taxonID	type	description	language	source
01355F73A0DA52D5B33EDBE47A4A9AA0.taxon	description	Figs 2, 3, 4, 5 A Franz's Piping Frog / Rana gaitera de Franz	en	Gruenwald, Christoph I., Montano-Ruvalcaba, Carlos, Jones, Jason M., Ahumada-Carrillo, Ivan, Gruenwald, Andre J., Zheng, Jiacheng, Strickland, Jason L., Reyes-Velasco, Jacobo (2023): A novel species of piping frog Eleutherodactylus (Anura, Eleutherodactylidae) from southern Mexico. Herpetozoa 36: 95-111, DOI: http://dx.doi.org/10.3897/herpetozoa.36.e104707, URL: http://dx.doi.org/10.3897/herpetozoa.36.e104707
01355F73A0DA52D5B33EDBE47A4A9AA0.taxon	diagnosis	Diagnosis. Based on our phylogenetic analysis, this is a member of the genus Eleutherodactylus, subgenus Eleutherodactylus Syrrhophus, as defined by Hedges et al. (2008). In the Eleutherodactylus (Syrrhophus) nitidus species series and the Eleutherodactylus (Syrrhophus) nitidus species group as defined by Gruenwald et al. (2018), based on the condition of the tympanic annuli, ventral epidermis and visceral peritoneum. A small frog, but relatively large Syrrhophus, adult males measure 25.6 - 29.5 mm SVL; vocal slits are present in males, readily visible under partially translucent ventral epidermis; digital tips are widely expanded, 1.8 - 2.6 times the width of the narrowest part of the finger on the third and fourth fingers; fingers moderately long, finger lengths are I-II-IV-III with third finger length ranging from 15 - 18 % of SVL; compact lumbo-inguinal gland above the inguinal region present, raised, indistinct, but visible in live specimen; ventral epidermis is partially translucent and visceral peritoneum is clear, not white, thus abdominal vein is not clearly visible against a white background on the venter of live specimens and viscera is partially visible through translucent dark grey ventral epidermis; limbs moderate, TL / SVL ratio is 0.40 - 0.50, FeL / SVL ratio is 0.31 - 0.38 and TotFL / SVL ratio is 0.50 - 0.60; snout relatively short, END / SVL ratio is 0.08 - 0.11; tympanum small, indistinct and round, tympanic annuli not visible in live specimen; TW / ED ratio is 0.40 - 0.60. The dorsal and lateral skin is slightly shagreened, whereas the ventral skin is smooth. Dorsal colouration cream or tan, with darker brown reticulations on dorsal and lateral portions of head and back; loreal and region dark brown, from snout to behind tympanum, with some pale spots and speckling present on labial region same colour as pale dorsal colouration; pale interorbital bar variable, may be present or absent, when present same colour as pale dorsal colouration of head and body; distinct irregular-edged dark brown transverse bands present on legs; upper arms same colouration and pattern as forearms; no pale mid-dorsal stripe; upper flanks same colour as dorsum, lower flanks whitish with some grey marbling; venter pale lavender with some sparse white spotting and pale grey on throat. No inguinal flash colours present on thighs and groin. The mating call of adult males is a short low-pitched pipe (see below; Fig. 4).	en	Gruenwald, Christoph I., Montano-Ruvalcaba, Carlos, Jones, Jason M., Ahumada-Carrillo, Ivan, Gruenwald, Andre J., Zheng, Jiacheng, Strickland, Jason L., Reyes-Velasco, Jacobo (2023): A novel species of piping frog Eleutherodactylus (Anura, Eleutherodactylidae) from southern Mexico. Herpetozoa 36: 95-111, DOI: http://dx.doi.org/10.3897/herpetozoa.36.e104707, URL: http://dx.doi.org/10.3897/herpetozoa.36.e104707
01355F73A0DA52D5B33EDBE47A4A9AA0.taxon	description	Description of the holotype. Adult male, relatively large (26.2 mm SVL); head as wide (9.5 mm) as long (9.5 mm), head wider than body; snout rounded from a dorsal view and rounded to slightly truncate from a lateral profile; tympanum indistinct, rounded with no supra-tympanic fold present; tympanum small, oval, greatest width of tympanum 1.4 mm; greatest diameter of eye 2.8 mm; tympanum width to eye-diameter 0.51; eyelid width 1.6 mm, approximately 38 % of the IOD; first finger shorter than second finger; finger lengths from shortest to longest I-II-IV-III; digital pads on fingers two, three and four expanded, 2.1 times the narrowest point of the digit on fingers three and four; expanded finger pads widely expanded, truncate, three palmar tubercles; inner palmar tubercle 70 % of middle palmar tubercle and outer palmar tubercles about 60 % as large as middle palmar tubercle, (Fig. 5 A); toe lengths from shortest to longest I-V-II-III-IV, TL 1 and TL 5 very similar; outer metatarsal conical with a round base moderate, approximately 56 % of inner metatarsal tubercle; inner metatarsal tubercle spherical shape with oval base, large, approximately 1.1 mm in length. Dorsal skin smooth, lateral skin slightly shagreened with some low tubercles; ventral skin smooth. Vocal slits present. In life, the holotype had a yellowish-tan dorsal colouration on the back, with darker brown blotches on the back and flanks. Head yellowish-tan with some dark brown speckling, no pale interorbital bar; however, the interorbital region mostly lacks darker brown speckling giving it the resemblance of a pale interorbital bar. Labial region pale grey with some tan and some white speckling. Three and four white-tipped tubercles present at the rictal. A dark brown stripe present from the tip of the snout posteriorly through loreal region, eye and tympanum to right above rictal tubercles. Forearms, thighs, femur and tarsus tan with indistinct pale brown banding. The upper arms were unmarked, tan to slightly orange. No inguinal flash colouration was present on groin or thighs. Ventral colouration was lavender with some sparse white and on sides, ventral colouration on throat grey. Ventral skin was slightly translucent and visceral peritoneum clear, so no visible red abdominal vein and viscera were visible in life. Colouration in preservative is pale tan on dorsum, with darker brown reticulations. Pale tan interorbital bar present. Dark canthal bar is dark brown. Unmarked upper arms are cream to white. Limbs cream with dark brown cross-banding. The dorsal surfaces of the legs are light brown and the groin and posterior surfaces of the thighs are brown. Ventral surfaces yellowish-cream, unmarked, slightly darker brownish pigmentation on throat and chin. Ventral surfaces of hands and feet brown, with dark brown spots. (Fig. 2 D, E).	en	Gruenwald, Christoph I., Montano-Ruvalcaba, Carlos, Jones, Jason M., Ahumada-Carrillo, Ivan, Gruenwald, Andre J., Zheng, Jiacheng, Strickland, Jason L., Reyes-Velasco, Jacobo (2023): A novel species of piping frog Eleutherodactylus (Anura, Eleutherodactylidae) from southern Mexico. Herpetozoa 36: 95-111, DOI: http://dx.doi.org/10.3897/herpetozoa.36.e104707, URL: http://dx.doi.org/10.3897/herpetozoa.36.e104707
01355F73A0DA52D5B33EDBE47A4A9AA0.taxon	description	Measurements of the holotype (in millimetres). IND 2.2, IOD 4.2, END 2.5, ETD 0.9, UpL 6.4, FoL 7.5, PaL 2.3, HaL 6.7, F 1 L 2.3, F 1 PW 0.5, F 1 W 0.4, F 2 L 2.8, F 2 PW 0.8, F 2 W 0.4, F 3 L 4.7, F 3 PW 1.1, F 3 W 0.5, F 4 L 3.5, F 4 PW 1.1, F 4 W 0.5, IPTL 0.7, MPTL 1.0, OPTL 0.6, FeL 10.0, TL 11.0, TaL 7.1, TotFL 11.4, T 1 L 2.4, T 1 PW 0.62, T 1 W 0.5, T 2 L 3.6, T 2 PW 0.7, T 2 W 0.5, T 3 L 4.4, T 3 PW 0.7, T 3 W 0.5, T 4 L 6.0, T 4 PW 0.7, T 4 W 0.5, T 5 L 2.5, T 5 PW 0.6, T 5 W 0.4, IMTL 1.1, OMTL 0.6, FeL / SVL 38 %, TL / SVL 42 %, HaL / SVL 26 %, TotFL / SVL 56 %, HL / SVL 35 %, HW / SVL 36 %.	en	Gruenwald, Christoph I., Montano-Ruvalcaba, Carlos, Jones, Jason M., Ahumada-Carrillo, Ivan, Gruenwald, Andre J., Zheng, Jiacheng, Strickland, Jason L., Reyes-Velasco, Jacobo (2023): A novel species of piping frog Eleutherodactylus (Anura, Eleutherodactylidae) from southern Mexico. Herpetozoa 36: 95-111, DOI: http://dx.doi.org/10.3897/herpetozoa.36.e104707, URL: http://dx.doi.org/10.3897/herpetozoa.36.e104707
01355F73A0DA52D5B33EDBE47A4A9AA0.taxon	distribution	Distribution and ecology. Eleutherodactylus franzi appears to be endemic to the Sierra de Taxco Region of northern Guerrero (Fig. 6). This is an extension of the Mexican Transverse Volcanic Belt which extends south into the Balsas Basin. This frog occurs at high elevations between 2,000 - 2,400 m a. s. l. and has been collected in tropical deciduous forest, oak woodland and pine-oak forest, as well as mixtures of these assemblages. It is restricted to areas of karstic rock outcroppings and the associated sinkholes and caves systems. This species may also occur in nearby Estado de Mexico, as habitat is continuous and the State line is near the type locality. Old reports of E. pipilans from that State may be referable to this species; however, we have made attempts to examine these specimens at MZFC and were not able to find the specimens (C. Gruenwald, pers. obs.)	en	Gruenwald, Christoph I., Montano-Ruvalcaba, Carlos, Jones, Jason M., Ahumada-Carrillo, Ivan, Gruenwald, Andre J., Zheng, Jiacheng, Strickland, Jason L., Reyes-Velasco, Jacobo (2023): A novel species of piping frog Eleutherodactylus (Anura, Eleutherodactylidae) from southern Mexico. Herpetozoa 36: 95-111, DOI: http://dx.doi.org/10.3897/herpetozoa.36.e104707, URL: http://dx.doi.org/10.3897/herpetozoa.36.e104707
01355F73A0DA52D5B33EDBE47A4A9AA0.taxon	etymology	Etymology. This species is named after Hector Franz-Chavez, Mexican herpetologist and avid field collector who collected the type material and who helped collect an extensive sampling of the Eleutherodactylus specimens to be used in our succession of studies.	en	Gruenwald, Christoph I., Montano-Ruvalcaba, Carlos, Jones, Jason M., Ahumada-Carrillo, Ivan, Gruenwald, Andre J., Zheng, Jiacheng, Strickland, Jason L., Reyes-Velasco, Jacobo (2023): A novel species of piping frog Eleutherodactylus (Anura, Eleutherodactylidae) from southern Mexico. Herpetozoa 36: 95-111, DOI: http://dx.doi.org/10.3897/herpetozoa.36.e104707, URL: http://dx.doi.org/10.3897/herpetozoa.36.e104707
