taxonID	type	description	language	source
190C87C4D079FFD8FF24C3F04FA2FB05.taxon	discussion	Comment. We concur with Archibald et al. (2021) and Archibald & Cannings (2021), against Nel & Zheng (2021) and Nel & Jouault (2022), that the five “ dysagrionid ” taxa from Burmese amber (i. e., Burmadysagrion Zheng et al., 2016, Electrodysagrion Zheng et al., 2017, Palaeodysagrion, and possibly Pseudopalaeodysagrion gen. nov.) do not belong in Dysagrionidae, but to a separate clade of zygopterid damselflies. The main reason is that the alternative explanation of the very different head structures with a diagenetic artefact seems highly implausible, given the number of different taxa from different localities that show no diagenetic distortions in any other parts of the body or the wings. Based on the results of the cladistic analysis of Archibald et al. (2021), we here elevate the subfamily Burmadysagrioninae erected by Zheng et al. (2016 b) to the status of a family separate from Dysagrionidae, to include all all four genera. While Dysagrionidae belong to the suborder Cephalozygoptera, Burmadysagrionidae are closer to genuine Zygoptera. Revised diagnosis. We consider the diagnosis of Burmadysagrioninae by Zheng et al. (2016 b) as invalid because it includes numerous characters that are only known from Burmadysagrion zhangi Zheng et al., 2016 but not the other four species. Therefore, we tentatively suggest the following new diagnosis: dysagrionid-like wing venation, but zygopteroid transverse head (vs globular head in Dysagrionidae); ax 2 aligned with arculus in forewing but somewhat distal of arculus in hind wing (except in Pseudopalaeodysagrion youlini, and only very slightly so in Electrodysagrion); pterostigma with basal side of normal obliquity (vs very oblique in Dysagrionidae); pterostigma short and only covering a single cell (vs numerous cells in Dysagrionidae) (except in Palaeodysagrion cretacicus which has 3 cells); CuA straight and with a single row of cells in cubito-anal space (vs CuA curved and with several rows of cells in cubito-anal space in Dysagrionidae).	en	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
190C87C4D079FFD8FF24C6B449E4F951.taxon	type_taxon	Type species. Palaeodysagrion cretacicus Zheng et al., 2016	en	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
190C87C4D079FFD8FF24C6B449E4F951.taxon	diagnosis	Revised diagnosis. Wing length about 25 mm; dense wing venation with numerous cells; 15 – 17 postnodal crossveins that are well-aligned with the postsubnodals; pterostigma elongate and covering 3.5 cells; both side of pterostigma parallel and of normal obliquity; pterostigmal brace present; IR 1 very long with 2 rows of cells between it and RP 1 and RP 2 respectively; nodus (n) in a basal position at about 26 – 27 % of wing length; nodus and subnodus oblique; base of IR 2 aligned with subnodus (sn); no lestine oblique vein ‘ O’ between RP 2 and IR 2; midfork one cell (elongate bridge space) basal of nodus, closer to nodus than to arculus; 6 – 7 crossveins between RP 1 + 2 and IR 2; cells in postdiscoidal space numerous, short and high; cells in space between MP and CuA numerous, short, high (transverse), and only weakly pentagonal; CuA very long; cubito-anal space with a single row of cells (slightly widened in hind wing); cubital crossing (CuP) oblique and situated close to arculus; ax 2 aligned with arculus in forewing but slightly distal of arculus in hind wing; discoidal cell elongate (even more so in hind wing) with distal side somewhat longer than basal side (esp. in fore wing); no accessory antenodal crossveins.	en	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
190C87C4D079FFDFFF24C5004FCAFD21.taxon	description	(Figures 1 – 9)	en	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
190C87C4D079FFDFFF24C5004FCAFD21.taxon	materials_examined	Holotype. NIGP 163546 (basal third of a hind wing). Since the holotype shows sufficient characters to distinguish it from P. youlini we see no necessity to apply to ICZN for designation of the new specimen as a neotype. New specimen. Relatively complete specimen without number in the private collection of Hans Georg Müller (Gelsenkirchen, Germany). Since it is not a type specimen, we see no problem with this deposition in a private collection.	en	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
190C87C4D079FFDFFF24C5004FCAFD21.taxon	distribution	Locality and horizon. Burmese amber (Hukawng Valley, Kachin State, Myanmar). Mid-Cretaceous: earliest Cenomanian.	en	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
190C87C4D079FFDFFF24C5004FCAFD21.taxon	diagnosis	Diagnosis. Same as for genus, since monotypic after the removal of P. youlini to a new genus (see below).	en	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
190C87C4D079FFDFFF24C5004FCAFD21.taxon	description	Description of new material. The new specimen is preserved in a 31 gram amber piece of 67 * 50 * 15 mm size with multiple syninclusions of other arthropods. The right pair of wings is completely preserved (forewing above, hind wing below in the photos), while the left pair of wings is broken at the base and missing. The wing tips are darker coloured from apex to the level of basal end of pterostigma (Figure 9), which clearly represents an original colour pattern. The pterostigmata have no discernible microsculpture. The wing veins (including crossveins) are equipped with medium length spines. Body (Figure 2). Head transverse as in Zygoptera (left eye fragmentary); prothorax small and saddle-shaped; pterothorax skewed as in Zygoptera, interpleural suture seems complete; base of abdomen (segments I – III) without visible secondary genital apparatus, but rest of abdomen missing; all six legs complete and very long, long femoral and tibial spines, femora distinctly longer than tibiae (pro-, meso- and metafemora 2.9, 3.2 and 3.9 mm, pro-, meso- and metatibiae 2.9, 2.9 and 4.6 mm), forelegs with cleaning comb along tip of tibia, tarsi 3 - segmented, the tarsal claws are not totally and clearly visible but may lack a hook. Forewing (Figures 4 – 6). Intact, hyaline, 25.0 mm long and 5.1 mm wide at widest point; only 2 primary antenodal crossveins ax 1 and ax 2 1.4 mm apart; ax 2 aligned with arculus; distance between wing base and arculus or nodus 3.0 or 6.6 mm respectively; nodus at 26.4 % of wing length, of normal structure with oblique nodal and less oblique subnodal veinlets aligned; 17 postnodal and 16 postsubnodal crossveins, aligned except for the distal 3 – 4; pterostigma 3.5 cells and 1.8 mm long, 0.6 mm wide; pterostigmal brace oblique; 8 crossveins distal of pterostigma; basal cell free; arculus angled; discoidal cell 1.4 mm long and 0.3 – 0.6 mm wide, subrectangular, and free; subdiscoidal cell free; RP midfork (branching RP 1 + 2 and RP 3 + 4) 6.0 mm distal of wing base and 1.3 basal of subnodus; bridge space free; IR 2 aligned with subnodus; branching of RP 1 and RP 2 6.3 mm distal of subnodus; IR 1 long and zigzagged, originating 3 cells distal of RP fork; distally 2 rows of cells between RP 1 and IR 1 and between IR 1 and RP 2; distally 2 – 3 rows of cells between RP 2 and IR 2, and 2 – 4 rows between IR 2 and RP 3 + 4; no lestine oblique vein ‘ O’ between RP 2 and IR 2; MA reaching to distal end of MA distally zigzagged and parallel to RP 3 + 4 with a single row of cells between them; about 9 cells between MA and MP along hind margin; MP without curvature at origin at discoidal cell; CuA zigzagged and parallel to MP, with a single row of cells in the field between CuA and hind margin; anal crossing (CuP) 0.4 mm basal of arculus; petiole 2.2 mm long; no accessory intercalary veins. Hind wing (Figures 7 – 8). Intact, hyaline, 24.6 mm long and 4.7 mm wide at widest point; only 2 primary antenodal crossveins ax 1 and ax 2 1.5 mm apart; ax 2 slightly distal of arculus; distance between wing base and arculus or nodus 3.0 or 6.7 mm respectively; distance between ax 2 and nodus 5.1 mm (in holotype 5.8 mm); nodus at 27.2 % of wing length, of normal structure with oblique nodal and less oblique subnodal veinlets aligned; 15 postnodal and 14 postsubnodal crossveins, more or less aligned with each other; pterostigma 3 cells and 2.3 mm long, 0.6 mm wide; pterostigmal brace oblique; 11 crossveins distal of pterostigma; basal cell free; arculus angled and sectors of arculus (RP and MA) widely separated at origin; discoidal cell 1.6 mm long and 0.26 - 0.48 mm wide, subrectangular, and free, much more elongate than in forewing; subdiscoidal cell free; RP midfork (branching RP 1 + 2 and RP 3 + 4) 5.9 mm distal of wing base and 1.3 basal of subnodus; bridge space free; IR 2 aligned with subnodus; branching of RP 1 and RP 2 5.2 mm distal of subnodus; IR 1 long and zigzagged, originating 3 cells distal of RP fork; distally 2 rows of cells between RP 1 and IR 1 and between IR 1 and RP 2; distally 2 – 3 rows of cells between RP 2 and IR 2, and 2 - 4 rows between IR 2 and RP 3 + 4; no lestine oblique vein ‘ O’ between RP 2 and IR 2; MA distally zigzagged and parallel to RP 3 + 4 with a single row of cells between them; 12 cells between MA and MP along hind margin; MP without curvature at origin at discoidal cell; CUA zigzagged and parallel to MP, with a single row of cells in the field between CuA and hind margin; anal crossing (CuP) 0.35 mm basal of arculus; petiole 2.3 mm long; no accessory intercalary veins.	en	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
190C87C4D07CFFDDFF24C19448DBFA18.taxon	type_taxon	Type species. P. youlini Zheng et al., 2017 Revised diagnosis. Same as type species since monotypic. Differing from Palaeodysagrion in the following characters: less dense wing venation; only 5 postnodal crossveins (vs 17); pterostigma very short and covering only a single cell (vs elongate and covering 3 cells); IR 1 very short with a single row of cells between it and RP 1 and RP 2 respectively (vs very long IR 1 with 2 rows of cells between it and RP 1 and RP 2 respectively); subnodus transverse (vs oblique); short bridge space (vs elongate bridge space); only 1 crossvein between RP 1 + 2 and IR 2 (vs 6); cells in postdiscoidal space more narrow, more elongate, and fewer in number; cells in space between MP and CuA more elongate and strongly pentagonal (vs higher and more rectangular); CuA much shorter; cubital crossing (CuP) situated more basal from arculus. Discoidal cell elongate and rectangular (much more elongate than in Electrodysagrion and Burmadysagrion, similar to Pseudopalaeodysagrion).	en	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
190C87C4D07CFFDDFF24C19448DBFA18.taxon	etymology	Etymology. The genus name refers to the assumed similarity of the wing base with the genus Palaeodysagrion.	en	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
