taxonID	type	description	language	source
DD17C27CFFF7FFBCFF00D7594FE0FF14.taxon	type_taxon	Type species. Chartocinus rarus n. sp., by original designation.	en	Ng, Peter K. L., Rahayu, Dwi Listyo (2022): A new genus and new species of Hymenosomatidae (Crustacea: Brachyura) from Indonesian Papua. Zootaxa 5092 (4): 480-486, DOI: https://doi.org/10.11646/zootaxa.5092.4.7
DD17C27CFFF7FFBCFF00D7594FE0FF14.taxon	diagnosis	Diagnosis. Carapace longitudinally subovate, distinctly dorsoventrally flattened; dorsal surface almost flat, surrounded by continuous distinct rim except at base of rostrum; regions not discernible, without any distinct grooves, H-shaped gastric groove shallow, joining cervical groove, no grooves on posterior part; carapace rim and margin of rostrum lined with dense short, hooked setae that partially obscure surface; rostrum unilobate, subtriangular, no lateral lobes present; anterolateral margin unarmed; posterolateral margin with 3 low, wide lobes above P 3 and P 4 coxae; orbits clearly visible in dorsal view, eyes mobile; epistome broad, not projecting anteriorly, posterior margin with gently sinuous lateral margins, with 2 low, subtruncate median lobes; third maxilliped short, subquadrate, operculiform, covering buccal cavity when closed, merus short, subtrapezoidal, anteroexternal angle distinct, produced; ambulatory legs slender, very long, meri and dactyli unarmed; male thoracic sternum with sternites 3 – 8 fused, without trace of sutures or depressions, forming wide plate; male sternopleonal cavity short, with distal tip reaching to imaginary line joining anterior edge of P 4 coxae; male pleon-pleotelson short, triangular, somites 1 and 2 free, somites 3 – 5 fused but sutures still visible, pleotelson semicircular, no intercalated plates visible; G 1 short, stout, distal half tapering to slender sharp tip, directed towards median part of sternum, inner margin appears bilobed, with deep median cleft.	en	Ng, Peter K. L., Rahayu, Dwi Listyo (2022): A new genus and new species of Hymenosomatidae (Crustacea: Brachyura) from Indonesian Papua. Zootaxa 5092 (4): 480-486, DOI: https://doi.org/10.11646/zootaxa.5092.4.7
DD17C27CFFF7FFBCFF00D7594FE0FF14.taxon	etymology	Etymology. From the Latin for paper (charta) in arbitrary combination with the genus name Carcinus; alluding to the very flat carapace of the type species. Gender masculine.	en	Ng, Peter K. L., Rahayu, Dwi Listyo (2022): A new genus and new species of Hymenosomatidae (Crustacea: Brachyura) from Indonesian Papua. Zootaxa 5092 (4): 480-486, DOI: https://doi.org/10.11646/zootaxa.5092.4.7
DD17C27CFFF7FFBCFF00D7594FE0FF14.taxon	discussion	Remarks. Although the general features of Chartocinus n. gen. do not stand out, the one character it has that is unusual in Hymenosomatidae is that its male sternopleonal cavity is short and prominently posterior in position, suggesting that thoracic sternites 3 and / or 4 may have been displaced posteriorly as well. This character easily distinguishes Chartocinus n. gen. from other genera where the male sternopleonal cavity is more elongate and more anteriorly positioned to different degrees. The male thoracic sternum in the new genus is completely smooth and other than sternites 1 and 2, there are no sutures or grooves that demarcate the boundaries of sternites 3 – 8 on the surface, even on the lateral margins. This is a relatively extreme condition but has precedence in the Hymenosomatidae for both sexes (cf. Guinot 1979: fig. 30 A, pl. 20; Ng & Chuang 1996: fig. 6 J; Guinot 2011 a: fig. 1 C; Guinot et al. 2013: 189). Chartocinus n. gen. otherwise most closely resembles members of the Pacific genus Amarinus Lucas, 1980, in possessing an ovate carapace without defined regions and grooves on the dorsal surface, a simple unilobate rostrum, without distinct lateral armature, relatively short and subquadrate third maxilliped with the anteroexternal part of the merus produced, simple chelipeds which are not armed or inflated, a relatively short male pleon, and a short and stout G 1 (cf. Ng & Chuang 1996; Ng & Richer de Forges 1996). Chartocinus n. gen., however, differs from Amarinus in that the ridge which demarcates the rim of the carapace only reaches the orbit, not being visible behind the rostrum (Figs. 1 A, 2 A) (versus rim is complete, clearly visible behind rostrum), ambulatory legs (P 2 – P 5) are proportionately much longer (Fig. 1 A, B) (versus short), the flexor margin of all ambulatory dactyli is unarmed (Fig. 2 G – J) (versus all ambulatory dactyli with a subterminal spine), the male pleotelson is semicircular without any intercalated plate at the articulation with somite 5 (Fig. 2 E) (versus triangular to rounded with distinct intercalated plates at the articulation with the pleotelson; cf. Guinot & Richer de Forges 1997: 460, fig. 4 B – D; Guinot 2011 a: 23, 26 – 28, fig. 3 A – C), and the G 1 has a prominent cleft on the inner margin giving it a bilobed appearance (Fig. 2 K, L) (versus with margins entire) (cf. Holthuis 1968; Lucas 1980; Lucas & Davie 1982; Ng & Chuang 1996; Ng & Richer de Forges 1996; Rahayu & Ng 2004; Naruse et al. 2008 b). The incomplete rim of the carapace is noteworthy as in known Amarinus and many other hymenosomatids, the rim is distinct even behind the rostrum. In the case of Chartocinus n. gen., the rim gets lower and weaker as it passes the orbit and is not obviously visible behind the rostrum (Fig. 1 A, 2 A). In addition, all Amarinus species are known from freshwater and low salinity habitats, including swamps at an altitude of 1600 m in New Guinea (Holthuis 1968); whereas Chartocinus n. gen. was collected from a subtidal marine site. One key feature of Chartocinus n. gen. is the prominent pattern of hooked setae that border the carapace, rostrum, margins of the pereopods and pleonal somite 1 which obscure the outline of the animal. This is a character similar to Odiomaris pilosus (A. Milne - Edwards, 1873) from New Caledonia, a species previously placed in Amarinus. The setae, however, are a bit different, as those in Chartocinus n. gen. are shorter, hooked and more densely arranged. Chartocinus n. gen. can easily be separated from Odiomaris Ng & Richer de Forges, 1996, by the same characters as for Amarinus, except that in Odiomaris, the anteroexternal angle of the merus of the third maxilliped is not produced and the G 1 is more elongate with the distal part with two tightly apposed processes (cf. Ng & Richer de Forges 1996; Davie & Richer de Forges 1996; Guinot & de Mazancourt 2020). The male pleotelson has two depressions on the ventral surface (and are homologous to the pleonal sockets matching the intercalated plates in Amarinus and Odiomaris, representing the complementary pleonal parts for the locking of the pleon. Chartocinus represents the completely fused state of these small plates with the pleotelson (see Guinot 1979, 2011 a).	en	Ng, Peter K. L., Rahayu, Dwi Listyo (2022): A new genus and new species of Hymenosomatidae (Crustacea: Brachyura) from Indonesian Papua. Zootaxa 5092 (4): 480-486, DOI: https://doi.org/10.11646/zootaxa.5092.4.7
DD17C27CFFF5FFBBFF00D12A4A9FFE64.taxon	description	(Figs. 1, 2)	en	Ng, Peter K. L., Rahayu, Dwi Listyo (2022): A new genus and new species of Hymenosomatidae (Crustacea: Brachyura) from Indonesian Papua. Zootaxa 5092 (4): 480-486, DOI: https://doi.org/10.11646/zootaxa.5092.4.7
DD17C27CFFF5FFBBFF00D12A4A9FFE64.taxon	materials_examined	Material examined. Holotype: male (2.9 x 3.1 mm) (MZB Cru 5342), station EM 330, sandy mud substrate, Kamora estuary, 4 ° 48 ’ 48 ’’ S 136 ° 38 ’ 56 ’’ S, Papua, Indonesia, coll. trawl from survey, 7 November 2003.	en	Ng, Peter K. L., Rahayu, Dwi Listyo (2022): A new genus and new species of Hymenosomatidae (Crustacea: Brachyura) from Indonesian Papua. Zootaxa 5092 (4): 480-486, DOI: https://doi.org/10.11646/zootaxa.5092.4.7
DD17C27CFFF5FFBBFF00D12A4A9FFE64.taxon	description	Description of male holotype. Carapace longitudinally subovate, slightly longer than wide, width to length ratio 1.07; carapace distinctly dorsoventraly flattened; dorsal surface almost flat, surrounded by almost distinct continuous rim except behind rostrum; regions not discernible, without any visible grooves, H-shaped gastric groove shallow, joining cervical groove, no visible grooves on posterior part (Figs. 1 A, 2 A). Carapace rim and margin of rostrum lined with dense short, hooked setae that partially obscure surface (Fig. 1 A). Rostrum unilobate, subtriangular, short, horizontal, no lateral lobes present; joining entire concave supraorbital margin (Figs. 1 A, 2 A). Lateral margin of carapace (including branchiostegal surface) smooth, unarmed; anterolateral margin unarmed; posterolateral margin with 3 low, wide lobes above P 3 and P 4 coxae (Figs. 1 A, 2 A). Posterior carapace margin gently convex (Figs. 1 A, 2 A). Orbits clearly visible in dorsal view; eyes well developed, mobile, with prominent peduncle, visible dorsally, cornea rounded, fully pigmented (Figs. 1 A, 2 A). Antennular fossa shallow, surface gently concave, separated by narrow triangular septum without spine or tooth; basal antennular article quadrate, not swollen, free, positioned at base of fossa, second and third articles elongated, unable to fold into fossa (Fig. 2 B). Antenna with short flagellum, positioned between base of ocular peduncle and outer margin of antennular fossa; first article (osmoregulatory opening) present as round pore on each side of epistome; basal article fused with epistome (Fig. 2 B). Epistome distinct, broad, not projecting anteriorly, almost vertical from lateral view; posterior margin with gently sinuous lateral margins, with 2 low, subtruncate median lobes; not projecting anteriorly, not visible from dorsal view, area between lobes low, weakly triangular, sloping inwards to endostome (Fig. 2 B). Suborbital tooth low, indistinct, not visible from dorsal view (Fig. 2 B). Third maxilliped short, subquadrate; operculiform, with most of median part of buccal cavity covered when closed; merus short, subtrapezoidal, longer than ischium, anteroexternal angle distinct, produced; ischium with mesial margin gently convex, without visible sulcus; palp as long as merus, carpus and dactylus subequal in length, carpus rounded; exopod relatively stout, reaching to distal margin of merus, with distinct flagellum (Fig. 2 C). Chelipeds symmetrical, slender, relatively short; margins lined with dense short hooked setae which partially obscure surfaces; basis and ischium separate, short, unarmed; merus long, subcylindrical, margins unarmed; carpus gently curved, unarmed; chela with smooth rounded palm, outer surface gently convex, covered with long dense plumose setae which partially obscures surface and margins; fingers as long as palm, setose on outer surfaces; cutting edge of dactylus with prominent submedian tooth, cutting edge of propodus 2 low teeth and numerous denticles, tips gently curved (Figs. 1, 2 F). Ambulatory legs slender, long, P 2 longest; margins lined with dense short, hooked setae along margins that do not obscure surface; all meri with dorso-distal angle unarmed; carpus gently curved, unarmed; propodus subrectangular, elongated, unarmed; dactylus gently falciform with gently curving sharp tip, flexor margin unarmed; lengths of dactyli P 4> P 5> P 3> P 2 (Figs. 1, 2 G – J). Thoracic sternum wide, almost flat (Figs. 1 B, 2 D); sternites 1 and 2 fused, no suture visible but demarcated by a fold, with sternite 1 gently bent inwards towards mandibles; sternite 2 separated from sternite 3 by distinct groove; sternites 3 – 8 fused, without trace of sutures or depressions, sternite 4 strongly displaced posteriorly; sternopleonal cavity short, deep, reaching to imaginary line joining anterior edge of P 4 coxae (Fig. 2 D); margins of sternopleonal cavity with rim, low knob on subdistal edge forming sternal part of pleonal locking mechanism; pterygostome gently produced as low convexity, joining anterolateral extension of sternite 4, separating Milne Edwards openings from base of chelipeds; Milne Edwards openings relatively broad, ovate, margins lined with dense setae. Pleon-pleotelson short, triangular; somite 1 longitudinally narrow, laterally broad, subrectangular, reaching to base of P 5, somite 1 visible from dorsal view, lateral margins gently convex; somite 2 as wide as somite 1, longitudinally wide, mobile; somites 3 – 6 fused but still demarcated by shallow but visible sutures, especially laterally, forming trapezoidal plate; pleotelson free, almost semicircular elongated, lateral margins gently distinctly convex, with depression on each side of ventral surface forming socket of pleonal locking mechanism (Fig. 2 E). G 1 short, stout, distal half tapering to slender sharp tip, directed towards median part of sternum; outer margin distinctly convex; inner margin appears bilobed, with deep median cleft, each lobe with strongly convex margin; groove for G 2 submarginal; subdistal surface with row of long setae (Fig. 2 K, L). G 2 short, half length of G 1; tip subspatuliform tip; base elongate (Fig. 2 M). Females. Not known.	en	Ng, Peter K. L., Rahayu, Dwi Listyo (2022): A new genus and new species of Hymenosomatidae (Crustacea: Brachyura) from Indonesian Papua. Zootaxa 5092 (4): 480-486, DOI: https://doi.org/10.11646/zootaxa.5092.4.7
DD17C27CFFF5FFBBFF00D12A4A9FFE64.taxon	etymology	Etymology. From the Latin noun (rarus) for rare and delicate.	en	Ng, Peter K. L., Rahayu, Dwi Listyo (2022): A new genus and new species of Hymenosomatidae (Crustacea: Brachyura) from Indonesian Papua. Zootaxa 5092 (4): 480-486, DOI: https://doi.org/10.11646/zootaxa.5092.4.7
DD17C27CFFF5FFBBFF00D12A4A9FFE64.taxon	discussion	Remarks. The only known specimen was trawled from soft substrates on estuary of Kamora, Timika, in Indonesian Papua, in shallow waters. No other data is available. Five other species of hymenosomatids are known around Timika — Amarinus pristes Rahayu & Ng, 2004, Neorhynchoplax aspinifera (Lucas, 1980), N. elongata Rahayu & Ng, 2004, Lucascinus bedfordi (Montgomery, 1931) and Elamenopsis gracilipes Rahayu & Ng, 2019 (Rahayu & Ng, 2004, 2019). All of them, including A. pristes, occur in intertidal mangrove habitats.	en	Ng, Peter K. L., Rahayu, Dwi Listyo (2022): A new genus and new species of Hymenosomatidae (Crustacea: Brachyura) from Indonesian Papua. Zootaxa 5092 (4): 480-486, DOI: https://doi.org/10.11646/zootaxa.5092.4.7
