taxonID	type	description	language	source
A20FF296692C52CBA46D56B87934B936.taxon	description	Figs 2, 3, 4, 5, 6, 7	en	Lopez-Perilla, Yeny Rocio, Fernandez-Roldan, Juan David, Meza-Joya, Fabio Leonardo, Medina-Rangel, Guido Fabian (2023): A new Bolitoglossa (Amphibia, Caudata, Plethodontidae) from the Cordillera Oriental of Colombia. ZooKeys 1158: 27-48, DOI: http://dx.doi.org/10.3897/zookeys.1158.99077, URL: http://dx.doi.org/10.3897/zookeys.1158.99077
A20FF296692C52CBA46D56B87934B936.taxon	diagnosis	Diagnosis. Bolitoglossa muisca is a member the subgenus Bolitoglossa Eladinea and of the Bolitoglossa adspersa species group. The new species is characterized by the following morphological characters: a large-size body; a broad head; a rounded snout in dorsal and ventral views; a very thick postocular fold; a moderate subgular fold; smooth skin texture; moderately long limbs; moderate webbing on third finger and toe; and a short, robust tail. Even though Bolitoglossa adspersa and B. muisca share their moderate webbing on hands and feet, we regard the former as having less webbing than the latter. Moreover, the tips of the fingers and toes are separated from the distal margin of the webbing, exposing their subcircular-shaped digits; unlike those of B. muisca (Fig. 4). The new species is slightly larger on average than B. adspersa (mean SVL 52.8 + / - 3.4 mm; range 33.0 - 72.1 mm; n = 22 vs. 45.0 + / - 3.4 mm; range 30.7 - 66.3 mm; n = 26), additionally, the tail of B. adspersa is thin and long in relation to the trunk but thick and short in relation to the trunk in B. muisca (Table 1). Bolitoglossa muisca differs from B. capitana by having moderately webbed hands and feet (vs. almost fully webbed hands and feet in B. capitana), by being overall smaller in size (mean SVL 52.8 + / - 3.4 mm; range 33.0 - 72.1 mm; n = 22 vs. 75.2 + / - 10.9 mm; range 59.2 - 85.5 mm; n = 5), by having fewer maxillary teeth (mean MT 24.4 + / - 3.0; range 14 - 36; n = 17 vs. 30.4 + / - 5.1; range 22 - 37; n = 5) by having fewer vomerine teeth (mean VT 35.1 + / - 4.0; range 24 - 53; n = 17 vs. 66.4 + / - 14.8; range 46 - 87; n = 5), and because the new species bears a faint, very small gular fold that is large, thick and notable in B. capitana (Cruz-Rodriguez et al. 2021: figs 13, 14). Bolitoglossa muisca differs from B. pandi because the tips of the third digit and toe of the latter are triangular and pointed in outline (vs. third digits and toes oval and webbed to a higher degree in B. muisca), by being a slightly larger species (mean SVL 52.8 + / - 3.4 mm; range 33.0 - 72.1 mm; n = 22 vs. 44.0 + / - 2.9 mm; range 35.9 - 52 mm; n = 12), and by having mostly smooth skin (vs. coarse skin in B. pandi); moreover, the tail of B. pandi tapers gradually and symmetrically from broad to slender antero-posteriorly, unlike that of the new species, which is slightly rectangular in outline, becoming abruptly wider than the base, and ending in a rounded tip (Table 1).	en	Lopez-Perilla, Yeny Rocio, Fernandez-Roldan, Juan David, Meza-Joya, Fabio Leonardo, Medina-Rangel, Guido Fabian (2023): A new Bolitoglossa (Amphibia, Caudata, Plethodontidae) from the Cordillera Oriental of Colombia. ZooKeys 1158: 27-48, DOI: http://dx.doi.org/10.3897/zookeys.1158.99077, URL: http://dx.doi.org/10.3897/zookeys.1158.99077
A20FF296692C52CBA46D56B87934B936.taxon	description	Description of the holotype. An adult female (SVL = 61.3 mm) with a broad head (HW / SVL = 0.16); head longer than wide (HW / HL = 0.90); neck with a small, faint gular fold; snout short and truncated in profile, and dorsal view, but less so in ventral view (SNL = 3.1 mm); large eyes that do not extend beyond the outline of the head in dorsal view and smaller than interorbital distance (EYD = 3.18 mm, IOD = 3.37 mm); with a thick post ocular fold that extends past the posterior commissure of the eye onto the anterior margin of the gular fold; canthus rostralis subtle, small, rounded in outline; 33 maxillary teeth, 16 to the right and 17 to the left; vomerine teeth 25, these are not arranged in a single row but grouped towards the margins of the parasphenoid bone; with three premaxillary teeth that pierce the upper lip in males; nasolabial grooves well developed; moderate interdigital webbing on hands but third finger extends slightly further than the other fingers; toes with less interdigital webbing than fingers, toes II-V with less membrane than toe I; with subterminal pads on digits, digits in order of increasing length I <II <IV> III; toes I <II <III <IV> V; longest digits of hand and feet are subcircular (L 3 T and L 3 F), limbs relatively long (FL / SVL = 0.23, HLL / SVL = 0.23); tail not exceeding standard length (TL / SVL 0.93), narrower than the body at the base (posterior to the vent), slightly rectangular in outline, becoming abruptly wider than the base and ending in a rounded tip, but this condition is artefactual because the tip of the tail is missing; a long trunk (52.5 mm); with 13 costal grooves (Fig. 3). See Table 2 for meristic data of all type specimens.	en	Lopez-Perilla, Yeny Rocio, Fernandez-Roldan, Juan David, Meza-Joya, Fabio Leonardo, Medina-Rangel, Guido Fabian (2023): A new Bolitoglossa (Amphibia, Caudata, Plethodontidae) from the Cordillera Oriental of Colombia. ZooKeys 1158: 27-48, DOI: http://dx.doi.org/10.3897/zookeys.1158.99077, URL: http://dx.doi.org/10.3897/zookeys.1158.99077
A20FF296692C52CBA46D56B87934B936.taxon	etymology	Etymology. Named after the native human inhabitants of the Altiplano Cundiboyacense and Sabana de Bogota. The Muiscas regarded amphibians as sacred creatures associated with sex, fertility, and the arrival of the rainy season. The specific epithet is used as a noun in apposition.	en	Lopez-Perilla, Yeny Rocio, Fernandez-Roldan, Juan David, Meza-Joya, Fabio Leonardo, Medina-Rangel, Guido Fabian (2023): A new Bolitoglossa (Amphibia, Caudata, Plethodontidae) from the Cordillera Oriental of Colombia. ZooKeys 1158: 27-48, DOI: http://dx.doi.org/10.3897/zookeys.1158.99077, URL: http://dx.doi.org/10.3897/zookeys.1158.99077
A20FF296692C52CBA46D56B87934B936.taxon	distribution	Distribution. At present, Bolitoglossa muisca is known only from small cloud forest remnants on the western slopes of the Cordillera Oriental of Colombia in Bojaca, Granada, San Antonio del Tequendama, Silvania, and Soacha municipalities, Cundinamarca department. All specimens collected between 2390 - 2700 m a. s. l. (Fig. 2).	en	Lopez-Perilla, Yeny Rocio, Fernandez-Roldan, Juan David, Meza-Joya, Fabio Leonardo, Medina-Rangel, Guido Fabian (2023): A new Bolitoglossa (Amphibia, Caudata, Plethodontidae) from the Cordillera Oriental of Colombia. ZooKeys 1158: 27-48, DOI: http://dx.doi.org/10.3897/zookeys.1158.99077, URL: http://dx.doi.org/10.3897/zookeys.1158.99077
A20FF296692C52CBA46D56B87934B936.taxon	materials_examined	Other material examined. Countries are indicated in bold capitals, departments in regular capitals, municipalities, and localities in plain text. * Denotes specimens examined via photographs. Bolitoglossa adspersa (n = 29). Colombia: Boyaca: Duitama, Paramo de la Rusia: ICN 4301, 4310; Toquilla, Paramo de Toquilla: ICN 9487. Cundinamarca: Bogota, D. C., Paramo de Cruz Verde: IAvH-Am- 8917, San Cristobal, Tanques de Vitelma, finca La Marranera: IAvH-Am- 696, 2947, 2954, 2957 - 58, 2964, 2967, 2972, Usme, represa El Hato: ICN 37555; Cabrera: vereda de Hoyerias: IAvH-Am- 13253 Fomeque, Laguna de Chingaza: ICN 4884, 4891; Guasca, Paramo de Guasca: ICN 4521, 4525, 4546; Fomeque, laguna Chingaza: ICN 4885; Fusagasuga, km 50 carretera Fusagasuga-La Florida: ICN 39096; Guatavita, vereda Montenquiva: IAvH-Am- 13174 - 75, ICN 55420; Guayabetal: Paramo Atravezado: IAvH-Am- 14808; Paramo de Palacio, P. N. N. Chingaza: IAvH-Am- 3101, 3096, Quetame, km 22 carretera central Villavicencio-Alto del Tigre: ICN 7123; Ubaque, reserva ecologica Matarredonda: IAvH-Am- 13254. Bolitoglossa capitana (n = 4). Colombia: Cundinamarca: Alban, Granjas del Padre Luna: ICN 9221 & MLS 182 - 184 *. Bolitoglossa guaneae (n = 27). Colombia: Santander: Charala, Virolin, Canaverales, sector El Reloj: ICN 5197, 8555, 8557, 12770 - 72, 34230, Cuchilla del Fara: ICN 47980, Hacienda La Sierra: ICN 34229 - 30, km 56: ICN 19558, UIS-A 1369, UIS-A 2078, UIS-A 2082, UIS-A 2179, UIS-A 2317, UIS-A 2325 - 6, UIS-A 2891, UIS-A 2893, UIS-A 2895, UIS-A 2898 - 9, paratypes UIS-A 2203, UIS-A 2320, UIS-A 2324, UIS-A 2897. Bolitoglossa muisca (n = 54). Colombia: Cundinamarca: Bojaca, vereda Roble Hueco, predio La Esmeralda: IAvH-Am- 17413 - 22, Predio Penas Blancas: IAvH-Am- 17423 - 28; Granada, hacienda El Soche: ICN 3544 - 48, MVZ 167997 *; San Antonio del Tequendama, vereda Roquemonte, cerca de entrada Parque Chicaque: ICN 60319 - 25; Silvania, vereda Noruega Alta, Hacienda La Tribuna: ICN 58245 - 58268; Soacha, vereda Cascajal: IAvH-Am- 17429. Bolitoglossa nicefori (n = 13). Colombia: Santander: Floridablanca, El Mortino, quebrada Torrentosa: ICN 50000, 58227, 58231; Zapatoca, finca Los Puentes, quebrada Uchuvala: ICN 58223 - 24, 58226; Piedecuesta, vereda Los Monos: UIS-A 2987, UIS-A 2991; Los Santos, vereda El Carrizal, finca Utopia: UIS-A 5273 - 4; Guapota, vereda Las Flores, finca La Chocolatera: UIS-A 5264; San Gil, vereda San Jose, finca La Esperanza: UIS-A 5270 - 1. Bolitoglossa pandi (n = 20). Colombia: Cundinamarca: Pandi, vereda Buenos Aires Alta: ICN 45500, Supata, vereda Las Lajas, reserva Cuzcungos: ICN 58492 - 05, Villeta, La Esmeralda: IAvH-Am- 10303 - 08.	en	Lopez-Perilla, Yeny Rocio, Fernandez-Roldan, Juan David, Meza-Joya, Fabio Leonardo, Medina-Rangel, Guido Fabian (2023): A new Bolitoglossa (Amphibia, Caudata, Plethodontidae) from the Cordillera Oriental of Colombia. ZooKeys 1158: 27-48, DOI: http://dx.doi.org/10.3897/zookeys.1158.99077, URL: http://dx.doi.org/10.3897/zookeys.1158.99077
