taxonID	type	description	language	source
03AD7622FFB2B64028C7979F58F9FB79.taxon	type_taxon	— Type: A. bracteatus Schltr. & K. Krause.	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB2B64028C7979F58F9FB79.taxon	distribution	Atractocarpus is currently a genus of c. 40 species that occur in Australia, the Federated States of Micronesia, Fiji, Indonesia, New Caledonia, Papua New Guinea, Philippines, Solomon Islands, Society Islands, Tonga and Vanuatu, with the highest diversity in New Caledonia. The New Caledonian species are mostly riparian, living in mesic to humid forests, or more rarely in bushes. The Australian and other Pacific islands representatives occupy various mesic forests, swamp forests and maritime semi-deciduous thickets.	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB2B64028C7979F58F9FB79.taxon	diagnosis	The diagnostic combination of characters for the genus, sensu Puttock (1999) is: flowers with subsessile anthers; styles completely included in the tube; patent corolla lobes; porate pollen grains in monads; ovary bilocular, placentation axile; berries moderate to large, spherical to fusiform, many-seeded; and seeds large with an extensive peripheral hilum. An interesting characteristic feature also noted by Puttock (1999) consists of the placental mass degenerating into a watery pulp, enclosing seeds. At the fruit stage, the locule limits and the placentation type become consequently not obvious in fruit-section (see Figs 2; 3).	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB2B64428AE93AA5F8FFBB8.taxon	description	(Figs 1; 2)	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB2B64428AE93AA5F8FFBB8.taxon	materials_examined	— Type: New Caledonia. Couvelée, fl., 1900 - 1906, Le Rat 537 (neo-, P, hic designatus). — Previous type: New Caledonia. Süd-Bezirk, Couvelée, 1900 - 1906, Le Rat 90 (holo-, B †).	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB2B64428AE93AA5F8FFBB8.taxon	materials_examined	— Type: New Caledonia, Baie de Prony, fr., 1862, Pancher s. n. (holo-, P). OTHER SPECIMENS EXAMINATED. — New Caledonia. Baie de Prony, fl., 1910, d’Alleizette (leg. Godefroy) 354 (P). — Forêts de la baie de Prony, fl. fr., IX. 1868, Balansa 353 (P). — Île Casy, dans la baie de Prony, fr., 24. VI. 1869, Balansa 2014 (P). — Collines éruptives situées vers le cours supérieur de la Tamoa, fl. fr., 11. IV. 1870, Balansa 2896 (P). — Vallée du Dothio, fl., XII. 1871, Balansa 3419 (P, P). — Environs de Mamarie, dans la partie supérieure du bassin du Tio, fl., I. 1872, Balansa 3650 (P). — Thio-Canala, Dothio, 21 ° 35 ’ S, 166 ° 6 ’ E, 170 m, fl. fr., 29. XII. 1977, Bamps 6103 (BR, NOU, P). — Col d’Amieu, réserve forestière, 21 ° 37 ’ S, 165 ° 49 ’ E, 390 m, fl., 30. XII. 1977, Bamps 6116 (BR, P). — Mt Mou, 300 m, 10. IX. 1950, Baumann-Bodenheim 5859 (P). — Île des Pins, E. Baie d’Oupi, 25. V. 1951, Baumann-Bodenheim 13412 (P). — Île des Pins, Kuto, 26. V. 1951, Baumann-Bodenheim 13459 (P). — Île des Pins, Kuto, fr., 27. V. 1951, Baumann-Bodenheim 13591 (P). — Île des Pins, île dans l’anse de Kanuméra, fr., 27. V. 1951, Baumann-Bodenheim 13616 (P). — Baie des Pirogues, plage, fr., 2. XII. 1951, Baumann-Bodenheim 15818 (P). — Creek Tao, fl., 26. XII. 1951, Baumann-Bodenheim 15958 (P). — Île des Pins (Kunié), fr., 17. VII. 1965, Bernardi 9698 (G, P). — S. loc., fr., I. 1980, Bourret 2210 (NOU). — Rivière des Pirogues, fr., 4. I. 1978, Cabalion NC 352 (P). — Bonne Anse, Baie du Sud, 1903, Cribb 1363 (NOU, P). — Plum, creek Lucky, fr., 21. II. 2001, Dagostini & Rigault 289 (NOU). — Forêt Nord, fr., 7. IX. 2006, Dagostini 1201 (NOU, P). — Wenig verzweigtes Stammschen im Unterwuchs des Waldes im Malgrunde des Kalouéholatales, fr., II. 1924, Däniker 3137 (P, Z). — Mont Dore, fl., 26. III. 1974, Debray 2202 (P). — Île des Pins, fl., 24. I. 1978, Debray 2707 (P). — Île des Pins, bord de chemin, fl., 26. I. 1978, Debray 2744 (P). — Port Boisé, bord de rivière, fl., 1861, Deplanche 245 (P). — S. loc., fl., 1861, Deplanche 245 bis (P). — Nouméa, littoral entre la Baie Nord et le Cap N’Dua, fl., II. 1909, Franc 759 (P). — Prony, lieux ombragés et frais, fl., I. 1914, Franc 1639 a (P). — Prony, littoral, fl., I. 1914, Franc 1766 (P). — Port Boisé, littoral, fl., I. 1914, Franc 1766 a (P). — Île des Pins, fl. fr., 1874 - 1876, Germain s. n. (P). — Base du Mt Mou, 15. X. 1950, Guillaumin & Baumann-Bodenheim 6889 (P). — Base du Mt Dore, 8. II. 1950, Guillaumin & Baumann-Bodenheim 7947 (P). — Base du Mt Dore, 8. II. 1951, Guillaumin & Baumann-Bodenheim 7956 (P). — Vallée des Pirogues, fl., 29. III. 1951, Guillaumin & Baumann-Bodenheim 11528 (P). — Port Boisé, fr., 10. V. 1979, Hoff 844 (NOU). — Port Boisé, fr., 10. VI. 1980, Hoff 2209 (NOU). — Port Boisé, fr., 10. VI. 1980, Hoff 2247 (NOU). — Baie des Pirogues, 120 m, 16. XII. 1950, Hürlimann 371 (P). — Tonaouron-Goro, 10 m, fl., 9. I. 1951, Hürlimann 567 (P). — “ Sill ” serpentineux de St André Yallein, fl., 24. VII. 1968, Jaffré 64 (NOU). — Route de Prony, fr., 31. X. 1972, Jaffré 998 (NOU, P). — Route de Prony, fl., 4. I. 1978, Jaffré 2164 (NOU, P). — Route de Prony, baie N’go, sur éboulis péridotitiques et colluvions ferralitiques, lisière de forêt dense, fr., 27. VIII. 2000, Jaffré 3445 (P). — Prony, 300 m, fl., 1900 - 1910, Le Rat 1731 (P). — Plum, just above beach, fr., 14. I. 1956, MacKee 3778 (P). — Île des Pins, Kuto, raised coral rock on sandy beach, 5 - 10 m, fr., 4. VIII. 1956, MacKee 5021 (P). — Mokoué, région de Canala, fl., 17. I. 1961, MacKee 8156 (P). — Crête à l’Ouest du Col de Petchicara, versant de Dothio, 500 m, fl., 7. V. 1965, MacKee 12531 (P). — Nakéty, 30 m, fl. fr., 8. V. 1965, MacKee 12548 (NOU, P). — Île des Pins, presqu’île d’Oro, forêt à l’intérieur de la presqu’île, 5 m, fr., 19. XII. 1965, MacKee 14096 (P). — Prony, Baie Nord, terrain serpentineux, 0 - 30 m, fr., 10. IV. 1966, MacKee (leg. Corbasson) 14708 (P). — Baie des Pirogues, maquis près de la plage, fl., 15. IV. 1967, MacKee 16574 (P). — Plum, Baie Ngo, maquis sur terrain rocheux serpentineux, 50 m, 28. I. 1970, MacKee (leg. Corbasson) 21487 (P). — Base S. E. du Mt Dore, fl., V. 1970, MacKee 21874 (NOU). — Kuebini, Île Nou, fl., 29. X. 1970, MacKee 22785 (NOU). — Île des Pins, plateau central, bandes de forêt en terrain rocheux serpentineux, 100 m, fl. fr., 1. IV. 1971, MacKee 23571 (P). — Nakety, Mokoué, forêt-galerie, 20 m, fl. fr., 13. XI. 1971, MacKee 24686 (P). — Port Boisé, littoral Est, forêt sur le terrain rocheux serpentineux, 0 - 10 m, fl., 13. VII. 1973, MacKee 26940 (P). — Port Boisé, littoral rocheux serpentineux, fr., 27. X. 1973, MacKee (leg. Aymard) 27613 (P). — Prony, Île Casy, terrain serpentineux, 0 - 40 m, fl. fr., 15. IV. 1974, MacKee 28504 (P). — Île des Pins, Baie des Crabes, fl. fr., 8. IX. 1977, MacKee 33821 (NOU, P). — Cap Reine Charlotte, forêt basse côtière, terrain rocheux serpentineux, 2 m, fl., 29. XI. 1978, MacKee 36143 (P). — Dothio, Haute Ouen Koura, fl., II. 1979, MacKee 36583 (NOU, P). — Doti, terrain serpentineux, 50 m, fr., 17. VIII. 1984, MacKee (leg. Nasi) 42122 (P). — Yaté, 15 m, fl. fr., 22. IV. 1993, MacKee 46196 (NOU, P). — Port Boisy, c. 20 air-km S of Yaté, coastal forest of Port Boisy, fr., 12. VI. 1980, McPherson 2795 (NOU, P). — Dothio River valley, near Thio, along road and trails by river, leaving highway just short of one-way portion, forest, c. 50 m, fr., 27. I. 1983, McPherson 5431 (MO, P). — Province Sud, Port Boisé, 20 - 50 m, fl. fr. 21. XI. 2003, Mouly 149 (P). — Province Sud, Île des Pins, sentier forestier entre la baie d’Oro et la baie d’Upi, 0 - 10 m, fl. 4. I. 2004, Mouly & Innocente 269 (P). — Col de Pétchécara, 100 m à gauche en suivant le début de la piste, forêt en face de la piste, fr., 14. IV. 1999, Munzinger & Cabalion 479 (P). — Forêt Anaïs, Prony, 22 ° 18 ’ 06 ’’ S, 166 ° 42 ’ 56 ’’ E, fr., 16. VI. 2004, Munzinger 2130 (NOU, P, UPS). — Île des Pins, îlot Moro à l’Ouest de l’Île des Pins, 22 ° 39 ’ 6 ’’ S, 167 ° 23 ’ 39 ’’ E, fr., 27. VII. 2004, Munzinger (leg. Cunkelman) 2278 (P). — Tontouta, 21 ° 58 ’ 20 ’’ S., 166 ° 14 ’ 56 ’’ E., fr., 26. II. 2006, Munzinger 3348 (NOU, P). — Col de Mouirange, versant NO, fl. (fr. doubtful), 4. II. 1966, Nothis 8 (NOU, P). — Lisières des massifs dans le voisinage de la mer, 1862, Pancher s. n. (P). — Baie de Prony, fr., Pancher s. n. (P). — Passe entre Thio-Nakéty, c. 400 m, fl., 2. I. 1912, Sarasin 371 (P). — Creek Pernod, rive droite, 250 m, fl., 15. VI. 1965, Schmid 330 (NOU, P). — Kouakoué, bord de ruisseau, fr., 18. XI. 1965, Schmid 772 (NOU). — Île des Pins, vers la Baie de la Corbeille, fr., 10. VI. 1967, Schmid 2138 (NOU). — Île des Pins, forêt sur épandage au pied du Pic N’Ga, fl., 9. VI. 1967, Schmid 2138 bis (NOU). — Île des Pins, Est, fr., 3. V. 1974, Schmid 4969 (NOU). — Ilôt Casy, Baie de Prony, fr., 15. IV. 1974, Sévenet 637 (NOU). — Vers la Baie d’Oupi, Île des Pins, fr., 2. V. 1974, Sévenet 652 (NOU). — N. E. Baie Ngo, N. Pt. 16, fl., 7. III. 1964, Stauffer 5768 (NOU, P). — Petite forêt sèche, Mont-Dore, fr., 11. V. 1978, Suprin 403 (NOU). — Secteur de Ngo, Rivière des Pirogues, forêt de thalweg, fr., 17. VII. 1980, Suprin 582 (P). — Baie de Ngo, forêt sur éboulis et sol brun, c. 200 m, fr., 18. VIII. 1978, Tirel 1202 (P). — Bord du Dothio, route des Colons, fl. fr., 4. I. 1965, Veillon 5 (NOU, P). — Au-dessus de Baie N’Go, c. 200 m, fl., 4. II. 1982, Veillon 4814 (NOU, P). — Base du Pic du Pin, près de la mine Anna Madeleine, exploitation forestière Lafleur, clairière de la zone marginale forestière, terrain serpentineux, fl., 1. IV. 1942, Virot 604 (P). — Littoral près de Goro, au sud-est du village, c. 5 m, fl., 28. III. 1942, Virot 644 (P). — Presqu’île de Kuto, Île des Pins, bois littoraux humides, plateaux madréporiques émergés, c. 5 m, fr., 5. II. 1943, Virot 985 (P).	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB2B64428AE93AA5F8FFBB8.taxon	description	DESCRIPTION Dioecious, slender tree 1 - 3 m high, monocaul or few stemmed. Stipules paired, imbricate, lanceolate, 12 - 21 × 3 - 5 mm. Leaves of non-flowering shoots elliptic to oblanceolate, glabrous; petioles 16 - 22 mm long; lamina 160 - 320 × 50 - 160 mm, apex acuminate, base attenuated, glossy dark green, dull grey-green below, coriaceous; secondary veins 12 - 14 pairs at 50 - 60 ° to the midvein. Inflorescences axillary, branch-like to 250 mm long, perennial in male individuals, flowers arranged in cymes, up to 5 - florous in male, and solitary to 3 - florous in female individuals, flowers subtended by modified leaves, cordate, 30 - 40 mm long; flowers ± sessile, 5 - merous, with a slightly sexual differentiation. Hypanthium 9 - 12 mm long and linear in male, ovules aborted; 12 - 16 × 3 - 5 × 3 - 5 mm, centrally inflated in female individuals. Calyx tube obconical, 3 - 4 mm long; lobes triangular, 1.5 - 2 mm long. Corolla tube 18 - 28 mm long, urceolate, c. 5 mm diameter at the widest point, glabrescent outside, glabrous inside except where it is bearded below the anthers. Corolla lobes lanceolate, c. 18 × 7 - 10 mm, glabrous. Anthers bilocular, 7 - 10 mm long, dorsifixed, supramedifixed, attached 4 - 5 mm from their apices, subsessile, inserted 5 - 6 mm below the sinuses of the corolla lobes. Style included, c. 25 mm long; stigmatic lobes 2, c. 9 mm long. Ovary bilocular, elongate, c. 9 × 2 × 2 mm, placentation axile, each bearing many ovules in a longitudinal row. Fruit berry, solitary, fusiform, 120 - 310 × 30 - 35 mm, smooth narrowing towards the calyx tube; exocarp smooth, green when mature; placentas extending in a whitish pulp. Seeds c. 5 × 5 × 0.6 mm; hilum occupying c. 0.6 of the perimeter of seed; exotesta cells with a thickened ring around the inner tangential wall and occasional interstitial thickenings. NOTE ON TYPIFICATION Herbaria where A. Le Rat collections are deposited (according to HUH Index of Botanists: http: // kiki. huh. harvard. edu / databases / botanist _ index. html) were consulted, as well as institutions where F. R. R. Schlechter usually sent herbarium material. The holotype Le Rat 90 has been destroyed in B (B herbarium communication), and any duplicate has been found nor in P where most of Le Rat collection is, neither in other herbaria (A, BM, BR, E, G, H, K, L, S, U, W, WRSL). The gathering Le Rat 537, collected in the type locality by the same botanist and deposited at P, is designated as a neotype.	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB2B64428AE93AA5F8FFBB8.taxon	distribution	DISTRIBUTION, ECOLOGY, AND HABITAT Atractocarpus bracteatus is endemic to southeastern New Caledonia and the Isle of Pines (Fig. 4). The species is a non obligatory riparian, and it grows mainly on ultramafic soils, calcareous rocks and occasionally on eroded elevated corals. The ant indicated in Figure 1, as observed in the field, is not an evidence of ant-pollination, but an indication that A. bracteatus nectar secretions attract ants, forming temporary colonies on plant individuals. Fruiting takes long time in the species, as observed on cultivated specimens in New Caledonia, necessitating more than two years to reach maturity.	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB2B64428AE93AA5F8FFBB8.taxon	discussion	NOTE Two unpublished names, Genipa semperflorens and Genipa xylopioides, are present in annotations by Baillon on herbarium sheets in P. Genipa was at that time one of the few considered genera in Gardenieae, together with Gardenia and Randia. Both Randia and Genipa are now restricted to the Neotropics (Persson 2002; Gustafsson & Persson 2002), and New Caledonian Gardenieae species were never considered as potential Genipa species since then.	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB6B649290A93E95972F975.taxon	description	(Fig. 3)	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB6B649290A93E95972F975.taxon	materials_examined	— Type: New Caledonia. Île Art, 1859, Montrouzier 179 (lecto-, MPU, hicdesignatus). — Previous syntypes: New Caledonia. Île Art, 1859, Montrouzier 94 (LYJB †). — Île Art, 1859, Montrouzier 95 (LYJB †).	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB6B649290A93E95972F975.taxon	materials_examined	— Type: New Caledonia. Île Yandé, fr., 23. VI. 1871, Balansa 3212 (lecto-, P, hic designatus; isolecto-, P). — Previous syntype: New Caledonia. “ Rade de Canala, sur les rochers maritimes ” (doubtful locality, see note below), fr., 29. VI. 1869, Balansa 2012 (P). O THER SPECIMENS EXAMINATED. — New Caledonia. Mt Yandé, fr., 23. VIII. 1978, Cabalion 644 (P). — Insel Yande, fl., 7. IV. 1925, Däniker 1574 (P). — Vallée de Voh sup. 250 m, 12. IV. 1951, Guillaumin & Baumann-Bodenheim 12122 (P). — Boulinda 200 m, fr., 1. VII. 1972, Jaffré 844 (NOU, P). — Base du Massif du Boulinda, fr., 8. VII. 1975, Jaffré 1346 (NOU); idem, Jaffré 1352 (NOU). — Île Pott, Panan, 10 - 60 m, fr., 29 Jul 1968, MacKee 19355 (NOU, P). — Île Yandé, fl. fr., 6. X. 1970, MacKee 22621 (NOU). — Népoui, presqu’île de Muéo, fl., 10. XII. 1970, MacKee 22997 (NOU, P). — Baaba, secteur S. O., fr., 6. I. 1971, MacKee 23194 (NOU). — Plateau de Tiéa, pente Nord, maquis sur terrain rocheux serpentineux, 200 - 300 m, fr., 2. XI. 1971, MacKee 24536 (P). — Amos, fl., 4. XII. 1972, MacKee 26041 (NOU). — Gomen, Ouéholle, maquis sur pente serpentineuse, 300 m, fl., 28. XII. 1972, MacKee 26124 (P). — Col d’Arama, versant Ouest, maquis ouvert sur terrain serpentineux, 80 m, fr., 23. III. 1973, MacKee 26464 (P). — Kopéto, crête est du Mt Vert, maquis haut sur pente raide rocheuse serpentineuse, 700 m, fr., 24. VI. 1973, MacKee 26888 (P). — Vallée de Poya, Ndokoa, maquis sur terrain rocheux serpentineux, 100 m, fr., 31. X. 1973, MacKee 27703 (P). — Poya, Avangui, 300 m, fl., 2. I. 1976, MacKee 30611 (NOU, P). — Poindimié, Povila, forêt dense basse, terrain serpentineux rocheux, 200 m, fr., 13. IX. 1976, MacKee 31522 (P). — Poya, Nékoro, 200 m, fr., 21. V. 1977, MacKee 33208 (NOU, P). — Paagoumène, maquis haut sur terrain serpentineux, 10 m, fl., 6. II. 1980, MacKee 37809 (P). — Poya, Ndokoa, fr., 11. VII. 1981, MacKee 39214 (NOU, P). — Dent de Poya, 500 m, fr., 17. IX. 1981, MacKee 39569 (NOU, P). — Poya, restes de forêt sur alluvions, 40 m, fl., 1. XII. 1990, MacKee (leg. Cherrier) 45197 (P). — Poya, restes de forêt sur alluvions, 40 m, fr., 1. XII. 1990, MacKee (leg. Cherrier) 45202 (P). — Île Yandé, fr., 23. VIII. 1978, Morat 6131 (NOU). — Province Nord, Poya, Nétéa, le long du creek, 50 - 150 m, fr., 8. VI. 2007, Mouly & Barrabé 850 (BR, K, NOU, P). — Île Yandé, face Est, 200 m, fr., 5. X. 2005, Munzinger & Swenson 3037 (NOU, P). — Poya, Nétéa, 21 ° 15 ’ 6 ’’ S., 165 ° 11 ’ 27 ’’ E., fr., 3. III. 2007, Munzinger 4190 (NOU). — Gorges de N’Dokoa, Poya, fr., 20. X. 1974, Sévenet 731 (P). — Île Yandé, N. O., fr., 23. VIII. 1978, Veillon 3657 (NOU, P). — Montagnes de Balade, fl., 1855 - 1860, Vieillard 649 (P). — Wagap, 1861 - 1867, Vieillard 2752 (P).	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB6B649290A93E95972F975.taxon	description	DESCRIPTION Dioecious, small slender tree 1 - 2.5 m high, monocaul, or few stemmed. Stipules paired, imbricate, lanceolate, 12 - 15 × 3 - 4 mm. Leaves opposite, of nonflowering shoots oblong-elliptic, glabrous; petioles 26 - 42 mm long; lamina 140 - 220 × 50 - 100 mm, apex acuminate, base rounded, glossy dark green, dull grey-green below, coriaceous; secondary veins 5 - 10 pairs at 55 - 65 ° to the midvein. Inflorescences axillary, branch-like to 140 mm long, consisting of 2 internodes, flowers arranged in cymes, 3 - florous in male, and solitary in female individuals; flowers ± sessile, 5 - merous, with a slight sexual differentiation, flowers substended by modified leaves, cordate, 20 - 45 mm long. Hypanthium 9 - 10 mm long and linear in male, 11 - 14 × 3 - 5 × 3 - 5 mm, centrally inflated in female individuals. Calyx tube obconical, 2 - 3 mm long; lobes triangular, 1.5 - 2 mm long. Corolla tube c. 20 mm long, urceolate, c. 4 mm diameter at the widest point, glabrescent outside, glabrous inside, except where it is bearded below the anthers. Corolla lobes lanceolate, c. 15 × 7 - 10 mm, glabrous. Anthers bilocular, 7 - 10 mm long, dorsifixed, supramedifixed, attached 4 - 5 mm from their apices, subsessile, inserted 5 - 6 mm below the sinuses of the corolla lobes. Style included, c. 19 mm long; stigmatic lobes 2, c. 7 mm long. Ovary bilocular, elongate, c. 8 × 2 × 2 mm, placentation axile, each bearing many ovules. Fruit solitary, oblong-ovoid, 30 - 60 × 25 - 50 mm, smooth narrowing towards the calyx tube; exocarp smooth, green when mature; placentas extending in a whitish pulp. Seeds c. 4 × 4 × 0.5 mm; hilum occupying c. 0.5 mm of the perimeter of seed; exotesta cells with a thickened ring around the inner tangential wall and occasional interstitial thickenings.	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB6B649290A93E95972F975.taxon	discussion	NOTE ON TYPIFICATIONS Since Montrouzier never cited the original material in his Flora, possible syntypes are accepted from citations in Guillaumin & Beauvisage (1913). As two of the three syntypes of A. heterophyllus (Montrouzier 94, 95) have been destroyed in LYJB, only one Montrouzier’s gathering (Montrouzier 179) remains available at MPU. Though without any annotation from Montrouzier as an author, the MPU gathering cannot be rejected as original material, being in agreement with the protologue and collected before the publication of the protologue. The collection Montrouzier 179 is thus designated here as lectotype for A. heterophyllus. The lectotype of Atractocarpus balansaeanus was chosen from two original gatherings cited by Guillaumin (1930). The collection Balansa 3212, designated as lectotype, is well preserved and representative of the protologue. The other syntype collection Balansa 2012 consists of only one sheet in worse condition than the lectotype.	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
03AD7622FFB6B649290A93E95972F975.taxon	distribution	DISTRIBUTION, ECOLOGY, AND HABITAT Atractocarpus heterophyllus is endemic to northern New Caledonia and Belep Islands (Fig. 4). This species is exclusively riparian, restricted to limestone and serpentine soils, on rocks and rock slides. On the main island, most of specimens collected belong to the western coast populations. The gathering Balansa 2012 bears the label information “ Canala ” (southeastern coast), a locality poorly reliable for the species. Besides the gathering Balansa 2012, the two old collections made by Vieillard from the east are unclearly localized. Indeed, Vieillard indicated on labels the locality where he was based instead of the locality of the collection. The eastern coast is now well explored, and only one recent collection can be attributed to A. heterophyllus, in the Poindimié area. The few eastern populations are probably smaller and more threatened than the western ones.	en	Mouly, Arnaud (2012): On the distinction of Atractocarpus heterophyllus (Montrouz.) Guillaumin & Beauvis. and A. bracteatus Schltr. & K. Krause (Rubiaceae): in search of the correct name for the type species of Atractocarpus Schltr. & K. Krause. Adansonia (3) 34 (1): 103-114, DOI: 10.5252/a2012n1a12
