taxonID	type	description	language	source
03E487E3CA6BC51BFC9D54C6CCE7C1EC.taxon	materials_examined	Material examined: Turkey: 1 ♂, Nevşehir, Göre, 21. IV. 1992, leg. Kocian (cKoc). Iran: 1 ♀, Ardabil province, 20 km NE Khalkhal, 37 ° 41 ' N, 48 ° 23 ' E, 1480 m, 13. X. 2011, leg. Frisch (MNHUB); 1 ♀, Gilan province, Bandar-e Anzali, Bashm, 37 ° 30 ' N, 49 ° 21 ' E, 0 m, marshland, 1. V. 2012, leg. Weipert (NME). Kyrgyzstan: 1 ♀, Issyk-Kul, Tyup, 42 ° 45 ' N, 78 ° 23 ' E, 1620 m, 28. VI. 2011, leg. Frisch (MNHUB); 1 ♀, IssykKul, Balykchy - Cholpon-Ata, Semenovka, 42 ° 46 ' N, 77 ° 32 ' E, 1860 m, 30. VI. 2011, leg. Frisch (cAss). Japan: 1 ex., Hokkaido, Sapporo env., Nopporo virgin forest, 15. V. 2010, leg. Lackner (cAss); 2 exs., Hokkaido, Sapporo, Hokudai Campus, VII. 2011, leg. Lackner (cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA6BC51BFC9D54C6CCE7C1EC.taxon	discussion	Comment: Drusilla puncticollis, whose descriptions is based on two syntypes from “ Kamtschatka ” (MOTSCHULSKY 1845), was previously treated as a subspecies of D. canaliculata (SMETANA 2004). According to MOTSCHULSKY (1845), who already expressed doubts regarding the status of this taxon (“ Je propose cette espèce avec doute ”), D. puncticollis is distinguished from D. canaliculata by larger body size, darker coloration, and a more deeply impressed pronotum. Examined material from the eastern East Palaearctic region indeed tends to be darker and of slightly larger size than material from West Europe; for previous records see ASSING (2005 a, b, 2006 b, 2008 a, 2009, 2010). However, similar character conditions have been observed also in specimens from some regions in Middle Asia and the West Palaearctic region (particularly Iran), suggesting that these differences should be interpreted as intraspecific, possibly clinal variation. Moreover, the presence of a distinct subspecies in Kamchatka appears most unlikely. Hence the synonymy proposed above. The previously known trans-Palaearctic distribution of D. canaliculata extended from West Europe to the Russian Far East (SMETANA 2004). The above specimens from Hokkaido represent the first records from Japan.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFCC75386CA48C62C.taxon	materials_examined	Material examined: Lebanon: 10 ♂♂, 14 ♀♀, 7 exs., Rayfoun, 33 ° 58 ' N, 35 ° 42 ' E, 990 m, mixed oak forest, 15. III. 2013, leg. Reuter (cAss, cFel); 2 exs., same data, but pitfall trap, 18. XI. 2012 (cFel); 11 exs., same data, but pitfall trap, 30. III – 5. IV. 2013 (cFel); 1 ex., 27 km NE Beirut, Kafr Debian env., 1200 m, oak forest, pitfall trap, 9. XI. 2012, leg. Reuter (cFel).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFCC75386CA48C62C.taxon	discussion	Comment: This species has been recorded only from Lebanon (ASSING 2005 b).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFCC755C6CA72C58C.taxon	materials_examined	Material examined: Turkey: 12 exs., Hatay, 15 km WSW Antakya, Batiayaz, Musa Dağ, ca. 500 m, pitfall trap, 6. – 23. IV. 2014, leg. Reuter (cFel).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFCC755C6CA72C58C.taxon	discussion	Comment: The known distribution of D. denigrata is confined to the Turkish province Hatay and northern Syria (ASSING 2005 b, 2010).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFF4156A6CFC6CB6C.taxon	materials_examined	Material examined: Tunisia: 23 exs., Béja, 11 km W Teboursouk pass, 36 ° 30 ' N, 9 ° 10 ' E, 700 m, 1. III. 2012, leg. Meybohm (cAss); 3 exs., Béja, W Djebba, Teboursuk, 36 ° 28 ' N, 9 ° 05 ' E, 640 m, wet fields, under stones, 4. III. 2012, leg. Assmann (cFel).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFF4156A6CFC6CB6C.taxon	discussion	Comment: This anophthalmous representative of the subgenus Apteranopsis JEANNEL, 1960 is locally endemic in Tunisia (ASSING 2005 b).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFCC752E6CAD5C0EC.taxon	materials_examined	Material examined: Italy: 1 ex., Salerno, Montecorvino Rovella, M. Corvino, garden, 27. VI. 2009, leg. Frenzel (cApf).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFCC752E6CAD5C0EC.taxon	discussion	Comment: The distribution of D. italica is confined to Italy, where it is widespread and rather common. For a distribution map see ASSING (2006 b).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFF415486CCBCC5CC.taxon	materials_examined	Material examined: Tunisia: 1 ex., Jendouba, 8 km S Aïn Draham, 36 ° 43 ' N, 8 ° 42 ' E, 610 m, oak forest, 2. III. 2012, leg. Meybohm (cAss); 1 ex., same data, but leg. Hetzel (cFel); 14 exs., Jendouba, National Park El Feija, 36 ° 28 ' N, 8 ° 14 ' E, 590 m, arable land, 3. III. 2012, leg. Meybohm (cAss); 9 exs., same data, but leg. Hetzel (cFel); 2 exs., same data, but 36 ° 29 ' N, 8 ° 20 ' E, 520 m, clearing in oak forest, under stones, leg. Hetzel (cFel).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFF415486CCBCC5CC.taxon	discussion	Comment: The distribution of D. kabyliana is confined to Algeria and Tunisia (ASSING 2005 c, 2008 a).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFF415286CCB4C7EC.taxon	materials_examined	Material examined: Tunisia: 3 exs., Béja, Djebba, 36 ° 28 ' N, 9 ° 05 ' E, 640 m, 3. III. 2012, leg. Th. Schmidt (cAss); 15 exs., Béja, 11 km W Teboursouk pass, 36 ° 30 ' N, 9 ° 10 ' E, 700 m, under stones, 1. III. 2012, leg. Meybohm (cAss); 10 exs., same data, but leg. Hetzel (cFel); 2 exs., Béja, Djebba, 36 ° 28 ' N, 9 ° 06 ' E, 630 m, near rocks, sifted, 4. III. 2012, leg. Meybohm (cAss); 7 exs., Béja, Djebba, 36 ° 28 ' N, 9 ° 05 ' E, 640 m, moist arable land, 4. III. 2012, leg. Meybohm (cAss); 15 exs., same data, but leg. Hetzel (cFel).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFF415286CCB4C7EC.taxon	discussion	Comment: This species is distributed in North Africa and Sicily (ASSING 2005 b, 2005 c, 2006 b, 2010).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFCC75086CB3CC10C.taxon	materials_examined	Material examined: Greece, Pelopónnisos: 1 ♂, 1 ♀, Ahaia, Oros Aroania, 1550 m, 4. VI. 2007, leg. Giachino & Vailati (cAss); 1 ex., NW Panapoulos, 37 ° 50 ' N, 21 ° 40 ' E, 540 m, oak forest, 3. V. 2007, leg. Schnitter & Arndt (cApf); 1 ex., Korinthia, Feneós, oak forest, pitfall, 28. VII. 2005, leg. Arndt (cApf).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51BFCC75086CB3CC10C.taxon	discussion	Comment: Drusilla meridiana is endemic to the Pelopónnisos, southern Greece (ASSING 2005 b, 2010).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51AFCC75766CC08C7AC.taxon	materials_examined	Material examined: Kyrgyzstan: 2 ♂♂, Issyk-Kul, Kek Mojnok-Vtoroe - Balykchy, 42 ° 27 ' N, 75 ° 51 ' E, 1580 m, 19. VI. 2011, leg. Frisch (MNHUB, cAss); 1 ♂, Issyk-Kul, Tyup, Ak Bulak, 42 ° 49 ' N, 78 ° 14 ' E, 1910 m, 29. VI. 2011, leg. Frisch (MNHUB); 1 ♂, Chui, SE Bishkek, 11 km W Yurevka, 42 ° 44 ' N, 74 ° 55 ' E, 1100 m, 4. VII. 2011, leg. Frisch (MNHUB); 1 ♂, Chui, SW Tokmok, Kegety, 42 ° 37 ' N, 75 ° 08 ' E, 1500 m, 3. VII. 2011, leg. Frisch (cAss); 1 ♂, 1 ex., Chui, S Bishkek, SW Koytash, Tatyr, 42 ° 40 ' N, 74 ° 39 ' E, 1500 m, 5. VII. 2011, leg. Frisch (MNHUB, cAss); 2 exs., Batken, Isfana-Isfara, WSW Ozgorush, tributary of Lyailek river, 39 ° 44 ' N, 70 ° 00 ' E, 1560 m, 23. VI. 2012, leg. Frisch (MNHUB, cAss); 30 exs., Batken, S Quvasoy, Majdan, 40 ° 05 ' N, 72 ° 04 ' E, 1370 m, 17. VI. 2012, leg. Frisch (MNHUB); 1 ex., Batken, S Quvasoy, S Karaul, Austan, 39 ° 58 ' N, 72 ° 07 ' E, 1380 m, 18. VI. 2012, leg. Frisch (MNHUB); 1 ex., Batken, S Quvasoy, 2 km N Langar, 39 ° 52 ' N, 72 ° 05 ' E, 1610 m, 18. VI. 2012 (MNHUB); 20 exs., Batken, SW Isfana, Dinau, 39 ° 42 ' N, 69 ° 22 ' N, 1770 m, 24. VI. 2012, leg. Frisch (MNHUB, cAss); 1 ex., Batken, Isfana-Isfara, WSW Ozgorush, Lyailek river, 39 ° 44 ' N, 70 ° 00 ' E, 1520 m, 22. VI. 2012, leg. Frisch (cAss); 6 exs., Batken, Isfana-Isfara, WSW Ozgorush, tributary of Lyailek river, 39 ° 44 ' N, 70 ° 00 ' E, 1560 m, 23. VI. 2012, leg. Frisch (MNHUB, cAss); 1 ex., Batken, E Sovetsky, 40 ° 07 ' N, 71 ° 23 ' E, 1410 m, 20. VI. 2012, leg. Frisch (MNHUB); 2 exs., Batken, ESE Sovetsky, 40 ° 5 ' N, 71 ° 23 ' E, 1790 m, 20. VI. 2012, leg. Frisch (MNHUB, cAss); 7 exs., Jalal Abad, ESE Toktogul, Torkent, 41 ° 51 ' N, 73 ° 10 ' E, 960 m, 2. VII. 2012, leg. Frisch (MNHUB, cAss); 5 exs., Jalal Abad, NE Karakul, N Kekbel, Kekbel pass, 41 ° 43 ' N, 72 ° 55 ' E, 1420 m, 2. VII. 2012, leg. Frisch (MNHUB, cAss). Tajikistan: 1 ♂, Varzob district, 48 km N Dushanbe, Khodja Obi Gharm, 1900 m, 3. – 4. VII. 2012, leg. Valainis (cSha).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA68C51AFCC75766CC08C7AC.taxon	discussion	Comment: Drusilla heydeni is widespread in Middle Asia, where it is the most common representative of the genus (ASSING 2005 a, 2008 a, 2011).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA69C51AFC9D5266CA83C02C.taxon	materials_examined	Type material examined: Holotype ♀: “ China: Sichuan, Qingcheng Shan, 65 km NW Chengdu, 103.33 E, 30.53 N, 18. V. / 3. – 4. VI. 1997, 800 km W Taiping, 800 – 1000 m, leg. A. Pütz / Holotypus Drusilla gilva mihi, det. R. Pace 2009 / Drusilla gilva n. sp., det. R. Pace 2009 ” (cPüt).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA69C51AFC9D5266CA83C02C.taxon	discussion	Comment: The original description is based on one female specimen from “ Sichuan, Qingcheng Shan, 65 km NW Chengdu ” (PACE 2012).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA69C51AFF1F555ACB2EC28C.taxon	description	Twelve species of Drusilla had been recorded from China (exclusive of Taiwan and Hong Kong): Drusilla aino NAKANE, 1963 (Japan; doubtfully recorded from Jilin), D. cenwanglaomontis PACE, 2012 (Guangxi); D. damingensis PACE, 2012 (Guangxi), D. dendrolasii MARUYAMA & KISHIMOTO, 2002 (Hunan), D. gibberella PACE, 1998 (China: Hainan; Hong Kong; Nepal), D. gilva PACE, 2012 (Sichuan), D. imurai MARUYAMA & KISHIMOTO, 2002 (Hubei), D. philippina (BERNHAUER, 1915) (Hainan), D. sinicola PACE, 2012 (Sichuan), D. watanabei MARUYAMA & KISHIMOTO, 2002 (Hubei, Hunan), D. yunnanensis PACE, 1993 (Hubei, Hunan, Yunnan), and D. zhejiangensis PACE, 1998 (Zhejiang) (MARUYAMA & KISHIMOTO 2002 a, b; PACE 1993, 1998, 2006, 2012). Unlike most Drusilla species from the West Palaearctic region, at least some of the species recorded from China are remarkably widespread. This even applies to the micropterous D. watanabei. The distributions of the Chinese representatives of the genus are illustrated in Maps 1 – 2. The examined recently collected Drusilla material from the Chinese province Yunnan is composed of four species, two of them described previously and two undescribed. In order to facilitate future identification of D. watanabei and D. yunnanensis, both species are redescribed and newly illustrated. The two new species are distinguished from the three Drusilla species recorded from Burma (D. assamensis (CAMERON, 1939), D. feae (CAMERON, 1939), D. malaisei (SCHEERPELTZ, 1965 )) by different coloration alone.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA69C51FFC9D53C6CDF5C2AC.taxon	discussion	(Figs 1 – 13)	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA69C51FFC9D53C6CDF5C2AC.taxon	materials_examined	Material examined: China: Yunnan: 3 ♂♂, 1 ♀, mountains W Dongchuan, Sedan Snow Mountain Scenic Resort, 26 ° 06 ' N, 102 ° 55 ' E, 2620 m, secondary pine forest, litter, moss, and roots of herbs sifted, 14. VIII. 2014, leg. Assing (cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA69C51FFC9D53C6CDF5C2AC.taxon	discussion	Comment: The original description is based on 16 type specimens from two localities in Hubei and Hunan provinces (MARUYAMA & KISHIMOTO 2002 a). The authors did not find (and illustrate) a spermatheca, although the type series includes seven females. A full redescription is provided below.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA69C51FFC9D53C6CDF5C2AC.taxon	description	Description: Relatively large species; body length 7.0 – 7.8 mm; length of forebody 2.6 – 3.0 mm. Coloration: body black, with the posterior margins of the abdominal segments paler; legs with the tibiae and tarsi yellowish to dark-yellowish, the profemora blackish, the mesofemora blackish with the basal third yellowish, and the metafemora yellowish in basal half and infuscate in apical half; antennae reddish-yellow, with antennomere I weakly to distinctly infuscate; maxillary palpi dark-brown, with the terminal palpomere reddish-yellow. Head (Figs 1 – 2) with sexually dimorphic punctation and with fine and shallow microreticulation. Eyes distinctly shorter than distance from posterior margin of eye to posterior constriction of head. Antenna (Fig. 3) 2.6 – 2.9 mm long and slender; all antennomeres at least weakly oblong. Pronotum (Figs 1 – 2) 1.00 – 1.05 times as long as broad and 1.17 – 1.18 times as broad as head, broadest in anterior half; posterior angles weakly marked; near posterior margin with pronounced transverse impression; other impressions and punctation sexually dimorphic; interstices without microsculpture and glossy. Elytra (Figs 1 – 2) short, approximately 0.65 times as long as pronotum; humeral angles moderately marked; punctation sexually dimorphic; interstices without microsculpture. Hind wings reduced to short stubs extending slightly beyond posterior margin of elytra. Metatarsomere I elongate, approximately as long as the combined length of II – IV. Abdomen (Fig. 4) distinctly broader than elytra; tergites III – VIII with sexual dimorphism; punctation sexually dimorphic; anterior impressions of tergites III – VI impunctate; tergites with very weak, nearly obsolete transverse microsculpture; posterior margin of tergite VII with palisade fringe. ♂: lateral portions of dorsal surface of head with dense and rather coarse punctation, with the interstices narrower than diameter of punctures; frons, posterior portion, and the narrow midline of the head impunctate or nearly so (Fig. 1); pronotum in posterior half with pronounced longitudinal elevation on either side, between these elevations with broadly flattened impression; median portion of pronotum somewhat shiny and with moderately dense and moderately coarse punctation, with the interstices as broad as, or broader than, diameter of punctures; lateral portions of pronotum practically matt, with very dense and coarsely granulose punctation, with the interstices narrower than diameter of punctures (Fig. 1); elytra with dense and coarsely granulose punctation (Fig. 1); abdominal tergite III (Fig. 4) with a pronounced median tubercle of somewhat variable shape and with scattered fine punctures; tergites IV – VI each with a median pair of distinct granules near posterior margin, impunctate (or nearly so), except for some punctures at posterior margins; tergite VII with a pronounced median granule and numerous additional smaller granules in posterior half (Fig. 4); tergite VIII (Fig. 5) with densely granulose punctation and with distinct microreticulation, posterior margin broadly concavely excavate; sternite VIII (Fig. 6) weakly transverse and with nearly truncate posterior margin; median lobe of aedeagus large, 1.0 – 1.1 mm long, shaped as in Figs 7 – 8; paramere 0.9 mm long and shaped as in Fig. 9. ♀: head (Fig. 2) with sparse and very fine punctation, interstices much broader than diameter of punctures; pronotum (Fig. 2) with weakly pronounced elevations in posterior half; punctation moderately fine, moderately dense, and non-granulose; elytra (Fig. 2) with weakly granulose puncation; abdominal tergites III – V with fine and sparse punctation; tergite VI impunctate except for a median pair of setiferous punctures bearing long dark setae and additional punctures at posterior margin; tergite VII with a median pair of setiferous punctures bearing long dark setae and with additional fine and sparse punctation in posterior half; posterior margin of tergite VIII somewhat truncate, only indistinctly concave in the middle (Fig. 10); sternite VIII (Fig. 11) distinctly shorter and more transverse than that of male, posterior margin truncate; spermatheca very small in relation to body size, 0.25 mm long, and shaped as in Figs 12 – 13. Comparative notes: This highly distinctive species differs from other Drusilla species known from China by numerous characters, particularly its large body size, the long and slender antennae, the short elytra, the reduced hind wings, the male secondary sexual characters, and the male and female primary sexual characters.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA69C51FFC9D53C6CDF5C2AC.taxon	distribution	Distribution and natural history: The known distribution now includes a total of three localities in Hubei, Hunan, and Yunnan provinces. The examined specimens were sifted from litter, moss, and the roots of herbs in a terraced secondary pine forest at an altitude of 2620 m. MARUYAMA & KISHIMOTO (2002 a) found D. watanabei associated with the ant Lasius (Dendrolasius) spathepus WHEELER, 1910.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA6CC511FCC75246CD03C6EC.taxon	description	(Figs 14 – 25)	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA6CC511FCC75246CD03C6EC.taxon	materials_examined	Type material: Holotype ♂: “ CHINA (Yunnan) Dali Bai Aut. Pref., Wuliang Shan, 20 km NW Weishan 1900 m (creek valley, under bank vegetation, stones, gravel) 25 ° 19 ' 58 " N, 100 ° 07 ' 59 " E, 17. IX. 2009 D. W. Wrase [58] / Holotypus ♂ Drusilla flagellata spec. nov. det. V. Assing 2015 ” (cAss). Paratypes: 4 ♂♂, 2 ♀♀: same data as holotype (cSch, cAss); 1 ♂, 1 ♀: same data, but leg. Schülke (cSch); 1 ♂: “ CHINA (Yunnan) Lincang Pref., Xue Shan, 48 km N Lincang 2070 m, 24 ° 19 ' 03 " N, 100 ° 07 ' 13 " E (slope close to small pond, in loamy soil, plant roots) 12. IX. 2009, D. W. Wrase [45] ” (cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA6CC511FCC75246CD03C6EC.taxon	etymology	Etymology: The specific epithet (Latin, adjective) alludes to the conspicuously long flagellum in the internal sac of the aedeagus.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA6CC511FCC75246CD03C6EC.taxon	description	Description: Body size variable; body length 4.5 – 6.5 mm; length of forebody 2.2 – 2.7 mm. Coloration: head, pronotum, and abdomen blackish; elytra dark-yellowish, with the scutellar region and the postero-lateral angles extensively infuscate; legs pale-yellowish, with the apices of the femora more or less distinctly infuscate; antennae black; maxillary palpi reddish to reddish-brown, with the terminal palpomere yellow. Head (Figs 14 – 15) approximately 1.2 times as broad as long; dorsal surface with scattered and very fine punctation, without microsculpture, and glossy. Eyes large and bulging, much longer than distance from posterior margin of eye to posterior constriction of head. Antenna (Fig. 16) 2.0 – 2.3 mm long; antennomeres III – X gradually increasing in width; antennomere X approximately as long as broad. Pronotum (Figs 14 – 15) 1.02 – 1.06 times as broad as long and 1.06 – 1.09 times as broad as head, broadest at anterior angles; posterior angles obtusely marked; posteriorly with small impression of more or less triangular shape; midline with narrow, deep, and sharply delimited furrow extending from posterior impression cephalad, but not reaching anterior margin of pronotum; punctation dense and distinct, not granulose, subject to weakly pronounced sexual dimorphism; interstices without microsculpture and glossy. Elytra (Figs 14 – 15) approximately 0.9 times as long as pronotum; humeral angles marked; punctation dense, similar to that of pronotum. Hind wings fully developed. Metatarsomere I nearly as long as the combined length of II – IV. Abdomen (Fig. 17) narrower than elytra; tergites III – VII without sexual dimorphism; anterior impressions of tergites III – V and posterior margins of tergites III – VI with fine punctation, remainder of tergal surfaces with scattered and conspicuously sparse punctation, nearly impunctate; posterior margin of tergite VII with palisade fringe. ♂: pronotum (Fig. 14) with extensive, but not very deep median impression with somewhat denser punctation; elytra in large males each with conspicuous oblong and oblique elevation (Fig. 14); tergite VIII (Fig. 18) strongly transverse and with distinctly serrate posterior margin; sternite VIII (Fig. 19) transverse, posterior margin truncate in the middle; median lobe of aedeagus (Figs 20 – 21) 1.0 mm long and of highly distinctive morphology, laterally somewhat flattened, with short ventral process, and with long flagellum in internal sac; paramere (Fig. 22) very small, approximately 0.55 mm long, little more than half as long as median lobe. ♀: pronotum (Fig. 15) flattened, but not impressed in the middle; tergite VIII (Fig. 23) strongly transverse, posterior margin with a triangular projection on either side; sternite VIII (Fig. 24) distinctly shorter and more transverse than that of male, posterior margin convex; spermatheca long and of distinctive shape (Fig. 25). Intraspecific variation: Body size and the modifications of the male elytra are subject to remarkable intraspecific variation. The elevations on the male elytra are fully pronounced only in the holotype, weakly pronounced in one, barely noticeable in two, and absent in three males. Comparative notes: This species is characterized particularly by the modifications of the male elytra, the shapes of the male and female tergites VIII, and above all by the conspicuous morphology of the aedeagus and the distinctive shape of the spermatheca. For additional characters separating it from the sympatric and syntopic D. palata see the comparative notes in the following section.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA6CC511FCC75246CD03C6EC.taxon	distribution	Distribution and natural history: The known distribution is confined to two localities in West Yunnan. The specimens were collected on and near the bank of a stream and near a pond at altitudes of 1900 and 2070 m, in one locality together with D. palata. Two paratypes are slightly teneral.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA62C510FF415586CFAAC2CC.taxon	description	(Figs 26 – 33)	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA62C510FF415586CFAAC2CC.taxon	materials_examined	Type material: Holotype ♂: “ CHINA (Yunnan) Dali Bai Aut. Pref., Wuliang Shan, 20 km NW Weishan 1900 m (creek valley, under bank vegetation, stones, gravel) 25 ° 19 ' 58 " N, 100 ° 07 ' 59 " E, 17. IX. 2009 D. W. Wrase [58] / Holotypus ♂ Drusilla palata spec. nov. det. V. Assing 2015 ” (cAss). Paratypes: 2 ♂♂: “ CHINA (Yunnan) Dali Bai Aut. Pref., Hóu Hé river bank, 10 km S Weishan, 1650 m (in gravel close to water) 25 ° 08 ' 36.8 " N, 100 ° 19 ' 34.5 " E, 14. IX. 2009 D. W. Wrase [53] ” (cSch).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA62C510FF415586CFAAC2CC.taxon	etymology	Etymology: The specific epithet is an adjective derived from the Latin noun pala (shovel) and alludes to the conspicuous shape of the posterior projection of the male sternite VIII.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA62C510FF415586CFAAC2CC.taxon	description	Description: Body length 5.1 – 6.3 mm; length of forebody 2.5 – 2.6 mm. Coloration: head, pronotum, and abdomen blackish; elytra dark-yellowish, with the scutellar region and an extensive lateral spot infuscate; legs uniformly pale-yellowish; antennae blackish-brown; maxillary palpi reddish to dark-brown, with the terminal palpomere yellow. Head (Fig. 26) approximately 1.2 times as broad as long; dorsal surface with median portion flattened or shallowly impressed; punctation fine to moderately coarse and of variable density, sparse to dense; interstices without microsculpture and glossy. Eyes large and bulging, much longer than distance from posterior margin of eye to posterior constriction of head. Antenna (Fig. 27) 2.1 mm long; antennomeres III – X gradually and very weakly increasing in width; preapical antennomeres weakly transverse. Pronotum (Fig. 26) 1.06 – 1.10 times as broad as long and 1.06 times as broad as head, broadest at anterior angles; posterior angles obtusely marked; posteriorly with a very small and indistinct impression; midline with narrow, deep, and sharply delimited furrow extending from posterior impression cephalad, but not reaching anterior margin of pronotum; punctation dense and coarse, not granulose; interstices without microsculpture and glossy. Elytra (Fig. 26) approximately 0.9 times as long as pronotum; humeral angles marked; punctation dense, similar to that of pronotum. Hind wings fully developed. Metatarsomere I shorter than the combined length of II – IV. Abdomen (Fig. 28) narrower than elytra; tergites III – VII without sexual dimorphism; anterior impressions of tergites III – V and posterior margins of tergites III – VII with fine punctation, remainder of tergal surfaces with very sparse punctation; posterior margin of tergite VII with palisade fringe. ♂: pronotum (Fig. 26) with extensive and pronounced median impression in posterior half; tergite VIII (Fig. 29) strongly transverse and with pronounced posterior concavity of nearly semi-circular shape; sternite VIII (Fig. 30) of highly distinctive shape, posteriorly with a pronounced truncate projection and with acutely projecting posterior angles; median lobe of aedeagus (Figs 31 – 32) 0.9 mm long and of weakly derived morphology, with pronounced crista apicalis, and with short flagellum and additional dark structures in internal sac; paramere (Fig. 33) approximately 0.75 mm long, only slightly shorter than median lobe. ♀: unknown. Comparative notes: From the externally (size, coloration, proportions, punctation) similar and syntopic D. flagellata, this species is readily distinguished by the uniformly yellowish legs, the deeper impression on the male pronotum, the slightly different coloration of the elytra, the absence of a sexual dimorphism of the elytra, the competely different shapes of the male tergite and sternite VIII, the completely different morphology of the median lobe of the aedeagus, and the distinctly larger and differently shaped paramere.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA62C510FF415586CFAAC2CC.taxon	distribution	Distribution and natural history: Drusilla palata is known from two localities in West Yunnan. The specimens were collected from gravel on the banks of a stream and a river at altitudes of 1650 and 1900 m, in one locality together with D. flagellata.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA63C512FF1F51A6CA06C10C.taxon	description	(Figs 34 – 45)	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA63C512FF1F51A6CA06C10C.taxon	materials_examined	Material examined: China: Yunnan: 3 ♂♂, 2 ♀♀, Yunnan, mountains S Jianshui, 23 ° 25 ' N, 102 ° 51 ' E, 1810 m, secondary forest margin, litter and various debris sifted, 22. VIII. 2014, leg. Assing & Schülke (cSch, cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA63C512FF1F51A6CA06C10C.taxon	discussion	Comment: In the original description, which is based on a male holotype from “ Yunnan, Dali ” and a female paratype from “ Yunnan, Kunming ”, PACE (1993) describes the coloration as “ ...; uriti liberi primo e secondo giallo-rossicci; antenne brune con i due articoli basali di un rossiccio scuro; zampe gialle con ginocchia medie e posteriori di un rossiccio scuro ”. The other characters specified in the very short original description are too vague to be of taxonomic use. Despite the differences in the coloration (see the redescription below), the examined material is undoubtedly conspecific with the holotype, as can be inferred from the identical shape of the median lobe of the aedeagus (compare Figs 40 – 41 and figures 144 – 145 in PACE 1993). The type specimens may be teneral. The illustration of the spermatheca (PACE 1993: figure 141) of the paratype is misleading. In order to allow a reliable identification of this species, a redescription and new illustrations are provided.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA63C512FF1F51A6CA06C10C.taxon	description	Redescription: Body length 4.8 – 6.3 mm; length of forebody 2.3 – 2.6 mm. Coloration: body black, elytra with or without faint bronze hue; legs uniformly yellowish, rarely with the femoral apices indistinctly darker; antennae black, with antennomeres I – III slightly paler blackishbrown; maxillary palpi pale-brown to dark-brown, with the terminal palpomere yellow. Head (Figs 34 – 35) approximately 1.1 times as broad as long; punctation of dorsal surface subject to sexual dimorphism; interstices without microsculpture and glossy. Eyes large and bulging, much longer than distance from posterior margin of eye to posterior constriction of head. Antenna (Fig. 36) 2.0 – 2.1 mm long; antennomeres III – X gradually increasing in width; antennomeres VI – X transverse. Pronotum (Figs 34 – 35) approximately 1.06 times as broad as long and 1.10 – 1.13 times as broad as head, broadest at anterior angles; posterior angles obtusely marked; posteriorly with small transverse impression; midline with very fine line extending cephalad from posterior impression, but not reaching anterior margin of pronotum; punctation sexually dimorphic; interstices without microsculpture and glossy. Elytra (Figs 34 – 35) approximately 0.9 times as long as pronotum; humeral angles marked; punctation very dense, not very coarse, but much more distinct than that of pronotum. Hind wings fully developed. Metatarsomere I shorter than the combined length of II – IV. Abdomen (Fig. 37) approximately as broad as elytra; tergites III – VII without sexual dimorphism; anterior impressions of tergites III – V and anterior portion of tergite VI impunctate; tergite III with moderately sparse to very sparse fine punctation; tergites IV – VI with setiferous punctures at posterior margin, remainder of tergal surfaces with very sparse and minute punctures; tergite VII with sparse and fine punctation, posterior margin with palisade fringe. ♂: postero-median portion of head with moderately sparse, somewhat coarser punctures, remainder of dorsal surface with fine and sparse punctation; pronotum (Fig. 34) with extensive, but not very deep median impression in posterior half; punctation of pronotum fine and moderately dense, postero-lateral edges of postero-median impression with coarser granulose sculpture; tergite VIII (Fig. 38) strongly transverse and with distinctly serrate posterior margin, postero-lateral angles with more pronounced tooth on either side; sternite VIII (Fig. 39) weakly transverse, posterior margin weakly convex in the middle; median lobe of aedeagus (Figs 40 – 41) relatively small, approximately 0.65 mm long; paramere 0.6 mm long and shaped as in Fig. 42. ♀: head (Fig. 35) with extemely fine, barely noticeable sparse punctation; pronotum (Fig. 35) somewhat depressed in posterior half, but without distinct impression (aside from the small posterior transverse impression); punctation of pronotum rather dense and fine, but more distinct than that of head; tergite VIII (Fig. 43) strongly transverse, posterior margin weakly concave in the middle; sternite VIII (Fig. 44) distinctly shorter and more transverse than that of male, posterior margin weakly convex; spermatheca 0.33 mm long, with large, apically truncate, and strongly sclerotized distal portion (Fig. 45). Comparative notes: Among the Drusilla species recorded from China, D. yunnanensis is characterized particularly by the dense punctation of the elytra, the fine punctation of the head and the pronotum, the impunctate anterior impressions of the abdominal tergites III – V, the male secondary sexual characters, the shape of the female tergite VIII, the morphology of the median lobe of the aedeagus, and the shape of the spermatheca.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA63C512FF1F51A6CA06C10C.taxon	distribution	Distribution and natural history: The currently known distribution is confined to three localities in Yunnan. MARUYAMA & KISHIMOTO (2002 a) collected the species from trails of Lasius (Dendrolasius) spathepus WHEELER, 1910 and L. (D.) capitatus KUZNETSOV-UGAMSKY, 1927. The examined specimens were sifted from litter and various debris at the margin of a secondary mixed forest at an altitude of 1810 m.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA61C512FC9D5646CB6DC46C.taxon	materials_examined	Material examined: Russia: 5 exs., N-Caucasus, 80 km SW Maykop, 950 m, 6. – 9. V. 1991, leg. Medvedev (NHMB, cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA61C512FC9D5646CB6DC46C.taxon	discussion	Comment: The distribution of this species is confined to the Caucasus region, from Turkey and Armenia across Georgia to South Russia and Azerbaijan (SMETANA 2004).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA61C512FC9D52A6CB6FC0EC.taxon	materials_examined	Material examined: Ukraine: 5 exs., Crimea, Alushta distr., m. Kazu-Kaya, 30. IV. 2009, leg. Gontarenko (cGon, cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA61C512FC9D52A6CB6FC0EC.taxon	discussion	Comment: The trans-Palaearctic distribution of P. limbata ranges from West Europe to East Siberia (SMETANA 2004).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA61C512FC9D5386CA71C6AC.taxon	description	(Figs 46 – 47)	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA61C512FC9D5386CA71C6AC.taxon	materials_examined	Material examined: Iraq: 1 ♂, Amedi env., 1200 m, V. 2007, leg. Reuter (cFel).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA61C512FC9D5386CA71C6AC.taxon	discussion	Comment: The original description of this species is based on two females from northern Iraq (ASSING 2008 b). The male sexual characters were previously unknown. Based on the above male, they are as follows: posterior margin of tergite VIII weakly concave in the middle; posterior margin of sternite VIII broadly convex; median lobe of aedeagus 0.72 mm long and shaped as in Figs 46 – 47. The male sexual characters are highly similar to those of P. ruficollis (GRIMM, 1845), except for the slightly longer ventral process of the aedaegus.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA61C515FC9C5706CB17C26C.taxon	description	(Figs 48 – 49)	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA61C515FC9C5706CB17C26C.taxon	materials_examined	Material examined: China: Gansu: 4 exs. [1 teneral], Lenglong Ling mountains, 60 km NNW Honggu, Jin Sha Gorge, 36 ° 52 ' N, 102 ° 39 ' E, 2330 m, mixed forest, litter and moss sifted, 12. VII. 2011, leg. Schülke (cSch, cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA61C515FC9C5706CB17C26C.taxon	discussion	Comment: The previously known distribution of P. lugens was confined to the West Palaearctic region (including Middle Asia) (MARUYAMA 2006). The above specimens represent the first record from the East Palaearctic region east of Middle Asia and considerably expand the range eastwards. The median lobe of the aedeagus of the material from Gansu is illustrated in Figs 48 – 49.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA66C514FCC75106CC21C52C.taxon	description	(Figs 50 – 57)	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA66C514FCC75106CC21C52C.taxon	materials_examined	Material examined: China: Gansu: 2 ♂♂ [1 teneral], mountains SE Longnan, 33 ° 11 ' N, 105 ° 14 ' E, 2130 m, N-slope with scree, stony soil sifted, 7. VIII. 2012, leg. Assing & Wrase (cAss, cSch); 7 exs., Lenglong Ling mountains, 60 km NNW Honggu, Jin Sha Gorge, 36 ° 52 ' N, 102 ° 39 ' E, 2330 m, mixed forest, litter and moss sifted, 12. VII. 2011, leg. Schülke (cSch, cAss); 2 exs., same data, but 14. – 15. VII. 2011 (cSch, cAss); 2 exs., same data, but pitfall traps, 12. – 13. VII. 2011, leg. Wrase (cSch, cAss). Qinghai: 20 exs., Daban Shan, 62 km NNW Honggum, 36 ° 51 ' N, 102 ° 37 ' E, 2240 m, stream valley with mixed forest, pitfall traps, 26. – 29. VI. 2011, leg. Wrase (cSch, cAss); 31 exs., same data, but 11. – 15. VII. 2011 (cSch, cAss); 10 exs., same data, but loamy slope, 26. & 29. VI. 2011 (cSch, cAss); 13 exs., same data, but 2240 – 2350 m, litter and moss sifted, 10. VII. 2011, leg. Schülke (cSch, cAss); 8 exs., same data, but 26. VI. 2011, leg. Schülke (cSch, cAss); 1 ex., same data, but 26. VI. – 10. VII. 2011, leg. Wrase (cAss); 4 exs., same data, but 2275 m, litter sifted, 10. VI. 2011, leg. Wrase (cSch, cAss); 1 ex., same data, but 26. VI. 2011 (cAss); 1 ex., Daban Shan, 80 km NNW Honggu, 36 ° 59 ' N, 102 ° 26 ' E, 2350 – 2450 m, stream valley with mixed forest, litter and moss sifted, 27. VI. 2011, leg. Schülke (cSch). Yunnan: 3 ♂♂, 1 ♀, mountains W Dongchuan, Sedan Snow Mountain Scenic Resort, 26 ° 06 ' N, 102 ° 55 ' E, 2620 m, secondary pine forest, litter, moss, and roots of herbs sifted, 14. VIII. 2014, leg. Assing (cAss); 1 ♂, 1 ♀, NE Kunming, 25 ° 09 ' N, 102 ° 54 ' E, 2280 m, secondary pine forest, with scattered old alder, litter sifted, 11. VIII. 2014, leg. Assing & Schülke (cSch).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA66C514FCC75106CC21C52C.taxon	discussion	Comment: This species was previously known only from the Chinese provinces Beijing and Yunnan (MARUYAMA 2006). The above specimens are brachypterous, with the hind wings of strongly reduced length (just projecting beyond posterior margin of elytra). The specimens from Yunnan are distinguished from those from Qinghai and Gansu by an on average more transverse pronotum. Moreover, remarkable variation in the shape of the spermatheca was observed (Figs 54 – 57). Since no additional distinguishing characters were found and since the median lobe of the aedeagus is practically identical (Figs 50 – 53), however, these differences are interpreted as intraspecific variation. Many of the specimens from Gansu and Qinghai were collected with pitfall traps with vinegar.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C514FC9D5546CDF6C56C.taxon	materials_examined	Material examined: Japan: 3 exs., Hokkaido, Sapporo env., Nopporo virgin forest, 5. VI. 2011, leg. Lackner (cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C514FC9D5546CDF6C56C.taxon	discussion	Comment: This rather widespread species has been recorded from Japan, the Russian Far East and Korea (MARUYAMA 2006).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C514FF1F56C6CDE3C18C.taxon	description	(Figs 58 – 60)	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C514FF1F56C6CDE3C18C.taxon	materials_examined	Material examined: China: Gansu: 57 exs., Lenglong Ling mountains, 60 km NNW Honggu, Jin Sha Gorge, 36 ° 52 ' N, 102 ° 39 ' E, 2330 m, mixed forest, litter and moss sifted, 12. VII. 2011, leg. Schülke & Wrase (cSch, cAss); 3 exs., same data, but 14. – 15. VII. 2011 (cSch, cAss); 6 exs., same data, but pitfall traps, 12. – 13. VII. 2011, leg. Wrase (cSch, cAss). Qinghai: 1 ex., Daban Shan, 62 km NNW Honggum, 36 ° 51 ' N, 102 ° 37 ' E, 2240 – 2350 m, litter and moss sifted, 10. VII. 2011, leg. Schülke (cSch); 49 exs., same data, but 26. VI. 2011, leg. Schülke (cSch, cAss); 9 exs., same data, but 2350 m (cSch, cAss); 26 exs., same data, but 2275 m, litter sifted, 26. VI. 2011, leg. Wrase (cSch, cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C514FF1F56C6CDE3C18C.taxon	discussion	Comment: Pella jureceki was previously known from the Russian Far East, Korea, and the Chinese province Beijing. According to the redescription by MARUYAMA (2006), the species is “ blackish brown in ground colour ”, but all the examined specimens are completely black. The primary sexual characters of the material from Gansu are illustrated in Figs 58 – 60. The specimens listed above were collected together with P. cooterorum (in all localities), in Qinghai also with P. zhoui and in Gansu with P. lugens. Unlike P. cooterorum, P. jureceki was almost exclusively sifted from leaf litter.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C514FC9D5366CB1BC7AC.taxon	description	(Figs 61 – 64)	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C514FC9D5366CB1BC7AC.taxon	materials_examined	Material examined: China: Qinghai: 6 exs., Daban Shan, 62 km NNW Honggum, 36 ° 51 ' N, 102 ° 37 ' E, 2275 m, stream valley with mixed forest, litter and moss sifted, 26. VI. 2011, leg. Wrase (cSch, cAss); 1 ex., same data, but 2230 – 2250 m, leg. Schülke (cSch); 1 ex., same data, but 2350 m, leg. Schülke (cSch).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C514FC9D5366CB1BC7AC.taxon	discussion	Comment: Pella zhoui was previously recorded from the Chinese province Beijing and from the Russian Far East (MARUYAMA 2006). The above specimens were collected together with P. cooterorum and P. jureceki. Their primary sexual characters are illustrated in Figs 61 – 64.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C516FC9D56C6CCE8C5AC.taxon	description	(Figs 65 – 72)	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C516FC9D56C6CCE8C5AC.taxon	materials_examined	Type material: Holotype ♂: ” INDIA W. Bengal, Darjeeling distr., Teesta-Rangpo 350 m, Besuchet-Löbl 12. X. 78 / Holotypus ♂ Amaurodera angularis spec. nov., det. V. Assing 2015 ” (MHNG). Paratypes: 2 ♂♂ [1 teneral]: same data as holotype (MHNG, cAss); 1 ♀: ” INDIA W. Bengal, Darjeeling distr., Sevoke 200 m, 7. X. 78 Besuchet-Löbl ” (cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C516FC9D56C6CCE8C5AC.taxon	discussion	Comment: The specific epithet (Latin, adjective: angled) alludes to the shape of the ventral process of the aedeagus in lateral view.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C516FC9D56C6CCE8C5AC.taxon	description	Description: Body length 3.8 – 4.2 mm; length of forebody 1.85 – 2.05 mm. Coloration: head reddish; pronotum reddish to reddish-brown; elytra reddish; abdomen yellowish, with segments VI – VII more or less distinctly and more or less extensively infuscate; legs and antennae yellowish. Head (Fig. 65) as long as broad or weakly oblong, broadest across eyes, and with sexual dimorphism; punctation sparse and extremely fine; interstices with or without very shallow, nearly obsolete microreticulation. Eyes 0.7 – 0.8 times as long as the distance from posterior margin of eye to posterior constriction of head. Antenna (Fig. 66) approximately 2.4 mm long and very slender. Pronotum (Fig. 65) 1.30 – 1.35 times as long as broad and approximately 1.05 times as broad as head; dorsal surface matt; dorsal and lateral surface meeting at an angle posteriorly; microreticulation reaching lateral carinae only posteriorly; narrow median furrow of reduced, but variable length, extending along anterior half to anterior four fifths, far from reaching anterior or posterior margins. Elytra (Fig. 65) approximately 0.55 times as long as pronotum; punctation sparse and extremely fine; pubescence whitish and depressed. Hind wings present. Metatarsomere I longer than combined length of II and III, but shorter than combined length of II – IV. Abdomen (Fig. 67) narrower than elytra; tergites III – VI impunctate except for some setiferous punctures at posterior margin and a median pair of setiferous punctures bearing long brown setae; posterior margin of tergite VII with palisade fringe; posterior margin of tergite VIII (Fig. 68) convex and denticulate. ♂: dorsal surface of head with distinct and extensive impression (Fig. 65); sternite VIII (Fig. 69) nearly as long as broad and with weakly convex posterior margin; median lobe of aedeagus (Figs 70 – 71) 0.6 mm long; ventral process strongly angled in lateral view; paramere 0.5 mm long. ♀: surface of head not impressed; spermatheca as in Fig. 72. Comparative notes: This species is distinguished from other species of the A. veluticollis group (see ASSING 2003) particularly by the conspicuous shape of the ventral process of the aedeagus and by the sexual dimorphism of the head. It additionally differs from the sympatric A. cameroni by the shorter median furrow of the pronotum, the longer elytra, and the differently shaped spermatheca (A. cameroni: spermatheca proximally truncate). The new species is distinguished from A. veluticollis (MOTSCHULSKY, 1858), whose male sexual characters are still unknown and whose original description is based on an unspecified number of syntypes from ” Indes orientales ” (probably Burma today), by the different shape and sculpture of the pronotum (A. veluticollis: antero-lateral projections less pronounced; dorsal and lateral surfaces posteriorly meeting at a more pronounced angle; microreticulation extending to lateral carinae), the shorter median sulcus on the pronotum, and the differently shaped apical cuticular intrusion of the spermatheca. The general shape of the spermatheca is more or less similar also in other species of the A. veluticollis group, e. g., A. thailandensis PACE, 1986 and A. birmana PACE, 1986. For illustrations of A. veluticollis, A. cameroni, and other Himalayan representatives of the genus see ASSING (2003, 2005 d, 2006 a, 2010).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA67C516FC9D56C6CCE8C5AC.taxon	distribution	Distribution and natural history: The known distribution is confined to two localities in Darjeeling district, West Bengal province, North India. The type specimens were collected at low altitudes (200 and 350 m), in one locality together with A. cameroni. One of the paratypes is teneral.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA65C516FC9D54A6CA4EC46C.taxon	materials_examined	Material examined: India: Uttarakhand: 20 exs., NE Govind Ghat, road to Ghangaria, 9. – 10. VI. 2011, leg. Shavrin (cSha, cAss); 4 exs., 14 km E Uttarkashi, 30 ° 45 ' N, 78 ° 34 ' E, 1450 m, 10. – 12. IV. 2012, leg. Shavrin (cAss, cSha). Uttar Pradesh: 1 ex., Kumaon, Rangarh env., 2000 m, 9. X. 1979, leg. Löbl (MHNG). Nepal: 11 exs., Manaslu W Gupchi Danda, 2200 – 2300 m, 28. V. 2006, leg. Schmidt (NME, cAss); 1 ex., Manaslu, E-slope, Machha Khola valley, Gumda to Lapsibot, 28 ° 12 ' N, 84 ° 50 ' E, 1500 – 1900 m, 23. V. 2006, leg. Schmidt (cAss); 1 ex., W-Dhaulagiri, Pelma Khola valley, 28 ° 42 ' N, 82 ° 56 ' E, 2400 – 2700 m, 17. IX. 2012, leg. Schmidt (NME).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA65C516FC9D54A6CA4EC46C.taxon	discussion	Comment: This species is widespread and the most common representative of the genus in the Himalayan region (ASSING 2003, 2005 d, 2006 a, 2010). For a distribution map see ASSING (2006 a).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA65C516FC9D5306CDB0C7CC.taxon	materials_examined	Material examined: India: West Bengal: 1 ♂, Darjeeling District, Sevoke, 200 m, 7. X. 1978, leg. Besuchet & Löbl (MHNG).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA65C516FC9D5306CDB0C7CC.taxon	discussion	Comment: The known distribution of A. cameroni extends from Uttarakhand to Assam (ASSING 2003, 2005 d, 2009).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA65C516FF1F5746CD98C06C.taxon	materials_examined	Material examined: China: 1 ♂, Yunnan, Xishuanbanna, 60 km N Jinghong, Guanping env., 1000 m, 19. – 27. I. 2003, leg. Murzin (cAss); 2 exs., Hainan, Limu Shan, forest above forest administration centre, 19 ° 11 ' N, 109 ° 44 – 45 ' E, 650 – 900 m, secondary forest, sifted from moist leaf litter along trail, 6. V. 2011, leg. Fikáček (NMP, cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA65C516FF1F5746CD98C06C.taxon	discussion	Comment: Amaurodera smetanai was originally described from East Nepal and subsequently twice recorded from the Chinese island Hainan (ASSING 2003; PACE 2012; YAN & LI 2015). The original description of A. yaoana is based on three females from two localities in Thailand. Based on the distinctive shape of the spermatheca, these specimens are conspecific with the widespread A. smetanai. The description of A. smetanai was published in March, whereas the exact date of the publication of the description of A. yaoana is not indicated, so that the latter is the junior synonym. The known distribution of A. smetanai now ranges from Nepal across southern China (Yunnan, Hainan) to Thailand.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA65C509FC9D5706CD00C1AC.taxon	materials_examined	Material examined: India: Uttarakhand: 17 exs., Kosi river valley, 5 km N Ramnagar, 29 ° 26 ' N, 79 ° 08 ' E, 23. – 26. IV. 2012, leg. Shavrin (cAss, cSha); 7 exs., left side of Kosi river, 5 km N Ramnagar, “ N 29 * 32 E 79 * 149 ”, 19. – 23. VI. 2011, leg. Shavrin (cSha, cAss); 7 exs., 14 km E Uttarkashi, 30 ° 45 ' N, 78 ° 34 ' E, 1450 m, 10. - 12. IV. 2012, leg. Shavrin (cAss, cSha); 10 exs., 15 km SW New Tehri, 30 ° 16 ' N, 78 ° 22 ' E, 870 m, 18. – 20. IV. 2012, leg. Shavrin (cSha, cAss); 4 exs., 10 km NW New Tehri, 30 ° 27 ' N, 78 ° 32 ' E, 1200 m, 9. – 10. IV. 2012, leg. Anichtchenko & Shavrin (cSha, cAss); 1 ex., Uttarkashi district, 14 km NW New Tehri, 30 ° 28 ' N, 78 ° 21 ' E, 950 m, 18. IV. 2012, leg. Anichtchenko (cAss). Uttar Pradesh: 1 ex., Garhwal, 16 km from Srinagar, 550 m, 29. X. 1979, leg. Löbl (MHNG). Nepal: 1 ex., S-Dhaulagiri, Phedi to Phalegaon, 28 ° 28 ' N, 83 ° 21 ' E, 1100 – 1800 m, 9. V. 2012, leg. Schmidt (NME).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA65C509FC9D5706CD00C1AC.taxon	discussion	Comment: Like the preceding species, A. soror is widespread in the Himalayan region (ASSING 2003, 2005 d, 2006 a, 2009). For a distribution map see ASSING (2006 a).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FF4154E6CD01C68C.taxon	materials_examined	Material examined: Nepal: 3 exs., Manaslu, SE-slope, Gupchi Danda, 28 ° 09 ' N, 84 ° 47 ' E, 2900 – 3000 m, 21. V. 2006, leg. Schmidt (NME, cAss); 4 exs., Manaslu W Gupchi Danda, 28 ° 09 ' N, 84 ° 46 ' E, 2500 – 2800 m, 19. – 20. V. 2006, leg. Schmidt (NME, cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FF4154E6CD01C68C.taxon	discussion	Comment: The known distribution of A. coriacea is confined to the Manaslu range (ASSING 2003, 2005 d, 2009).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FF415666CC3EC46C.taxon	materials_examined	Material examined: Nepal: 1 ex., Khandbari District, Induwa Khola valley, 2850 m, 15. IV. 1984, leg. Smetana & Löbl (MHNG).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FF415666CC3EC46C.taxon	discussion	Comment: Amaurodera granata has been recorded from Sankhua Saba, Khandbari, and Solukhumbu Districts in East Nepal (ASSING 2003, 2005 d).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FF415706CA5CC2CC.taxon	description	(Figs 73 – 75)	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FF415706CA5CC2CC.taxon	materials_examined	Material examined: India: West Bengal: 4 exs., Darjeeling District, Algarah – Labha, 1900 m, 11. X. 1978, leg. Besuchet & Löbl (MHNG, cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FF415706CA5CC2CC.taxon	discussion	Comment: Previously, only the holotype from Ilam District, East Nepal, was known (ASSING 2005 d). The above specimens represent the first record from India. The previously unknown female sexual characters are illustrated in Figs 73 – 75.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FF415346CF51C70C.taxon	materials_examined	Material examined: Nepal: 5 exs., SW-Dhaulagiri, Maraini, 28 ° 31 ' N, 83 ° 16 ' E, 2600 m, 24. IX. 2012, leg. Schmidt (NME, cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FF415346CF51C70C.taxon	discussion	Comment: This species is endemic to the Annapurna and Dhaulagiri ranges in Nepal (ASSING 2003, 2005 d, 2006 a, 2009).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FCC751A6CB6FC7CC.taxon	discussion	Comment: In a recent article on miscellaneous aleocharines from Nepal, PACE (2013) synonymized A. verrucosa with A. martensi COIFFAIT, 1982 and A. fulgens with A. silvana PACE, 1992 stating that he did not see any differences in the aedeagal morphology, evidently because he had seen material of neither A. verrucosa nor A. fulgens. True, as emphasized by ASSING (2003), A. fulgens is very similar to A. martensi, most likely its adelphotaxon, both in external and sexual characters, but it differs by constant differences of the aedeagal shape (ventral view) and in external characters (particularly elytral punctation). Moreover, the distributions of both species are allopatric. In the case of A. silvana and A. fulgens, the external and sexual characters distinguishing both species are so obvious and so numerous (see the key, descriptions, and illustrations in ASSING (2003 )) that considering these names synonyms appears somewhat bizarre. In conclusion, both A. verrucosa and A. fulgens are formally revalidated.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FCC756A6CDEDC48C.taxon	materials_examined	Material examined: Mongolia: 4 exs., Ulan Bator env., Zajsan, 28. V. 1989, leg. Bocák (NHMB, cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FCC756A6CDEDC48C.taxon	discussion	Comment: This species has been recorded from East Siberia, the Russian Far East, Mongolia, and North China (SMETANA 2004).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FCC75566CADAC50C.taxon	materials_examined	Material examined: Nepal: 1 ex., Kosi, Arun valley, S Num, 1050 m, 21. IV. 1984, leg. Löbl & Smetana (MHNG); 1 ex., Khandbari District, Arun valley at Num main bridge, 1050 m, 20. IV. 1984, leg. Smetana (cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7AC509FCC75566CADAC50C.taxon	discussion	Comment: This species has been recorded from central and eastern Nepal, as well as from North India (West Bengal: Darjeeling) (ASSING 2006 c).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7BC508FF1F5086CA91C56C.taxon	description	(Figs 76 – 81)	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7BC508FF1F5086CA91C56C.taxon	materials_examined	Type material: Holotype ♂ [with worker of Formica chinensis attached to the pin]: “ China [18 a] – S-Gansu, mts. SE Longnan, nest of Formica, 33 ° 11 ' 17 " N, 105 ° 14 ' 12 " E, 2060 m, 7. VIII. 2012, V. Assing / Holotypus ♂ Lomechusoides penicillatus spec. nov. det. V. Assing 2014 ” (cAss). Paratypes: 2 ♂♂ [each with worker of Formica chinensis attached to the pin]: same data as holotype (cAss); 2 ♀♀, 1 sex?: same data as holotype, but leg. Wrase (cSch, cAss).	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7BC508FF1F5086CA91C56C.taxon	etymology	Etymology: The specific epithet (Latin, adjective: with brushes) alludes to the pronounced tufts of golden glandular setae on the anterior segments of the abdomen.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7BC508FF1F5086CA91C56C.taxon	description	Description: Large and robust species; body length 7.1 – 8.0 mm; length of forebody 3.4 – 3.6 mm; width of pronotum (across posterior angles) 2.0 – 2.2 mm; combined width of elytra 2.3 – 2.5 mm. Coloration: head dark-brown to blackish-brown; pronotum brown to dark-brown; elytra reddish to reddish-brown; abdomen dark-reddish, with segment VI and anterior portion of segment VII more or less distinctly darker; legs darkbrown; antennae blackish-brown. Head (Fig. 76) with deep, shallowly microreticulate, and somewhat glossy median groove; remainder of dorsal surface with pronounced microreticulation and matt; punctation relatively fine and barely noticeable in the microsculpture; pubescence short, depressed to suberect, silvery to golden, and directed anteriad. Antenna (Fig. 77) 3.4 – 3.7 mm long; antennomere I large; II much smaller than I and weakly oblong (length including basal portion); III club-shaped and nearly twice as long as broad; IV very weakly oblong; V – X distinctly oblong; XI coniform and apically acute, slightly more than three times as long as broad and (without basal portion) approximately as long as combined length of IX and X; III – XI with very fine and dense, II with stouter and sparser pubescence; inner faces of IV and of anterior half of III with brush-like clusters of short erect setae in addition to the fine pubescence. Pronotum (Fig. 76) approximately 1.6 times as broad as long and 2.2 times as broad as head, distinctly narrowed in anterior half and weakly dilated in posterior half, widest across posterior angles; lateral margins weakly sinuate in posterior half in dorsal view; lateral margins relatively weakly thickened; posterior margin strongly convex in the middle; punctation moderately sparse, distinct, and granulose; lateral margin separated from disc by longitudinal impression; disc, including anterior and posterior margins, with shallow microreticulation and somewhat glossy; lateral margins with pronounced microreticulation and matt. Elytra (Fig. 76) slightly broader than, and nearly as long as pronotum; anteriorly with 5 – 6 long black setae each; posterior margin weakly sinuate near posterior angle; punctation much finer than that of pronotum; interstices with fine microreticulation. Hind wings present. Abdomen (Fig. 78) broader than elytra, broadest at segment V; tergites II – V laterally with pronounced tufts of long golden glandular setae and in posterior halves with scattered punctures with long black setae; tergite VI with sparse setiferous punctures bearing long black setae at posterior margin, otherwise impunctate; tergites VII – VIII with micropunctation; all segments with shallow microreticulation; posterior margin of segment VIII broadly and deeply concave posteriorly, postero-lateral angles each with tuft of golden setae; posterior margin of tergite VII with palisade fringe. ♂: median lobe of aedeagus 1.2 mm long and shaped as in Figs 79 – 80. ♀: spermatheca as in Fig. 81. Comparative notes: Based on the characters specified by JÁSZAY & HLAVÁČ (2013), the new species belongs to the L. strumosus group; which previously included five species: L. inflatus (ZETTERSTEDT, 1828), L. mongolicus, L. sibiricus (MOTSCHULSKY, 1860), L. strumosus (FABRICIUS, 1792), and L. teres (EPPELSHEIM, 1884). Among these species, L. penicillatus is most similar to L. strumosus, together with which it would key out using the key in WASMANN (1897) and from which it differs by the broader and more transverse pronotum, the more oblong preapical antennomeres, the much less thickened lateral margins of the pronotum, the more extensively glossy pronotum (L. strumosus: anterior and posterior margins with pronounced microreticulation and matt), the shorter tibiae, the much more pronounced lateral tufts of glandular setae on the abdominal segments II – V, the posteriorly much more deeply concave tergite VIII, and by the different sexual characters.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
03E487E3CA7BC508FF1F5086CA91C56C.taxon	distribution	Distribution and natural history: The type locality is situated in a mountain range to the southeast of Longnan in southern Gansu province, China. All the specimens were collected from the same nest of Formica chinensis WHEELER, 1913 (Formica truncorum group) on a slope with scree, shrubs, and small trees.	en	Assing, Volker (2015): New species and additional records of Lomechusini from the Palaearctic region, primarily from China (Coleoptera: Staphylinidae: Aleocharinae). Beiträge Zur Entomologie = Contributions to Entomology 65 (2): 243-262, DOI: 10.21248/contrib.entomol.65.2.243-262, URL: https://www.contributions-to-entomology.org/article/view/1882
