taxonID	type	description	language	source
4383E1E7B7305F67A5F593683DED7ACE.taxon	materials_examined	Material examined. Japan - Hokkaido [Hokkaido] • 1 ♀; Yamato, Erimo; 15 Jul. 2002; H. Kogi leg.; Host: Quercus dentata; TO • 1 ♂; same locality and collector; 19 Jul. 2003; Host: Quercus dentata • 1 ♂; Syoya, Erimo; 30 Jun. 2002; H. Kogi leg. • 1 ♂; Tomikawa, Monbetu; 21 Jul. 2004; H. Kogi leg.; Host: Quercus dentata; TO • 3 ♀♀; same locality and collector; 23 - 25 Jul. 2006; 6 vii 2007 • 2 ♀♀; Higasihayakita, Hayakita; 20 Jul. 2005; H. Kogi leg. • 1 ♂; Tiyosibetu, Hamamasu; 13 Jul. 2002; H. Kogi leg. • 1 ♂; Katsuranosawa, Atuta; 21 Jul. 2002; H. Kogi leg. • 1 ♂; Uenai, Tomakomai; 26 Jul. 2002; H. Kogi leg. - Honshu [Iwate] • 1 ♀; Iwayama, Morioka; 2 Jul. 2009; T. Oku leg.; Host: Quercus mongolica; 13 Jul. 2009 em.; TO. - Honshu [Nagano] • 1 ♀; Kojiro, Tenryu-mura; N. Hirano leg.; 5 Jul. 2008; gen. slide no. KM- 118 • 1 ♂; same locality and collector; 28 Aug. 2009; gen slide no. KM- 106. - Honshu [Aichi] • 1 ♂; Asahikogen, Asahi-cho; 7 Jul. 2001; T. Mano leg.; OPU. - Honshu [Mie] • 1 ♂; Hijiki 250 m, Ueno-city; 27 Jun. 1997; T. Mano leg.; gen. slide no. KM- 1; OPU • 1 ♂; Hudodani, Miyama-tyo; 30 Jun. 2001; T. Mano leg.; gen. slide no. KM- 96; OPU. - Honshu [Kyoto] • 1 ♂; Mt. Ponpon; 1 Jul. 2000; N. H. Ahn leg.; gen. slide no. KM- 2; OPU. - Honshu [Nara] • 1 ♀; Wasamatayama; 9 - 10 Aug. 1989; S. Moriuti leg.; gen. slide no. KM- 51; OPU.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
4383E1E7B7305F67A5F593683DED7ACE.taxon	diagnosis	Diagnosis. The forewing is white with pale brown in the distal part, without an apical black spot. The anellus lobe of the male genitalia is a pear-shaped lobe basally, long, strongly sclerotized, and spine-like apically; the valva is narrow and elongate. The apophysis posterioris of the female genitalia is approx. two times of the length of papilla analis and approx. three times longer than the apophysis anterioris; the signum is long, narrow, and arch-shaped.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
4383E1E7B7305F67A5F593683DED7ACE.taxon	description	Description. Male (Figs 4 C, 7 B). Forewing length 2.7 - 3.4 mm. Wing expanse 6.0 - 7.3 mm. Head: shiny creamy white with appressed scales. Antennae filiform, basal segment elongate without pecten and creamy white; flagellum creamy white on dorsal surface before middle, then grayish brown beyond, with extraordinarily long and fine cilia on its ventral surface. Labial palpus white, long, and recurved; first segment shortest, creamy white suffused with brown scales on outer surface; second segment thickened, up to 2.5 times the length of the first and white; third segment nearly as long as second, creamy white evenly on both surfaces, apex sharply acute. Thorax: creamy white. Tegula shiny, creamy white dorsally, ornamented with bronze-brown scales along anterior margin. Legs: white; forefemur, tibia, and tarsus suffused inwardly with brown; hind tibia creamy white, with a row of long, stiff, stout white bristles at approx. the midpoint anteriorly, with dark brown bristles at ca. 1 / 4 posteriorly on dorsal surface, with white bristles ventrally. Forewing: eleven veins, R 4 + M 1 stalked, R 5 absent, anal vein furcate (Fig. 7 B). Forewing ground color shiny creamy white, somewhat rounded and pointed apically, with a brownish hue apically 1 / 5 of the way beyond costal margin; cilia well-fringed, brown from costal area before apex and brownish white along outer margin to inner base of wing. Hindwing: narrower than forewing, white to whitish brown; cilia well-fringed, white to brownish white; apex produced conspicuously. Male genitalia: (Figs 8 C, D, 9 B) eighth abdominal sternite triangular, long, slightly broadened at base, and then tapered toward posterior with a blunt tip. Uncus swollen, like a small tubercle, short and fine hairs on its top. Gnathos short and stout, its posterior margin closer to base with a hump-like outgrowth; closer to top, somewhat expanded, with a blunt tip. Tegumen much longer than uncus, with dense hairs at approx. the midpoint of its length on dorsal surface. Anellus lobe quite long and pear-shaped basally, bearing a rather long and strongly sclerotized spine-like process at apex, slightly acute and curved inward. Valva elongate, slender, slightly expanded basally, narrow to 3 / 4 of its length, moderately dilated, and lobate with fine, dense hairs on its inner surface apically, moderately curved inwardly, exceeded apex of tegumen. Vinculum narrow, bearing a few rather long spines on median process of vinculum. Saccus broad basally and U-shaped. Phallus long, small, and spherical at base, and extended distally. Female (Fig. 4 D). Forewing length 3.2 - 3.4 mm. Wing expanse 7.1 - 7.5 mm. Similar to male. Female genitalia: (Fig. 10 B) papillae anales nearly half the length of the apophysis posterioris, with two lobes and long and short fine setae on its entire surface. Apophysis posterioris as much as three times the length of apophysis anterioris. Ostium opening near posterior margin of 8 th sternite. Ductus bursae narrow, nearly equal in length to corpus bursae and moderately sclerotized. Ductus seminalis arising from the posterior third of ductus bursae. Corpus bursae oblong; signum long and a narrow arch shaped at left side wall of posterior end.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
4383E1E7B7305F67A5F593683DED7ACE.taxon	distribution	Distribution. Japan (Hokkaido, Honshu), Russia, Korea.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
4383E1E7B7305F67A5F593683DED7ACE.taxon	biology_ecology	Biology. Kogi (2008) reported that the larvae of T. angustella occur in August until the following June and make a portable case with fragments of a host plant leaf. The mature larvae use a larger piece of leaf like a hat. Dr. T. Oku collected a case from Q. mongolica in Iwate Prefecture in July.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
F3B4DBC7943850BCB0DD53F105DA7CCF.taxon	materials_examined	Material examined. Japan - Hokkaido [Hokkaido] • 1 ♂, 1 ♀; Katsuranosawa, Uryu-cho, Uryu-gun; 20 Jul. 2018; S. Yagi leg. • 1 ♂; Katsuranosawa, Ishikari city; 15 Jul. 2007; H. Kogi leg. • 1 ♂; Manzi, Kurisawa; 8 Jul. 2001; H. Kogi leg.; gen. slide no. KM- 143 • 1 ♂, 1 ♀; same locality and collector; 28 Jul. 2003; gen. slide no. KM- 144 (♂) • 1 ♂; Kotan Atuta; 10 Jul. 2003; H. Kogi leg. • 1 ♀; Nopporo (Atsubetsu); 27 Jul. 1993; T. Hirowatari leg. • 2 ♀♀; Kamisibun, Iwamizawa; 9 Jul 2011; 8 Aug. 2012; H. Kogi leg.; gen. slide no. KM- 145 • 1 ♀; Hukui, Niseko; 4 Aug. 2006; H. Kogi leg. • 1 ♀; Tiyosibetu, Hamamasu; 14 Jul. 2000; H. Kogi leg. • 1 ♀; Asari Pass, Otaru; 28 Jul. 2012; H. Kogi leg.; gen. slide no. KM- 146. - Honshu [Iwate] • 1 ♀; Atei-Sanso Niisato vill.; 6 Jul. 2002; T. Oku leg. (TO).	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
F3B4DBC7943850BCB0DD53F105DA7CCF.taxon	diagnosis	Diagnosis. The forewing is white with broad, dark brown fascia, a rather large yellow patch before the apex, and a black rounded apical spot demarcated by a white line. The anellus lobe of the male genitalia is a short and heavily sclerotized beak-shaped lobe basally, armed with a small claw-like process apically, which is a unique characteristic of this species. The apopyhsis posterioris of the female genitalia is ca. 1 / 2 the length of the papilla analis and approx. two times longer than the apophysis anterioris; the signum is absent in the corpus bursae.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
F3B4DBC7943850BCB0DD53F105DA7CCF.taxon	description	Description. Male. (Figs 4 A, 7 A). Forewing length 3.2 - 3.7 mm. Wing expanse 7.0 - 8.1 mm. Head: covered with shiny, creamy white appressed scales. Antennae filiform; basal segment (scape) elongate without pecten, creamy white; flagellum creamy white on dorsal surface before middle, then entirely grayish brown beyond on its dorsal and ventral surface, with rather long and fine cilia ventrally. Labial palpus white, long, and recurved; first segment shortest with creamy white scales; second segment thickened, ca. 2.5 the thickness times of first, covered entirely with white scales; third segment white on dorsal surface, sparse brown scales medially on ventral surface, as long as second segment, apex sharply acute. Thorax: creamy white. Tegula shiny, creamy white dorsally, with brown scales along anterior margin. Legs: white; forefemur, tibia, and tarsus suffused inwardly with brownish tinge, white on outer surface; mid legs entirely white; hind femur and tibia white, with a row of long, stiff, stout white bristles on upper and lower surfaces; all tarsal segments brownish gray. Forewing: eleven veins, R 3 + R 4 stalked, M 1 separate, R 5 absent, anal veins furcate (Fig. 7 A). Forewing broader in this genus, ground color white to middle, followed by a dark brown fascia, rather large yellow patch before apex not protruding to costal margin; small brown area in costal margin, intercepted by oblique white lines with a rounded black apical spot; small scattering of yellow scales below apical point at outer margin, small brown scale in tornal area; cilia blackish brown at inner margin of apex, outer margin with brown through inner base of wing. Hindwing: darker grayish brown with tiny black apical dot at apex; cilia well-fringed, grayish brown, with dark brown at tip of wing. Male genitalia: (Figs 8 A, B, 9 A) eighth abdominal sternite rather semicircular, broadly concave with strongly sclerotized margin. Uncus semicircular with a few hairs on top. Gnathos sickle-shaped, short and stout, wide and flattened posteriorly, and then curved at middle toward apex. Tegumen long, slightly concave medially, bearing dense hairs on dorsal surface at approx. the midpoint. Anellus lobe, a short and heavily sclerotized beak-shaped lobe, approx. half the length of valva, slightly narrow at base, and then abruptly dilate with short and small claw-like process apically, with a few sclerotized spines at approx. the midpoint on inner dorsal surface. Valva elongate, slender, and curved inwardly, somewhat broad basally, narrowly elongate from base to 2 / 3 of its length, then distended again at apex, bearing thin and fine setae on its inner surface. Vinculum moderately broad, median process rising to a pair of transverse ridges of short, thorn-like spines on edge of posterior surface, rather long and short fine setae emerging around and below the ridge surface. Saccus broad basally, somewhat triangularly produced. Phallus short and sclerotized, clavate basally and straightly elongate with a round tip distally. Female. (Figs 1, 4 B). Forewing length 2.5 - 3.9 mm. Wing expanse 5.5 - 8.2 mm. Similar to male. Female genitalia: (Fig. 10 A) papilla analis long and bilobed with short and long fine setae on its entire surface. Apophyses short; apophysis anterioris shorter, nearly 1 / 2 length of apophysis posterioris. Ostium opening near anterior margin of 8 th sternite. Ductus bursae partially sclerotized, narrow, and slightly elongate, as long as corpus bursae. Corpus bursae large and rounded; signum absent.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
F3B4DBC7943850BCB0DD53F105DA7CCF.taxon	distribution	Distribution. Japan (Hokkaido, Honshu), Russia, Korea.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
A1D2408ECD4B5B80B43012FE5C330042.taxon	materials_examined	Material examined. Japan - Honshu [Chiba] • 1 ♀; Kayano, Orikisawa, Kimitsu-shi; 4 Sep. 2013; O. Saito leg. • 1 ♂; Otake, Narita-shi; 23 Jul. 2016; O. Saito leg. • 1 ♀; same locality and collector; 20 Aug. 2016 • 1 ♂; same locality and collector; 3 Jun. 2017 • 1 ♀; same locality and collector; 9 Sep. 2017 • 1 ♀; same locality and collector; 7 Oct. 2017. - Honshu [Ishikawa] • 1 ♀; Hodatsushimizu cho, Shikinami; 16 Jun. 2018; S. Tomura leg. - Honshu [Nagano] • 2 ♀♀; Kojiro, Tenryu-mura; 19 Jun. 2009; 28 Aug. 2009; N. Hirano leg. • 1 ♂; Hiraoka, Tenryu-mura; 16 Jun. 2007; N. Hirano leg. [Gifu] • 1 ♀; Yamagata-gun, Miyama-cho, Iodo; 17 Jun. 1994; T. Mano leg.; gen. slide no. KM- 140; OPU. - Honshu [Shizuoka] • 1 ♂; Oonogi Umegasima, Shizuoka-city; 25 Aug. 2001; T. Mano leg.; gen. slide no. KM- 32; OPU • 1 ♀; Shizuoka city, Hirano, Abe river; 23 Aug. 1997; T. Mano leg.; OPU • 1 ♀; Hirano, Shizuoka city; 26 Aug. 1995; T. Mano leg.; OPU • 1 ♀; Konya Spa, Shizuoka city; 27 Jul. 2002; T. Oku leg.; TO. - Honshu [Aichi] • 3 ♂♂; Zaikaji-temple, Toyokawa city; 4 Jun. 1994; T. Mano leg.; gen. slide no. KM- 18; OPU • 1 ♂; Somasaka Pass, Toyokawa city; 21 May. 1992; T. Mano leg.; gen. slide no. KM- 20; OPU • 2 ♀♀; same label; gen. slide no. KM- 79; OPU • 1 ♀; Matsumine-cho, Toyota city; 13 Jun. 2001; T. Mano leg.; gen. slide no. KM- 77; OPU. - Honshu [Mie] • 1 ♂; Yamadano (Hakusan-tyo); 18 Apr. 2001; Host: Mallotus japonica (" shinme " [= a shoot]); 30 Apr. 2001 em.; T. Saito leg.; OPU • 1 ♂, 1 ♀; same locality and collector; 23 Apr. 2002; gen. slide no. KM- 12 (♂); OPU • 6 ♂♂, 2 ♀♀; same label; 23 Apr. 2002; Host: Mallotus japonica (" shinme " [= a shoot]); 30 Apr. - 6 May. 2002 em.; gen. slide no. KM- 65; OPU • 1 ♀; Obara-ishiki, Hokusei-cyo; 7 Sep. 1997; T. Mano leg.; OPU • 1 ♀; Mikuni valley alt 430 m, Hujiwara-cho; 27 Aug. 1998; T. Mano leg; OPU • 1 ♂; Hijiki 250 m, Ueno City; 12 Sep. 1997; T. Mano leg.; gen. slide no. KM- 31; OPU • 1 ♀; Inabe-gun, Fujiwara-cho, Shinodachi; 29 Aug. 1994; T. Mano leg.; OPU. - Honshu [Shiga] • 1 ♂; Makino, Takashima-shi; 13 May. 2015; H. Shimizu leg. - Honshu [Kyoto] • 1 ♂; Mt. Ponpon; 1 Jul. 2000; N. H. Ahn leg.; gen. slide no. KM- 34; OPU • 1 ♂, 1 ♀; Yamanouchi-cho, Seikanji, Higashiyama-ku, Kyoto-shi; 1 Jun. 2013; H. Shimizu leg. • 4 ♀♀; Kanmurijima I., Miyazu-shi; 27 - 28 Sep. 1982; M. Sasakawa; Y. Yoshiyasu; N. Nishida & T. Kamura leg.; gen. slide no. KM- 63; OPU. - Honshu [Nara] • 1 ♀; Kasugayama (Nara-si); 26 Aug. 2000; T. Saito leg.; OPU • 1 ♂; Mt. Takatoriyama (Takatori) ca 580 m; 25 Aug. 1993; Kadohara leg.; OPU • 1 ♀; Katuragi-shi, Taima; 7 Aug. 2009; H. Shimizu leg. • 1 ♀; Sannoko, Kawakami village; 18 Jun. 1991; T. Ueda leg.; OPU • 1 ♂; Kawakami village, Kitamata; 15 Jun. 1993; T. Ueda leg.; gen. slide no. KM- 30; OPU. - Honshu [Hyogo] • 1 ♂; Tamida (Inagawa-tyo); 29 Apr. 2002; T. Saito leg.; Host: Mallotus japonica; 3 May. 2003 em.; gen. slide no. KM- 33; OPU. - Honshu [Osaka] • 1 ♀; Aokaiyama (Toyono-tyo); 10 Jun. 1999; T. Saito leg.; gen. slide no. KM- 62; OPU • 1 ♀; Minou River, Minou City; 5 Aug. 1991; M. Aoyagi & T. Ueda leg.; OPU • 1 ♂; Higasiosaska-shi, Rokumanji-cho; 23 May. 2015; H. Shimizu leg. • 1 ♂; same locality and collector; 18 May. 2017 • 1 ♂; Yao-shi, Kodachi, Jyusan-toge, Fumin no mori; 16 Jul. 2011; H. Shimizu leg. • 1 ♀; same locality and collector; 17 May. 2016 • 1 ♀; Hatigamine; 4 Jun. 1993; S. Koshino leg.; gen. slide no. KM- 66; OPU • 1 ♂, 1 ♀; same locality and collector; 15 Jul. 1993; gen. slide no. KM- 19 (♂), 126 (♀); OPU • 1 ♀; Minamikawachi-gun, Mizukoshi-toge, Mt. Kongo; 7 Jun. 2015; H. Shimizu leg. • 2 ♀♀; Tondabayasi Dakeyama ca. 260 m; 4 Sep. 1992; Kadohara leg.; OPU • 1 ♀; Kawachinagano city, Iwawaki temple; 3 May. 2001; T. Saito leg.; Host: Mallotus japonica; 20 May. 2001 em.; OPU • 1 ♀; Kawati, Iwawakisan; 29 May. 1954; T. Yasuda leg.; OPU • 1 ♀; Kawakubo; 19 Sep. 1995; S. Kosino leg.; gen. slide no. KM- 72; OPU • 1 ♀; Tottori (Hannan); 18 May. 1997; S. Kosino leg.; OPU • 3 ♀♀; Izumi-katuragisan (Kisiwada-si); 17 Jul. 2004; T. Saito leg.; gen. slide no. KM- 42; KM- 64; OPU • 1 ♀; same locality and collector; 21 Jul. 2004; OPU • 1 ♀; Kyosi, 21 May. 1998; S. Kosino leg.; gen. slide no. KM- 128; OPU. - Honshu [Wakayama] • 1 ♀; Wakayama-shi, Nougawa; 11 Jun. 2007; M. Murase leg.; OPU • 1 ♂, 1 ♀; Wakayama-shi, Yata; 3 Jul. 2001; T. Hirowatari; B. W. Lee; N. H. Ahn; Y. Miyamoto & K. Yamada leg.; gen. slide no. KM- 35 (♂), 60 (♀); OPU. - Shikoku [Ehime] • 1 ♀; Matsuyama; 16 May 1957; M. Okada leg.; gen. slide no, KM- 71; OPU. - Kyushu [Fukuoka] • 1 ♂; Orio; 4 Jul. 1958; T. Kawamura leg.; gen. slide no. KM- 21 • 2 ♀♀; Orio; 24 Jun. 1959; T. Kawamura leg.; gen. slide no. KM- 78 • 1 ♂; Kasii, Fukuoka City; 25 May. 1958; K. Yano leg.; gen. slide no. KM- 37 • 1 ♂; Aburayama, Fukuoka; 24 May. 1959; T. Kawarabata leg.; gen. slide no. KM- 26 • 1 ♂; Atago, Nishi-ku, Fukuoka shi; 26 May. 2018 larva; Host: Mallotus japonica; 9 Jun. 2018 em.; S. Yagi leg. • 1 ♀; same locality, collector and host; 9 Jun. 2018 larva; 9 Jul. 2018 em • 2 ♂♂; Ito campus, Nishi-ku; 26 May. 2017; K. M. M. Kyaw; gen. slide no. KM- 43, 44 • 1 ♂; Mt. Hikosan; 18 Jun. 1962; H. Kuroko leg.; OPU • 1 ♀; same locality and collector; 1 Sep. 1953; OPU • 1 ♀; same locality; 20 Aug. 2013; S. Yagi leg.; gen. slide no. KM- 70 • 1 ♀; same locality and collector; 25 Jul. 2014; LT; gen. slide no. KM- 69 • 1 ♀; same locality; 31 Jul. 2014; LT; T. Hirowatari & S. Yagi leg. • 1 ♂; same locality; 4 Sep. 2014; LT, S. Yagi leg.; gen. slide no. KM- 15 • 1 ♂, 4 ♀♀; same locality and collector; 27 Jul. 2015; LT; gen. slide no. KM- 17 (♂), 73 (♀), 75 (♀) • 1 ♂; same locality and collector; 13 Aug. 2016; LT; gen. slide no. KM- 36. - Kyushu [Saga] • 1 ♂; Hokuzan, Saga-shi; 3 Jun. 2017; S. Tomura leg. - Kyushu [Kagoshima] • 1 ♀; Sata; 18 May. 1952; T. Kodama leg.; OPU. - Ryukyus [Kagoshima] • 1 ♀; Nakama, Yakusima Is.; 20 Sep. 1978; S. Moriuti leg.; OPU • 1 ♂; Okawa Rindo, Kurio, Yakushima, 120 m; 21 Jun. 2017; S. Yagi leg. • 2 ♂♂, 2 ♀♀; Hatsuno, Amamioshima Is.; 3 - 5 Apr. 1996; T. Ueda leg.; gen. slide no. TU- 751 (♂), KM- 14 (♂), 68 (♀); OPU • 1 ♀; Mt. Yuwan-dake (Lower), Uken-son vill.; 18 May. 2015; S. Sameshima leg.; gen. slide no. KM- 57; KGU • 1 ♀; Akatsuchiyama, Yuwan, Uken-son; 20 Aug. 2014; S. Sameshima leg.; KGU • 1 ♀; same locality; 6 Jul. 2016; LT 245 m; S. Yagi leg.; gen. slide no. KM- 83 • 1 ♂, 3 ♀♀; Chuorindo, Amamioshima Is.; 6 Apr. 1996; T. Ueda leg.; gen. slide no. KM- 25 (♂), 67 (♀); OPU • 1 ♀; Nishinakama, Amamioshima Is.; 6 Jun. 1996; T. Ueda leg.; OPU • 1 ♂; Fureainomori, Uken vill.; 25 Sep. 2002; gen. slide no. KM- 27; Host: Mallotus japonica; OPU • 1 ♂; Mt. Yui-dake, Setouchi-cho, Amamioshima Is.; 30 Jun. 2006; U. Jinbo leg.; NSMT-I-L- 35780; • 1 ♀; Tokuno-shima, San Tokunoshima; 9 Jul. 2016; LT 230 m; S. Yagi leg.; gen. slide no. KM- 59 • 1 ♀; Tokuno-shima, Fugusuku; 11 Jul. 2016; S. Yagi leg. - Ryukyus [Okinawa] • 1 ♂; Mt. Terukubi 330 m; Benoki, Kynigami-son; 5 Aug. 2015; L. T; S. Yagi leg.; gen. slide no. KM- 29 • 1 ♀; Yona, Kunigami vill.; 15 - 18 May. 1998; T. Ueda leg.; OPU • 1 ♀; Mt. Fuenchijisan, Kunigami vill.; 8 Jun. 1997; T. Ueda leg.; gen. slide no. KM- 74. • 1 ♀; Hentona; 9 Aug. 2016; LT 60 m; S. Yagi leg. • 1 ♀; Uka, Kunigami-son, Kunigami-gun; 31 May. 2015; 250 m LT; S. Yagi leg. • 1 ♀; Banna Park, Ishigaki Is.; 31 Mar. 2002; B. W. Lee leg.; OPU; gen. slide no. KM- 87 • 1 ♀; Mt Omoto, Ishigaki Is.; 11 Nov. 2003; Host: Mallotus japonica; 18 Nov. 2003; S. Shimizu leg.; OPU; gen slide no. KM- 76.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
A1D2408ECD4B5B80B43012FE5C330042.taxon	diagnosis	Diagnosis. This species is similar to T. biformis, which is known in the Russian Far East and Japan but can be distinguished by markings in the distal yellow zone of the forewing with a distinct black streak below the middle of the yellow zone. The male genitalia is similar to those of T. epiclista Meyrick, 1908 described from Khasi Hills, India, but can be distinguished by the basally broadened valva and the presence of a thumb-like basal process bearing numerous setae, the lack of a pre-apical spine on the costal process of the valva, and the slender posterior part of the phallus.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
A1D2408ECD4B5B80B43012FE5C330042.taxon	description	Description. Male (Figs 4 G, 7 D). Forewing length 2.7 - 4.3 mm. Wing expanse 5.8 - 8.8 mm. Male genitalia: (Figs 8 G, H, 9 D). Female (Fig. 4 H). Forewing length 3.0 - 4.4 mm. Wing expanse 5.9 - 9.1 mm. Similar to male. Female genitalia: (Fig. 10 D) see Park and Kim (2016) for detailed descriptions of the adults and genitalia. In the present study, intraspecific variation observed was the presence or absence of a blackish streak on the yellowish zone of the forewing markings in all examined specimens. Additionally, in contrast, the biggest size of the adult moth of Japanese specimens (wing span 8.8 mm) is quite smaller than that of Korean specimens (wing span 12 mm in Park and Kim's description). In the female genitalia also, Park and Kim described the shape of the signum forming as an elongate plate with dense spicules and located at posterior end; however, that of the Japanese specimens is developing as a pentagonal shape at the posterior end in our observation.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
A1D2408ECD4B5B80B43012FE5C330042.taxon	distribution	Distribution. Japan (Honshu, Shikoku, Kyushu, Ryukyus), Korea.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
A1D2408ECD4B5B80B43012FE5C330042.taxon	biology_ecology	Biology (Fig. 11). Although the host plant of T. chujaensis is unknown, we found that some specimens of this species preserved in OPU were labeled as having been reared on Mallotus japonica (Euphorbiaceae). In the present study, we confirmed that the larvae feed on this plant in the field. The larvae also make portable cases with flower buds or attack the leaf bud (young shoot) of the host plant. The larva uses the flower buds at flowering time to make a case from the plant. After making the portable case, the larva moves from one place to another and attaches the case to the lower surface of the leaf for pupation (Fig. 11 C). When there is an early leaf bud (young shoot) on the host plant, the larva penetrates the petiole of the young, newly emerged leaf and feeds inside (Fig. 11 E-G). There is no external injury during the feeding period until the leaf is fully grown. Before pupation, the larva cuts the petiole and makes a portable case. After that, it fixes the portable case sideways. Pupation also takes place inside the cases and pupal exuvia is left after the adult emerges.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
EB01F302B41C555194E3114082D4FF0F.taxon	diagnosis	Diagnosis. At a glance, the external features are similar to those of T. chujaensis (Park, 2016) comb. nov. but it can be distinguished by wing markings in the distal yellow zone of the forewing, which lacks a distinct blackish streak below the middle of the yellow zone and features grayish scales at the costal margin before the apex and the area beyond the tornus. Additionally, it can easily be distinguished based on the male genitalia; the uncus is more rounded apically; the gnathos is U-shaped and acute apically; the valva is narrowly elongate with a sharped pre-apical process ca. 1 / 4 of the way along its length and the vinculum lacks thumb-like lobes posteriorly; the saccus has a rounded base. The shape of the phallus is also different. However, the male genitalia are similar to those of Thiotricha clidias Meyrick, 1918, which was described from Khasi Hills, India, although they differ in the shape of the phallus. In T. clidias Meyrick, 1918, the phallus is rounded basally, abruptly sinuate and slender in distally but as a cucurbit-shaped in T. elaeocarpiella sp. nov.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
EB01F302B41C555194E3114082D4FF0F.taxon	description	Description. Male (Figs 5 A, 6 A, 7 E) Forewing length 2.9 mm in holotype, 2.6 - 3.4 mm in paratypes. Wing expanse 6.4 mm in holotype, 5.2 - 7.1 mm in paratypes. Head: shiny creamy white with appressed scales. Antennae filiform, basal segment elongate and creamy white, sparsely speckled with brown scales; flagellum grayish white on dorsal surface before midpoint, then brownish gray beyond on its dorsal and ventral surfaces, with extraordinarily long and fine cilia ventrally. Labial palpus white, moderately long and recurved; first segment approximately half the length of the second, with blackish gray scales on lateral surface; second segment as much as 1.5 times the length of the first, creamy white throughout on outer surface; bundle of hair pencils arising from apex of first and second segment, appressed on dorsal surface to near the end of the third segment; third segment as thick as second, with blackish gray scales medially on lower surface ventrally, shiny creamy white evenly on both surfaces, apex sharply acute (Fig. 6 A). Thorax: creamy white. Tegula shiny, creamy white dorsally, ornamented with blackish gray scales along anterior margin. Legs: white; forefemur, tibia, and tarsus suffused inwardly with blackish brown; scattered with white scales on outer surface; mid femur entirely white; mid tibia and tarsus white but slightly speckled with blackish brown scales on outer surface; hind femur white; hind tibia creamy white, with a row of long, stiff, stout, creamy white bristles above and below, suffused with a small blackish gray scale on lateral outer surface posteriorly; first tarsal segment entirely blackish gray; second and third segment white with blackish gray apical ring; last two segments white. Forewing: eleven veins, R 4 + M 1 stalked, R 5 absent, anal vein furcate (Fig. 7 E). Forewing ground color shiny grayish white to white to ca. 3 / 4 of the way from base; distinct orange zone in distal 1 / 4, deeply concave along costal margin; costal margin and area beyond tornus grayish colored; small black spot at apex, narrowly connected to another black spot in tornus; cilia before apex to tornus brown, creamy yellow from tornus to inner base of wing. Hindwing: narrower than forewing, creamy white to grayish white, with pale orange apical zone; apex sharply produced, with small apical black spot; cilia well-fringed to base, fringe around apex creamy white, with broad, dark brown median band. Male genitalia: (Figs 8 I, J, 9 E) eighth abdominal sternite more or less triangular, emarginate at the tip, slightly broadened basally with moderately sclerotized margin anteriorly, gradually narrow toward apex. Uncus directed backwards, with broad basal expansion, then narrowly elongate, forming a furrow on lower surface medially, bearing short spines on its lateral margin and abruptly rounded with short and fine setae evenly on its apical dorsal surface. Gnathos U-shaped, stout, strongly bent at basal 1 / 3, sharply acute apically. Tegumen longer than uncus, slightly concave medially on lateral margins with dense hairs on dorsal surface beyond middle. Anellus lobe, a large process, as much as 1 / 2 the length of process of valva, ovate membranous pouch at base, with short sclerotized apical spine and short fine setae around apical spine. Valva slender, elongate, broad basally, narrow along 2 / 3 of length, then dilated apically with dense, long, fine hairs hanging down from its inner surface and developing a sclerotized point, spine-like pre-apical process arising from its base, nearly 1 / 4 of apex. Vinculum long and slightly narrow, with few rather long setae on its lateral margin. Saccus roundly produced basally. Phallus cucurbit-shaped in basal half, slightly sinuate, slender and recurved upwardly in distal half. Female (Figs 5 B, 6 B). Forewing length 2.5 - 3.3 mm. Wing expanse 5.3 - 7.1 mm. Similar to male but differs as follows: Labial palps of first segment shortest, with creamy white scales, as thick as second segment; second segment as long as third segment and with white scales on lower surface and grayish or grayish brown scales on upper surface; third segment slender and acute with gray to grayish-brown scales on both surfaces (Fig. 2 B) Female genitalia: (Fig. 10 E) papillae anales with long and short fine setae on its entire surface. Apophysis posterioris longer than apophysis anterioris; apophysis anterioris ca. 1 / 3 the length of posterioris. Ductus bursae rather long, narrow, slightly sclerotized along the posterior half of its length. Corpus bursae clavate in shape; signum rounded at center.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
EB01F302B41C555194E3114082D4FF0F.taxon	distribution	Distribution. Japan (Kyushu, Ryukyus).	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
EB01F302B41C555194E3114082D4FF0F.taxon	etymology	Etymology. The name refers to its main host plant, Elaeocarpus zollingeri.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
EB01F302B41C555194E3114082D4FF0F.taxon	biology_ecology	Biology (Fig. 12). The larva uses the flower bud or the young shoot of its host plant to construct portable cases. When it utilizes a flower bud, at first, the larva penetrates the bud and then lives and feeds within it. After that, it moves from one flower to another by carrying the bud and attaching it to other flower buds to complete its life cycle (Fig. 12 D, E). When the larva is ready to pupate, it attaches the case to the underside of a leaf with silk. When it utilizes a young shoot (Fig. 12 C), the larva leaves small dot-like traces of feeding after making cases by the shoot. Pupation also takes place inside the portable cases. After completing development, the adult emerges from the case, leaving the pupal exuvia inside.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
2DAD0FF80CAB50E6B1E5B91FAD140889.taxon	diagnosis	Diagnosis. The external morphological character of this new species is quite similar to that of T. angustella; the wings of both species are shaded brown apically. However, the two can be differentiated based on the brightness of the color of the apical wing markings. In the new species, the wings feature a huge dark brown area distally, so it can be recognized easily at a glance. Likewise, in the male genitalia, the anellus lobe is a small membranous spherical lobe basally with a delicate, thread-like spine apically. Also, the size of apophyses and shape of the signum in the female genitalia are different to those of T. angustella. On the other hand, the male genitalia are quite similar to those of Thiotricha xanthodora Meyrick, 1923, which was described from Pyinmana, Myanmar, but differ in terms of the uniformly elongate valva, spherical-shaped and straight phallus. In T. xanthodora, the shape of the valva is dilated along 1 / 3 of its length apically whereas the phallus is slightly rounded basally and twisted forward.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
2DAD0FF80CAB50E6B1E5B91FAD140889.taxon	description	Description. Male (Figs 5 C, E, 7 F). Forewing length 3.3 mm in holotype, 2.5 - 3.2 mm in paratypes. Wing span 6.2 mm in holotype, 5.6 - 6.9 mm in paratypes Head: shiny creamy white with appressed scales. Antennae filiform, basal segment rather large and elongate, white, sparsely speckled with brown scales on dorsal surface; flagellum creamy white on dorsal surface before middle, then entirely grayish brown beyond on both surfaces with extraordinarily long and fine cilia ventrally. Labial palps white, long, and recurved; first segment shortest, creamy white with brown scales medially on outer surface; second segment thickened with white scales evenly on both surfaces, as much as 2.5 times the length of the first; third segment as long as second segment, entirely grayish brown, considerably acute and slender. Thorax and tegula: creamy white. Legs: white; forefemur and tibia inwardly suffused with brown and white on outer surface; fore tarsus completely brown; mid femur and tibia entirely white; mid tarsus with brown; hind femur and tibia creamy white, with long, stiff, white bristles until the midpoint anteriorly, brown bristles on upper surface at ca. 1 / 4 beyond half of its length posteriorly, with white bristles ventrally; all tarsal segments dark grayish in color. Forewing: eleven veins, R 3 + R 4 stalked, M 1 separate, R 5 absent, anal veins furcate. (Fig. 7 F). Forewing rather broad, rounded and slightly pointed apically, ground color creamy white from base along 2 / 3 of wing, significantly occupied with a huge dark brown or pale-yellow area along costal margin to apex at ca. 1 / 3 apically; cilia well-fringed and dark brown or yellowish brown before apex to inner base of wing. Hindwing: narrower than forewing, brownish white, pale brown; cilia well-fringed around apex and then white to anterior rim of base. Male genitalia: (Figs 8 K, L, 9 F) eighth abdominal sternite obtuse, slightly emarginate at the tip, short, and broadened from base toward apex. Uncus swollen and rather small, like a tubercle, with long and short fine setae on its dorsal surface. Gnathos short and stout, slightly flattened posteriorly, then moderately curved apically. Tegumen extremely long and larger than uncus, with a cluster of dense hairs at approximately its midpoint dorsally. Anellus lobe, a small membranous rounded lobe basally, bearing a flexible and weakly sclerotized thread-like spine, reaching toward nearly 2 / 3 of valva, slightly curved inwardly, a few spines arising around it. Valva simple, uniformly elongate, broad basally, narrowly elongate from base toward apex, with numerous long fine hairs on its ventral surface. Vinculum narrow and elongate, with long and short fine hairs on rim of its surface posteriorly. Saccus somewhat rounded and triangular in shape. Phallus spherical basally, becoming narrow and straight, then slender in distal half. Female (Fig. 5 D, F). Forewing length 2.5 - 3.1 mm. Wing expanse 5.6 - 6.5 mm. Similar to male. Female genitalia: (Fig. 10 F) papillae anales nearly equal in length to posterior apophysis, with long and short fine setae on its entire surface. Apophysis posterioris longer and apophysis anterioris nearly half the length of posterior. Ostium opening close to anterior margin of 8 th sternite. Ductus bursae rather broad, uniformly elongate, nearly same length as corpus bursae. Ductus seminalis arising ca. 1 / 3 of posterior of ductus bursae. Corpus bursae oblong in shape with narrow; signum short and arch-shaped at left side wall of posterior end.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
2DAD0FF80CAB50E6B1E5B91FAD140889.taxon	distribution	Distribution. Japan (Ryukyus).	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
2DAD0FF80CAB50E6B1E5B91FAD140889.taxon	etymology	Etymology. The name refers to the coloration of the forewing (yellow apically).	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
C33C34F7B36855CA93AB379B67EF3C27.taxon	materials_examined	Material examined. Japan - Hokkaido [Hokkaido] • 2 ♂♂; Katsuranosawa, Uryu-cho, Uryu-gun; 20 Jul. 2018; S. Yagi leg. • 1 ♂, 1 ♀; Tomuraushi, Shintoku town; 20 Aug. 2000; H. Kogi leg.; gen. slide no. KM- 148 (♂) • 2 ♂♂; Tokachigaoka, Otofuke-cho; 13 Jul. 2000; T. Hirowatari; N. H. Ahn; Y. Miyamoto; H. Okamoto; K. Yamada leg.; gen. slide no. KM- 4, 47; OPU • 1 ♀; Fukuyama, Hobetu Town; 12 Jul. 2005; H. Kogi leg. • 1 ♀; Siratukari, Atuta; 21 Apr. 2003 em.; H. Kogi leg. • 1 ♀; Sibi Isikari; 5 Jul. 2016; H. Kogi leg. • 1 ♂, same locality, 8 Jul. 2007, H. Kogi; 1 ♂; Ishikari-hama, Ishikari-shi; 18 Jul. 2018; S. Tomura leg. • 3 ♂♂, 1 ♀; Oyafunebochi, Oyafune-cho, Ishikari; 18 Jul. 2018; S. Yagi leg. • 1 ♂; Sinkoh, Isikari; 15 Jun. 2003 em.; Host: Quercus dentata; H. Kogi leg.; gen. slide no. KM- 142 • 1 ♀; same locality and collector; 25 Jul. 2005 em; Host: Quercus crispula; H. Kogi leg. • 1 ♂; Moiwa, Tomari; 24 Jul. 2006; H. Kogi leg.; gen. slide no. KM- 141 • 1 ♀; Asari-pass, Otaru; 22 Jul. 2002; H. Kogi leg. • 1 ♀; Tomakomai shi, Kashiwabara; 21 Jul. 2018; S. Tomura leg. - Honshu [Iwate] • 1 ♂; Dogamori, Morioka; 13 Jul. 1994; N. Doi leg.; TO. - Honshu [Chiba] • 1 ♂; Otake, Narita-shi; 18 Jun. 2016; O. Saito leg. - Honshu [Gifu] • 1 ♀; Oniiwa-onsen, Hiyoshi, Mizumani; 1 Jun. 2017; S. Yagi leg. - Honshu [Nagano] • 1 ♂; Kojiro, Tenryu mura; 5 Jul. 2008; N. Hirano leg.; gen. slide no. KM- 108 • 1 ♀; Reisengoya 2260 m, Mt. Norikura, Azumi-vill; 20 Jun. 2001; T. Mano leg.; OPU. - Honshu [Aichi] • 1 ♂; Asahi-highland, Asahi-cho; 13 Jul. 1996; T. Mano leg.; gen. slide no. KM- 50; OPU • 1 ♀; same locality and collector; 7 Jul. 2001; OPU. - Honshu [Kyoto] • 1 ♂; Mt. Ponpon; 1 Jul. 2000; N. H. Ahn leg.; gen. slide no. KM- 117; OPU • 2 ♀♀; same label; gen. slide no. KM- 48; OPU - Honshu [Osaka] • 1 ♂; Ikoma, 30 Jun. 1995; S. Kosino leg.; gen. slide no. KM- 135; OPU • 1 ♂; Rokumanji-cho, Higashiosaka-shi; 3 Sep. 2017; H. Shimizu leg. • 1 ♂, 1 ♀; Aokaiyama (Toyono-tyo); 7 Jul. 1999; T. Saito leg.; gen. slide no. KM- 5 (♂), KM- 49 (♀); OPU • 1 ♀; same label; 21 Jun. 1999; gen. slide no. KM- 52; OPU • 1 ♀; Iwawaki; 28 Jun. 1952; T. Kodama leg.; OPU • 1 ♀; Izumi-katuragisan (Kisiwada-si); 17 Aug. 2004; T. Saito leg.; gen. slide no. KM- 115; OPU. - Honshu [Hiroshima] • 1 ♂, 2 ♀♀; Yoshiwa, Hatsukaichi-shi; 15 - 16 Jul. 2017; S. Tomura leg. - Kyushu [Fukuoka] • 1 ♂; Nokonoshima, Fukuoka; 3 Jun. 2013; S. Yagi leg.; gen. slide no. KM- 101 • 1 ♂; Hikosan, Soeda-machi; 1 Jul. 2016; S. Yagi leg.; gen. slide no. KM- 13 • 1 ♂; same label; gen. slide no. KM- 28 • 1 ♂; same label; gen. slide no. KM- 97. - Kyushu [Kumamoto] • 1 ♂; Taziri Ubuyama; 14 Jul. 2013; S. Yagi leg.; gen. slide no. SY- 1.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
C33C34F7B36855CA93AB379B67EF3C27.taxon	diagnosis	Diagnosis. The forewing is white with a small orange patch and a black rounded spot apically. The anellus lobe of the male genitalia is narrow, slender, and longer than the valva, with a sharp and thorn-like spine at the apex, which is a unique characteristic of this species. The apopyhsis posterioris of the female genitalia is ca. 1 / 3 of the length of papilla analis and approx. half the length of the apophysis anterioris; the signum is absent in the corpus bursae.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
C33C34F7B36855CA93AB379B67EF3C27.taxon	description	Description. Male (Figs 4 E, 7 C). Forewing length 2.6 - 3.6 mm. Wingspan 5.5 - 7.6 mm. Head: shiny, creamy white with appressed scales. Antennae filiform, basal segment elongate and white; flagellum whitish brown on dorsal surface before middle, then entirely grayish brown beyond, with rather long and fine cilia ventrally. Labial palpus white, long, and recurved; first segment shortest, creamy white; second segment thickened, as much as 2.5 times the length of the first, covered evenly with creamy white scales; third segment as long as second, creamy white, apex sharply acute. Thorax and tegula: creamy white. Legs: white; forefemur, tibia, and tarsus suffused inwardly with dark brown, white on outer surface; mid legs entirely white; hind femur and tibia white, dispersed inwardly with brown; with compact ventral and dorsal rows of long, stiff, stout white bristles; all tarsal segments grayish brown. Forewing: eleven veins, R 3 + R 4 stalked, M 1 separate, R 5 absent, anal vein furcate (Fig. 7 C). Forewing ground color creamy white to white to ca. 3 / 4 of the way from base, large orange patch of more or less inverted triangular shape along costal margin, extending to apex of wing; large round black spot apically, bordered by a white line with an orange patch; outer margin blackish brown; small brown scales below apical point; diffused brown scales below orange patch at tornal area; cilia well-fringed, brown-white on costal margin before apex, dark brown with fuscous median band from apex to termen, grayish brown through inner base of the wing. Hindwing: narrower than forewing, brown with tiny dark brown apical dot; fringe around apex darker in color, long, and brown, cilia well-fringed on inner region of hind wing. Male genitalia: (Figs 8 E, F, 9 C) eight abdominal sternite mucronate, short and wide surfaces, anterior 2 / 3 broadly concave, and then narrow beyond, slightly sharpened basally. Uncus swollen and rounded with short, fine setae on apex. Gnathos sickle-shaped, short and stout, flattened at base posteriorly and then slightly curved toward apex. Tegumen long, nearly three times length of uncus, with dense hairs at approximately its midpoint on dorsal surface. Anellus lobe, a pair of slender processes, slightly longer than valva, slightly broadened and elbowed basally, then narrowly elongate and slightly swollen 1 / 4 of the way before apex, with rather long and weakly sclerotized thorn-like spine at tip. Valva enlarged at base, with long fine setae on rim of inner surface, gradually narrowing from base to 2 / 3 of length, slightly curved inwardly, a few setae at corner of anterior margin, rhomboid with a blunt tip apically, short and fine setae on inner and outer surfaces. Vinculum slightly narrow and enlarged surface, a few setae on its median process. Saccus broad basally, somewhat inflate and convex. Phallus large and clavate basally, narrowly elongate in distal half. Female (Fig. 4 F). Forewing length 2.6 - 3.5 mm. Wing expanse 6.2 - 7.2 mm. Similar to male. Female genitalia: (Fig. 10 C) papilla analis bilobed, ca. twice length of apophysis anterioris, with long and short fine setae on its entire surface. Apophyses short but apophysis anterioris ca. two times longer than apophysis posterioris. Ostium opening near anterior margin of 8 th sternite. Ductus bursae nearly as long as corpus bursae and weakly sclerotized. Corpus bursae slightly oblong; signum absent.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
C33C34F7B36855CA93AB379B67EF3C27.taxon	distribution	Distribution. Japan (Hokkaido, Honshu, Kyushu), Russia, Korea.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
C33C34F7B36855CA93AB379B67EF3C27.taxon	biology_ecology	Biology. Kogi (2004) reported that adults of this species fly in July in Hokkaido and larvae live in portable cases in summer until the following spring. In late fall, the larvae move to the underside of twigs of the host plant for hibernation. The next spring, the larvae create triangular cases like hats with fragments of host plant leaves and skeletonize the leaves.	en	Kyaw, Khine Mon Mon, Yagi, Sadahisa, Oku, Jouhei, Sakamaki, Yositaka, Hirowatari, Toshiya (2019): Taxonomic study of Thiotricha Meyrick (Lepidoptera, Gelechiidae) in Japan, with the description of two new species. ZooKeys 897: 67-99, DOI: http://dx.doi.org/10.3897/zookeys.897.38529, URL: http://dx.doi.org/10.3897/zookeys.897.38529
