identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
190C87C4D079FFD8FF24C3F04FA2FB05.text	190C87C4D079FFD8FF24C3F04FA2FB05.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Burmadysagrionidae Zheng 2016	<div><p>Family Burmadysagrionidae Zheng et al., 2016 stat. nov.</p> <p>Comment. We concur with Archibald et al. (2021) and Archibald &amp; Cannings (2021), against Nel &amp; Zheng (2021) and Nel &amp; Jouault (2022), that the five “dysagrionid” taxa from Burmese amber (i.e., Burmadysagrion Zheng et al., 2016, Electrodysagrion Zheng et al., 2017, Palaeodysagrion, and possibly Pseudopalaeodysagrion gen. nov.) do not belong in Dysagrionidae, but to a separate clade of zygopterid damselflies. The main reason is that the alternative explanation of the very different head structures with a diagenetic artefact seems highly implausible, given the number of different taxa from different localities that show no diagenetic distortions in any other parts of the body or the wings. Based on the results of the cladistic analysis of Archibald et al. (2021), we here elevate the subfamily Burmadysagrioninae erected by Zheng et al. (2016b) to the status of a family separate from Dysagrionidae, to include all all four genera. While Dysagrionidae belong to the suborder Cephalozygoptera, Burmadysagrionidae are closer to genuine Zygoptera.</p> <p>Revised diagnosis. We consider the diagnosis of Burmadysagrioninae by Zheng et al. (2016b) as invalid because it includes numerous characters that are only known from Burmadysagrion zhangi Zheng et al., 2016 but not the other four species. Therefore, we tentatively suggest the following new diagnosis: dysagrionid-like wing venation, but zygopteroid transverse head (vs globular head in Dysagrionidae); ax2 aligned with arculus in forewing but somewhat distal of arculus in hind wing (except in Pseudopalaeodysagrion youlini, and only very slightly so in Electrodysagrion); pterostigma with basal side of normal obliquity (vs very oblique in Dysagrionidae); pterostigma short and only covering a single cell (vs numerous cells in Dysagrionidae) (except in Palaeodysagrion cretacicus which has 3 cells); CuA straight and with a single row of cells in cubito-anal space (vs CuA curved and with several rows of cells in cubito-anal space in Dysagrionidae).</p> </div>	https://treatment.plazi.org/id/190C87C4D079FFD8FF24C3F04FA2FB05	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bechly, Günter;Velten, Jürgen	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
190C87C4D079FFD8FF24C6B449E4F951.text	190C87C4D079FFD8FF24C6B449E4F951.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Palaeodysagrion Zheng 2016	<div><p>Palaeodysagrion Zheng et al., 2016</p> <p>Type species. Palaeodysagrion cretacicus Zheng et al., 2016</p> <p>Revised diagnosis. Wing length about 25 mm; dense wing venation with numerous cells; 15–17 postnodal crossveins that are well-aligned with the postsubnodals; pterostigma elongate and covering 3.5 cells; both side of pterostigma parallel and of normal obliquity; pterostigmal brace present; IR1 very long with 2 rows of cells between it and RP1 and RP2 respectively; nodus (n) in a basal position at about 26–27 % of wing length; nodus and subnodus oblique; base of IR2 aligned with subnodus (sn); no lestine oblique vein ‘O’ between RP2 and IR2; midfork one cell (elongate bridge space) basal of nodus, closer to nodus than to arculus; 6–7 crossveins between RP1+2 and IR2; cells in postdiscoidal space numerous, short and high; cells in space between MP and CuA numerous, short, high (transverse), and only weakly pentagonal; CuA very long; cubito-anal space with a single row of cells (slightly widened in hind wing); cubital crossing (CuP) oblique and situated close to arculus; ax2 aligned with arculus in forewing but slightly distal of arculus in hind wing; discoidal cell elongate (even more so in hind wing) with distal side somewhat longer than basal side (esp. in fore wing); no accessory antenodal crossveins.</p></div> 	https://treatment.plazi.org/id/190C87C4D079FFD8FF24C6B449E4F951	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bechly, Günter;Velten, Jürgen	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
190C87C4D079FFDFFF24C5004FCAFD21.text	190C87C4D079FFDFFF24C5004FCAFD21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Palaeodysagrion cretacicus Zheng 2016	<div><p>Palaeodysagrion cretacicus Zheng et al., 2016</p> <p>(Figures 1–9)</p> <p>Holotype. NIGP163546 (basal third of a hind wing). Since the holotype shows sufficient characters to distinguish it from P. youlini we see no necessity to apply to ICZN for designation of the new specimen as a neotype.</p> <p>New specimen. Relatively complete specimen without number in the private collection of Hans Georg Müller (Gelsenkirchen, Germany). Since it is not a type specimen, we see no problem with this deposition in a private collection.</p> <p>Locality and horizon. Burmese amber (Hukawng Valley, Kachin State, Myanmar). Mid-Cretaceous: earliest Cenomanian.</p> <p>Diagnosis. Same as for genus, since monotypic after the removal of P. youlini to a new genus (see below).</p> <p>Description of new material. The new specimen is preserved in a 31 gram amber piece of 67*50* 15 mm size with multiple syninclusions of other arthropods. The right pair of wings is completely preserved (forewing above, hind wing below in the photos), while the left pair of wings is broken at the base and missing. The wing tips are darker coloured from apex to the level of basal end of pterostigma (Figure 9), which clearly represents an original colour pattern. The pterostigmata have no discernible microsculpture. The wing veins (including crossveins) are equipped with medium length spines.</p> <p>Body (Figure 2). Head transverse as in Zygoptera (left eye fragmentary); prothorax small and saddle-shaped; pterothorax skewed as in Zygoptera, interpleural suture seems complete; base of abdomen (segments I–III) without visible secondary genital apparatus, but rest of abdomen missing; all six legs complete and very long, long femoral and tibial spines, femora distinctly longer than tibiae (pro-, meso- and metafemora 2.9, 3.2 and 3.9 mm, pro-, meso- and metatibiae 2.9, 2.9 and 4.6 mm), forelegs with cleaning comb along tip of tibia, tarsi 3-segmented, the tarsal claws are not totally and clearly visible but may lack a hook.</p> <p>Forewing (Figures 4–6). Intact, hyaline, 25.0 mm long and 5.1 mm wide at widest point; only 2 primary antenodal crossveins ax1 and ax2 1.4 mm apart; ax2 aligned with arculus; distance between wing base and arculus or nodus 3.0 or 6.6 mm respectively; nodus at 26.4 % of wing length, of normal structure with oblique nodal and less oblique subnodal veinlets aligned; 17 postnodal and 16 postsubnodal crossveins, aligned except for the distal 3–4; pterostigma 3.5 cells and 1.8 mm long, 0.6 mm wide; pterostigmal brace oblique; 8 crossveins distal of pterostigma; basal cell free; arculus angled; discoidal cell 1.4 mm long and 0.3–0.6 mm wide, subrectangular, and free; subdiscoidal cell free; RP midfork (branching RP1+2 and RP3+4) 6.0 mm distal of wing base and 1.3 basal of subnodus; bridge space free; IR2 aligned with subnodus; branching of RP1 and RP2 6.3 mm distal of subnodus; IR1 long and zigzagged, originating 3 cells distal of RP fork; distally 2 rows of cells between RP1 and IR1 and between IR1 and RP2; distally 2–3 rows of cells between RP2 and IR2, and 2–4 rows between IR2 and RP3+4; no lestine oblique vein ‘O’ between RP2 and IR2; MA reaching to distal end of MA distally zigzagged and parallel to RP3+4 with a single row of cells between them; about 9 cells between MA and MP along hind margin; MP without curvature at origin at discoidal cell; CuA zigzagged and parallel to MP, with a single row of cells in the field between CuA and hind margin; anal crossing (CuP) 0.4 mm basal of arculus; petiole 2.2 mm long; no accessory intercalary veins.</p> <p>Hind wing (Figures 7–8). Intact, hyaline, 24.6 mm long and 4.7 mm wide at widest point; only 2 primary antenodal crossveins ax1 and ax2 1.5 mm apart; ax2 slightly distal of arculus; distance between wing base and arculus or nodus 3.0 or 6.7 mm respectively; distance between ax2 and nodus 5.1 mm (in holotype 5.8 mm); nodus at 27.2 % of wing length, of normal structure with oblique nodal and less oblique subnodal veinlets aligned; 15 postnodal and 14 postsubnodal crossveins, more or less aligned with each other; pterostigma 3 cells and 2.3 mm long, 0.6 mm wide; pterostigmal brace oblique; 11 crossveins distal of pterostigma; basal cell free; arculus angled and sectors of arculus (RP and MA) widely separated at origin; discoidal cell 1.6 mm long and 0.26-0.48 mm wide, subrectangular, and free, much more elongate than in forewing; subdiscoidal cell free; RP midfork (branching RP1 +2 and RP3 +4) 5.9 mm distal of wing base and 1.3 basal of subnodus; bridge space free; IR2 aligned with subnodus; branching of RP1 and RP2 5.2 mm distal of subnodus; IR1 long and zigzagged, originating 3 cells distal of RP fork; distally 2 rows of cells between RP1 and IR1 and between IR1 and RP2; distally 2–3 rows of cells between RP2 and IR2, and 2-4 rows between IR2 and RP3 +4; no lestine oblique vein ‘O’ between RP2 and IR2; MA distally zigzagged and parallel to RP3 +4 with a single row of cells between them; 12 cells between MA and MP along hind margin; MP without curvature at origin at discoidal cell; CUA zigzagged and parallel to MP, with a single row of cells in the field between CuA and hind margin; anal crossing (CuP) 0.35 mm basal of arculus; petiole 2.3 mm long; no accessory intercalary veins.</p> </div>	https://treatment.plazi.org/id/190C87C4D079FFDFFF24C5004FCAFD21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bechly, Günter;Velten, Jürgen	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
190C87C4D07CFFDDFF24C19448DBFA18.text	190C87C4D07CFFDDFF24C19448DBFA18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudopalaeodysagrion Bechly & Velten 2023	<div><p>Pseudopalaeodysagrion n. gen.</p> <p>Type species. P. youlini Zheng et al., 2017</p> <p>Revised diagnosis. Same as type species since monotypic. Differing from Palaeodysagrion in the following characters: less dense wing venation; only 5 postnodal crossveins (vs 17); pterostigma very short and covering only a single cell (vs elongate and covering 3 cells); IR1 very short with a single row of cells between it and RP1 and RP2 respectively (vs very long IR1 with 2 rows of cells between it and RP1 and RP2 respectively); subnodus transverse (vs oblique); short bridge space (vs elongate bridge space); only 1 crossvein between RP1+2 and IR2 (vs 6); cells in postdiscoidal space more narrow, more elongate, and fewer in number; cells in space between MP and CuA more elongate and strongly pentagonal (vs higher and more rectangular); CuA much shorter; cubital crossing (CuP) situated more basal from arculus. Discoidal cell elongate and rectangular (much more elongate than in Electrodysagrion and Burmadysagrion, similar to Pseudopalaeodysagrion).</p> <p>Etymology. The genus name refers to the assumed similarity of the wing base with the genus Palaeodysagrion.</p> </div>	https://treatment.plazi.org/id/190C87C4D07CFFDDFF24C19448DBFA18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bechly, Günter;Velten, Jürgen	Bechly, Günter, Velten, Jürgen (2023): A Revised Diagnosis of Palaeodysagrion cretacicus Zheng et al., 2016 (Insecta: Odonata) from mid-Cretaceous Burmese Amber, with erection of a new genus of fossil damselflies. Zootaxa 5263 (4): 547-556, DOI: 10.11646/zootaxa.5263.4.6, URL: http://dx.doi.org/10.11646/zootaxa.5263.4.6
