taxonID	type	description	language	source
03AB87F4FFB58B2FFF21FD09FBB333F0.taxon	type_taxon	[type species: Andrikothelyna papuana Pace, 2000].	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB58B2FFF21FD09FBB333F0.taxon	diagnosis	Diagnosis. Andrikothelyna may be distinguished from other genera based on the following characteristics: strongly convex body; posterior angle of head round and neck almost absent; two-segmented labial palpi; short and unilobed ligula; two medial setae distant from each other; strongly transverse mentum; tarsal formula 4 - 4 - 5; extremely long segment I of meso- and metatarsi (fig. 15 in Pace 2013; elytra with a pair of a longitudinal row of weak or distinct granules along the suture.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB58B2FFF21FD09FBB333F0.taxon	description	Redescription. Body fusiform (Figs 1 – 8); dorsal surface microreticulation indistinct but that on abdominal tergite II distinct; pronotum, elytra, and abdominal tergites covered with rough reticulation sometimes forming small granules around pores. Head transverse; almost all microsetae directed anterolaterally; infraorbital carina well developed; posterior angle round; neck almost absent. Antennae moderately long; segments IV – XI with dense microsetae. Mouthparts. Labrum (Fig. 9). Anterior margin weakly concave, surface with about 17 setae; epipharynx with 2 pairs of small sensillae on anterior margin, and 3 pairs of micro setae on lateral margin; basal region with transverse row of sensory pores. Mandible (Figs 11 – 12) short, right with large subapical teeth; prostheca not fringed with tooth or setulae, basal area forming tongue like lobe; spinules of molar dorsal area not in distinct row; ventral molar area without denticles. Maxilla (Fig. 13), palpi without pseudosegment; apex of galea moderately brushy; lacinia widened apicad, spines on adoral margin distributed at only apical half. Labium (Fig. 14), palpi with two segments, apical segment longer than basal one; apex of ligula unilobed and rounded. Prementum with two medial setae with pores distant from each other, with many pseudopores, one real pore, and one setalpore on each side; lateral margin of hypopharynx with setae entirely; apodeme with two medial projections. Mentum (Fig. 15) strongly transverse, 3 times as wide as long, with about 10 setae. Thorax. Pronotum strongly transverse, postero-lateral angle gently sinuate, microsetae directed posteriorly. Elytra transverse; surface along suture and lateral margin with longitudinal row of strong to weak granules. Mesoventral processes (Fig. 16) without medial carina, apex round or truncated, extending middle of mesocoxal cavities; mesocoxal cavities mostly separated; apex of metaventral process round. Abdomen elongate, slightly narrowed posteriad; posterior margin of tergite VIII with teeth. Legs. Tarsal formula 4 - 4 - 5; segment I of meso- and metatarsi very long. Male. Median lobe of aedeagus narrow; apical lobe of paramerite short. Female. Spermatheca short and simple.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB58B2FFF21FD09FBB333F0.taxon	discussion	Remarks. Speiraphallusa completely matches Andrikothelyna in character states which are essential for genus classification such as mouth parts, tarsi, and granules on elytra. Thus, it is reasonable to consider Speiraphallusa as a junior synonym of Andrikothelyna. Despite examining only males, Pace (2013) described the rows of granules of elytra, along the suture and the lateral margin of elytra, as a characteristic of the males of Speiraphallusa. However, no sex-specific differences were present in the elytra in all the new species studied in this paper. Distinct sexual dimorphism was observed on the tergite VII and sternite VIII of Andrikothelyna limbata sp. nov. and Andrikothelyna naomichii sp. nov. In the original description of the genus Speiraphallusa, the head was described as “ neck moderate ” (Pace 2013), following which, Santiago-Jiménez & Santiago-Navarro (2016) recognized the state “ with neck ” as one of the characteristics of the genus in key to the genera of the tribe Placusini (p. 49). However, the neck was absent in all the new species examined in this paper. Perhaps the presence of the “ neck ” was incorrectly recognized by Pace (2013) when he viewed the specimen in the dorsal view. Pires-Silva (2023) mentioned the diagnostic characteristic of Speiraphallusa in the key to genera of Placusini (p. 469) as “ tergite VIII transverse, not modified ”, but all species of this genus have teeth on posterior margin of the tergite VIII. In Andrikothelyna orientis (Pace 2013) comb. nov. only a flagellum of the median lobe of the aedeagus appears to be coiled (as shown in fig. 12 in Pace 2013), but in A. limbata sp. nov., which has a similar shape of the median lobe to that of A. orientis comb. nov. not only flagellum but also bulb is spiraled (Figs 23 – 25). Therefore, it is likely that the figure shown in Pace (2013) is drawn based on a misunderstanding of the morphology and the bulb is coiled or spiraled in A. orientis comb. nov. The morphology of the flagellum and the bulb itself being spiral shaped in A. limbata sp. nov., possibly also in A. orientis comb. nov., is unique, and no similar morphology is known within the subfamily. The genus is recorded newly in Taiwan and Japan in this study.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB58B2FFF21FD09FBB333F0.taxon	distribution	Distribution. Japan, Taiwan, Malaysia, and Papua New Guinea.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB58B2FFF21FD09FBB333F0.taxon	discussion	Bionomics. The following three new species were collected from dead vines or dead branches on living trunks. Talipariti tiliaceum and Machilus thunbergii are recorded as the plant species below.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB68B2FFF21FA27FE1A3733.taxon	materials_examined	Type material. Not examined.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB68B2FFF21FA27FE1A3733.taxon	diagnosis	Diagnosis. Andrikothelyna orientis comb. nov. can be distinguished from other species of the genus based on the following characteristics: abdominal tergite VIII having four teeth at posterior margin, two medial teeth very short; median lobe of male aedeagus very narrow, flagellum spiraled.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB68B2FFF21FA27FE1A3733.taxon	distribution	Distribution. Malaysia (Hulu Perak).	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB68B2FFF21FA27FE1A3733.taxon	discussion	Remarks. Andrikothelyna orientis was described based on a single male holotype and the female sexual character states are unknown.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB68B2FFF21FC38FBC635C2.taxon	materials_examined	Type material. Not examined.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB68B2FFF21FC38FBC635C2.taxon	diagnosis	Diagnosis. A. papuana can be distinguished from other species of the genus based on the following characteristics: pronotum having posterior medial depression; posterior margin of abdominal tergite VIII having four teeth.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB68B2FFF21FC38FBC635C2.taxon	distribution	Distribution. Papua New Guinea.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB68B2FFF21FC38FBC635C2.taxon	discussion	Remarks. Andrikothelyna papuana was described based on a single female holotype. The original description mentioned that the female spermatheca of the holotype was not visible because it might not be sclerotized (Pace 2000). However, the spermatheca may have been failed to be observed, considering the following three new species have clearly sclerotized spermatheca. The male sexual characteristics are unknown.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB08B28FF21FF59FCFD3428.taxon	description	(Figs 1, 5, 9 – 19) (Japanese name: sabiiro-toguro-hanekakushi)	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB08B28FF21FF59FCFD3428.taxon	diagnosis	Diagnosis. Andrikothelyna rubiginosa sp. nov. is similar to A. orientis comb. nov. but can be distinguished owing to its reddish body color (Fig. 1) and the posterior margin of tergite VIII (Fig. 17) having four prominent teeth and two tiny teeth, in contrast to only four teeth found in A. orientis comb. nov. This new species also can be recognized from other species of the genus based on the following characteristics: suboctagonal pronotum with its posterior margin slightly angulated at two points; elytra sharply carinate near its lateral margin; the elongated apical part of female spermatheca (Fig. 19).	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB08B28FF21FF59FCFD3428.taxon	materials_examined	Material examined. Holotype. Female, “ tđfi ỆḒḏñü / Musha, Renai, / Nantou Hsien, TAIWAN / 7 - 11. III. 2018, / Naomichi Tsuji ” (KUM). Paratypes. TAIWAN: Nantou: 3 ♀, same data as holotype (KUM).	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB08B28FF21FF59FCFD3428.taxon	description	Description. Body (Figs 1, 5) robust; dorsal surface moderately glossy, and distinctly reticulated. Color. Body reddish brown, pronotum dark red, antennae dark red; with segments I and II reddish yellow and III red, maxillary palpi, forelegs reddish yellow, meso- and meta- legs dark red. Head. Eyes large, as long as 1 / 3 of head. Antennae longer than head and pronotum combined; segments II – X dilated; segments I – III elongate, segments VIII – X distinctly transverse; segment XI elongate, round at apex; approximate relative length of segments from basal to apex: 16: 17: 16: 10: 10: 10: 11: 10: 11: 12: 39. Thorax. Pronotum, suboctagonal; middle of lateral margin slightly angulated; postero-lateral angle obtuse, posterior margin straight on medial 1 / 3, slightly angulated on two points. Elytra, postero-lateral angle strongly sinuate; longitudinal row of granules along suture moderate; near lateral margin sharply carinate. Mesoventral processes, apex round (Fig. 16). Abdomen. Tergite VIII (Fig. 17) covered with rough small granules; posterior margin with 4 major teeth; tiny teeth present between medial and lateral teeth; apex of each tiny tooth with microseta; lateral teeth short, less curved medially, medial teeth slightly shorter than lateral teeth. Legs. Slender in general; tibiae covered with fine setae. Approximate relative length of mesotarsal segments from basal to apex: 31: 10: 10: 27, and that of metatarsal segments: 54: 15: 15: 10: 32. Male. Unknown. Female. Tergite VII with one posteromedial granule and several small granules on medial area; posterior margin of sternite VIII (Fig. 18) finely sinuate, slightly emarginate medially. Spermatheca (Fig. 19), apical part and eggshaped, slightly longer than basal part; basal part decorated with rough structure. Measurements (n = 4). BL ≈ 2.97 – 3.28; AL, 0.95 – 1.03; PL, 0.4 – 0.47; PW, 0.8 – 0.99; EL, 0.49 – 0.55; EW, 0.9 – 1.08; HTL, 0.55 – 0.63; PW / PL, 1.91 – 2.12; EW / EL, 1.83 – 2.12; AL / PL, 2.1 – 2.43.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB08B28FF21FF59FCFD3428.taxon	etymology	Etymology. The specific name is derived from the Latin adjective “ rűbîginôsus ” which means rust-colored and refers to the body color of this new species.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB08B28FF21FF59FCFD3428.taxon	distribution	Distribution. Taiwan (Nantou). Bionomics. According to the collector, Naomichi Tsuji, individuals of the type series were collected by beating a dead vine approximately 1.5 m above the forest floor.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB18B2AFF21FB01FEB83620.taxon	description	(Figs 2 – 3, 6 – 7, 20 – 28) (Japanese name: tsumaki-toguro-hanekakushi)	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB18B2AFF21FB01FEB83620.taxon	diagnosis	Diagnosis. Andrikothelyna limbata sp. nov. is similar to A. orientis but can be distinguished based on yellow semicircular maculation on the posterior margin of elytra (Figs 2 – 3). This new species also may be distinguished from other species of the genus based on the following characteristics: posterior margin of abdominal tergite VIII having four teeth (Fig. 20); median lobe of male aedeagus, bulb, and flagellum spiraled three-dimensionally (Figs 23 – 25), flagellum having cluster of spines at apex; extremely narrow paramere (Fig. 26); basal part of female spermatheca (Figs 27 – 28) bent at the middle.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB18B2AFF21FB01FEB83620.taxon	materials_examined	Material examined. Holotype. Male, “ JAPAN Kyushu / Kaijin Shrine / TSUSHIMA Is. / 3. VI. 2016 / Tateo ITO leg. ” (KUM). Paratypes. JAPAN: Honshu: Gifu-ken: 1 ♂, 1 ♀, Sengen-yama, Kukuri, Kani-shi [33.38 ° N, 137.12 ° E] (alt. 200 – 370 m), 7. VIII. 2018, Masato Ando leg. (KUM). Kyoto-fu: 2 ♂, 2 ♀, Kawakita Nature & Athletic Park, Hayashinomoto, 12, Yawata-shi, 5. V. 2015, Tateo Ito leg. (KUM); 2 ♂, 1 ♀, same locality, 11. VII. 2015, Tateo Ito leg. (KUM); 3 ♂, 4 ♀, Kawakita Nature & Athletic Park, Motohiraki, Oyamazaki-cho, Otokuni-gun, 19. VIII. 2015, Tateo Ito leg. (cI); 1 ♂, 1 ♀, same locality, 22. VIII. 2015, Tateo Ito leg. (cI); 1 ♀, same locality, 26. VIII. 2015, Tateo Ito leg. (KUM); 1 ♂, same locality, 2. XI. 2015, Tateo Ito leg. (KUM); 1 ♀, same locality, 28. IX. 2015, Tateo Ito leg. (KUM); 2 ♂, same locality, 27. X. 2015, Tateo Ito leg. (KUM); 1 ♂, same locality, 17. III. 2016, Tateo Ito leg. (KUM); 1 ♂, same locality, 14. V. 2020, Tateo Ito leg. (cI); 6 ♂, 2 ♀, same locality, 1. VI. 2020, Tateo Ito leg. (cI); 1 unsexed, 1 ♂, 2 ♀, same locality, 17. VI. 2020, Tateo Ito leg. (KUM); 1 ♂, 1 ♀, same locality, 16. VII. 2020, Tateo Ito leg. (KUM); 1 ♀, Kawara, Maidzuru-shi, (from fallen tree), 27. VI. 2007, Yuzo Kuroda leg. (cI). Okayama-ken: 2 ♀, Handayama, Hokaiin, Kita-ku, Okayama-shi, 2. VI. 2004, Seidai Nagashima leg. (KUM); 1 ♀, Kasaiyama, Hataayu, Kita-ku, Okayama-shi, 15. I. 1974, Osamu Yamaji leg. (KUM). Shikoku: Kochi-ken: 1 ♀, Tôwa Uragoshi, Kubogawa, Shiman-chô, 8. V. 2010, Takasuke Miyata & Toshie Miyata leg. (car-net) (KUM). Kyushu: Nagasakiken: 3 ♂, Kaijin Shinto Shrine, Minemachikisaka, Tsushima-shi, 3. VI. 2016, Tateo Ito leg. (same data as holotype) (KUM). Kagoshima-ken: 1 ♂, 1 ♀, Ôura river estuary, Ôura-chô, Minamisatsuma-shi, 4. V. 1994, Kaoru Haga leg. (KUM). Ryukyu: Kagoshima-ken: Amami-Ôshima Is.: 1 ♀, Mt. Yuwan. 19. IV. 1971, M. Sakai leg. (EUM); 1 ♂, Hatsuno, 4. V. 1977, M. Sakai & A. Oda leg. (EUM). Okinawa-ken: Ishigaki Is.: 10 ♂, 4 ♀, Reverbed of Tôro Riv., Shiraho, Ishigaki-shi., 28 – 29. V. 2021, N. Tsuji & S. Imada leg. (KUM, one pair for cI); 2 ♂, Inoda Auto Camping Ground, Tôzato, Ishigaki-shi, 19. III. 2020, Takuto Hashizume leg. (KUM); 1 ♀, same locality, 22. III. 2020, Takuto Hashizume leg. (KUM); 1 unsexed, same locality, 23. III. 2020, Takuto Hashizume leg. (KUM); 1 ♀, same locality, 26. X. 2020, Tsubasa Nozaki leg. (KUM); 6 ♂, 2 ♀, Inoda, Ishigaki-shi [24.46 ° N, 124.25 ° E], 30. V. 2021, N. Tsuji & S. Imada leg. (KUM).	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB18B2AFF21FB01FEB83620.taxon	description	Description. Body (Figs 2 – 3, 6 – 7) robust; dorsal surface glossy, distinctly reticulated. Color. Body dark brown to black, pronotum brownish red to dark brown, posterior margin of each elytron with yellow semicircular maculation; antennae dark brown with segments I and II brownish yellow and the third yellowish brown, maxillary palpi dark brown to brownish yellow; forelegs reddish yellow to yellow, meso- and metalegs reddish brown to yellowish brown. Head. Eyes large, as long as 1 / 3 of head; infraorbital carina developed. Antennae longer than head and pronotum combined; segments II – X slightly dilated; segments V – X distinctly transverse; segment XI elongate, round at apex; approximate relative length of segments from basal to apex: 16: 18: 13: 10: 11: 10: 11: 10: 10: 11: 38. Thorax. Pronotum, postero-lateral angle obtuse. Elytra, postero-lateral angle gently sinuate; longitudinal row of granules along suture strong; near lateral margin gently carinate, granulated, and curved upward (Figs 6 – 7). Mesoventral process, apex varying truncated to round. Abdomen. Tergite VIII (Fig. 20) weakly granulated with two medial granules; posterior margin with four teeth; lateral teeth long, less curved, medial teeth slightly shorter than lateral teeth; medial grub distinctly shallower than lateral grub, depth 1 / 3 of lateral grub. Legs. Moderately slender in general; tibiae covered with fine setae. Approximate relative length of mesotarsal segments from basal to apex: 32: 10: 11: 34, and that of metatarsal segments: 42: 11: 10: 10: 28. Male. Tergite VII with two posteromedial granules; posterior margin of sternite VIII (Fig. 21) without medial notch and finely sinuate. Median lobe of aedeagus (Figs 23 – 25) length about 0.75 mm, very narrow; bulb itself spiraled three-dimensionally; flagellum spiraled, having flag-like appendage at base and cluster of spines at apex. Paramere (Fig. 26) very narrow; velum remarkably elongate; apical lobe of paramerite digitate. Female. Tergite VII without or with one posteromedial granule; posterior margin of sternite VIII (Fig. 22) with medial notch. Spermatheca (Figs 27 – 28), apical part short and subspherical; basal part smooth and bent at middle. Measurements. (population of Honshu and Kyushu, n = 5): BL ≈ 1.36 – 2.96; AL, 0.67 – 0.86; PL, 0.3 – 0.42; PW, 0.59 – 0.8; EL, 0.32 – 0.46; EW, 0.66 – 0.94; HTL, 0.36 – 0.52; PW / PL, 1.85 – 1.95; EW / EL, 2 – 2.08; AL / PL, 1.88 – 2.21. Measurement (population of Ishigaki Is., n = 5): BL ≈ 1.84 – 2.32; AL, 0.62 – 0.8; PL, 0.32 – 0.42; PW, 0.56 – 0.74; EL, 0.33 – 0.44; EW, 0.62 – 0.88; HTL, 0.34 – 0.48; PW / PL, 1.75 – 1.98; EW / EL, 1.88 – 2; AL / PL, 1.92 – 2.07. Variation. Sexual dimorphism was evident in the number of posteromedial granules on tergite VII and a presence of a notch on sternite VIII. Geographical variations were detected in body coloration, the degree of gloss on the dorsal surface, and the shape of the spermatheca in this species. The population from Honshu and Kyushu (Figs 2, 6, 27) revealed the following characteristics: body color dark brown; pronotum brownish red; maxillary palpi dark brown; forelegs reddish yellow; meso- and metalegs reddish brown; body surface less glossy; spermatheca weakly sclerotized, whereas the population from Ryukyu (Figs 3, 7, 28) exhibited the following characteristics: body black; pronotum dark brown; maxillary palpi brownish yellow; forelegs yellow; meso- and metalegs yellowish brown; body surface glossier compared to that of the other populations, antennae longer than that in the other. Regardless of the region, intra-species variations were found in the following characters: the apical shape of the mesoventral process and the presence of a posteromedial granule of female tergite VII.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB18B2AFF21FB01FEB83620.taxon	etymology	Etymology. The specific name of the species is derived from the Latin adjective “ limbâtus, ” meaning “ bordered, ” and refers to yellow maculation on the posterior edge of the elytra.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB18B2AFF21FB01FEB83620.taxon	distribution	Distribution. Japan: Honshu, Shikoku, Kyushu, Ryukyu (Amami Ôshima Is., Ishigaki Is.). Bionomics. On Ishigaki Island, this new species was spotted walking on the dead branches of the living trunks of beach hibiscus, Talipariti tiliaceum (L.) Fryxell (Figs 29 – 30) and observed walking on the branches at night and were collected.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB38B24FF21F970FE3E376A.taxon	description	(Figs 4, 8, 31 – 36, 38) (Japanese name: tsuji-toguro-hanekakushi)	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB38B24FF21F970FE3E376A.taxon	diagnosis	Diagnosis. Andrikothelyna naomichii sp. nov. can be easily distinguished from any other species of the genus owing to its orange-colored body. In addition, characteristics unique to this new species are as follows: elytra bearing weak granules along the suture (Figs 4, 8) and hardly carinate near the lateral margin; median lobe of male aedeagus (Fig. 34), bulb and flagellum not coiled; velum of paramere wing-shaped (Fig. 35); basal part of female spermatheca rough and not bent (Fig. 36).	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB38B24FF21F970FE3E376A.taxon	materials_examined	Material examined. Holotype. Male, “ ṖDZẹƃḇffiǎḛff / Mt. Maese-dake, Ishigaki, / [GPS] N: 24. 37, E: 124.15 / Ishigaki Is., Ryûkyû, JAPAN. / 30 - 31. V. 2021. N. Tsuji leg. ” (KUM). Paratypes. JAPAN: Ryukyu: Okinawa-ken: Ishigaki Is.: 2 ♂, 3 ♀, same data as holotype, (KUM); 6 ♂, 8 ♀, same locality, T. NOZAKI & K. Goino leg., 29. III. 2022, T. Nozaki & K. Goino leg. (KUM, one pair for cI);	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB38B24FF21F970FE3E376A.taxon	description	Description. Body (Figs 4, 8) small; dorsal surface less glossy, weakly reticulated. Color. Body orange, abdomen reddish brown; antennae orange with segments I to III pale yellow; maxillary palpi, forelegs yellow, meso- and metalegs reddish orange. Head. Eyes large, as long as 1 / 4 of head; infraorbital carina developed. Antennae (Fig. 4) longer than head and pronotum combined; segments II – X dilated; segments I – III elongate, segments IX – X distinctly transverse; segment XI elongate, round at apex; approximate relative length of segments from basal to apex: 18: 19: 16: 10: 10: 10: 11: 11: 11: 11: 29. Thorax. Pronotum, lateral margin round; postero-lateral angle obtuse. Elytra, postero-lateral angle strongly sinuate; longitudinal row of granules along suture weak (Figs 4, 8); near lateral margin hardly carinate. Mesoventral process, apex varying truncated to round. Abdomen. Posterior margin of tergite VIII (Fig. 31) with 4 – 6 short teeth, apex of each tooth with microseta; medial grub deepest; other grubs varying and frequently becoming asymmetric. Legs. Moderately slender in general; tibiae covered with fine setae. Approximate relative length of mesotarsal segments from basal to apex: 28: 12: 10: 34, and that of metatarsal segments: 37: 11: 10: 10: 36. Male. Tergite VII with two posteromedial granules; posterior margin of sternite VIII (Fig. 32) round and smooth. Median lobe of aedeagus (Fig. 34) length about 0.47 mm, elongate, bulb itself not coiled; flagellum not coiled, base oar shaped. Paramere (Fig. 35), velum wing-shaped; apical lobe of paramerite short and subtrapezoid. Female. Tergite VII without posteromedial granule; posterior margin of sternite VIII (Fig. 33) varying round to shallowly emarginated. Spermatheca (Fig. 36), apical part subspherical; basal part unsmooth. Measurements (n = 10). BL ≈ 1.56 – 2.16; AL, 0.59 – 0.65; PL, 0.32 – 0.36; PW, 0.62 – 0.69; EL, 0.34 – 0.39; EW, 0.69 – 0.76; HTL, 0.39 – 0.44; PW / PL, 1.87 – 2.02; EW / EL, 1.86 – 2.26; AL / PL, 1.73 – 1.95. Variation. Distinct sexual dimorphism was evident in the posteromedial granules of tergite VII. Intra-species variations exist in the following characters: teeth number and shape of the posterior margin of tergite VIII; the apical shape of the mesoventral process; and the posterior margin of female sternite VIII.	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB38B24FF21F970FE3E376A.taxon	etymology	Etymology. The specific name is dedicated to the first person to discover the new species, Mr. Naomichi Tsuji (KUM, Fukuoka).	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
03AB87F4FFB38B24FF21F970FE3E376A.taxon	distribution	Distribution. Japan: Ryukyu (Ishigaki Island). Bionomics. Individuals of the type series were collected from freshly dead or weakened branches on the living trunk of the Japanese bay tree, Machilus thunbergii Siebold et Zucc. (Fig. 37) and observed walking on these branches at night (Fig. 38).	en	Nozaki, Tsubasa (2023): Taxonomy of the genus Andrikothelyna Pace, 2000 (Coleoptera: Staphylinidae Aleocharinae), with description of three new species from Taiwan and Japan. Zootaxa 5264 (1): 64-76, DOI: 10.11646/zootaxa.5264.1.4, URL: http://dx.doi.org/10.11646/zootaxa.5264.1.4
