter of prosternum, with central part between carinae slightly depressed.</p> <p>Elytra 0.85 mm long, about as long a wide, 3.3 times as long and 1.2 times as wide as pronotum (Fig. 5G); lateral margins visible from above at basal two-thirds (Figs 3B, 5G). Mesoventral process about 1.1 times as wide as mesocoxal diameter (Fig. 3D). Metaventrite with fine setiferous punctures (Fig. 5H).</p> <p>Legs. Meso- and metatrochanters flattened, weakly angulately produced posteriorly (Fig. 5H).</p> <p>Abdomen with male genital segment as in Fig. 3E.</p> <p>Male genitalia. Aedeagus with penis long and comparatively narrow, sclerotized, sinuate, curved near base and at apical third, with apex not ramificate (Fig. 3E). Tegmen large, submembranous, long; tegminal strut large, flattened.</p> <p>Female unknown.</p> <p>Distribution: Brazil (Fig. 20B).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFF4FF953D470DB6FDBA47D3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFF7FF953EC10DE5FB2144EF.text	03C2CE00FFF7FF953EC10DE5FB2144EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cerasommatidia rotundata (Pakaluk & Slipinski 1990) COMB. NOV.	<div><p>CERASOMMATIDIA ROTUNDATA (PAKALUK &amp; ŚLIPIŃSKI, 1990) COMB. NOV.</p> <p>Ibicarella rotundata Pakaluk &amp; Ślipiński, 1990: 719. – Shockley et al., 2009: 28.</p> <p>Material examined: BRAZIL: Santa Catarina: Ibicare, 600 m, 27.09’ x 51.18’/ Sept. 1960, F. Plaumann, X.72/ Ibicarella rotundata sp. n., det. S. A. Ślipiński / Holotypus (ANIC).</p> <p>Diagnosis: Cerasommatidia rotundata can be distinguished from C. arroaei and C. plaumanni by its larger body size, the anterior pronotal margin with complete bordering line (vanished medially in other species of the genus); and the lateral margins of the elytra visible from above only for basal half length (for basal two-thirds in C. arroaei and C. plaumanni).</p> <p>Description: Body: length 1.70 mm, 1.3 times as long as wide, 1.2 times as long as high, short oval and moderately convex, black with dark-brown legs, antennae and palpi.</p> <p>Antenna nearly 0.35 of length of body; antennomeres 1–5 longer than wide; antennomeres 6, 7, 9 subquadrate and antennomere 8 slightly transverse. Apical labial palpomere short and rounded apically.</p> <p>Pronotum 3.0 times as wide as long. Anterior margin with complete bordering line. Pronotal sides narrowly bordered. Base not bordered. Prosternal process wide, with apex about 0.75 of width of procoxal cavity, narrowest near half-length, weakly widened apically, with lateral carinae raised, almost reaching apical quarter of prosternum, central part between carinae slightly depressed.</p> <p>Elytra 0.9 mm long, about as long a wide. Elytral lateral margins visible from above at basal half. Mesoventral process about 1.1 times as wide as mesocoxal diameter. Metaventrite with fine setiferous punctures.</p> <p>Legs. Meso- and metatrochanters flattened, weakly angulately produced posteriorly.</p> <p>Male genitalia. Unknown.</p> <p>Female genitalia. Not studied.</p> <p>Distribution: Brazil (Fig. 20B).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFF7FF953EC10DE5FB2144EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFF7FF8E3D7C0ECEFD47457E.text	03C2CE00FFF7FF8E3D7C0ECEFD47457E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Karumbe Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček 2023	<div><p>KARUMBE ARRIAGA- VARELA, TOMASZEWSKA &amp; SZAWARYN GEN. NOV.</p> <p>(FIGS 6–11)</p> <p>Zoobank registration: urn: lsid: zoobank. org:act: 50E490DD-3714-457E-9E27-69DABA334956:</p> <p>Type species: Karumbe geiseri sp. nov. by present designation.</p> <p>Diagnosis: Karumbe can be distinguished from other genera in Cerasommatidiidae by the following combination of characters: dorsal surfaces of head, pronotum and elytra homogeneously covered by moderately long suberect setae (Figs 6A–F, 9E, 10C, 11B, D); lateral margins of pronotum with an internal subparallel carina demarcating a wide area, and anterior pronotal margin bordered with internally crenulate bordering line (Figs 9C, 10C, 11B); prosternal process moderately to distinctly wide, possessing long lateral carinae extending anteriorly much beyond prosternal process (Figs 7F, 9D, 10B, 11C); trochanters flattened and expanded posteriorly to cover the tibio-tarsal joint (Figs 7I, K, 9F, 10E, F, 11F); abdominal ventrite 1 with complete laterally, semicircular postcoxal lines (Figs 7M, 11F); aedeagus with tegmen fused at base with penis (Fig. 8B, E). The crenulate bordering line of the anterior margin of the pronotum (Figs 9C, 10C, 11B) (simple in Cerasommatidia, Yamuy and MahaƲelo), distinctly and comparatively widely bordered pronotal base and the homogeneous moderately long suberect setae on dorsal surface are peculiar to this genus (tiny inconspicuous setae are present in Cerasommatidia and MahaƲelo, while Yamuy has two kinds of vestiture: tiny decumbent hairs and sparse, long erect setae).</p> <p>Description: Length 0.90–1.45 mm. Body short oval and moderately convex, 1.25–1.30 times as long as wide, around 2.0 times as long as high. Coloration light to dark brown, usually with paler legs, antennae and palpi (Fig. 6A–F).</p> <p>Head with dorsal surface homogeneously covered with moderately long setae (Fig. 9A). Clypeus large, rectangular (Fig. 9A). Frontoclypeal suture distinct, straight. Ventral antennal grooves short, not extending to posterior eye level (Fig. 9A). Antenna approximately 0.35–0.40 of body length; antennal club (Figs 7A, 11A) one quarter of total antennal length; antennomeres 1–5 longer than wide, with antennomere 5 longer than neighbouring antennomeres 4 and 6; antennomeres 6–9 variable. Eyes comparatively large, moderately prominent (Figs 7A, 9A). Galea large, densely setose at apex (Fig. 7C). Lacinia weakly narrower than galea with dense setae at apex and inner margin. Labium with mentum widest near midlength or basal third; terminal palpomere short and rounded to elongate and acuminate, 1.2–2.2 times as long as wide at base (Figs 7D, 9B, 10A).</p> <p>Prothorax. Pronotum 2.1–2.2 times as wide as long, widest at base and strongly convergent anteriorly, 1.7– 1.9 times wider at widest part than on front angles (Figs 9C, 10C, 11B). Anterior margin bordered with densely crenulate bordering line; base of pronotum sinuate with margin comparatively widely bordered with less distinctly crenulate, well-developed bordering line. Pronotal sides scarcely rounded, area between lateral edge and carina weakly to distinctly concave, wide and continuously weakly narrowing posteriorly; hind angles with small, oblique indentation to receive humeral corner of elytron. Pronotal disc moderately convex, with sparse punctures provided with moderately long, thick setae. Prosternum with anterior margin scarcely arcuate posteriorly (Figs 7F, 9D, 10B, 11C); prosternal process comparatively wide, at apex about 0.7–0.9 of width of procoxal cavity, with raised lateral carinae extending anteriorly well beyond level of anterior margin of procoxal cavities; with area between carinae weakly concave; prosternal process extending posteriorly to hind level of procoxae. Hypomeron with deep and long, weakly sinuate antennal grooves (Figs 7F, 9D, 10B, 11C).</p> <p>Pterothorax.Scutellar shield small, transverse, weakly rounded at apex (Fig. 7G). Mesoventrite with intercoxal process smooth, almost flat with anterior raised border incomplete medially (Figs 7I, 9F, 10E, 11C), 1.1–1.4 times as wide as mesocoxal diameter. Metaventrite as long as abdominal ventrites 1–3 together; with remnants of short discrimen (Fig. 7I); anterior margin bordered with carina weekly widening towards lateral corners, at intercoxal process bordering line weakly crenulate; central area with small to large and deep setiferous punctures. Anterior part of metanepisternum with small outer blunt projection (Fig. 7I). Elytra 0.65–1.35 mm long, about as long as wide, 2.7–3.1 times as long and 1.4 times as wide as pronotum, with lateral margins scarcely to not explanate (Fig. 6A, C, E). Surface with moderately dense and deep setiferous punctures (Figs 9E, 11D), deeper and larger than those on pronotum, bearing moderately long suberect setae; with additional intermixed shallow foveolate punctures. Epipleura moderately wide, narrowing to apex, complete at apex, with internal bordering line narrow, present from the level of mid coxae to apex. Hindwings well developed, without anal lobe, with single reduced anal vein, and median fleck present, undivided (Fig. 7H).</p> <p>Legs. Trochanters roundly or angulately produced posteriorly (Figs 9D, F, 10E, F, 11F). Meso- and metatrochanters with posterior margin as a laminar expansion that covers tibio-tarsal joint when retracted. Femora flattened (mid and hind femora more distinctly than fore femora) with grooves for tibiae present throughout whole length. Claws with distinct quadrate tooth at base (Fig. 7L).</p> <p>Abdomen with ventrite 1 anteriorly with marginal carina forming distinct, rounded and complete laterally postcoxal lines, extending posteriorly to nearly midlength of ventrite 1 (Figs 7M, 11F).</p> <p>Male genitalia (Fig. 8B, E). Aedeagus with penis short and stout, sclerotized, curved apically, asymmetric, with ramificate apex (Fig. 8B, E). Tegmen fused to penis, submembranous, throneshaped, short or basally elongate; tegminal strut short, reduced.</p> <p>Female genitalia. Spermatheca moderately large, elongate-reniform, submembranous; sperm duct short; accessory gland small membranous, of irregular shape (Fig. 8C, F, G).</p> <p>Etymology: Karumbe is the Guaraní word for ‘turtle’. Gender feminine.</p> <p>Species included: Karumbe brethesi, K. geiseri and K. pakaluki.</p> <p>Distribution: Brazil, Grenada, Venezuela (Fig. 20A, B).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFF7FF8E3D7C0ECEFD47457E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFECFF8D3EFB0F68FDBA41B4.text	03C2CE00FFECFF8D3EFB0F68FDBA41B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Karumbe brethesi Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček 2023	<div><p>KARUMBE BRETHESI ARRIAGA- VARELA, TOMASZEWSKA &amp; SZAWARYN SP. NOV.</p> <p>(FIGS 6A, B, 8G, 9A–F, 20B)</p> <p>Zoobank registration: urn: lsid: zoobank. org:act: A0499A2C-83A5-4C42-8422-B22FF60AB8F4</p> <p>Type material: Holotype, female, BRAZIL: São Paulo, Bras. Mráz Lgt, Mus. Pragense (NMPC). Paratypes: Same data as holotype (two females: 1, NMPC; 1, MIZ); Rio de Janeiro, Alte Bondlin, ex coll. Kessel, Mus. Zool. Polonicum Warszawa 19/46 (one female, MIZ); Bocaina, ex coll. Kessel, Mus. Zool. Polonicum Warszawa 19/46 (one female, MIZ).</p> <p>Diagnosis: Karumbe brethesi can be easily distinguished from K.pakaluki and K.geiseri by having larger body (Fig. 6A, B), peculiar punctations on dorsal surface of the body (Fig. 9E) (vs. setiferous punctures and additional round depressions in K.geiseri and K.pakaluki), the central area of the metaventrite covered with setiferous punctures inserted in wide foveate impressions (Fig. 9F) (simple setiferous punctures in K. pakaluki and K. geiseri), the pronotal disc with elongate, narrow concave area along the internal lateral carina (Fig. 9C), prosternal process with carinae convergent anteriorly (Fig. 9D) (subparallel in K. pakaluki and K. geiseri), and the terminal labial palpomere elongate and acuminate, 2.2 times as long as wide (Fig. 9B) (in K. pakaluki and K. geiseri: short and apically rounded, 1.2–1.4 times as long as wide).</p> <p>Description: Body: length 1.40–1.45 mm, 1.3 times as long as wide, 2.0 times as long as high, short oval and moderately convex, dark brown with infuscateyellowish legs, antennae and palpi (Fig. 6A, B).</p> <p>Antenna nearly 0.35 of length of body; antennomeres 1–7 longer than wide; antennomeres 8–9 subquadrate. Apical labial palpomere elongate and acuminate, 2.2 times as long as wide at its base (Fig. 9B).</p> <p>Pronotum 2.0 times as wide as long, 1.9 times wider at widest part than on front angles (Fig. 9C). Area between lateral edge and internal lateral carina concave, wide and continuously weakly narrowing posteriorly. Anterior crenulate margin moderately wide. Pronotal disc deeply concave along internal lateral carina. Basal margin bordered with weakly crenulate bordering line. Prosternal process, narrowest near half-length, weakly widened apically with apex about 0.7 of the width of procoxal cavity (Fig. 9D); with lateral carinae convergent anteriorly (comparatively narrowly separated throughout most of their length) almost reaching anterior quarter length of prosternum, central part depressed.</p> <p>Elytra 1.00– 1.35 mm long, about as long a wide, 2.6– 2.7 times as long and 1.4 times as wide as pronotum; lateral margins visible from above. Elytral dorsal surface with most punctures with peculiar raised sculpture consisting of a small ring and an accessory point posteriorly (Fig. 9E). Mesoventral process about 1.1 times as wide as mesocoxal diameter. Metaventrite with setiferous punctures at central area placed in large and shallow foveate impressions (Fig. 9F). Hindwings present, well developed (Fig. 7H).</p> <p>Legs. Pro-trochanters rounded; meso- and metatrochanters flattened and angulately produced posteriorly (Fig. 9F).</p> <p>Abdomen with ventrite 1 with comparatively shallow, rounded postcoxal lines (Fig. 9F). Ventrite 5 with posterior margin straight.</p> <p>Female genitalia (Fig. 8G). Bursa copulatrix large, with apical outlet of sperm duct. Spermatheca submembranous, moderately large, elongatesubcylindrical, oblong; sperm duct short; accessory gland small, membranous, of irregular shape.</p> <p>Male unknown.</p> <p>Etymology: This species is dedicated to Jean Brèthes, French/Argentinian entomologist who established the family Cerasommatidiidae in 1925.</p> <p>Distribution: Brazil (Fig. 20B).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFECFF8D3EFB0F68FDBA41B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFEFFF8D3EB10A07FB014550.text	03C2CE00FFEFFF8D3EB10A07FB014550.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Karumbe geiseri Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček 2023	<div><p>KARUMBE GEISERI ARRIAGA- VARELA, TOMASZEWSKA &amp; SZAWARYN SP. NOV.</p> <p>(FIGS 6C, D, 7F, I, L, 8A–C, 10A–F, 20A)</p> <p>Zoobank registration: urn: lsid: zoobank. org:act: 57FC49EB-6890-4AF8-BB33-70F66B4D466C</p> <p>Type material: Holotype, male, GRENADA: Balthasar (Windward side), Grenada, W.I., H. H. Smith (BMNH). Paratypes: Same data as for holotype (one male, one female and 1 ex., BMNH; 2 ex, MIZ).</p> <p>Diagnosis: Karumbe geiseri is most similar to K. pakaluki, but it can be distinguished from the later by the wider prosternal process being about 0.90 of width of procoxa (Fig. 10B) (0.67 in K. pakaluki); pronotum with postero-lateral indentation shorter and more rectangular (Fig. 10C) (longer and more oblique in K. geiseri) and by the shape of the aedeagus. From K. brethesi it can be easily distinguished by having dorsal surface of the body covered with setiferous punctures and additional foveolate depressions (in K. brethesi peculiar punctations as in Fig. 9E), the central area of the metaventrite covered with simple setiferous punctures without wide foveate impressions (Fig. 10E), the pronotal disc only weakly concave along the internal lateral carina, and the terminal labial palpomere short and apically rounded, 1.4 times as long as wide (Fig. 10A) (in K. brethesi: elongate and acuminate, 2.2 times as long as wide at base).</p> <p>Description: Body: length 0.90–0.96 mm, 1.25 times as long as wide, 2.0 times as long as high, short oval and moderately convex, brown with yellowish legs, antennae and palpi (Fig. 6C, D).</p> <p>Antenna nearly 0.40 times as long as body; antennomeres 1–5 longer than wide; antennomeres 6–9 subquadrate. Apical labial palpomere short and rounded, 1.4 as long as wide (Fig. 10A).</p> <p>Pronotum 2.2 times as wide as long, 1.85 times wider at widest part than on front angles (Fig. 10C). Anterior crenulate margin comparatively wide. Area between lateral edge and internal lateral carina weakly concave, wide and continuously weakly narrowing posteriorly; pronotal disc not concave along internal lateral carina. Basal pronotal bordering line shallowly and irregularly crenulate. Posterolateral indentations comparatively deep, oblique. Prosternal process broad (Figs 7F, 10B), with apex about 0.90 of width of procoxal cavity, narrowest near half-length, weakly widened apically, with lateral carina raised, almost reaching anterior quarter of prosternum, central part between carinae depressed.</p> <p>Elytra 0.67–0.76 mm long, about as long a wide, 3.1 times as long and 1.4 times as wide as pronotum; lateral margins visible from above (Fig. 6C). Elytral dorsal surface covered with simple setiferous punctures and additional foveolate depressions without setae. Mesoventral process about 1.4 times as wide as mesocoxal diameter (Figs 7I, 10E). Metaventrite with fine setiferous punctures (Fig. 10E). Elytral lateral margins visible from above. Wings well developed.</p> <p>Legs. Meso- and metatrochanters flattened, weakly roundly produced posteriorly (Fig. 10E, F).</p> <p>Abdomen. Ventrite 1 with rounded postcoxal lines. Ventrite 5 with posterior margin straight in both sexes. Male genital segment as in Fig. 8A.</p> <p>Male genitalia (Fig. 8B). Aedeagus with penis short and stout, sclerotized, narrow at base, widening distally with lateral ramus extending beyond. Tegmen throneshaped, short, submembranous, basal; tegminal strut short, reduced.</p> <p>Femalegenitalia (Fig.8C). Ovipositorweaklysclerotized. Spermatheca moderately large, irregular-subcylindrical, submembranous; sperm duct short; accessory gland small membranous, of irregular-cordate shape.</p> <p>Etymology: This species is dedicated to Michael Geiser, curator in the Natural History Museum, London, for kindly pointing out to us the existence of the specimens belonging to the species here described.</p> <p>Distribution: Grenada (Fig. 20A).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFEFFF8D3EB10A07FB014550	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFEFFF8C3D940F63FB094054.text	03C2CE00FFEFFF8C3D940F63FB094054.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Karumbe pakaluki Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček 2023	<div><p>KARUMBE PAKALUKI ARRIAGA- VARELA,</p> <p>TOMASZEWSKA &amp; SZAWARYN SP. NOV.</p> <p>(FIGS 6E, F, 7A–E, G, J, K, M, 8D–F, 11A–F)</p> <p>Zoobank registration: urn: lsid: zoobank. org:act: 349822CB-0F6B-4737-A717-D6CD788AD9CC</p> <p>Type material: Holotype, male, VENEZUELA: VEN: Miranda: 400 m, 35 km N Altagracia, Guatopo NP, Agua Blanca, 31.V-7.VI.87 -2, S&amp;J Peck, ravine FITs (CMN). Paratypes: same data as holotype (one male and 2 ex, CMN; one female and 1 ex, MIZ).</p> <p>Diagnosis: Karumbe pakaluki is most similar to K. geiseri but can be distinguished from it by having the narrower prosternal process being about 0.67 as wide as procoxal diameter (Fig. 11C) (0.9 in K. geiseri); pronotum with posterolateral indentation shorter and more rectangular (Fig. 11B) (longer and more oblique in K. geiseri) and by the shape of the aedeagus (Fig. 10E). From K. brethesi, the third species of the genus, K. pakaluki can be easily distinguished by having dorsal surface covered with setiferous punctures and additional foveolate depressions without setae (Fig.11D) (in K.brethesi peculiar punctations as in Fig. 9E), the central area of the metaventrite covered only with simple setiferous punctures (Fig. 11F), the pronotal disc much less concave along the internal lateral carina (Fig. 11B) and the terminal labial palpomere short and apically rounded, 1.2 longer than wide (Fig. 7D) (in K. brethesi: elongate and acuminate, 2.2 as long as wide as in Fig. 9B).</p> <p>Description: Body: length 0.91–0.95 mm, 1.30 times as long as wide, 2.0 times as long as high, short oval and moderately convex, dark brown with yellowish legs, antennae and palpi (Fig. 6E, F).</p> <p>Antenna nearly 0.35 of length of body (Figs 7A, 11A); antennomeres 1–5 longer than wide; antennomeres 6, 7, 9 subquadrate and antennomere 8 transverse. Apical labial palpomere short and rounded, 1.2 times as long as wide (Fig. 7D).</p> <p>Pronotum 2.1 times as wide as long, 1.7 times wider at widest part than on front angles (Fig. 11B). Anterior crenulate margin comparatively wide. Area between lateral edge and internal lateral carina weakly concave, wide and continuously weakly narrowing posteriorly; pronotal disc not concave along internal lateral carina. Basal pronotal bordering line shallowly and irregularly crenulate. Posterolateral indentations comparatively shallow, rectangular. Prosternal process wide (Fig. 11C), with apex about 0.67 of width of procoxal cavity, narrowest near half-length, weakly widened apically, with lateral carinae raised, reaching almost anterior quarter of prosternum, central part between carinae slightly depressed.</p> <p>Elytra 0.70– 0.75 mm long, about as long a wide, 3.1 times as long and 1.4 times as wide as pronotum; lateral margins visible from above (Fig. 6E). Elytral dorsal surface covered with simple setiferous punctures and additional foveolate depressions without setae (Fig. 11D). Metaventrite with fine setiferous punctures at centre (Fig. 11F). Mesoventral process about 1.3 times as wide as mesocoxal diameter. Wings well developed.</p> <p>Legs. Meso- and metatrochanters flattened, roundly produced posteriorly.</p> <p>Abdomen. Ventrite 1 with rounded and laterally complete postcoxal lines (Fig. 11F). Ventrite 5 in male triangularly produced posteriorly at middle, in female straight. Male genital segment as in Fig. 8D.</p> <p>Male genitalia (Fig. 8E). Aedeagus with penis short and stout, narrow at base, widening towards curved, ramificate apex. Tegmen throne-shaped, long; tegminal strut short, reduced.</p> <p>Female genitalia. Bursa copulatrix large, with apical outlet of sperm duct. Spermatheca (Fig. 8F) submembranous, moderately large, elongatesubcylindrical, oblong; sperm duct long, about as long as spermatheca; accessory gland small, membranous, of irregular shape.</p> <p>Etymology: This species is dedicated to the late James Pakaluk, who had keen interest in small brown beetles.</p> <p>Distribution: Venezuela (Fig. 20A).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFEFFF8C3D940F63FB094054	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFEEFF823D7D0A72FAFE463C.text	03C2CE00FFEEFF823D7D0A72FAFE463C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Yamuy Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček 2023	<div><p>YAMUY ARRIAGA- VARELA, TOMASZEWSKA &amp; SZAWARYN GEN. NOV.</p> <p>(FIGS 12–16)</p> <p>Zoobank registration: urn: lsid: zoobank. org:act: ACB24628-6F19-494F-8866-3CFA28015395</p> <p>Type species: Yamuy marginatus b y p r e s e n t designation.</p> <p>Diagnosis: Yamuy can be distinguished from other genera of Cerasommatidiidae by the following combination of characters: dorsal surface of the body covered with two kinds of vestiture (Figs 12, 15F, 16A, B): minuscule decumbent setae and sparse long erect setae/ spines; lateral margins of the pronotum with complete (Fig.16B) or reduced/short and present only basally (Fig. 15C) internal subparallel carina demarcating a wide area; anterior margin bordered narrowly with simple bordering line; prosternal process strongly narrowed preapically with lateral carinae short, anteriorly not extending beyond prosternal process (Figs 15D, 16C); trochanters simple, not covering the tibio-tarsal joint when retracted (Figs 15D, 16D); abdominal ventrite 1 without postcoxal lines (Figs 15H, 16F); aedeagus with tegmen sclerotized, fused to base of penis (Fig. 14B, E). The mixed vestiture (vestiture homogeneous in Cerasommatidia, Karumbe and MahaƲelo) and the simple not flattened trochanters are unique within the family (in Cerasommatidia, Karumbe and MahaƲelo trochanters are flattened and expanded posteriorly to cover the tibio-tarsal joint in repose).</p> <p>Description: Length 0.90–0.95 mm. Body short oval and convex, 1.6 times as long as wide, around 2.0–2.2 times as long as high. Coloration light brown to yellow, sometimes with contrasting markings on elytra, and whitish legs (Fig. 12A–D).</p> <p>Head with dorsal surface with two kinds of vestiture: sparse tiny decumbent hairs and a pair of long erect setae in vertex near posterior margin of each eye (Fig. 16A). Clypeus large, rectangular (Fig. 15B). Frontoclypeal suture distinct, straight. Ventral antennal grooves short, not extending to posterior eye level (Fig. 15A). Antenna approximately 0.4 of length of body (Figs 13A, 15A); antennal club one quarter of total antennal length; antennomeres 1–5 longer than wide, with antennomere 5 longer than neighbouring antennomeres 4 and 6; antennomeres 6–9 variable in species. Eyes comparatively large, moderately prominent (Figs 15B, 16A). Galea large, densely setose at apex (Fig. 13C). Lacinia weakly narrower than galea with dense setae at apex and inner margin. Labium with mentum widest near midlength (Fig. 15A); with palpomere 2 and 3 large, bulbous, transverse and subtriangular respectively; terminal palpomere short and rounded to elongate acuminate, around 2 times as long as wide at base.</p> <p>Prothorax. Pronotum 1.8–2.0 times wider than long, widest between base and basal third, weakly to moderately strongly convergent anteriorly, 1.3– 1.6 times wider at widest part than on front angles. Anterior margin narrowly bordered with regular bordering line; base of pronotum weakly sinuate with margin narrowly bordered with faint bordering line or unbordered (Figs 15C, 16B). Pronotal sides scarcely rounded, with complete (Fig. 16B) or short/reduced internal lateral carina present only at base (Fig. 15C), with area between carina and the lateral edge weakly concave, wider near basal third; hind angles with or without small, oblique indentation to receive humeral corner of elytron. Pronotal disc moderately to weakly convex, with surface covered with two kinds of vestiture: sparse small punctures bearing tiny decumbent hairs, and a pair of long erect setae. Prosternum with anterior margin scarcely arcuate posteriorly (Figs 15D, 16C); prosternal process narrow, at apex about 0.25–0.30 of width of procoxal cavity, with raised lateral carinae reaching anteriorly before or briefly beyond level of anterior margin of procoxal cavities; area between carinae concave; prosternal process extending posteriorly to hind level of procoxae. Hypomeron with deep and comparatively long, sinuate antennal grooves (Figs 15D, 16C).</p> <p>Pterothorax. Mesonotum with scutellar shield small, transverse, weakly rounded at apex (Fig. 16B). Mesoventrite with intercoxal process smooth, almost flat with anterior raised border incomplete medially; 1.0–1.4 times as wide as mesocoxal diameter (Figs 13D, 15G, 16D). Metaventrite as long as abdominal ventrites 1–3 together; with complete discrimen (visible only in the internal part of the ventrite after dissection, Fig. 13D); anterior margin with bordering line slightly widened towards lateral corners, with or without small longitudinal carina below each coxa; central area with fine setiferous punctures (Figs 15G, 16D). Anterior part of metanepisternum with small outer angulate projection. Elytra about 0.6 mm long, about as long a wide, 2.5 times as long and 1.3 times as wide as pronotum, with lateral margins narrowly explanate (Figs 12A, C, 15F). Surface with two kinds of vestiture: sparse small punctures bearing tiny decumbent hairs, and long setae: sparse in the centre of elytral disc (about eight to 14 setae) and arranged in a row near lateral margins (around eight to ten setae). Epipleura comparatively narrow, narrowing toward apex reaching nearly elytral apex but incomplete, with internal bordering line narrow, present from the level of mid-coxae to apex. Hindwings well developed, without anal lobe, with single reduced anal vein, and median fleck present, undivided.</p> <p>Legs. Trochanters not modified (Figs 15G, 16D). Femora weakly flattened (mid and hind femora more distinctly than fore femora); with grooves for tibiae present throughout whole length. Claws with comparatively large, sharp subquadrate tooth at base (Figs 13F, 15E, 16E).</p> <p>Abdomen with ventrite 1 with anterior marginal bordering, without postcoxal lines, with or without small longitudinal carina below each coxae (Figs 15H, 16F).</p> <p>Male genitalia (Fig. 14B, E). Aedeagus with penis short and stout, sclerotized, weakly curved, asymmetric, with ramificate apex. Tegmen sclerotized, short, ring-like, fused to penis at base, with long ramus extending anteriorly; tegminal strut short, reduced.</p> <p>Female genitalia (Fig. 14C, F). Spermatheca large, elongate, irregular ‘infundibuliform’, submembranous; sperm duct short; accessory gland small membranous, of irregular shape.</p> <p>Etymology: Yamuy is the Taíno word for ‘cat’, referring to the whisker-like setae on the pronotum and elytra, characteristic of this genus. Gender masculine.</p> <p>Species included: Yamuy constratus, Y. marginatus.</p> <p>Distribution: Puerto Rico, Venezuela (Fig. 20A).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFEEFF823D7D0A72FAFE463C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFE3FF863EEB0F99FD0F450B.text	03C2CE00FFE3FF863EEB0F99FD0F450B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Yamuy constratus Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček 2023	<div><p>YAMUY CONSTRATUS ARRIAGA- VARELA,</p> <p>TOMASZEWSKA &amp; SZAWARYN SP. NOV.</p> <p>(FIGS 12A, B, 13B–F, 14A–C, 15A–H, 19A–B, 20A)</p> <p>Zoobank registration: urn: lsid: zoobank. org:act: 5CD938AC-7821-46A6-ADB3-C64CC573D46F:</p> <p>Type material: Holotype, male, PUERTO RICO: Portorico Mor. [green label]/ 55669/ Zool. Mus. Berlin/ frontalis m. [handwriting] (MNB). Paratypes: PUERTO RICO: Portorico Moritz Nr. 55 669 [green label]/ Zool. Mus. Berlin (one female, MNB); Naguabo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.78833&amp;materialsCitation.latitude=18.263334" title="Search Plazi for locations around (long -65.78833/lat 18.263334)">El Yunque Nat. Forest</a>, S. part, 4.9 km N of Río Blanco, 18°15.8′N, 65°47.3′W; 495 m, 24.vi.-2.vii.2016; Fikáček &amp; Seidel lgt. PR11// flight intercept trap at the margin of the rainforest in an area with many flowering Etlingera elatior plants (one female, totally dissected, NMPC); Puerto Rico, El Verde Res. Sta., 250 m, Carib Nat. For., Rio Grande, 19 July 1994, slime mold fruit. body, M. A. Ivie (one female, MTEC; one female, MIZ); VENEZUELA: Caracas Moritz Nr. 55 670 [green label]/ Zool. Mus. Berlin (one female, MNB); same and 55670/ luridellus m. [handwriting]/ Zool. Mus. Berlin (one, MNB).</p> <p>Diagnosis: Yamuyconstratus canbeeasilydistinguished from Y. marginatus by having uniformly coloured elytra (Fig. 12A, B), the lateral margins of the pronotum narrowly bordered by a simple, thin bordering line and with short internal subparallel carinae at base (Fig. 15C) (in Y. marginatus internal carina demarcating a wide area is present throughout the length of pronotum), the anterior margin of metaventrite and abdominal ventrite 1 with irregularly crenulate bordering line, without longitudinal carinae (Fig. 15G, H) (in Y. marginatus anterior margin of metaventrite and abdominal ventrite 1 with short postcoxal longitudinal carinae beyond each coxal cavity) and by peculiar punctation on dorsal body surface: apart from long setae present in both species, a tiny short hairs are growing from conspicuous tubercles set in round foveolate punctae in Y. constratus (Fig. 15C, F), while in Y. marginatus short hairs are growing from simple punctae.</p> <p>Description: Body: length 0.9 mm, 1.3 times as long as wide, 2.0 times as long as high, short oval and moderately convex. Coloration uniformly brown to pale-yellow (Fig. 12A, B).</p> <p>Antenna nearly 0.4 of length of body (Fig. 15A); antennomeres 1–6 as long as wide; antennomeres 7–9 subquadrate. Apical labial palpomere short and acuminate (Fig. 15A), 1.8 times as long as wide.</p> <p>Pronotum 2.0 times as wide as long, 1.3 times wider at widest part than on front angles (Fig. 15C). Anterior margin with bordering line vanishing medially. Lateral edges bordered with simple thin bordering line, and with internal lateral carina reaching at most about basal third length of pronotum. Pronotal disc not concave near lateral margins. Base of pronotum not bordered. Posterolateral corners without indentation. Prosternal process with apex about 0.3 of width of procoxal cavity (Fig. 15D), narrowest near half-length, distinctly widened apically, lateral carinae reaching beyond half-length of prosternum, with area between carinae distinctly depressed.</p> <p>Elytra (Figs 12A, 15F) 0.6 mm long, about as long a wide, 2.5 times as long and 1.4 times as wide as pronotum; lateral margins visible from above (Fig. 12A). Mesoventral process about as wide as mesocoxal diameter (Figs 13D, 15G). Metaventrite with a few fine setiferous punctures mainly on anterior half; anterior margin without longitudinal carinae (Figs 13D, 15G).</p> <p>Legs. Meso- and metatrochanters weakly flattened, not produced posteriorly (Figs 13D, 15D, G).</p> <p>Abdomen. Ventrite 1 with anterior margin distinctly, simply margined, without postcoxal lines (Fig. 15H); longitudinal carina below each metacoxa absent. Male genital segment as in Fig. 14A.</p> <p>Male genitalia (Fig. 14B). Aedeagus with penis short and stout, sclerotized, narrow at base, weakly widening towards ramificate apex. Tegmen sclerotized short ring, with long ramificate projection extending anteriorly beyond apex of penis; tegminal strut short.</p> <p>Female genitalia (Fig. 14C). Ovipositor weakly sclerotized. Spermatheca comparatively large, irregular – ‘infundibuliform’, submembranous; sperm duct short; accessory gland small membranous, of irregular-cordate shape. Proctiger large, weakly sclerotized.</p> <p>Etymology: From the Latin, constratus, flat, plane, referring to the mostly flat lateral areas of the pronotum, having only short lateral carinae at base.</p> <p>Biology: One of the studied specimens was collected from a slime mold fruiting body at an elevation of around 250 m, while another specimen was collected using a flight intercept trap installed at the margin of a rainforest at elevation of 495 m (Fig. 19A, B).</p> <p>Distribution: Puerto Rico, Venezuela (Fig. 20A).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFE3FF863EEB0F99FD0F450B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFE4FF853EF80FAAFDFE4524.text	03C2CE00FFE4FF853EF80FAAFDFE4524.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Yamuy marginatus Arriaga-Varela & Tomaszewska & Szawaryn & Robertson & Seidel & Ślipiński & Fikáček 2023	<div><p>YAMUY MARGINATUS ARRIAGA- VARELA,</p> <p>TOMASZEWSKA &amp; SZAWARYN SP. NOV.</p> <p>(FIGS 12C, D, 13A, 14D–F, 16A–F, 20A)</p> <p>Zoobank registration: urn: lsid: zoobank. org:act: 2D30BBFE-4F4D-4C1D-9C97-C0BBB4F6B2F6</p> <p>Type material: Holotype, male, PUERTO RICO: Orocovis Bosque Estatal Toro Negro, sector <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.495&amp;materialsCitation.latitude=18.176666" title="Search Plazi for locations around (long -66.495/lat 18.176666)">Doña Juana</a>, 5.6 km N of Villalba 18°10.6′N, 66°29.7′W; 810 m, 29.vi.2016; Deler, Fikáček &amp; Seidel lgt., PR18// sifting of small accumulations of leaf litter and rotten palm leaves and fruit in sparse montane forest at the stream bank (NMPC). Paratypes: PUERTO RICO: San German, Bosque Estatal Maricao, 8.6 km N of Sabana Grande 18°8.0′N, 66°59.5′W; 820 m, 28.vi.2016; Fikáček &amp; Seidel lgt., PR16 (one, MIZ); Puerto Rico: Jayuya, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-66.591736&amp;materialsCitation.latitude=18.17225" title="Search Plazi for locations around (long -66.591736/lat 18.17225)">Toro Negro</a> at 18°10.335′N, 66°35.504′W; 1350 m, WP-479, Arboreal moss sifting; 3 Sept. 2014, A.S. Konstantinov (one female, MTEC).</p> <p>Diagnosis: Yamuymarginatus canbeeasilydistinguished from Y. constratus by having the elytra pale yellow decorated with dark-infuscate transverse markings (Fig. 12C, D) (uniformly coloured in Y. constratus), the lateral margins of the pronotum provided with long/ complete subparallel internal carinae demarcating a wide area (Fig. 16B) (internal lateral carinae present only basally in Y. constratus), the anterior margin of the metaventrite and abdominal ventrite 1 with a pair of short postcoxal longitudinal carinae (Fig. 16D, F) (in Y. constratus anterior margin of metaventrite and abdominal ventrite 1 bordered with crenulate bordering line, without carinae), and by punctation on dorsal surface: apart from long setae present in both species, tiny short hairs are growing from simple comparatively large punctae/depressions in Y. marginatus (Fig. 16B) and in Y. constratus they are growing from conspicuous tubercles set in round punctae.</p> <p>Description: Body: length 0.90–0.94 mm, 1.6 times as long as wide, 2.2 times as long as high, short oval and moderately convex (Fig. 12C, D). Coloration pale-yellow with whitish legs; elytra with dark-infuscate transverse markings on basal eighth and a large, transverse, irregularly oval one on each elytron at apical threequarters; antenna with apical three antennomeres dark infuscate (Fig. 12C, D).</p> <p>Antenna nearly 0.4 of the length of the body; antennomeres 1–6 as long as wide (Fig. 13A); antennomeres 7–9 subquadrate. Apical labial palpomere short and rounded, 1.3 times as long as wide.</p> <p>Pronotum 1.8 times as wide as long, 1.6 times wider at widest part than at front angles (Fig. 16B). Anterior pronotal bordering narrow and regular. Area between lateral edge and internal lateral carina weakly concave, wider at two-thirds and continuously weakly narrowing posteriorly and anteriorly; pronotal disc not concave along internal lateral carina. Base of pronotum not bordered. Posterolateral indentations comparatively deep, briefly acute. Prosternal process with apex about 0.3 of width of procoxal cavity, narrowest near half-length, distinctly widened apically, lateral carinae reaching anteriorly beyond half-length of prosternum, central area distinctly depressed (Fig. 16C).</p> <p>Elytra 0.6 mm long, about as long a wide, 2.5 times as long and 1.3 times as wide as pronotum. Elytral lateral margins visible from above (Fig. 12C). Mesoventral process about as wide as mesocoxal diameter (Fig. 16D). Metaventrite with few fine setiferous punctures on sides; with small longitudinal carina near the posterior-most part of each mesocoxal cavity almost reaching mid-length of metaventrite (Fig. 16D).</p> <p>Legs. Meso- and metatrochanters weakly flattened, not produced posteriorly (Fig. 16D).</p> <p>Abdomen. Ventrite 1 with anterior margin with a small longitudinal carina near the internal-most margin of each metacoxal cavity reaching mid-length of ventrite (Fig. 16F). Male genital segment as in Fig. 14D.</p> <p>Male genitalia (Fig. 14E). Aedeagus with penis short and stout, sclerotized, curved, weakly narrowing towards apex. Tegmen sclerotized, short ring, with long ramus extending anteriorly beyond apex of penis; tegminal strut short, reduced.</p> <p>Female genitalia (Fig. 14F). Ovipositor weakly sclerotized. Spermatheca comparatively large, elongate, irregular – ‘infundibuliform’, submembranous; sperm duct of medium length, about half the length of spermatheca; accessory gland small membranous, of irregular shape. Proctiger large, narrow, weakly sclerotized.</p> <p>Etymology: From the Latin, marginatus, edged, based on the characteristically bordered lateral pronotal margins that distinguish it from the other species in the genus.</p> <p>Biology: One of the specimens was collected by sifting small accumulations of leaf litter and rotten palm leaves and fruits in sparse montane forest at elevation of 821 m. The second one, by sifting arboreal mosses in montane area at elevation of 1350 m.</p> <p>Distribution: Puerto Rico (Fig. 20A).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFE4FF853EF80FAAFDFE4524	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFE7FFBA3EAA0F98FD8E4732.text	03C2CE00FFE7FFBA3EAA0F98FD8E4732.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mahavelo SZAWARYN, TOMASZEWSKA & ARRIAGAVARELA 2023	<div><p>MAHAVELO SZAWARYN, TOMASZEWSKA &amp; ARRIAGA- VARELA GEN. NOV.</p> <p>(FIGS 17, 18, 19C, D, 20C)</p> <p>Zoobank registration: urn: lsid: zoobank. org:act: DF9A1833-7A95-4B99-9148-AFCB9248D39B</p> <p>Type species: MahaƲelo madagasus by present designation.</p> <p>Diagnosis: MahaƲelo is a distinctive genus in Cerasommatidiidae by the lack of a scutellar shield (Fig. 18C, F), and by having the prosternal process provided with single, longitudinal, median carina (Fig. 18D). In general appearance it is most similar to Cerasommatidia, sharing the glabrous appearance of pronotal and elytral surfaces (covered with tiny inconspicuous hairs), pronotum without internal sublateral carinae, and pronotal and elytral surfaces covered with single size punctae (Fig. 18C, F). However, apart from its unique characters, MahaƲelo can also be separated from Cerasommatidia by having the pronotal hind angles with small, oblique indentation to receive the humeral corner of the elytron (Fig. 18C) (absent in Cerasommatidia).</p> <p>Description: Length 1.00 mm, width 0.79 mm. Body short oval and convex, 1.3 times as long as wide. Coloration uniformly brown (Fig. 17A–C).</p> <p>Head (Figs 17D, 18B) with dorsal surface uniformly covered with short fine setae. Clypeus large, rectangular. Frontoclypeal suture distinct but feebly marked, straight. Ventral antennal grooves deep and broad between eyes and mouthparts, long, extending beyond posterior eye level. Antenna (Fig. 18A) approximately 0.35 of length of body; antennal club one-quarter of total antennal length; antennomeres 1–5 much longer than wide, with antennomere 3 longer than antennomere 4 and 5; antennomeres 6–9 less elongate. Eyes comparatively small, moderately prominent, coarsely facetted (Figs 17D, 18B). Maxillary palpomere 1 narrow, elongate; palpomere 2 weakly trapezoidal and somewhat bulbous (Fig. 17E). Galea large, densely setose apically. Lacinia about half as wide as galea, with dense setae at apex and inner margin. Labium with mentum widest near basal third, palpomere 2 large, bulbous, somewhat oval; terminal palpomere short, subquadrate, about as long as penultimate, truncate apically.</p> <p>Prothorax. Pronotum 2.7 times wider than long, widest at base, and strongly convergent anteriorly (Fig. 18C). Anterior pronotal margin narrowly bordered with regular bordering line; base without distinct bordering line. Pronotal sides weakly rounded, narrowly bordered; hind angles with small, oblique indentation to receive humeral corner of elytron. Pronotal disc moderately convex, with uniform sparse fine punctures bearing tiny, thin hairs (Fig. 18C). Prosternum (Fig. 18D) with anterior margin scarcely arcuate posteriorly; prosternal process narrow, at apex about 0.60 of width of procoxal cavity, with long, single median carina but not reaching anterior prosternal margin or apex of prosternal process; prosternal process extending posteriorly to hind level of procoxae. Hypomeron with deep, long and straight antennal grooves.</p> <p>Pterothorax. Mesonotum lacking scutellar shield. Mesoventrite strongly transverse, with intercoxal process smooth, almost flat, with raised anterior border complete (Figs 17G, 18E); 1.2 times as wide as mesocoxal diameter. Metaventrite as long as abdominal ventrites 1 and 2 combined; discrimen absent; anterior margin with bordering carina simple, straight at intercoxal process and widening towards lateral corners (Fig. 18E); central area of metaventrite with fine setiferous punctures. Anterior part of metanepisternum with lateral sub-triangular projection on its external margin. Elytra about as long as wide, with lateral margins invisible from above (Figs 17A, 18F). Surface covered with uniform, small setiferous punctures bearing tiny, thin hairs. Epipleura incomplete but almost reaching elytral apex (Fig. 18G), narrow, with internal bordering line wide, present from the level of mid coxae to apex (Fig. 18E); basal part of epipleuron with short groove corresponding to projection of metanepisternum. Hindwings reduced.</p> <p>Legs. Trochanters produced posteriorly (Fig. 18E). Femora weakly flattened (mid and hind femora more distinctly than fore femora); with g r o o v e s f o r t i b i a e p r e s e n t t h r o u g h o u t w h o l e length. Claws with distinct sharp, subquadrate tooth at base.</p> <p>Abdomen with ventrite 1 with anterior bordering carina below coxae widening laterally (Fig. 18G), absent on intercoxal process; postcoxal lines absent.</p> <p>Male genitalia (Fig. 17H). Aedeagus with penis short and stout, sclerotized, weakly curved, wider near apex, asymmetric, with small acute apical projection. Tegmen moderately large, basal, throne-shaped, weakly sclerotized; tegminal strut short.</p> <p>Female genitalia (Fig. 17J). Spermatheca of moderate size, elongate, irregularly reniform, submembranous (Fig. 17K); bursa copulatrix large; sperm duct long; accessory gland small membranous, of irregular shape.</p> <p>Etymology: The name is derived from the Mahavelo Forest, Madgascar, where this genus was first found. Gender masculine.</p> <p>Distribution: Madagascar (Fig. 20C).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFE7FFBA3EAA0F98FD8E4732	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFD8FFB93E870D76FB60460E.text	03C2CE00FFD8FFB93E870D76FB60460E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Mahavelo MADAGASUS SZAWARYN, TOMASZEWSKA	<div><p>MAHAVELO MADAGASUS SZAWARYN, TOMASZEWSKA</p> <p>&amp; ARRIAGA- VARELA SP. NOV.</p> <p>(FIGS 17A–K, 18A–G, 19C, D, 20C)</p> <p>Zoobank registration: urn: lsid: zoobank. org:act: 1B1B5269-C265-4C28-B446-9D644006CCBA</p> <p>Type material: Holotype, male, MADAGASCAR: Toliara, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=46.15717&amp;materialsCitation.latitude=-24.75833" title="Search Plazi for locations around (long 46.15717/lat -24.75833)">Forêt</a> de Mahavelo, Isantoria River, 5.2 km 44°NE Ifotaka, 24°46′S, 46°09′E [–24.75833, 46.15717], 110 m a.s.l., 28.01- 1.02.2002, coll. Fisher-Griswold Arthropod Team, coll. code BLF05238 (CAS). Paratypes: same data as for holotype (23, CAS; 7, MIZ).</p> <p>Diagnosis: As for the genus.</p> <p>Description: Body: length 1.00 mm, width 0.79 mm, 1.3 times as long as high, short oval and moderately convex, uniformly brown, covered with inconspicuous hairs (Fig. 17A–C).</p> <p>Antenna nearly 0.35 of length of body (Fig. 18A); antennomere 3 about 3.4 times as long as wide; antennomeres 4 and 5 subequal in length, about 2.2 times as long as wide; antennomeres 6-9 subquadrate. Apical labial palpomere short.</p> <p>Pronotum 2.7 times as wide as long (Fig. 18C). Anterior margin with fine, regular bordering line vanishing medially; lateral margins narrowly bordered. Pronotal disc not concave near lateral margins. Base without distinct border. Posterolateral corners with minuscule indentation. Prosternal process wide, with apex about 0.6 of width of procoxal cavity, continuously weakly widened apically, bordered laterally nearly to level of anterior margin of procoxal cavities (Fig. 18D).</p> <p>Elytra about as long as wide, 3.9 times as long and 1.35 times as wide as pronotum; lateral margins not visible from above (Figs 17A, 18F). Mesoventral process about 1.1 times as wide as mesocoxal diameter (Fig. 18E). Metaventrite sparsely setose.</p> <p>Legs. Meso- and metatrochanters flattened, weakly to distinctly angulately produced posteriorly (Fig. 18G).</p> <p>Abdomen with male genital segment as in Figure 17I.</p> <p>Male genitalia (Fig. 17H). Aedeagus with penis short and stout, sclerotized, weakly curved, wider near apex, asymmetric, with branched apex. Tegmen sclerotized, basal, throne-shaped, moderately large; tegminal strut short, reduced.</p> <p>Female genitalia (Fig. 17J). Spermatheca (Fig. 17K) of moderate size, elongate, irregularly reniform, submembranous; sperm duct comparatively long; accessory gland small membranous, of irregular shape.</p> <p>Etymology: The name madagasus is an arbitrary combination of letters referring to Madagascar, where this species occurs.</p> <p>Biology: All specimens were sifted from leaf litter in dry spiny forest biotope at elevation of 110 m (Figs 1D, 19C).</p> <p>Distribution: Madagascar (Fig. 20C).</p></div> 	https://treatment.plazi.org/id/03C2CE00FFD8FFB93E870D76FB60460E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFDAFFB83FC70BD2FAC3448E.text	03C2CE00FFDAFFB83FC70BD2FAC3448E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cerasommatidiidae BRETHES 1925	<div>KEY TO GENERA AND SPECIES OF CERASOMMATIDIIDAE1. Trochanters flattened, projected to cover tibio-tarsal joint in repose (Figs 4E, 4, 5H, 7K, 9F, 10F, 11F, 18E); vestiture of dorsal body surface uniform (tiny decumbent hairs (often missing)) (Figs 4B, C, 5C, G, 18C, F) or moderately long suberect setae (Figs 9E, 10C, 11B, D)....................................................................................... 2 - Trochanters normal, not flattened or produced to cover tibio-tarsal joint in repose (Figs 13D, 15D, G, 16D); vestiture of dorsal surface of body composed of minuscule decumbent hairs and sparse long erect setae (Figs 15F, 16B) [Puerto Rico, Venezuela]........................................................................................................ Yamuy …4 2. Pronotum with lateral and anterior borders simply margined (Figs 4B, 5C, 18C); dorsal surface with tiny decumbent hairs (often missing) (Figs 4B, C, 5C, G, 18C, F)............................................................................... 3 - Pronotum with lateral margins bordered with an internal subparallel carinae (Figs 9C, 10C, 11B); anterior margin of pronotum with crenulate bordering line (Figs 9C, 10C, 11B); dorsal surface with moderately long suberect pubescence (often missing) (Figs 6A–F, 9E, 10C)…........................................................... Karumbe.... 5 3. Abdominal ventrite 1 with complete semicircular postcoxal lines (Figs 3D, 4G, 5H); prosternal process with long lateral carinae reaching anterior third of prosternum, without median carina (Fig. 5D); Brazil....................................................................................................................................... Cerasommatidia …7 - Abdominal ventrite 1 without postcoxal lines, only with anterior bordering margin thickened posterior to coxae and widening laterally (Fig. 18G); prosternum with raised, single, median carina, (Fig. 18D); Madagascar.......................................................................................................................... MahaƲelo madagasus 4. Pronotum with lateral margins bordered with an internal carina (Figs 12C, D, 16B); anterior margins of metaventrite and abdominal ventrite 1 with small longitudinal carina below each coxa (Fig 16D, F); Puerto Rico............................................................................................................................................ Yamuy marginatus - Pronotum with lateral margins only narrowly bordered with simple bordering line, with internal carina short, weakly marked, present only basally (Fig. 15C); anterior margins of metaventrite and abdominal ventrite 1 without longitudinal carinae beyond coxae (Figs 13D, 15G, H); Puerto Rico, Venezuela.................................................................................................................................... Yamuy constratus 5. Body more than 1.40 mm long; metaventrite with setiferous punctures inserted in wide foveate impressions (Fig. 9F); terminal labial palpomere elongate and acuminate, 2.2 times as long as wide (Fig. 9B); Brazil........................................................................................................................................... Karumbe brethesi- Body less than 1.00 mm long; metaventrite with simple setiferous punctures (Figs 10E, 11F); terminal labial palpomere short and apically rounded, 1.2–1.4 times as long as wide (Figs 7D, 10A); Venezuela, Grenada..6 6. Prosternal process narrower, about 0.67 of width of procoxal diameter (Fig. 11C); pronotum with posterolateral indentation short and rectangular (Fig. 11B); Venezuela.......................................... Karumbe pakaluki - Prosternal process wider, about 0.90 of width of procoxal diameter (Fig. 10B); pronotum with postero-lateral indentation large and oblique (Fig. 10C); Grenada..................................................................... Karumbe geiseri 7. Pronotum 2.0 times as wide as long, with bordering line at anterior margin complete; body size about 1.7 mm..................................................................................................................................... Cerasommatidia rotundata- Pronotum at least 2.2 times as wide as long, with bordering line at anterior margin vanishing medially; body size 1.1–1.4 mm...................................................................................................................................................... 8 8. Body 1.15 mm long; posterior margin of pronotum narrowly bordered at least medially (Fig. 5C); aedeagus as in Fig. 3E.................................................................................................................... Cerasommatidia plaumanni - Body 1.4 mm long; posterior margin of pronotum not bordered (Fig. 4B); aedeagus as in Fig. 3F........................................................................................................................................... Cerasommatidia arroaei</div>	https://treatment.plazi.org/id/03C2CE00FFDAFFB83FC70BD2FAC3448E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
03C2CE00FFDAFFBF3F1A0D3FFDE44194.text	03C2CE00FFDAFFBF3F1A0D3FFDE44194.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cerasommatidiidae BRETHES 1925	<div><p>BIOLOGY OF CERASOMMATIDIIDAE</p> <p>Information about the biology of this group of beetles is scarce, but some data can be gleaned from the habitats and methods by which they were collected. Specimens of Yamuy and MahaƲelo species were mostly collected by sifting leaf litter. One specimen was collected directly from a slime mould fruiting body; another one from a flight intercept trap. These observations can indicate that with high probability, the food source of Cerasommatidiidae is various species of slime moulds or other fungi-like organisms. Cerasommatidiidae are also active fliers. The coarsely facetted eyes of all taxa indicate that they probably have a nocturnal lifestyle. Yamuy constratus was collected in rainforest habitat, Y. marginatus in sparse montane forest and MahaƲelo madagasus in dry spiny forest biotope at low elevation. That indicates that these beetles inhabit different types of forest habitat, both rainforests (Puerto Rico), as well as dry spiny forest (Madagascar), at a variety of elevations ranging from lowland (110 m) to montane areas (1350 m). Although incomplete, this information can help locate additional habitats and new taxa of Cerasommatidiidae in the future.</p> </div>	https://treatment.plazi.org/id/03C2CE00FFDAFFBF3F1A0D3FFDE44194	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriaga-Varela, Emmanuel;Tomaszewska, Wioletta;Szawaryn, Karol;Robertson, James;Seidel, Matthias;Ślipiński, Adam;Fikáček, Martin	Arriaga-Varela, Emmanuel, Tomaszewska, Wioletta, Szawaryn, Karol, Robertson, James, Seidel, Matthias, Ślipiński, Adam, Fikáček, Martin (2023): The resurrection of Cerasommatidiidae, an enigmatic group of coccinelloid beetles (Coleoptera: Coccinelloidea) based on molecular and morphological evidence. Zoological Journal of the Linnean Society 197 (4): 1078-1115, DOI: 10.1093/zoolinnean/zlac082, URL: http://dx.doi.org/10.1093/zoolinnean/zlac082
